1356 lines
105 KiB
XML
1356 lines
105 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.926.48671" ID-GBIF-Dataset="495ab48b-1fe1-45f9-b5b8-5af790a7cfba" ID-PMC="PMC7170969" ID-Pensoft-Pub="1313-2970-926-133" ID-Pensoft-UUID="B7448F031B605E54AECA6AC3BB498B6A" ID-PubMed="32336923" ID-ZooBank="1D4D9F014F874E4991134B0D72A273D6" ModsDocID="1313-2970-926-133" checkinTime="1586878979185" checkinUser="pensoft" docAuthor="Neang, Thy, Henson, Adam & Stuart, Bryan L." docDate="2020" docId="2E84EE8AC1665CDAB3C1E18270D1D3E0" docLanguage="en" docName="ZooKeys 926: 133-158" docOrigin="ZooKeys 926" docSource="http://dx.doi.org/10.3897/zookeys.926.48671" docTitle="Cyrtodactylus phnomchiensis Neang & Henson & Stuart 2020, sp. nov." docType="treatment" docUuid="103B12F4-6D6F-4928-85A5-A927A81225FE" docUuidSource="ZooBank" docVersion="4" id="B7448F031B605E54AECA6AC3BB498B6A" lastPageNumber="133" masterDocId="B7448F031B605E54AECA6AC3BB498B6A" masterDocTitle="A new species of Cyrtodactylus (Squamata, Gekkonidae) from Cambodia's Prey Lang Wildlife Sanctuary" masterLastPageNumber="158" masterPageNumber="133" pageNumber="133" updateTime="1668168710194" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Cyrtodactylus (Squamata, Gekkonidae) from Cambodia's Prey Lang Wildlife Sanctuary</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neang, Thy</mods:namePart>
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<mods:affiliation>Wild Earth Allies, 77 a Street Beton, Bayap Village, Sangkat Phnom Penh Thmei, Khan Sen Sok, Phnom Penh, Cambodia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Henson, Adam</mods:namePart>
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<mods:affiliation>Wild Earth Allies, 2 Wisconsin Circle, Suite 900, Chevy Chase, Maryland 20815, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stuart, Bryan L.</mods:namePart>
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<mods:affiliation>North Carolina Museum of Natural Sciences, 11 West Jones Street, Raleigh, North Carolina 27601, USA</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>926</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>133</mods:start>
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<mods:end>158</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.926.48671</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.926.48671</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-926-133</mods:identifier>
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<mods:identifier type="ZooBank">1D4D9F014F874E4991134B0D72A273D6</mods:identifier>
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<mods:identifier type="Pensoft-UUID">B7448F031B605E54AECA6AC3BB498B6A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="163626691" LSID="urn:lsid:zoobank.org:act:103B12F4-6D6F-4928-85A5-A927A81225FE" httpUri="http://treatment.plazi.org/id/2E84EE8AC1665CDAB3C1E18270D1D3E0" lastPageNumber="133" pageId="0" pageNumber="133">
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<subSubSection pageId="0" pageNumber="133" type="nomenclature">
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<paragraph pageId="0" pageNumber="133">
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<taxonomicName LSID="2E84EE8A-C166-5CDA-B3C1-E18270D1D3E0" authority="Neang & Henson & Stuart, 2020" authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis" status="sp. nov.">Cyrtodactylus phnomchiensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="133">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Cyrtodactylus phnomchiensis sp. nov. in life. A Male holotype CBC 03012 and B female paratype CBC 03013." figureDoi="10.3897/zookeys.926.48671.figure3" httpUri="https://binary.pensoft.net/fig/399860" pageId="0" pageNumber="133">Figures 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F5" captionText="Figure 4. Head of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative. A Dorsal view illustrating the rostral, supranasal and internsupranasal scales B ventral view illustrating the mental and postmental scales." figureDoi="10.3897/zookeys.926.48671.figure4" httpUri="https://binary.pensoft.net/fig/399861" pageId="0" pageNumber="133">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F6" captionText="Figure 5. Cloacal region of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative illustrating the enlarged precloacal scales, enlarged femoral scales, and not enlarged subcaudal scales." figureDoi="10.3897/zookeys.926.48671.figure5" httpUri="https://binary.pensoft.net/fig/399862" pageId="0" pageNumber="133">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F7" captionText="Figure 6. Dorsal view of the type series of Cyrtodactylus phnomchiensis sp. nov. in preservative." figureDoi="10.3897/zookeys.926.48671.figure6" httpUri="https://binary.pensoft.net/fig/399863" pageId="0" pageNumber="133">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="holotype">
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<paragraph pageId="0" pageNumber="133">Holotype.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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CBC 03012, adult male (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Cyrtodactylus phnomchiensis sp. nov. in life. A Male holotype CBC 03012 and B female paratype CBC 03013." figureDoi="10.3897/zookeys.926.48671.figure3" httpUri="https://binary.pensoft.net/fig/399860" pageId="0" pageNumber="133">3</figureCitation>
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), Cambodia, Kampong Thom Province, Sandan District, Phnom Chi, Prey Lang Wildlife Sanctuary,
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<geoCoordinate degrees="12" direction="north" minutes="56" orientation="latitude" precision="1" seconds="11.6" value="12.936556">12°56'11.6"N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="39" orientation="longitude" precision="1" seconds="17.1" value="105.654755">105°39'17.1"E</geoCoordinate>
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, 237 m elevation, collected on 18 July 2019 by Thy Neang and En E.
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</paragraph>
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<caption doi="10.3897/zookeys.926.48671.figure3" httpUri="https://binary.pensoft.net/fig/399860" pageId="0" pageNumber="133" start="Figure 3" startId="F4">
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<paragraph pageId="0" pageNumber="133">
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<emphasis bold="true" pageId="0" pageNumber="133">Figure 3.</emphasis>
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<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
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<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
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</taxonomicName>
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sp. nov. in life.
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<emphasis bold="true" pageId="0" pageNumber="133">A</emphasis>
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Male holotype CBC 03012 and
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<emphasis bold="true" pageId="0" pageNumber="133">B</emphasis>
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female paratype CBC 03013.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="paratypes">
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<paragraph pageId="0" pageNumber="133">Paratypes.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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All from Cambodia, Kampong Thom Province, Sandan District, Phnom Chi, Prey Lang Wildlife Sanctuary: CBC 03003, adult male,
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<geoCoordinate degrees="12" direction="north" minutes="56" orientation="latitude" precision="1" seconds="09.2" value="12.935889">12°56'09.2"N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="39" orientation="longitude" precision="1" seconds="12.7" value="105.65353">105°39'12.7"E</geoCoordinate>
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, 269 m elevation, coll. 13 June 2019 by Thy Neang; CBC 03004, adult female,
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<geoCoordinate degrees="12" direction="north" minutes="56" orientation="latitude" precision="1" seconds="09.7" value="12.936028">12°56'09.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="39" orientation="longitude" precision="1" seconds="14.4" value="105.654">105°39'14.4"E</geoCoordinate>
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, 271 m elevation, coll. 13 June 2019 by Thy Neang; CBC 03013, adult female, same data as holotype; CBC 03014, adult female, same data as holotype except
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<geoCoordinate degrees="12" direction="north" minutes="56" orientation="latitude" precision="1" seconds="08.7" value="12.93575">12°56'08.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="39" orientation="longitude" precision="1" seconds="12.6" value="105.6535">105°39'12.6"E</geoCoordinate>
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, 284 m elevation.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="etymology">
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<paragraph pageId="0" pageNumber="133">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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The specific epithet is taken from the type locality of Phnom Chi and the Latin suffix -
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<emphasis italics="true" pageId="0" pageNumber="133">ensis</emphasis>
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meaning "originating from." The specific epithet is masculine in agreement with the gender of
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<taxonomicName authorityName="Gray" authorityYear="1827" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="diagnosis">
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<paragraph pageId="0" pageNumber="133">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
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<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
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</taxonomicName>
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sp. nov. is distinguished from the 19 other named species in the
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<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
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<emphasis italics="true" pageId="0" pageNumber="133">C. irregularis</emphasis>
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</taxonomicName>
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group (
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2504.1.4" author="Ngo, TV" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="47 - 60" refId="B20" refString="Ngo, TV, Chan, OK, 2010. A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam. Zootaxa 2504 (1): 47 - 60, DOI: https://doi.org/10.11646/zootaxa.2504.1.4" title="A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam." url="https://doi.org/10.11646/zootaxa.2504.1.4" volume="2504" year="2010">Ngo and Chan 2010</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3737.4.4" author="Nguyen, SN" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="399 - 414" refId="B21" refString="Nguyen, SN, Le, TT, Tran, TAD, Orlov, NL, Lathrop, A, MacCulloch, RD, Le, TT, Jin, J, Nguyen, LT, Nguyen, TT, Hoang, DD, Che, J, Murphy, RW, Zhang, Y, 2013. Phylogeny of the Cyrtodactylus irregularis species complex (Squamata: Gekkonidae) from Vietnam with the description of two new species. Zootaxa 3737 (4): 399 - 414, DOI: https://doi.org/10.11646/zootaxa.3737.4.4" title="Phylogeny of the Cyrtodactylus irregularis species complex (Squamata: Gekkonidae) from Vietnam with the description of two new species." url="https://doi.org/10.11646/zootaxa.3737.4.4" volume="3737" year="2013">Nguyen et al. 2013</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.30906/1026-2296-2019-24-2-133-154" author="Nguyen, SN" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="133" pagination="133 - 154" refId="B23" refString="Nguyen, SN, Zhou, WW, Le, TT, Tran, TA, Jin, J, Vo, BD, Nguyen, LT, Nguyen, TT, Nguyen, TQ, Hoang, DD, Orlov, NL, Che, J, Murphy, RW, Zhang, YP, 2017. Cytonuclear discordance, cryptic diversity, complex histories, and conservation needs in Vietnamese bent-toed geckos of the Cyrtodactylus irregularis species complex. Russian Journal of Herpetology 24 (2): 133 - 154, DOI: https://doi.org/10.30906/1026-2296-2019-24-2-133-154" title="Cytonuclear discordance, cryptic diversity, complex histories, and conservation needs in Vietnamese bent-toed geckos of the Cyrtodactylus irregularis species complex." url="https://doi.org/10.30906/1026-2296-2019-24-2-133-154" volume="24" year="2017">2017</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4403.2.5" author="Pauwels, OSG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="307 - 335" refId="B25" refString="Pauwels, OSG, Nazarov, RA, Bobrov, VV, Poyarkov, NA, 2018. Taxonomic status of two populations of Bent-toed Geckos of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam. Zootaxa 4403 (2): 307 - 335, DOI: https://doi.org/10.11646/zootaxa.4403.2.5" title="Taxonomic status of two populations of Bent-toed Geckos of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam." url="https://doi.org/10.11646/zootaxa.4403.2.5" volume="4403" year="2018">Pauwels et al. 2018</bibRefCitation>
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) by having the combination of SVL 76.1-80.7 mm; relatively long body, AG/SVL 0.451-0.481; relatively long tibia, CrusL/SVL 0.172-0.200; subdigital lamellae on fourth finger 18-20; subdigital lamellae on fourth toe 20-23; longitudinal dorsal rows of tubercles 18-20; paravertebral rows of tubercles 31-36; ventral scales 45-54; enlarged femoral scales 0-8, without pores; enlarged precloacal scales 7-10, bearing pores 4 or 5 in males, pits 1-7 in females; precloacal groove absent; median row of transverse subcaudal scales only slightly enlarged; posterior border of nuchal loop unbroken and pointed, bordered anteriorly and posteriorly by broad yellow or yellowish white band; dark brown body bands 3 or 4, the first intact, the second, third and fourth more irregular, alternating with two or three yellowish white or light brown body bands, about half the width of dark brown body bands; and yellow spots on top of head.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="133" type="description">
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<paragraph pageId="0" pageNumber="133">Description of holotype.</paragraph>
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<paragraph pageId="0" pageNumber="133">
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Adult male with SVL 76.1 mm; head slightly elongate, HeadL 22.1 mm, about 30% of SVL, moderately widened, HeadW 14.1 mm, HeadW/HeadL 0.64, slightly depressed, HeadD 9.4 mm, HeadD/HeadL 0.43, distinct from neck, triangular in dorsal profile; snout rather elongated, rounded in rostral region, ESD 9.0 mm, slightly less than HeadD, ESD/HeadL 0.41, frontonasal region flattened, prefrontal region slightly concave, forming elongated medial rostral groove, canthus rostralis flattened, slightly angled between loreal region and rostral groove; lores posterior to nostrals depressed, anterior to orbit flattened; eye large, eyeball rounded, slightly protruding, EyeD 5.1 mm, shorter than the distance between eye and ear, Eye-EarD 5.7 mm, pupil vertical, covered by crenellate supraciliaries; ear opening oval, deeply sunk, rather small, elongated in oblique position, EarDV 1.2 mm, almost twice longer than its diameter in horizontal position, EarDH 0.7 mm; rostral large, subrectangular, height 1.9 mm, shorter than its width 3.6 mm, medially divided dorsally by a suture, reaching to about half way of rostral height, in contact with 1stSL and nostrils laterally, supranasals and internasal dorsally (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F5" captionText="Figure 4. Head of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative. A Dorsal view illustrating the rostral, supranasal and internsupranasal scales B ventral view illustrating the mental and postmental scales." figureDoi="10.3897/zookeys.926.48671.figure4" httpUri="https://binary.pensoft.net/fig/399861" pageId="0" pageNumber="133">4</figureCitation>
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); nostrils pieced at anterior angle of snout, directed lateroposteriorly, surrounded by rostral anteriorly, 1stSL ventrally, supranasals dorsally, and three small postnasal scales; internarial distance narrow, IND 2.9 mm; supranasals subrectangular, separated by intersupranasal, slightly smaller in size, in contact with rostral anteriorly, nostrals laterally, four small scales posteriorly; intersupranasal single, subpentagonal, slightly protruding rostral, in contact with two small scales posteriorly; interorbital rather narrow, IOD 5.5 mm, longer than EyeD, slightly shorter than Eye-EarD 5.7 mm; supralabials (12R, 13L), subrectangular anteriorly, circular shape posteriorly, anterior SL separated from small scales on loreal region by row of slightly enlarged scales; infralabials (9R, 9L), larger than SL, first InL bordered by mental anteriorly, first postmental ventrally, second InL bordered by second enlarged postmental, enlarged chin shield scale ventrally, 3-7thInL bordered by a row of slightly enlarged chin shield scales ventrally; mental large, triangular, width 3.3 mm in width, 2.3 mm in length, in contact with first InL laterally, two pairs postmentals posteriorly; the first pair largest, subrectangular, in broad contact medially, second pair enlarged, half the size of the first pair, separated by four smaller gular scales medially, in contact with smaller scales posteriorly (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F5" captionText="Figure 4. Head of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative. A Dorsal view illustrating the rostral, supranasal and internsupranasal scales B ventral view illustrating the mental and postmental scales." figureDoi="10.3897/zookeys.926.48671.figure4" httpUri="https://binary.pensoft.net/fig/399861" pageId="0" pageNumber="133">4</figureCitation>
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). Scales on frontonasal, prefrontal, loreal regions small, almost homogenous, slightly larger than those on top of head; scales on occiput intermixed with scattered larger, more rounded, conical tubercles, more prominent tubercles on region between orbit and area above ear opening, a noticeably larger tubercle (compared to those surrounding) at the corner of jaw.
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</paragraph>
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<caption doi="10.3897/zookeys.926.48671.figure4" httpUri="https://binary.pensoft.net/fig/399861" pageId="0" pageNumber="133" start="Figure 4" startId="F5">
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<paragraph pageId="0" pageNumber="133">
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<emphasis bold="true" pageId="0" pageNumber="133">Figure 4.</emphasis>
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Head of male holotype CBC 03012 of
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<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
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<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
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</taxonomicName>
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sp. nov. in preservative.
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<emphasis bold="true" pageId="0" pageNumber="133">A</emphasis>
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Dorsal view illustrating the rostral, supranasal and internsupranasal scales
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<emphasis bold="true" pageId="0" pageNumber="133">B</emphasis>
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ventral view illustrating the mental and postmental scales.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="133">
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Body slightly slender, AG 36.6 mm, nearly half SVL, AG/SVL 0.481 with well-defined narrow vertebral furrow posteriorly; scales on dorsum small, mostly homogenous, granular, interspersed with larger, low, weakly keeled, irregularly arranged, tubercles; longitudinal dorsal rows of enlarged tubercles approximately 18; paravertebral tubercles 32; tubercles on nape within dark brown nuchal loop, anterior dorsal surface at level above shoulder smaller, more rounded, sparser than those on mid-dorsum and posterior dorsal surface, more prominent, being denser, weekly keeled, more regularly arranged on sacral and tail base region; tubercles on lateral body sparsely; ventral scales small, not imbricate, those near midline larger than lateral and dorsal scales; scales on throat and gular region the smallest; faint ventrolateral folds with few emerged tubercles; ventral scales at midbody between ventrolateral folds 47; precloacal region moderately enlarged, a few rows of enlarged precloacal non-pore bearing scales anterior to pore bearing precloacal scales; enlarged precloacal scales 7, in angular series, bearing 5 pores, terminal scale on each side poreless; post precloacal scale rows 3, the first row immediately posterior to enlarged precloacal pore-bearing scales with six scales in angular series, the second row with four scales in angular series, the third row with three scale in straight line, the medial scale largest; femoral scales slightly enlarged (8R, 8L), distal scales more than twice the size of proximal scales, all smaller than those of pore-bearing precloacal scales, separated from precloacal scales by diastema; precloacal groove absent; fully everted hemipenes thick, 5.9 mm in length, two penes at each sheath, two sockets posterior to hemipenal bases (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F6" captionText="Figure 5. Cloacal region of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative illustrating the enlarged precloacal scales, enlarged femoral scales, and not enlarged subcaudal scales." figureDoi="10.3897/zookeys.926.48671.figure5" httpUri="https://binary.pensoft.net/fig/399862" pageId="0" pageNumber="133">5</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">Limbs rather slender; digits with strongly inflected interphalangeal joints; forelimbs bearing five relatively slender fingers, moderately bowed, ending with curved claws, ForeL/SVL 0.162; expanded proximal subdigital lamellae on fourth finger 6, unmodified distal subdigital lamellae on fourth finger 12, total subdigital lamellae on fourth finger 18; hind limbs bearing five relatively slender toes, strongly bent, ending with curved claws, CrusL/SVL 0.176; expanded proximal subdigital lamellae on fourth toe 7, unmodified distal subdigital lamellae on fourth toe 15, total subdigital lamellae on fourth toe 22; all digits lacking scansorial setae on ventral surface; scales on limbs small, interspersed with larger, low, conical, weakly keeled tubercles; scales on palmar and plantar surfaces small.</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
Tail moderately wide anteriorly, TaW 5.5 mm, segmented, cylindrical, becoming slender toward tip, regenerated posteriorly; dorsal caudal longitudinal tubercle rows at base of tail 8; 2 transverse rows of dorsal caudal tubercles at posterior margin of third band on tail, 22.7 mm from tail base; vertebral caudal surface with scattered bump at approximate intervals of 3 mm; subcaudal scale rows smooth, small, differing in size and irregular in shape, usually alternating between a single slightly enlarged and two smaller scales, 2 or 3 times larger than neighboring lateral caudal scales (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F6" captionText="Figure 5. Cloacal region of male holotype CBC 03012 of Cyrtodactylus phnomchiensis sp. nov. in preservative illustrating the enlarged precloacal scales, enlarged femoral scales, and not enlarged subcaudal scales." figureDoi="10.3897/zookeys.926.48671.figure5" httpUri="https://binary.pensoft.net/fig/399862" pageId="0" pageNumber="133">5</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.926.48671.figure5" httpUri="https://binary.pensoft.net/fig/399862" pageId="0" pageNumber="133" start="Figure 5" startId="F6">
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<emphasis bold="true" pageId="0" pageNumber="133">Figure 5.</emphasis>
|
|
Cloacal region of male holotype CBC 03012 of
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
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|
sp. nov. in preservative illustrating the enlarged precloacal scales, enlarged femoral scales, and not enlarged subcaudal scales.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="133" type="color of holotype in life">
|
|
<paragraph pageId="0" pageNumber="133">Color of holotype in life</paragraph>
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<paragraph pageId="0" pageNumber="133">
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|
(Fig.
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|
<figureCitation captionStart="Figure 3" captionStartId="F4" captionText="Figure 3. Cyrtodactylus phnomchiensis sp. nov. in life. A Male holotype CBC 03012 and B female paratype CBC 03013." figureDoi="10.3897/zookeys.926.48671.figure3" httpUri="https://binary.pensoft.net/fig/399860" pageId="0" pageNumber="133">3</figureCitation>
|
|
). Dorsal surface, nape, and tail yellowish white to light brown; top of head with yellowish spots; interorbital region, rostral and loreal regions lighter brown with scattered yellowish scales; eye ring yellowish; rostral, mental lighter brown; supralabials, corner of jaw, and region extending through dorsal margin of ear opening to shoulder yellowish; nuchal loop with large dark brown band, pointed, extending between posterior margins of eyes, bordered anteriorly by broad yellow band along upper edge of dark brown nuchal loop, posteriorly by yellowish white band; three dorsal dark brown bands on body, the first more regular, the second and third bands irregular, interrupted by white irregular blotches; all dark brown bands ending near to mid-flank region, bordered below by lighter brown extending to lateral folds; dark brown body bands bordered by yellowish white or light brown bands about half the width of dark brown body bands, the last light brown band ending on tail base; anterior and posterior margins of body bands with darker brown coloration; dark brown bands 6 on regenerated tail, margins at tail base darker; white bands on tail 5, nearly encircling the tail except subcaudal scale row; subcaudal scales lighter brown than dorsal caudal scales; limbs lighter brown with orangish or yellowish on enlarged tubercles; ventral surfaces between ventrolateral folds, chin, throat, and limbs white with tiny black dots on tip of scales; ventral surfaces of fingers and toes dark brown. In preservative, all yellowish, yellowish white or orange coloration faded to white, cream, or light brown (Fig.
|
|
<figureCitation captionStart="Figure 6" captionStartId="F7" captionText="Figure 6. Dorsal view of the type series of Cyrtodactylus phnomchiensis sp. nov. in preservative." figureDoi="10.3897/zookeys.926.48671.figure6" httpUri="https://binary.pensoft.net/fig/399863" pageId="0" pageNumber="133">6</figureCitation>
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).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="133" type="variation">
|
|
<paragraph pageId="0" pageNumber="133">Variations.</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
Morphometric and meristic characters of the type series are presented in Table
|
|
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Mensural, meristic and color pattern characters of Cyrtodactylus phnomchiensis sp. nov. Abbreviations defined in the text. All specimens have regenerated portions of tails (*)." httpUri="http://table.plazi.org/id/D7F1617D4F0BA973CEFA57CC1AF10FE3" pageId="0" pageNumber="133" tableUuid="D7F1617D4F0BA973CEFA57CC1AF10FE3">1</tableCitation>
|
|
. The paratypes generally resemble the holotype (Fig.
|
|
<figureCitation captionStart="Figure 6" captionStartId="F7" captionText="Figure 6. Dorsal view of the type series of Cyrtodactylus phnomchiensis sp. nov. in preservative." figureDoi="10.3897/zookeys.926.48671.figure6" httpUri="https://binary.pensoft.net/fig/399863" pageId="0" pageNumber="133">6</figureCitation>
|
|
), except as follows. CBC 03003 has four enlarged post precloacal scale rows in an angular series, the last row with only a single enlarged scale. CBC 03004 and CBC 03014 have darker brown body bands. CBC 03003-03004 have a more pointed nuchal loop. CBC 03003 has more dense dark dots causing ventral surfaces to be darker brown. CBC 03003 has more slender, fully everted hemipenes. Females have enlarged precloacal scales with pits rather than pores.
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|
</paragraph>
|
|
<caption doi="10.3897/zookeys.926.48671.figure6" httpUri="https://binary.pensoft.net/fig/399863" pageId="0" pageNumber="133" start="Figure 6" startId="F7">
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|
<paragraph pageId="0" pageNumber="133">
|
|
<emphasis bold="true" pageId="0" pageNumber="133">Figure 6.</emphasis>
|
|
Dorsal view of the type series of
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in preservative.
|
|
</paragraph>
|
|
</caption>
|
|
<caption ID-Table-UUID="D7F1617D4F0BA973CEFA57CC1AF10FE3" httpUri="http://table.plazi.org/id/D7F1617D4F0BA973CEFA57CC1AF10FE3" pageId="0" pageNumber="133" start="Table 1" startId="T1">
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<emphasis bold="true" pageId="0" pageNumber="133">Table 1.</emphasis>
|
|
Mensural, meristic and color pattern characters of
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. Abbreviations defined in the text. All specimens have regenerated portions of tails (*).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<table pageId="0" pageNumber="133">
|
|
<tr pageId="0" pageNumber="133">
|
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<th colspan="1" pageId="0" pageNumber="133" rowspan="1">Voucher specimen</th>
|
|
<th colspan="1" pageId="0" pageNumber="133" rowspan="1">CBC 03012</th>
|
|
<th colspan="1" pageId="0" pageNumber="133" rowspan="1">CBC 03003</th>
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<th colspan="1" pageId="0" pageNumber="133" rowspan="1">CBC 03004</th>
|
|
<th colspan="1" pageId="0" pageNumber="133" rowspan="1">CBC 03013</th>
|
|
<th colspan="1" pageId="0" pageNumber="133" rowspan="1">CBC 03014</th>
|
|
<th colspan="1" pageId="0" pageNumber="133" rowspan="1">Range</th>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Type status</td>
|
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<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Holotype</td>
|
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<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Paratype</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Paratype</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Paratype</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Paratype</td>
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="133" rowspan="1">-</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Sex</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Female</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Female</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Female</td>
|
|
<td colspan="1" isEmpty="true" pageId="0" pageNumber="133" rowspan="1">-</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">76.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">79.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">77.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">80.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">76.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">76.1-80.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">TaL*</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">75.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">56.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">66.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">79.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">64.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">56.9-79.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">TaW</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.4-5.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">TaW/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.072</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.056</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.069</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.069</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.074</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.056-0.74</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">ForeL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">12.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">11.7-13.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">ForeL/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.162</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.167</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.157</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.170</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.153</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.153-0.170</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">CrusL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">14.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">16.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">14.2-16.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Crus/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.176</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.189</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.179</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.200</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.172</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.172-0.200</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">AG</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">36.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">36.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">35.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">36.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">35.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">35.3-36.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">AG/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.481</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.457</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.457</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.451</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.462</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.451-0.481</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">HeadL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">22.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">23.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">22.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">23.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">23.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">22.1-23.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">HeadL/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.290</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.297</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.287</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.292</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.305</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.287-0.305</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">HeadW</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">14.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">14.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">15.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">13.7-15.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">HeadD</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">8.6-9.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EyeD (eye diameter)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.5-5.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EyeD/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.067</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.065</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.062</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.059</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.059</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.059-0.067</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Ear-EyeD (eye-ear distance)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.7-6.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">ESD (eye-snout distance)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9.0-9.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">ESD/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.118</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.120</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.116</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.123</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.121</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.116-0.123</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">END (eye-nostrial distance)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">6.4-7.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">IO (interorbital distance)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4.8-5.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">IND (internarial distance)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2.6-2.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EarDV (vertical)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.2-1.3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EarDH (horizontal)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0.6-1.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Intersupranasal scales</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Supralabials (SL)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">12R/13L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">11R/11L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">11R/11L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">12R/12L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">12R/13L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">11-13</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Infralabials (InL)</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9R/9L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">10R/9L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">10R/10L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">10R/9L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">10R/10L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">8-10</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">PVT</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">32</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">31</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">36</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">34</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">32</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">31-36</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">LDRT</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">18</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">19</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">18-20</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">VS</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">47</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">47</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">52</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">45</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">54</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">45-54</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Median subcaudal scales slightly enlarged</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">SDLF4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">18</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">18</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">19</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">19</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">18-20</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">SDLT4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">22</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">23</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">21</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">21</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">20-23</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EFS</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">8R8L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3R/3L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7R/6L</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0-8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">FP</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">EPrecS</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">10</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">9</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7-10</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">PrecP</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">7</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">1-7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">PrecG</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">PostPSR</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3-4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">PostSP</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3-4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Number of dark brown body bands</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3-4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Femoral and precloacal scales continuous</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">no</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Yellowish spots on top of head</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Posterior border of nuchal loop pointed</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">First body band complete</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Second to fourth body bands more irregular</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Yellowish white or light brown bands about half the width of dark brown body bands</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Number of yellowish white or light brown body bands</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">2-3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Yellowish spot above ear opening</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="133">
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Enlarged tubercle at corner of jaw</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="133" rowspan="1">yes</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="133" type="distribution">
|
|
<paragraph pageId="0" pageNumber="133">Distribution and natural history.</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
The new species is known only from the type locality at Phnom Chi in Prey Lang Wildlife Sanctuary, Kampong Thom Province, Sandan District, Cambodia. All individuals were found at night between 2001-2147 hr in evergreen-large dipterocarp dominated forest associated with rocky terrain (Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F8" captionText="Figure 7. Habitat at Phnom Chi, the type locality of Cyrtodactylus phnomchiensis sp. nov." figureDoi="10.3897/zookeys.926.48671.figure7" httpUri="https://binary.pensoft.net/fig/399864" pageId="0" pageNumber="133">7</figureCitation>
|
|
). The holotype CBC 03012 was found on a rock face following evening rain, paratypes CBC 03013-14 were on boulders following evening rain, paratype CBC 03003 was on leaf litter along a forest trail, and paratype CBC 03004 was on a rock wall at the entrance to a cave. Only five individuals were found during five-survey nights, suggesting the species is relatively uncommon. None were encountered during a brief survey by NT in the wet season of 2014 (
|
|
<bibRefCitation author="Hayes, B" journalOrPublisher="Conservation International, Winrock International, USAID, Phnom Penh, Cambodia" pageId="0" pageNumber="133" refId="B7" refString="Hayes, B, Eang, HK, Neang, T, Furey, N, Chhin, S, Holden, J, Hun, S, Phen, S, La, P, Simpson, V, 2015. Biodiversity assessment of Prey Lang: Kratie, Kampong Thom, Stung Treng and Preah Vihear Provinces. Conservation International, Winrock International, USAID, Phnom Penh, Cambodia" title="Biodiversity assessment of Prey Lang: Kratie, Kampong Thom, Stung Treng and Preah Vihear Provinces." year="2015">Hayes et al. 2015</bibRefCitation>
|
|
). The new species is the only member of the C.
|
|
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1921" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="irregularis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
complex known to occur west of the Mekong River (
|
|
<bibRefCitation DOI="https://doi.org/10.30906/1026-2296-2019-24-2-133-154" author="Nguyen, SN" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="133" pagination="133 - 154" refId="B23" refString="Nguyen, SN, Zhou, WW, Le, TT, Tran, TA, Jin, J, Vo, BD, Nguyen, LT, Nguyen, TT, Nguyen, TQ, Hoang, DD, Orlov, NL, Che, J, Murphy, RW, Zhang, YP, 2017. Cytonuclear discordance, cryptic diversity, complex histories, and conservation needs in Vietnamese bent-toed geckos of the Cyrtodactylus irregularis species complex. Russian Journal of Herpetology 24 (2): 133 - 154, DOI: https://doi.org/10.30906/1026-2296-2019-24-2-133-154" title="Cytonuclear discordance, cryptic diversity, complex histories, and conservation needs in Vietnamese bent-toed geckos of the Cyrtodactylus irregularis species complex." url="https://doi.org/10.30906/1026-2296-2019-24-2-133-154" volume="24" year="2017">Nguyen et al. 2017</bibRefCitation>
|
|
;
|
|
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4403.2.5" author="Pauwels, OSG" journalOrPublisher="Zootaxa" pageId="0" pageNumber="133" pagination="307 - 335" refId="B25" refString="Pauwels, OSG, Nazarov, RA, Bobrov, VV, Poyarkov, NA, 2018. Taxonomic status of two populations of Bent-toed Geckos of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam. Zootaxa 4403 (2): 307 - 335, DOI: https://doi.org/10.11646/zootaxa.4403.2.5" title="Taxonomic status of two populations of Bent-toed Geckos of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam." url="https://doi.org/10.11646/zootaxa.4403.2.5" volume="4403" year="2018">Pauwels et al. 2018</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.926.48671.figure7" httpUri="https://binary.pensoft.net/fig/399864" pageId="0" pageNumber="133" start="Figure 7" startId="F8">
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<emphasis bold="true" pageId="0" pageNumber="133">Figure 7.</emphasis>
|
|
Habitat at Phnom Chi, the type locality of
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="133" type="comparisons">
|
|
<paragraph pageId="0" pageNumber="133">Comparisons.</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is distinguishable from all 19 other members of the
|
|
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. irregularis</emphasis>
|
|
</taxonomicName>
|
|
group by a unique combination of morphological characters (and in mitochondrial DNA; Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Upper (A) and lower (B) portions of a fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 717 aligned characters of the mitochondrial cytochrome c oxidase subunit I (COI) gene from geckos in the Cyrtodactylus irregularis group. The outgroup Hemidactylus frenatus (GenBank accession GQ 245970) and exemplars of other Cyrtodactylus clades including C. auribalteatus (GenBank accession AP 018116), C. badenensis (KF 929505), C. chanhomeae (MF 169908), C. interdigitalis (MF 169919), C. intermedius (MF 169920), C. jellesmae (MF 169923), C. peguensis (AP 018114), C. russelli (MF 169938), and C. thirakhupti (AP 018115) were also included in the analysis (not shown). Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, and open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95. Numbers at terminal tips are GenBank accession numbers." figureDoi="10.3897/zookeys.926.48671.figure2" httpUri="https://binary.pensoft.net/fig/399858" pageId="0" pageNumber="133">2</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012 by having ventral scales 45-54 (vs. 38-43 in
|
|
<taxonomicName lsidName="bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pits in females 1-7 (vs. absent in
|
|
<taxonomicName lsidName="bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
), dark brown body bands larger than yellowish white or light brown dorsal bands (vs. dark brown bands, when present, narrower than light yellow dorsal bands in
|
|
<taxonomicName lsidName="bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
), and distinct large yellow band on anterior margin of dark brown nuchal loop (vs. narrow light margin in
|
|
<taxonomicName lsidName="bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
), and yellow spots on top of head (vs. dark spots in
|
|
<taxonomicName lsidName="bidoupimontis" pageId="0" pageNumber="133" rank="species" species="bidoupimontis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bidoupimontis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. buchardi</emphasis>
|
|
</taxonomicName>
|
|
by having SVL 76.1-80.7 mm (vs. 60-65 mm in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 14 in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), SDLT4 20-23 (vs. 12-14 in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 30 in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), LDRT 18-20 (vs. 25 in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 9 in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
), and irregular dorsal body bands (vs. blotches in
|
|
<taxonomicName lsidName="buchardi" pageId="0" pageNumber="133" rank="species" species="buchardi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">buchardi</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler, 2012 by having LDRT 18-20 (vs. 20-24 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 36-46 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 7-11 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 15-17 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLT4 20-23 (vs. 17-20 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), CrusL/SVL in adult specimens 0.172-0.200 (vs. 0.144-0.157 in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), large nuchal loop bordered anteriorly and posteriorly by broad yellow bands (vs. narrow nuchal loop bordered by distinct narrow white lines in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), dark brown body bands 3-4 (vs. seven highly irregular dark blotches with light margins in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
), and top of head with yellowish spots (vs. distinct dark brown spots in
|
|
<taxonomicName lsidName="bugiamapensis" pageId="0" pageNumber="133" rank="species" species="bugiamapensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">bugiamapensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. caovansungi</emphasis>
|
|
</taxonomicName>
|
|
Orlov, Nguyen, Nazarov, Ananjeva & Nguyen, 2007 by having SVL 76.1-80.7 mm (vs. 90.4-94.0 mm in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 38-44 in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
), femoral pores absent (vs. 6 in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 9 in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 22 in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
), and enlarged subcaudals absent (vs. present in
|
|
<taxonomicName lsidName="caovansungi" pageId="0" pageNumber="133" rank="species" species="caovansungi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">caovansungi</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. cattienensis" pageId="0" pageNumber="133" rank="species" species="cattienensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. cattienensis</emphasis>
|
|
</taxonomicName>
|
|
Geissler, Nazarov, Orlov,
|
|
<normalizedToken originalValue="Böhme">Boehme</normalizedToken>
|
|
, Phung, Nguyen & Ziegler, 2009 by having SVL 76.1-80.7 mm (vs. 69.0 mm maximum in
|
|
<taxonomicName lsidName="cattienensis" pageId="0" pageNumber="133" rank="species" species="cattienensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cattienensis</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 28-42 in
|
|
<taxonomicName lsidName="cattienensis" pageId="0" pageNumber="133" rank="species" species="cattienensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cattienensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 12-16 in
|
|
<taxonomicName lsidName="cattienensis" pageId="0" pageNumber="133" rank="species" species="cattienensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cattienensis</emphasis>
|
|
</taxonomicName>
|
|
), and SDLT4 20-23 (vs. 14-19 in
|
|
<taxonomicName lsidName="cattienensis" pageId="0" pageNumber="133" rank="species" species="cattienensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cattienensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. cucdongensis" pageId="0" pageNumber="133" rank="species" species="cucdongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. cucdongensis</emphasis>
|
|
</taxonomicName>
|
|
Schneider, Phung, Le, Nguyen & Ziegler, 2014 by having SVL 76.1-80.7 mm (vs. 55.8-65.9 mm in
|
|
<taxonomicName lsidName="cucdongensis" pageId="0" pageNumber="133" rank="species" species="cucdongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cucdongensis</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 35-44 in
|
|
<taxonomicName lsidName="cucdongensis" pageId="0" pageNumber="133" rank="species" species="cucdongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cucdongensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 13-18 in
|
|
<taxonomicName lsidName="cucdongensis" pageId="0" pageNumber="133" rank="species" species="cucdongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cucdongensis</emphasis>
|
|
</taxonomicName>
|
|
), and SDLT4 20-23 (vs. 15-20 in
|
|
<taxonomicName lsidName="cucdongensis" pageId="0" pageNumber="133" rank="species" species="cucdongensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cucdongensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. cryptus" pageId="0" pageNumber="133" rank="species" species="cryptus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. cryptus</emphasis>
|
|
</taxonomicName>
|
|
Heidrich,
|
|
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
|
, Vu,
|
|
<normalizedToken originalValue="Böhme">Boehme</normalizedToken>
|
|
& Ziegler, 2007 by having precloacal pores in males 4-5 (vs. 9-11 in
|
|
<taxonomicName lsidName="cryptus" pageId="0" pageNumber="133" rank="species" species="cryptus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cryptus</emphasis>
|
|
</taxonomicName>
|
|
), and precloacal pits in females 1-7 (vs. absent in
|
|
<taxonomicName lsidName="cryptus" pageId="0" pageNumber="133" rank="species" species="cryptus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">cryptus</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. dati</emphasis>
|
|
</taxonomicName>
|
|
Ngo, 2013 by having SVL 76.1-80.7 mm (vs. 70.1 mm maximum in
|
|
<taxonomicName lsidName="dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">dati</emphasis>
|
|
</taxonomicName>
|
|
), regenerated TaL 56.9-79.1 mm vs. (vs. 50.3 mm maximum, non-regenerated TaL in
|
|
<taxonomicName lsidName="dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">dati</emphasis>
|
|
</taxonomicName>
|
|
), femoral pores in both sexes absent (vs. present in
|
|
<taxonomicName lsidName="dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">dati</emphasis>
|
|
</taxonomicName>
|
|
), nuchal loop continuous (vs. broken in
|
|
<taxonomicName lsidName="dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">dati</emphasis>
|
|
</taxonomicName>
|
|
), and dark brown body bands (vs. irregular dark brown blotches on body in
|
|
<taxonomicName lsidName="dati" pageId="0" pageNumber="133" rank="species" species="dati">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">dati</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. gialaiensis" pageId="0" pageNumber="133" rank="species" species="gialaiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. gialaiensis</emphasis>
|
|
</taxonomicName>
|
|
Luu, Dung, Nguyen, Le & Ziegler, 2017 by having SVL 76.1-80.7 mm (vs. 62.8 mm maximum in
|
|
<taxonomicName lsidName="gialaiensis" pageId="0" pageNumber="133" rank="species" species="gialaiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">gialaiensis</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 9-10 in
|
|
<taxonomicName lsidName="gialaiensis" pageId="0" pageNumber="133" rank="species" species="gialaiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">gialaiensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 14-15 in
|
|
<taxonomicName lsidName="gialaiensis" pageId="0" pageNumber="133" rank="species" species="gialaiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">gialaiensis</emphasis>
|
|
</taxonomicName>
|
|
), and SDLT4 20-23 (vs. 15-17 in
|
|
<taxonomicName lsidName="gialaiensis" pageId="0" pageNumber="133" rank="species" species="gialaiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">gialaiensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. huynhi</emphasis>
|
|
</taxonomicName>
|
|
Ngo & Bauer, 2008 by having SDLF4 18-20 (vs. 14-17 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
), AGL/SVL 0.451-0.481 (vs. 0.370-0.428 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 43-46 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 7-9 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
), dark brown body bands 3-4 (vs. 5-6 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
); femoral pores in both sexes absent (vs. 3-8 in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
), and nuchal loop bordered anteriorly and posteriorly by broad yellow bands (vs. narrow cream margin in
|
|
<taxonomicName lsidName="huynhi" pageId="0" pageNumber="133" rank="species" species="huynhi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">huynhi</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. irregularis</emphasis>
|
|
</taxonomicName>
|
|
by lacking enlarged triangular tubercles at base of tail (vs. present in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), CrusL/SVL 0.172-0.200 (vs. 0.138-0.156 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), LDRT 18-20 (vs. 22-24 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), paravertebral tubercles 31-36 (vs. 38-48 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
); ventral scales 45-54 (vs. 38-45 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 15-16 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), SDLT4 20-23 (vs. 18-20 in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), dark brown body bands 3-4 (vs. 5-7, mostly as irregular blotches in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
), and yellowish spots on top of head (vs. distinct dark brown spots in
|
|
<taxonomicName lsidName="irregularis" pageId="0" pageNumber="133" rank="species" species="irregularis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">irregularis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. kingsadai" pageId="0" pageNumber="133" rank="species" species="kingsadai">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. kingsadai</emphasis>
|
|
</taxonomicName>
|
|
Ziegler, Phung, Le & Nguyen, 2013 by having SVL 76.1-80.7 mm (vs. 83.0-94.0 mm in
|
|
<taxonomicName lsidName="kingsadai" pageId="0" pageNumber="133" rank="species" species="kingsadai">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">kingsadai</emphasis>
|
|
</taxonomicName>
|
|
), enlarged femoral scales 0-8 (vs. 9-12 in
|
|
<taxonomicName lsidName="kingsadai" pageId="0" pageNumber="133" rank="species" species="kingsadai">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">kingsadai</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pore in males 4-5 (vs. 7-9 in
|
|
<taxonomicName lsidName="kingsadai" pageId="0" pageNumber="133" rank="species" species="kingsadai">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">kingsadai</emphasis>
|
|
</taxonomicName>
|
|
), and subcaudal scales not enlarged (vs. enlarged in
|
|
<taxonomicName lsidName="kingsadai" pageId="0" pageNumber="133" rank="species" species="kingsadai">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">kingsadai</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. phuocbinhensis" pageId="0" pageNumber="133" rank="species" species="phuocbinhensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. phuocbinhensis</emphasis>
|
|
</taxonomicName>
|
|
Nguyen, Le, Tran, Orlov, Lathrop, MacCulloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang, 2013 by having SVL 76.1-80.7 mm (vs. 46.0-60.4 in
|
|
<taxonomicName lsidName="phuocbinhensis" pageId="0" pageNumber="133" rank="species" species="phuocbinhensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">phuocbinhensis</emphasis>
|
|
</taxonomicName>
|
|
); precloacal pits in females 1-7 (vs. absent in
|
|
<taxonomicName lsidName="phuocbinhensis" pageId="0" pageNumber="133" rank="species" species="phuocbinhensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">phuocbinhensis</emphasis>
|
|
</taxonomicName>
|
|
), top of head with yellow spots (vs. dark brown spots in
|
|
<taxonomicName lsidName="phuocbinhensis" pageId="0" pageNumber="133" rank="species" species="phuocbinhensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">phuocbinhensis</emphasis>
|
|
</taxonomicName>
|
|
), and dark brown body bands (vs. two dark brown longitudinal stripes or blotches in
|
|
<taxonomicName lsidName="phuocbinhensis" pageId="0" pageNumber="133" rank="species" species="phuocbinhensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">phuocbinhensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. pseudoquadrivirgatus" pageId="0" pageNumber="133" rank="species" species="pseudoquadrivirgatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. pseudoquadrivirgatus</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Rösler">Roesler</normalizedToken>
|
|
, Vu, Nguyen, Ngo & Ziegler, 2008 by having yellow spots on top of head (vs. dark blotches on top of head in
|
|
<taxonomicName lsidName="pseudoquadrivirgatus" pageId="0" pageNumber="133" rank="species" species="pseudoquadrivirgatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">pseudoquadrivirgatus</emphasis>
|
|
</taxonomicName>
|
|
) and dark brown body bands (vs. highly irregular body blotches in
|
|
<taxonomicName lsidName="pseudoquadrivirgatus" pageId="0" pageNumber="133" rank="species" species="pseudoquadrivirgatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">pseudoquadrivirgatus</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. sangi</emphasis>
|
|
</taxonomicName>
|
|
Pauwels, Nazarov, Bobrov & Poyarkov, 2018 by having SVL 76.1-80.7 mm (vs. 56.3 mm maximum in
|
|
<taxonomicName lsidName="sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">sangi</emphasis>
|
|
</taxonomicName>
|
|
), paravertebral tubercles 31-36 (vs. 27-29 in
|
|
<taxonomicName lsidName="sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">sangi</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 37 in
|
|
<taxonomicName lsidName="sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">sangi</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pores in males 4-5 (vs. 7 in
|
|
<taxonomicName lsidName="sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">sangi</emphasis>
|
|
</taxonomicName>
|
|
), and first dark brown body band complete, second, third, and fourth more irregular (vs. highly irregular bands in
|
|
<taxonomicName lsidName="sangi" pageId="0" pageNumber="133" rank="species" species="sangi">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">sangi</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. takouensis</emphasis>
|
|
</taxonomicName>
|
|
Ngo & Bauer, 2008 by having LDRT 18-20 (vs. 9-10 smooth tubercles in
|
|
<taxonomicName lsidName="takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">takouensis</emphasis>
|
|
</taxonomicName>
|
|
); ventral scales 45-54 (vs. 39-40 in
|
|
<taxonomicName lsidName="takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">takouensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF4 18-20 (vs. 16-17 in
|
|
<taxonomicName lsidName="takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">takouensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLT4 20-23 (vs. 18-20 in
|
|
<taxonomicName lsidName="takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">takouensis</emphasis>
|
|
</taxonomicName>
|
|
), and dark brown canthal stripe absent (vs. present in
|
|
<taxonomicName lsidName="takouensis" pageId="0" pageNumber="133" rank="species" species="takouensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">takouensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
Nguyen, Le, Tran, Orlov, Lathrop, MacCulloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang, 2013 by having supralabials 11-13 (vs. 8-9 in
|
|
<taxonomicName lsidName="taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
), precloacal pits in females present (vs. absent in
|
|
<taxonomicName lsidName="taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
), SDLF 18-20 (vs. 13-18 in
|
|
<taxonomicName lsidName="taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
), top of head with yellow spots (vs. dark brown blotches in
|
|
<taxonomicName lsidName="taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
), and dark brown body bands (vs. black irregular blotches margined by light brown in
|
|
<taxonomicName lsidName="taynguyenensis" pageId="0" pageNumber="133" rank="species" species="taynguyenensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">taynguyenensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="C. yangbayensis" pageId="0" pageNumber="133" rank="species" species="yangbayensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. yangbayensis</emphasis>
|
|
</taxonomicName>
|
|
Ngo & Chan, 2010 by having SDLT4 20-23 (vs. 15-17 in
|
|
<taxonomicName lsidName="yangbayensis" pageId="0" pageNumber="133" rank="species" species="yangbayensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">yangbayensis</emphasis>
|
|
</taxonomicName>
|
|
) and lacking enlarged subcaudal scales (vs. present in
|
|
<taxonomicName lsidName="yangbayensis" pageId="0" pageNumber="133" rank="species" species="yangbayensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">yangbayensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="133">
|
|
<taxonomicName authorityName="Neang & Henson & Stuart" authorityYear="2020" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtodactylus phnomchiensis" order="Squamata" pageId="0" pageNumber="133" phylum="Chordata" rank="species" species="phnomchiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">Cyrtodactylus phnomchiensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most closely related in mitochondrial DNA to
|
|
<taxonomicName lsidName="C. ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. ziegleri</emphasis>
|
|
</taxonomicName>
|
|
Nazarov, Orlov, Nguyen & Ho, 2008 (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Upper (A) and lower (B) portions of a fifty percent majority-rule consensus phylogram resulting from partitioned Bayesian analysis of 717 aligned characters of the mitochondrial cytochrome c oxidase subunit I (COI) gene from geckos in the Cyrtodactylus irregularis group. The outgroup Hemidactylus frenatus (GenBank accession GQ 245970) and exemplars of other Cyrtodactylus clades including C. auribalteatus (GenBank accession AP 018116), C. badenensis (KF 929505), C. chanhomeae (MF 169908), C. interdigitalis (MF 169919), C. intermedius (MF 169920), C. jellesmae (MF 169923), C. peguensis (AP 018114), C. russelli (MF 169938), and C. thirakhupti (AP 018115) were also included in the analysis (not shown). Black circles at nodes indicate Bayesian posterior probabilities ≥ 0.99, and open circles at nodes indicate Bayesian posterior probabilities ≥ 0.95. Numbers at terminal tips are GenBank accession numbers." figureDoi="10.3897/zookeys.926.48671.figure2" httpUri="https://binary.pensoft.net/fig/399858" pageId="0" pageNumber="133">2</figureCitation>
|
|
), but differs in morphology from
|
|
<taxonomicName lsidName="C. ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">C. ziegleri</emphasis>
|
|
</taxonomicName>
|
|
by having SVL 76.1-80.7 mm (vs. 84.6-93.0 mm in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), paravertebral tubercles 31-36 (vs. 38-46 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), ventral scales 45-54 (vs. 33-45 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), HeadL/SVL 0.287-0.305 (vs. 0.263-0.284 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), ESD/SVL 0.116-0.123 (vs. 0.103-0.113 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), CrusL/SVL 0.172-0.200 (vs. 0.140-0.168 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), AG/SVL 0.451-0.481 (vs. 0.390-0.444 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), eyeD/SVL 0.059-0.067 (vs. 0.053-0.057 in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), top of head with yellow spots (vs. dark brown spots in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), large dark brown nuchal loop (vs. narrow in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), distinct, broad yellow band on anterior margin of dark brown nuchal loop (vs. absent in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
), and dark brown body bands bordered by yellowish white or light brown bands about half the width of dark brown bands (vs. light yellow or light brown body bands about same width as dark brown body bands in
|
|
<taxonomicName lsidName="ziegleri" pageId="0" pageNumber="133" rank="species" species="ziegleri">
|
|
<emphasis italics="true" pageId="0" pageNumber="133">ziegleri</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |