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<document id="B65E31106F6E444233B7285740896315" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D6D9229FDC7F968C9FD2DF2" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius sturnus Arrow 1941" docType="treatment" docVersion="10" lastPageNumber="115" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="114" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BA83FB6602F5E33670B60EDE8A8C6601">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
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<mods:namePart id="9681CB4A5DAA5351BF7A69EBFEB4AA0F">Telnov, Dmitry</mods:namePart>
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<mods:title id="3F49D19DAB0B5E0EDC8DBC32FC538642">European Journal of Taxonomy</mods:title>
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<mods:date id="3EDB5F8760EC27D1592C36D2EF073B0F">2020</mods:date>
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<treatment id="1A3787F69D6D9229FDC7F968C9FD2DF2" ID-DOI="http://doi.org/10.5281/zenodo.4344237" ID-GBIF-Taxon="168648124" ID-Zenodo-Dep="4344237" LSID="urn:lsid:plazi:treatment:1A3787F69D6D9229FDC7F968C9FD2DF2" httpUri="http://treatment.plazi.org/id/1A3787F69D6D9229FDC7F968C9FD2DF2" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114">
<subSubSection id="DA84656B9D6D922AFDC7F968CBB62E93" box="[596,992,1750,1777]" pageId="113" pageNumber="114" type="nomenclature">
<paragraph id="922136E09D6D922AFDC7F968CBB62E93" blockId="113.[486,1101,1750,1812]" box="[596,992,1750,1777]" pageId="113" pageNumber="114">
<heading id="C969818C9D6D922AFDC7F968CBB62E93" box="[596,992,1750,1777]" centered="true" fontSize="11" level="2" pageId="113" pageNumber="114" reason="2">
<taxonomicName id="559E4D639D6D922AFDC7F968CBB62E93" ID-CoL="3XFLM" authority="Arrow, 1941" authorityName="Arrow" authorityYear="1941" box="[596,992,1750,1777]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="sturnus">
<emphasis id="A0EAEAF29D6D922AFDC7F968CB122E93" bold="true" box="[596,836,1750,1777]" italics="true" pageId="113" pageNumber="114">Maechidius sturnus</emphasis>
<bibRefCitation id="F60F4B119D6D922AFCDFF969CBB62E93" author="Arrow G. E." box="[844,992,1751,1777]" pageId="113" pageNumber="114" pagination="448 - 464" refId="ref97558" refString="Arrow G. E. 1941. XXIX. - Some undescribed species of melolonthid Coleoptera from Western New Guinea and the adjacent islands of Waigeu and Japen. Annals and Magazine of Natural History, Series 11 7 (41): 448 - 464. https: // doi. org / 10.1080 / 03745481.1941.9727945" type="journal article" year="1941">Arrow, 1941</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6D922AFE75F947CC1B2F71" box="[486,1101,1785,1812]" pageId="113" pageNumber="114" type="description">
<paragraph id="922136E09D6D922AFE75F947CC1B2F71" blockId="113.[486,1101,1750,1812]" box="[486,1101,1785,1812]" pageId="113" pageNumber="114">
<figureCitation id="0AA52A659D6D922AFE75F947CA162F71" box="[486,576,1785,1811]" captionStart="Figs86–89" captionStartId="112.[189,241,1928,1954]" captionTargetBox="[282,1293,267,1862]" captionTargetId="figure-66@112.[278,1296,259,1877]" captionTargetPageId="112" captionText="Figs86–89. PapuanandWallaceanMaechidiusMacleay, 1819,habitus, dorsalview.86.M.speciosus(Frey, 1969) comb. nov., ♀ (BMNH). 87. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 88. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 89. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122167" httpUri="https://zenodo.org/record/4122167/files/figure.png" pageId="113" pageNumber="114">Figs 87</figureCitation>
,
<figureCitation id="0AA52A659D6D922AFDDFF947CA2C2F71" box="[588,634,1785,1811]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="113" pageNumber="114">181</figureCitation>
,
<figureCitation id="0AA52A659D6D922AFD14F947CAE02F76" box="[647,694,1785,1812]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="113" pageNumber="114">254</figureCitation>
,
<figureCitation id="0AA52A659D6D922AFD52F947CAA62F71" box="[705,752,1785,1811]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="113" pageNumber="114">347</figureCitation>
, 419, 433, 523, 563, 721–723
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6D922AFF2EF884C90A2F8B" pageId="113" pageNumber="114" type="materials_examined">
<paragraph id="922136E09D6D922AFF2EF884C93A2F31" blockId="113.[189,364,1849,1876]" box="[189,364,1849,1876]" pageId="113" pageNumber="114">
<heading id="C969818C9D6D922AFF2EF884C93A2F31" bold="true" box="[189,364,1849,1876]" fontSize="11" level="3" pageId="113" pageNumber="114" reason="3">
<emphasis id="A0EAEAF29D6D922AFF2EF884C93A2F31" bold="true" box="[189,364,1849,1876]" pageId="113" pageNumber="114">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6D922AFF79F8D9C90A2FE3" blockId="113.[189,1399,1895,2025]" box="[234,348,1895,1921]" pageId="113" pageNumber="114">
<emphasis id="A0EAEAF29D6D922AFF79F8D9C90A2FE3" bold="true" box="[234,348,1895,1921]" pageId="113" pageNumber="114">
<typeStatus id="4D2588429D6D922AFF79F8D9C90A2FE3" box="[234,348,1895,1921]" pageId="113" pageNumber="114" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D6D922AFF2EF834C90A2F8B" blockId="113.[189,1399,1895,2025]" pageId="113" pageNumber="114">
<materialsCitation id="22F63CBD9D6D922AFF2EF834C90A2F8B" ID-GBIF-Occurrence="2898878559" collectingDate="1938-10" collectionCode="DUTCH, NEW, R, BMNH" collectorName="L. E. Cheesman." country="Indonesia" elevation="152" location="Type" municipality="Japen" pageId="113" pageNumber="114" specimenCount="2" typeStatus="holotype">
<collectingCountry id="EA8976709D6D922AFF2EF834C90D2FC6" box="[189,347,1930,1956]" name="Indonesia" pageId="113" pageNumber="114">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D6D922AFE13F834C9CE2FC6" box="[384,408,1930,1956]" pageId="113" pageNumber="114">♂</specimenCount>
; “
<typeStatus id="4D2588429D6D922AFE25F834C9A52FC6" box="[438,499,1930,1956]" pageId="113" pageNumber="114">
<location id="9741603B9D6D922AFE25F834C9A52FC6" LSID="urn:lsid:plazi:treatment:1A3787F69D6D9229FDC7F968C9FD2DF2:9741603B9D6D922AFE25F834C9A52FC6" box="[438,499,1930,1956]" country="Indonesia" municipality="Japen" name="Type" pageId="113" pageNumber="114">Type</location>
</typeStatus>
[p, label circular, red frame] //
<collectionCode id="F48FAE259D6D922AFCE9F834CB892FC6" box="[890,991,1930,1956]" pageId="113" pageNumber="114">DUTCH</collectionCode>
<collectionCode id="F48FAE259D6D922AFC7EF834CC792FC6" box="[1005,1071,1930,1956]" country="Australia" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="113" pageNumber="114">NEW</collectionCode>
<collectingCountry id="EA8976709D6D922AFBAFF834CCE72FC6" box="[1084,1201,1930,1956]" name="Guinea-Bissau" pageId="113" pageNumber="114">GUINEA</collectingCountry>
:
<collectingMunicipality id="7245AC9A9D6D922AFB51F834CD532FC6" box="[1218,1285,1930,1956]" pageId="113" pageNumber="114">Japen</collectingMunicipality>
[sic
<specimenCount id="8498FD699D6D922AFAD7F837CD1E2FC6" box="[1348,1352,1929,1956]" pageId="113" pageNumber="114">♂</specimenCount>
] I.,
<collectionCode id="F48FAE259D6D922AFF2EF812C8872FA4" box="[189,209,1964,1990]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="113" pageNumber="114">R</collectionCode>
.Manai–Undei.
<quantity id="55669B059D6D922AFE1AF812C98F2FA4" box="[393,473,1964,1991]" metricMagnitude="2" metricUnit="m" metricValue="1.524" pageId="113" pageNumber="114" unit="ft" value="500.0">
<elevation id="19B3D1D39D6D922AFE1AF812C98F2FA4" box="[393,473,1964,1991]" metricMagnitude="2" metricUnit="m" metricValue="1.524" pageId="113" pageNumber="114" unit="ft" value="500.0">500 ft.</elevation>
</quantity>
,
<date id="E62010209D6D922AFE74F813CA152FA4" box="[487,579,1964,1991]" pageId="113" pageNumber="114" value="1938-10">
<collectingDate id="F664E9C89D6D922AFE74F813CA152FA4" box="[487,579,1964,1991]" pageId="113" pageNumber="114" value="1938-10">x. 1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D6D922AFDC3F812CB532FA4" box="[592,773,1964,1990]" pageId="113" pageNumber="114">L.E.Cheesman.</collectorName>
B.M.1938–593 [p] //
<emphasis id="A0EAEAF29D6D922AFB92F812CCB62FA4" box="[1025,1248,1964,1990]" italics="true" pageId="113" pageNumber="114">
Maechidus
<taxonomicName id="559E4D639D6D922AFB19F813CCB62FA4" authorityName="Arrow" authorityYear="1941" box="[1162,1248,1965,1990]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="sturnus">sturnus</taxonomicName>
</emphasis>
,
<typeStatus id="4D2588429D6D922AFB63F813CD752FA5" box="[1264,1315,1965,1991]" pageId="113" pageNumber="114">type</typeStatus>
Arrow [h]”;
<collectionCode id="F48FAE259D6D922AFF6FF871C90E2F8B" box="[252,344,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="113" pageNumber="114">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D6E9229FF2EFEB6C9FD2DF2" pageId="114" pageNumber="115" type="description">
<paragraph id="922136E09D6E9229FF2EFEB6C9182940" blockId="114.[189,334,264,290]" box="[189,334,264,290]" pageId="114" pageNumber="115">
<heading id="C969818C9D6E9229FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="114" pageNumber="115" reason="3">
<emphasis id="A0EAEAF29D6E9229FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="114" pageNumber="115">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6E9229FF2EFE86CD072D7C" blockId="114.[189,1399,312,1311]" pageId="114" pageNumber="115">
Dorsum and venter uniformly black, edges of labroclypeus, lateral margins of pronotum, legs and antennae brown. Male labroclypeus (
<figureCitation id="0AA52A659D6E9229FDF9FEE2CA9B2914" box="[618,717,348,374]" captionStart="Figs 181–192" captionStartId="144.[189,241,1787,1813]" captionTargetBox="[338,1249,274,1742]" captionTargetId="figure-122@144.[338,1249,265,1749]" captionTargetPageId="144" captionText="Figs 181–192. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 181. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 182. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 183. Same, ♀ (SNSD). 184. M. suwawa sp. nov., holotype, ♂ (BMNH). 185. Same, paratype, ♀ (BMNH). 186. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 187. M. trivialis sp. nov., holotype, ♂ (BMNH). 188. M. ursus sp. nov., holotype, ♀ (NME). 189. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 190. Same, ♂ (SNSD). 191. M. weigeli sp. nov., holotype, ♂ (NME). 192. Same, paratype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122187" httpUri="https://zenodo.org/record/4122187/files/figure.png" pageId="114" pageNumber="115">Fig. 181</figureCitation>
) broadly emarginate on anterior margin, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, slightly protruding. Canthus nearly straight in dorsal view. Head punctures irregularly hexagonal, dense, moderately deep. Intervening spaces glossy, in part wrinkled. At least areas around either compound eye covered with microscopical velvety pubescence. Head setae minute, rise from anterior margin of corresponding punctures, not surpassing its length. Pronotum sinuous on anterior margin, anterolateral angles slightly protruding. Basal margin of pronotum somewhat obtuse angulate medially. Lateral margin of pronotum in dorsal view broadly rounded, gradually constricted postmedially towards base, crenulate all along (
<figureCitation id="0AA52A659D6E9229FF55FDC2C97B2AF5" box="[198,301,636,663]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="114" pageNumber="115">Fig. 254</figureCitation>
). Inconspicuous appressed scale-like seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Hypomeron slightly emarginate and long setose on anterior margin which is flange-like protruding. Antennal pocket deep. Pronotal punctures irregularly ovoid, very deep, dense, arranged in different directions. Intervening spaces glossy, in part wrinkled, variably large. Lateral margins, antero- and posterolateral angles completely covered with delicate microscopical velvety pubescence. Pronotal setae similar to those on head. Elytra cylindrical, subopaque. Elytral punctures ovoid, deep and dense, arranged irregularly (
<figureCitation id="0AA52A659D6E9229FD4AFCEACB152B0C" box="[729,835,852,878]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="114" pageNumber="115">Fig. 347</figureCitation>
). Intervening spaces in part subopaque, in part glossy, wrinkled, generally smaller than punctures. Elytral setae minute, rise from anterior margin of corresponding puncture, not surpassing its midlength. With few longer scale-like suberect clavate setae scattered over elytral disc; punctures bearing those longer setae partly or completely encircled with microscopical velvety pubescence (
<figureCitation id="0AA52A659D6E9229FDCFFC5ACA952B9C" box="[604,707,996,1022]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="114" pageNumber="115">Fig. 347</figureCitation>
). Male abdominal sternites medially with somewhat longer and curved scale-like setae. Posterior margin of male abdominal sternites 2 and 3 each with bunch of three long setae on either side near middle (
<figureCitation id="0AA52A659D6E9229FD4FFB92CB1E2C24" box="[732,840,1068,1094]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="114" pageNumber="115">Fig. 433</figureCitation>
). Male pygidium with ovoid deep moderately large punctures (Fig. 523). Intervening spaces glossy to subopaque, generally larger than punctures. Setae of pygidium minute on anterior part (not surpassing length of corresponding punctures), longer on median and posterior parts (hardly surpassing length of corresponding punctures), becoming suberect and longer along distal margin. Surface of pygidium perimeter and narrow longitudinal midline covered with microscopical velvety pubescence. External margin of male protibia arched, with single acute distal tooth on external margin (Fig. 419). Spiculum gastrale as in
<figureCitation id="0AA52A659D6E9229FC12FABACBB12D7D" box="[897,999,1284,1311]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="114" pageNumber="115">Fig. 563</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D6E9229FB3AFABACD182D7C" box="[1193,1358,1284,1310]" captionStart="Figs 712–725" captionStartId="179.[189,241,1892,1918]" captionTargetBox="[433,1144,277,1858]" captionTargetId="figure-72@179.[427,1160,265,1870]" captionTargetPageId="179" captionText="Figs 712–725. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 712– 714. M. skalei sp. nov., holotype, ♂ (NME). 715–717. M. sougb sp. nov., holotype, ♂ (NME). 718–720. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 721–723. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 724–725. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122252" httpUri="https://zenodo.org/record/4122252/files/figure.png" pageId="114" pageNumber="115">Figs 721–723</figureCitation>
.
</paragraph>
<paragraph id="922136E09D6E9229FF2EFAF8C9E62D02" blockId="114.[189,432,1350,1376]" box="[189,432,1350,1376]" pageId="114" pageNumber="115">
<heading id="C969818C9D6E9229FF2EFAF8C9E62D02" bold="true" box="[189,432,1350,1376]" fontSize="11" level="3" pageId="114" pageNumber="115" reason="3">
<emphasis id="A0EAEAF29D6E9229FF2EFAF8C9E62D02" bold="true" box="[189,432,1350,1376]" pageId="114" pageNumber="115">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D6E9229FF2EFAC8C9FD2DF2" blockId="114.[189,427,1398,1424]" box="[189,427,1398,1424]" pageId="114" pageNumber="115">Female is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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