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<document id="13919D74E429BDAC55B01A0DE2176F6E" ID-CLB-Dataset="39115" ID-DOI="10.11646/zootaxa.4109.2.3" ID-GBIF-Dataset="60a1e58f-6277-414a-a302-7fd2a6b2da1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="271419" ID-ZooBank="3DF4E74C-F422-48AD-AFEE-F3B0FEA8F443" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462515728922" checkinUser="plazi" docAuthor="Mens, Lotte P., Schütte, Kai, Stokvis, Frank R. &amp; Dijkstra, Klaas-Douwe B." docDate="2016" docId="244B882F23247621FF7125E5FD75A717" docLanguage="en" docName="zootaxa.4109.2.3.pdf.imf" docOrigin="Zootaxa 4109 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acisoma attenboroughi Mens, Schütte, Stokvis &amp; Dijkstra, 2016, sp. nov." docType="treatment" docVersion="11" lastPageNumber="166" masterDocId="D872F057232E762CFFE62149FFCFA027" masterDocTitle="Six, not two, species of Acisoma pintail dragonfly (Odonata: Libellulidae)" masterLastPageNumber="172" masterPageNumber="153" pageNumber="163" updateTime="1698662321709" updateUser="ExternalLinkService">
<mods:mods id="CE664E36F3CFEA01A0C87F9550CC129A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="85B51A3727B5655426A3451EA02341F5">
<mods:title id="714F0F26B18833298C6515B8DC2795B7">Six, not two, species of Acisoma pintail dragonfly (Odonata: Libellulidae)</mods:title>
</mods:titleInfo>
<mods:name id="8AECE1F5CB44631F6FB7ED07927C9EDC" type="personal">
<mods:role id="EED27394118FD2C3801A88E522CBFE7E">
<mods:roleTerm id="571A669576027BBD380BDBC055A73CF0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3E938F85AE2DB45A51DC2FB291B8AB7">Mens, Lotte P.</mods:namePart>
</mods:name>
<mods:name id="864FD7635EEDEC12C53DAAD3CB2705B4" type="personal">
<mods:role id="AEB8297CBD52603EBEB9B782ABD35505">
<mods:roleTerm id="5317A941EAF544C39FC581B169AB6E7E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C34B17F4788FC8C619F726D1CC020C3">Schütte, Kai</mods:namePart>
</mods:name>
<mods:name id="4CC8D1C54E2E8D4DA27F512A2D8B8D54" type="personal">
<mods:role id="604E2083AEA08C8EE8F5A48B76461626">
<mods:roleTerm id="72BC362A1D14979F4A721D07E69A628E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD823674745A2566FE0C5E8E42BD07A0">Stokvis, Frank R.</mods:namePart>
</mods:name>
<mods:name id="D9840F37FAE9AD38BBC224CEE04F103F" type="personal">
<mods:role id="80ECDE5F38DF9235C7D3EBC274A4198A">
<mods:roleTerm id="3F0F42FED21EFEC930DFFCA1213FF85D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7BE37834516E05E16FBFD60ECA6E50E">Dijkstra, Klaas-Douwe B.</mods:namePart>
</mods:name>
<mods:typeOfResource id="32BEF58643D87F56C8DFE74CBEEAFAAF">text</mods:typeOfResource>
<mods:relatedItem id="433A4DCB9A92C3D123D29789CA6F6F5C" type="host">
<mods:titleInfo id="B677D1533B8513E7EA12026902E04B84">
<mods:title id="F01622AD1612DC52B2C970E6E55C8E1A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7DA4C8D8D38340AFADCA15BA5E389C24">
<mods:date id="7658E768719224CDB4BAB5E709FB921E">2016</mods:date>
<mods:detail id="2F2851C5F7E4BCC20A53169568598E11" type="volume">
<mods:number id="A6A589AB32F715A7B7DC60AAD961516D">4109</mods:number>
</mods:detail>
<mods:detail id="4F475B4774399546B021B080661BA9FC" type="issue">
<mods:number id="4A06F39ED4464314BE679689674FE977">2</mods:number>
</mods:detail>
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<mods:start id="791EA04EB27A80774FF3BC3FF435D2A1">153</mods:start>
<mods:end id="BA7E3588E90E868B39B00C11C00D72FF">172</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6FD465808FE0D1BFDA8C31C55FFC1E07">journal article</mods:classification>
<mods:identifier id="6D9AF85A71B6721D98A1DCFFC1C7CD32" type="CLB-Dataset">39115</mods:identifier>
<mods:identifier id="E27593E4B716B3C9B86DE6BF32B0F691" type="DOI">10.11646/zootaxa.4109.2.3</mods:identifier>
<mods:identifier id="9AD7B88DCBD2A2CCA8002AB71AC043F4" type="GBIF-Dataset">60a1e58f-6277-414a-a302-7fd2a6b2da1a</mods:identifier>
<mods:identifier id="1945CEAE0B9384CB6F84389C57F7BE3B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="626F050761883956AEE28A7DAF4676B9" type="Zenodo-Dep">271419</mods:identifier>
<mods:identifier id="57883B9594C715BDFF93CB50EA5D60AF" type="ZooBank">3DF4E74C-F422-48AD-AFEE-F3B0FEA8F443</mods:identifier>
</mods:mods>
<treatment id="244B882F23247621FF7125E5FD75A717" ID-DOI="http://doi.org/10.5281/zenodo.5673007" ID-GBIF-Taxon="120674211" ID-Zenodo-Dep="5673007" LSID="urn:lsid:plazi:treatment:244B882F23247621FF7125E5FD75A717" httpUri="http://treatment.plazi.org/id/244B882F23247621FF7125E5FD75A717" lastPageId="13" lastPageNumber="166" pageId="10" pageNumber="163">
<subSubSection id="E4F86AB223247626FF7125E5FD7DA4C0" pageId="10" pageNumber="163" type="nomenclature">
<paragraph id="AC5D393923247626FF7125E5FC9EA4E1" blockId="10.[151,849,1196,1255]" box="[151,849,1196,1222]" pageId="10" pageNumber="163">
<heading id="F7158E5523247626FF7125E5FC9EA4E1" bold="true" box="[151,849,1196,1222]" fontSize="11" level="1" pageId="10" pageNumber="163" reason="1">
<emphasis id="9E96E52B23247626FF7125E5FC9EA4E1" bold="true" box="[151,849,1196,1222]" pageId="10" pageNumber="163">
<taxonomicName id="6BE242BA23247626FF7125E5FE7EA4E1" ID-CoL="9WT34" box="[151,433,1196,1222]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="9E96E52B23247626FF7125E5FE7EA4E1" bold="true" box="[151,433,1196,1222]" italics="true" pageId="10" pageNumber="163">Acisoma attenboroughi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="85A5585023247626FE5F25E5FDDAA4E1" box="[441,533,1196,1222]" pageId="10" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
—Attenborough’s Pintail
</emphasis>
</heading>
</paragraph>
<paragraph id="AC5D393923247626FF712587FD7DA4C0" blockId="10.[151,849,1196,1255]" box="[151,690,1230,1255]" pageId="10" pageNumber="163">
(
<figureCitation id="34D925BC23247626FF792587FF22A4C0" box="[159,237,1230,1255]" captionStart="FIGURE 4" captionStartId="7.[151,250,705,727]" captionTargetBox="[223,1345,203,654]" captionTargetId="figure@7.[216,1367,190,668]" captionTargetPageId="7" captionText="FIGURE 4. Head of Acisoma species in rostral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides (typical), D: A. panorpoides (pale), E: A. attenboroughi, F: A. ascalaphoides, G: A. trifidum." httpUri="https://zenodo.org/record/271423/files/figure.png" pageId="10" pageNumber="163">Figs. 4</figureCitation>
E, 5A–C, 5F–G, 6E–F, 7D, 8D, 10D, 11)
</paragraph>
</subSubSection>
<subSubSection id="E4F86AB223247626FF71245FFA49A668" pageId="10" pageNumber="163" type="materials_examined">
<paragraph id="AC5D393923247626FF71245FFAE8A508" blockId="10.[151,1437,1302,2014]" box="[151,1319,1302,1327]" pageId="10" pageNumber="163">
<emphasis id="9E96E52B23247626FF71245FFDE4A508" bold="true" box="[151,555,1302,1327]" pageId="10" pageNumber="163">
<typeStatus id="7359879B23247626FF71245FFF1DA508" box="[151,210,1302,1327]" pageId="10" pageNumber="163">Type</typeStatus>
material.
<typeStatus id="7359879B23247626FEAA245FFE76A508" box="[332,441,1302,1327]" pageId="10" pageNumber="163" type="holotype">Holotype</typeStatus>
(
<figureCitation id="34D925BC23247626FE21245FFDD4A508" box="[455,539,1302,1327]" captionStart="FIGURE 11" captionStartId="13.[151,250,1466,1488]" captionTargetBox="[150,1436,193,1444]" captionTargetId="figure@13.[150,1436,193,1445]" captionTargetPageId="13" captionText="FIGURE 11. Holotype of Acisoma attenboroughi sp. nov., ZMUH." httpUri="https://zenodo.org/record/271430/files/figure.png" pageId="10" pageNumber="163">Fig. 11</figureCitation>
):
</emphasis>
male, Andasibe, lake Toamasina,
<date id="D85C1FF923247626FC4C245FFBFEA508" box="[938,1073,1302,1327]" pageId="10" pageNumber="163" value="2006-12-12">12-xii-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FB24245EFAD4A508" box="[1218,1307,1303,1327]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
).
</paragraph>
<paragraph id="AC5D393923247626FF212473FA49A668" blockId="10.[151,1437,1302,2014]" pageId="10" pageNumber="163">
<emphasis id="9E96E52B23247626FF212473FE47A574" bold="true" box="[199,392,1338,1363]" pageId="10" pageNumber="163">Other material.</emphasis>
<collectingCountry id="D4F579A923247626FE702473FDEDA574" box="[406,546,1338,1363]" name="Madagascar" pageId="10" pageNumber="163">Madagascar</collectingCountry>
:
<specimenCount id="BAE4F2B023247626FDD52472FD47A574" box="[563,648,1338,1363]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Ankarafantsika National Park, Ampijoroa,
<date id="D85C1FF923247626FB7F2473FAD5A574" box="[1177,1306,1338,1363]" pageId="10" pageNumber="163" value="2001-03-06">06-iii-2001</date>
, G. Garcia (
<collectionCode id="CAF3A1FC23247626FF792416FF36A550" LSID="urn:lsid:biocol.org:col:34992" box="[159,249,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="163">RMNH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FEF72416FEACA550" box="[273,355,1374,1399]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Tananarive,
<date id="D85C1FF923247626FDE72417FD4DA550" box="[513,642,1374,1399]" pageId="10" pageNumber="163" value="1985-03-21">21-iii-1985</date>
, F. Keiser (
<collectionCode id="CAF3A1FC23247626FCE82416FCA7A550" LSID="urn:lsid:biocol.org:col:34992" box="[782,872,1375,1399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="10" pageNumber="163">RMNH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FC662416FC1DA550" box="[896,978,1374,1399]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, locality unknown, 1993, J.M. Elouard (
<collectionCode id="CAF3A1FC23247626FF7924CAFF34A5BC" LSID="urn:lsid:biocol.org:col:34988" box="[159,251,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="163">MNHN</collectionCode>
);
<specimenCount id="BAE4F2B023247626FEEB24CAFE94A5BC" box="[269,347,1410,1435]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
,
<collectingCountry id="D4F579A923247626FE8324CBFE21A5BC" box="[357,494,1410,1435]" name="Madagascar" pageId="10" pageNumber="163">Madagascar</collectingCountry>
, Coll. Martin (
<collectionCode id="CAF3A1FC23247626FD7724CAFD22A5BC" LSID="urn:lsid:biocol.org:col:34988" box="[657,749,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="163">MNHN</collectionCode>
);
<specimenCount id="BAE4F2B023247626FD1924CAFC97A5BC" box="[767,856,1410,1435]" pageId="10" pageNumber="163" type="male">7 males</specimenCount>
, Andasibe, lake Toamasina,
<date id="D85C1FF923247626FB7324CBFAD3A5BC" box="[1173,1308,1410,1435]" pageId="10" pageNumber="163" value="2006-12-12">12-xii-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FF7924EEFF37A598" box="[159,248,1447,1471]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FEF224EFFEB9A598" box="[276,374,1446,1471]" pageId="10" pageNumber="163" type="male">5 males</specimenCount>
, Andasibe, river Toamasina,
<date id="D85C1FF923247626FD3924EFFCA9A598" box="[735,870,1446,1471]" pageId="10" pageNumber="163" value="2006-12-12">12-xii-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FBF724EEFBA5A598" box="[1041,1130,1447,1471]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FB6124EEFB27A598" box="[1159,1256,1446,1471]" pageId="10" pageNumber="163" type="male">7 males</specimenCount>
, Fort-Dauphin, Amboavola,
<date id="D85C1FF923247626FECF2483FE67A5C4" box="[297,424,1482,1507]" pageId="10" pageNumber="163" value="2006-11-28">28-xi-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FDDD2482FD5BA5C4" box="[571,660,1483,1507]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FD4E2482FD37A5C4" box="[680,760,1482,1507]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Fort-Dauphin, Tsimelahy,
<quantity id="6B1A94DC23247626FBCB2483FBBBA5C4" box="[1069,1140,1482,1507]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="163" unit="km" value="1.5">1.5km</quantity>
de accueil,
<date id="D85C1FF923247626FB1B2483FABAA5C4" box="[1277,1397,1482,1507]" pageId="10" pageNumber="163" value="2006-10-01">01-x-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FEE324A6FE91A620" box="[261,350,1519,1543]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FE9A24A6FE1AA620" box="[380,469,1518,1543]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Fort-Dauphin, Enato,
<date id="D85C1FF923247626FD1124A7FCA0A620" box="[759,879,1518,1543]" pageId="10" pageNumber="163" value="2006-10-26">26-x-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FBC624A6FBB6A620" box="[1056,1145,1519,1543]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FB7124A6FB3FA620" box="[1175,1264,1518,1543]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Fianarantsoa, Ranomafana, ValBio field station next to river,
<date id="D85C1FF923247626FD2F275BFC87A60C" box="[713,840,1554,1579]" pageId="10" pageNumber="163" value="2006-11-01">01-xi-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FC01275AFB8FA60C" box="[999,1088,1555,1579]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FBB1275AFB7AA60C" box="[1111,1205,1554,1579]" pageId="10" pageNumber="163" type="male">3 males</specimenCount>
, Tolongoina, coffee plantation,
<date id="D85C1FF923247626FEF3277FFE5BA668" box="[277,404,1590,1615]" pageId="10" pageNumber="163" value="2006-11-10">10-xi-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FDC3277EFD4DA668" box="[549,642,1591,1615]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
);
<specimenCount id="BAE4F2B023247626FD68277EFD18A668" box="[654,727,1590,1615]" pageId="10" pageNumber="163" type="male">1 male</specimenCount>
, Tolongoina, tavy in forest,
<date id="D85C1FF923247626FBF4277FFB5EA668" box="[1042,1169,1590,1615]" pageId="10" pageNumber="163" value="2006-11-04">04-xi-2006</date>
, K. Schütte (
<collectionCode id="CAF3A1FC23247626FAC7277EFAB5A668" box="[1313,1402,1591,1615]" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="10" pageNumber="163">ZMUH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="E4F86AB223247626FF212714FC9BA6BD" pageId="10" pageNumber="163" type="etymology">
<paragraph id="AC5D393923247626FF212714FC9BA6BD" blockId="10.[151,1437,1302,2014]" pageId="10" pageNumber="163">
<emphasis id="9E96E52B23247626FF212714FE81A651" bold="true" box="[199,334,1629,1654]" pageId="10" pageNumber="163">Etymology.</emphasis>
The new species honours Sir David Attenborough on his 90th birthday. His documentaries inspired entire generations to love and conserve nature (
<bibRefCitation id="C87344C823247626FD4E27C8FC87A6BD" author="Dijkstra" box="[680,840,1665,1690]" pageId="10" pageNumber="163" refString="Dijkstra, K. D. B. (2016) Restore our sense of species. Nature, 533 (7602). [in press]" type="journal article" year="2016">Dijkstra 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E4F86AB223247621FF2127ECFD75A717" lastPageId="13" lastPageNumber="166" pageId="10" pageNumber="163" type="description">
<paragraph id="AC5D393923247621FF2127ECFE68A6E3" blockId="10.[151,1437,1302,2014]" lastBlockId="13.[151,1437,1564,1840]" lastPageId="13" lastPageNumber="166" pageId="10" pageNumber="163">
<emphasis id="9E96E52B23247626FF2127ECFE49A699" bold="true" box="[199,390,1701,1726]" pageId="10" pageNumber="163">Male diagnosis.</emphasis>
Confined to
<collectingCountry id="D4F579A923247626FDCF27ECFD7DA699" box="[553,690,1701,1726]" name="Madagascar" pageId="10" pageNumber="163">Madagascar</collectingCountry>
, where easily identified by the characters typical of the group lumped formerly under
<taxonomicName id="6BE242BA23247626FE452783FD80A6C6" box="[419,591,1737,1761]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23247626FE452783FD80A6C6" box="[419,591,1737,1761]" italics="true" pageId="10" pageNumber="163">A. panorpoides</emphasis>
</taxonomicName>
: (a) extensively white labrum, labium, thorax and underside of S3–7; (b) 1 cell in Fw triangle; (c) fairly long Fw Pt, about 11–14% as long as Hw; (d) abdomen that narrows abruptly on S
<quantity id="6B1A94DC23247626FA9427A4FA53A721" box="[1394,1436,1773,1798]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="163" unit="in" value="5.0">5 in</quantity>
lateral view, with S6–10 slender (
<figureCitation id="34D925BC23247626FDFA2658FDACA70D" box="[540,611,1809,1834]" captionStart="FIGURE 7. S 2 – 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 – 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="10" pageNumber="163">Fig. 7</figureCitation>
D); (e) fragmented and frayed white markings on S2–5 and large white lateral spots on S7 (
<figureCitation id="34D925BC23247626FE93267CFE77A769" box="[373,440,1845,1870]" captionStart="FIGURE 7. S 2 – 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 – 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="10" pageNumber="163">Fig. 7</figureCitation>
D); and (f) broad and triangular lobe of hamule (
<figureCitation id="34D925BC23247626FC3F267CFBD4A769" box="[985,1051,1845,1870]" captionStart="FIGURE 8" captionStartId="9.[151,250,911,933]" captionTargetBox="[157,1430,195,882]" captionTargetId="figure@9.[151,1435,193,890]" captionTargetPageId="9" captionText="FIGURE 8. Secondary genitalia of Acisoma species in ventral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides, D: A. attenboroughi, E: A. ascalaphoides, F: A. trifidum. Structure of hamule indicated." httpUri="https://zenodo.org/record/271427/files/figure.png" pageId="10" pageNumber="163">Fig. 8</figureCitation>
D). Unique in the genus by the (1) dorso-central black spot on the mesepisternum that is shifted toward and fused with the black humeral stripe (
<figureCitation id="34D925BC23247626FABC2610FA53A755" box="[1370,1436,1881,1906]" captionStart="FIGURE 6" captionStartId="7.[151,250,1802,1824]" captionTargetBox="[196,1382,1322,1745]" captionTargetId="figure@7.[193,1384,1321,1753]" captionTargetPageId="7" captionText="FIGURE 6. Thorax of Acisoma species in lateral view. A: A. variegatum, B: A. inflatum (typical), C: A. inflatum (dark), D: A. panorpoides, E: A. attenboroughi (typical), F: A. attenboroughi (dark), G: A. ascalaphoides, H: A. trifidum." httpUri="https://zenodo.org/record/271425/files/figure.png" pageId="10" pageNumber="163">Fig. 6</figureCitation>
E–F); and (2) S4 typically with all black ventral carina but partly white ventro-apical carina, although the latter is all black in about 10% of cases (
<figureCitation id="34D925BC23247626FDD026E8FD4FA79D" box="[566,640,1953,1978]" captionStart="FIGURE 7. S 2 – 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 – 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="10" pageNumber="163">Fig. 7</figureCitation>
D). The shape and general appearance is intermediate between
<taxonomicName id="6BE242BA23247626FA6226EBFEECA7FA" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23247626FA6226EBFEECA7FA" italics="true" pageId="10" pageNumber="163">A. panorpoides</emphasis>
</taxonomicName>
from Asia and
<taxonomicName id="6BE242BA23247626FE37268FFD82A7FA" box="[465,589,1989,2013]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="163" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23247626FE37268FFD82A7FA" box="[465,589,1989,2013]" italics="true" pageId="10" pageNumber="163">A. inflatum</emphasis>
</taxonomicName>
from Africa, with a (3) stout abdomen, with S6 about 1.5x to
<date id="D85C1FF923247626FAE3268FFAEFA7F9" box="[1285,1312,1990,2014]" pageId="10" pageNumber="163">2x</date>
as long as high, and a fairly distinct ‘step’ between the ventral borders of S5 and S6 (
<figureCitation id="34D925BC23237621FC372755FBDCA613" box="[977,1043,1564,1589]" captionStart="FIGURE 7. S 2 – 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 – 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="13" pageNumber="166">Fig. 7</figureCitation>
D). Differs from
<taxonomicName id="6BE242BA23237621FB292755FAB6A613" box="[1231,1401,1564,1588]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="166" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23237621FB292755FAB6A613" box="[1231,1401,1564,1588]" italics="true" pageId="13" pageNumber="166">A. panorpoides</emphasis>
</taxonomicName>
by the (4) wide black band across the antefrons, which is 40–80% as wide as the labrum is high (
<figureCitation id="34D925BC23237621FB2B2776FADDA67F" box="[1229,1298,1599,1624]" captionStart="FIGURE 4" captionStartId="7.[151,250,705,727]" captionTargetBox="[223,1345,203,654]" captionTargetId="figure@7.[216,1367,190,668]" captionTargetPageId="7" captionText="FIGURE 4. Head of Acisoma species in rostral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides (typical), D: A. panorpoides (pale), E: A. attenboroughi, F: A. ascalaphoides, G: A. trifidum." httpUri="https://zenodo.org/record/271423/files/figure.png" pageId="13" pageNumber="166">Fig. 4</figureCitation>
E); (5) large black spot on the labrum attached to its base (
<figureCitation id="34D925BC23237621FD4E272DFD21A65B" box="[680,750,1636,1661]" captionStart="FIGURE 4" captionStartId="7.[151,250,705,727]" captionTargetBox="[223,1345,203,654]" captionTargetId="figure@7.[216,1367,190,668]" captionTargetPageId="7" captionText="FIGURE 4. Head of Acisoma species in rostral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides (typical), D: A. panorpoides (pale), E: A. attenboroughi, F: A. ascalaphoides, G: A. trifidum." httpUri="https://zenodo.org/record/271423/files/figure.png" pageId="13" pageNumber="166">Fig. 4</figureCitation>
E); and (6) always incomplete distal Ax in Fw; and from
<taxonomicName id="6BE242BA23237621FA62272DFF3BA687" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="166" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23237621FA62272DFF3BA687" italics="true" pageId="13" pageNumber="166">A. inflatum</emphasis>
</taxonomicName>
by the (7) sublateral black marking concentrated on apical side of S4 (
<figureCitation id="34D925BC23237621FBEE27CEFB84A687" box="[1032,1099,1671,1696]" captionStart="FIGURE 7. S 2 – 10" captionStartId="8.[151,250,1858,1880]" captionTargetBox="[228,1399,193,1818]" captionTargetId="figure@8.[206,1415,190,1821]" captionTargetPageId="8" captionText="FIGURE 7. S 2 – 10 of Acisoma species in lateral view, scaled to their average size. Segment numbers and relevant characters are indicated." httpUri="https://zenodo.org/record/271426/files/figure.png" pageId="13" pageNumber="166">Fig. 7</figureCitation>
D); and (8) more slender hook of the hamule (
<figureCitation id="34D925BC23237621FEA627E5FE4CA6E3" box="[320,387,1708,1733]" captionStart="FIGURE 8" captionStartId="9.[151,250,911,933]" captionTargetBox="[157,1430,195,882]" captionTargetId="figure@9.[151,1435,193,890]" captionTargetPageId="9" captionText="FIGURE 8. Secondary genitalia of Acisoma species in ventral view. A: A. variegatum, B: A. inflatum, C: A. panorpoides, D: A. attenboroughi, E: A. ascalaphoides, F: A. trifidum. Structure of hamule indicated." httpUri="https://zenodo.org/record/271427/files/figure.png" pageId="13" pageNumber="166">Fig. 8</figureCitation>
D).
</paragraph>
<caption id="F89D69B123257627FF71261CFE7CA7C2" httpUri="https://zenodo.org/record/271428/files/figure.png" pageId="11" pageNumber="164" targetBox="[418,1156,196,1812]" targetPageId="11">
<paragraph id="AC5D393923257627FF71261CFE7CA7C2" blockId="11.[151,1437,1877,2021]" pageId="11" pageNumber="164">
<emphasis id="9E96E52B23257627FF71261CFED9A74D" bold="true" box="[151,278,1877,1899]" pageId="11" pageNumber="164">FIGURE 9.</emphasis>
Distribution of Afrotropical
<taxonomicName id="6BE242BA23257627FDAB261FFD6BA74C" box="[589,676,1878,1899]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="9E96E52B23257627FDAB261FFD6BA74C" box="[589,676,1878,1899]" italics="true" pageId="11" pageNumber="164">Acisoma</emphasis>
</taxonomicName>
species based on examined specimens and verified records in
<taxonomicName id="6BE242BA23257627FAA0261CFF3BA7AE" authority="Database" authorityName="Database" class="Insecta" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="order">Odonata Database</taxonomicName>
of Africa (see supplementary materials): A:
<taxonomicName id="6BE242BA23257627FD53263DFCEAA7AE" box="[693,805,1907,1929]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23257627FD53263DFCEAA7AE" box="[693,805,1907,1929]" italics="true" pageId="11" pageNumber="164">A. inflatum</emphasis>
</taxonomicName>
(circles),
<taxonomicName id="6BE242BA23257627FC6F263DFBFBA7AE" box="[905,1076,1907,1929]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="9E96E52B23257627FC6F263DFBFBA7AE" box="[905,1076,1907,1929]" italics="true" pageId="11" pageNumber="164">A. attenboroughi</emphasis>
</taxonomicName>
(triangles), unverified records of
<taxonomicName id="6BE242BA23257627FA60263DFF23A78F" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23257627FA60263DFF23A78F" italics="true" pageId="11" pageNumber="164">A. inflatum</emphasis>
</taxonomicName>
or
<taxonomicName id="6BE242BA23257627FEE826DAFE50A78F" box="[270,415,1939,1960]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="variegatum">
<emphasis id="9E96E52B23257627FEE826DAFE50A78F" box="[270,415,1939,1960]" italics="true" pageId="11" pageNumber="164">A. variegatum</emphasis>
</taxonomicName>
(grey dots); B:
<taxonomicName id="6BE242BA23257627FDA726DAFD1EA78F" box="[577,721,1939,1960]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="variegatum">
<emphasis id="9E96E52B23257627FDA726DAFD1EA78F" box="[577,721,1939,1960]" italics="true" pageId="11" pageNumber="164">A. variegatum</emphasis>
</taxonomicName>
(circles),
<taxonomicName id="6BE242BA23257627FCDF26DAFC29A78F" box="[825,998,1938,1960]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="ascalaphoides">
<emphasis id="9E96E52B23257627FCDF26DAFC29A78F" box="[825,998,1938,1960]" italics="true" pageId="11" pageNumber="164">A. ascalaphoides</emphasis>
</taxonomicName>
(squares),
<taxonomicName id="6BE242BA23257627FBB126DAFB0AA78F" box="[1111,1221,1938,1960]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="trifidum">
<emphasis id="9E96E52B23257627FBB126DAFB0AA78F" box="[1111,1221,1938,1960]" italics="true" pageId="11" pageNumber="164">A. trifidum</emphasis>
</taxonomicName>
(crosses), unverified records of
<taxonomicName id="6BE242BA23257627FEE926F8FE4AA7E0" box="[271,389,1969,1991]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23257627FEE926F8FE4AA7E0" box="[271,389,1969,1991]" italics="true" pageId="11" pageNumber="164">A. inflatum</emphasis>
</taxonomicName>
or
<taxonomicName id="6BE242BA23257627FE5526F8FD87A7E0" box="[435,584,1969,1991]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="164" phylum="Arthropoda" rank="species" species="variegatum">
<emphasis id="9E96E52B23257627FE5526F8FD87A7E0" box="[435,584,1969,1991]" italics="true" pageId="11" pageNumber="164">A. variegatum</emphasis>
</taxonomicName>
(grey dots). All records are provided in Suppl. Tab. 1 available from https:// science.naturalis.nl/dijkstra.
</paragraph>
</caption>
<caption id="F89D69B123227620FF7127B7FE5BA7EB" httpUri="https://zenodo.org/record/271429/files/figure.png" pageId="12" pageNumber="165" targetBox="[151,1436,193,1768]" targetPageId="12">
<paragraph id="AC5D393923227620FF7127B7FE5BA7EB" blockId="12.[151,1437,1790,1996]" pageId="12" pageNumber="165">
<emphasis id="9E96E52B23227620FF7127B7FEECA734" bold="true" box="[151,291,1790,1812]" pageId="12" pageNumber="165">FIGURE 10.</emphasis>
Male habitus of
<taxonomicName id="6BE242BA23227620FE3827B6FDFAA733" box="[478,565,1791,1812]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="9E96E52B23227620FE3827B6FDFAA733" box="[478,565,1791,1812]" italics="true" pageId="12" pageNumber="165">Acisoma</emphasis>
</taxonomicName>
species. A:
<taxonomicName id="6BE242BA23227620FD5B27B6FC9FA733" box="[701,848,1791,1812]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="variegatum">
<emphasis id="9E96E52B23227620FD5B27B6FC9FA733" box="[701,848,1791,1812]" italics="true" pageId="12" pageNumber="165">A. variegatum</emphasis>
</taxonomicName>
South Africa, 21-ii-2007 (mature), B:
<taxonomicName id="6BE242BA23227620FB1627B6FAABA733" box="[1264,1380,1790,1812]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="inflatum">
<emphasis id="9E96E52B23227620FB1627B6FAABA733" box="[1264,1380,1790,1812]" italics="true" pageId="12" pageNumber="165">A. inflatum</emphasis>
</taxonomicName>
, The Gambia, 30-xi-2010 (mature), C:
<taxonomicName id="6BE242BA23227620FE132654FD5DA715" box="[501,658,1820,1842]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23227620FE132654FD5DA715" box="[501,658,1820,1842]" italics="true" pageId="12" pageNumber="165">A. panorpoides</emphasis>
</taxonomicName>
, Hong Kong, China, 26-x-2011 (shortly after emergence, colours not fully developed), D:
<taxonomicName id="6BE242BA23227620FEDC2675FDA7A776" authority="Vohiparara" authorityName="Vohiparara" box="[314,616,1851,1873]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="9E96E52B23227620FEDC2675FE28A776" box="[314,487,1851,1873]" italics="true" pageId="12" pageNumber="165">A. attenboroughi</emphasis>
, Vohiparara
</taxonomicName>
, Madagascar, 13-i-2016, E:
<taxonomicName id="6BE242BA23227620FC742675FB00A776" authority="Sainte Luce" authorityName="Sainte Luce" box="[914,1231,1851,1873]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="ascalaphoides">
<emphasis id="9E96E52B23227620FC742675FB8DA776" box="[914,1090,1851,1873]" italics="true" pageId="12" pageNumber="165">A. ascalaphoides</emphasis>
, Sainte Luce
</taxonomicName>
, Madagascar, 19-x- 2006 (mature), F:
<taxonomicName id="6BE242BA23227620FEBB2613FDF8A757" authority="Moanda" authorityName="Moanda" box="[349,567,1882,1904]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="trifidum">
<emphasis id="9E96E52B23227620FEBB2613FE00A757" box="[349,463,1882,1904]" italics="true" pageId="12" pageNumber="165">A. trifidum</emphasis>
, Moanda
</taxonomicName>
, Gabon, 22-v-2013 (mature). Note characters such as the slender abdomen of
<taxonomicName id="6BE242BA23227620FA602613FEC5A7A9" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="variegatum">
<emphasis id="9E96E52B23227620FA602613FEC5A7A9" italics="true" pageId="12" pageNumber="165">A. variegatum</emphasis>
</taxonomicName>
, pale face and ventral ‘step’ between S5–6 in
<taxonomicName id="6BE242BA23227620FCC82630FC1DA7A9" box="[814,978,1912,1934]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="panorpoides">
<emphasis id="9E96E52B23227620FCC82630FC1DA7A9" box="[814,978,1912,1934]" italics="true" pageId="12" pageNumber="165">A. panorpoides</emphasis>
</taxonomicName>
, and shifted dorso-central black spot on mesepisternum in
<taxonomicName id="6BE242BA23227620FEB226D1FDCFA78A" box="[340,512,1943,1965]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="12" pageNumber="165" phylum="Arthropoda" rank="species" species="attenboroughi">
<emphasis id="9E96E52B23227620FEB226D1FDCFA78A" box="[340,512,1943,1965]" italics="true" pageId="12" pageNumber="165">A. attenboroughi</emphasis>
</taxonomicName>
. Photographs by Wil Leurs (A–B), Erland Nielsen (C), Kai Schütte (E), Netta Smith (D) and Nicolas Mézière (F).
</paragraph>
</caption>
<caption id="F89D69B123237621FF7124F3FC89A5F7" box="[151,838,1466,1488]" httpUri="https://zenodo.org/record/271430/files/figure.png" pageId="13" pageNumber="166" targetBox="[150,1436,193,1444]" targetPageId="13">
<paragraph id="AC5D393923237621FF7124F3FC89A5F7" blockId="13.[151,838,1466,1488]" box="[151,838,1466,1488]" pageId="13" pageNumber="166">
<emphasis id="9E96E52B23237621FF7124F3FED1A5E8" bold="true" box="[151,286,1466,1488]" pageId="13" pageNumber="166">FIGURE 11.</emphasis>
Holotype of
<taxonomicName id="6BE242BA23237621FE4224F2FD5FA5F7" box="[420,656,1466,1488]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="166" phylum="Arthropoda" rank="species" species="attenboroughi" status="sp. nov.">
<emphasis id="9E96E52B23237621FE4224F2FD5FA5F7" box="[420,656,1466,1488]" italics="true" pageId="13" pageNumber="166">Acisoma attenboroughi</emphasis>
</taxonomicName>
<emphasis id="9E96E52B23237621FD7024F2FD28A5F7" bold="true" box="[662,743,1467,1488]" pageId="13" pageNumber="166">
<taxonomicNameLabel id="85A5585023237621FD7024F2FD28A5F7" box="[662,743,1467,1488]" pageId="13" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ZMUH.
</paragraph>
</caption>
<paragraph id="AC5D393923237621FF212786FD75A717" blockId="13.[151,1437,1564,1840]" pageId="13" pageNumber="166">
<emphasis id="9E96E52B23237621FF212786FE63A6CF" bold="true" box="[199,428,1743,1768]" pageId="13" pageNumber="166">Range and ecology.</emphasis>
Widespread across
<collectingCountry id="D4F579A923237621FD6C2786FCDCA6CF" box="[650,787,1743,1768]" name="Madagascar" pageId="13" pageNumber="166">Madagascar</collectingCountry>
, including the interior, at weedy ponds such as abandoned rice fields. Found down to the coast in the east (Toamasina, Tolagnaro) and west (Mahajanga) but not in the littoral forests where
<taxonomicName id="6BE242BA23237621FED22651FE3CA717" box="[308,499,1816,1840]" class="Insecta" family="Libellulidae" genus="Acisoma" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="166" phylum="Arthropoda" rank="species" species="ascalaphoides">
<emphasis id="9E96E52B23237621FED22651FE8AA708" box="[308,325,1816,1839]" italics="true" pageId="13" pageNumber="166">A</emphasis>
.
<emphasis id="9E96E52B23237621FEB52651FE3CA717" box="[339,499,1816,1840]" italics="true" pageId="13" pageNumber="166">ascalaphoides</emphasis>
</taxonomicName>
occurs (
<figureCitation id="34D925BC23237621FDB5265EFD59A717" box="[595,662,1815,1840]" captionStart="FIGURE 9" captionStartId="11.[151,250,1877,1899]" captionTargetBox="[418,1156,196,1812]" captionTargetId="figure@11.[400,1172,193,1823]" captionTargetPageId="11" captionText="FIGURE 9. Distribution of Afrotropical Acisoma species based on examined specimens and verified records in Odonata Database of Africa (see supplementary materials): A: A. inflatum (circles), A. attenboroughi (triangles), unverified records of A. inflatum or A. variegatum (grey dots); B: A. variegatum (circles), A. ascalaphoides (squares), A. trifidum (crosses), unverified records of A. inflatum or A. variegatum (grey dots). All records are provided in Suppl. Tab. 1 available from https: // science. naturalis. nl / dijkstra." httpUri="https://zenodo.org/record/271428/files/figure.png" pageId="13" pageNumber="166">Fig. 9</figureCitation>
A).
</paragraph>
</subSubSection>
</treatment>
</document>
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