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<document id="1775BC9B3DC6C7F92D4A4FCF76BD479A" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D43923BFDBAFD86CA372A91" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius peregrinus Lansberge 1886" docType="treatment" docVersion="10" lastPageNumber="97" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="96" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="61771723F7A162C0249C49C69C2674ED">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="D2D2505A98F675ED71FD4C8378ADEB6C">Telnov, Dmitry</mods:namePart>
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<mods:title id="657E39134BBA4BCB1CE7D5C302E7F271">European Journal of Taxonomy</mods:title>
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<mods:date id="D6AEBE8A0DE88134D1F479D488A2D2A4">2020</mods:date>
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<mods:number id="A35ADFE01D6A780AFC920618873FD0E1">2020-10-19</mods:number>
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<mods:number id="D6FA2FD7580E06492761AA673E9267FB">721</mods:number>
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<mods:identifier id="AB4B3796ADCE40907FF61981A210A450" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D43923BFDBAFD86CA372A91" ID-DOI="http://doi.org/10.5281/zenodo.4344467" ID-GBIF-Taxon="168648155" ID-Zenodo-Dep="4344467" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91" httpUri="http://treatment.plazi.org/id/1A3787F69D43923BFDBAFD86CA372A91" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<subSubSection id="DA84656B9D439204FDBAFD86CC5D2A31" box="[553,1035,568,595]" pageId="95" pageNumber="96" type="nomenclature">
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<heading id="C969818C9D439204FDBAFD86CC5D2A31" box="[553,1035,568,595]" centered="true" fontSize="11" level="2" pageId="95" pageNumber="96" reason="2">
<taxonomicName id="559E4D639D439204FDBAFD86CC5D2A31" ID-CoL="7X3XG" authority="Lansberge, 1886" authorityName="Lansberge" authorityYear="1886" box="[553,1035,568,595]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="peregrinus">
<emphasis id="A0EAEAF29D439204FDBAFD86CB162A31" bold="true" box="[553,832,568,595]" italics="true" pageId="95" pageNumber="96">Maechidius peregrinus</emphasis>
<bibRefCitation id="F60F4B119D439204FCDBFD87CC5D2A31" author="Lansberge J. W. van" box="[840,1035,569,595]" pageId="95" pageNumber="96" pagination="131 - 137" refId="ref98256" refString="Lansberge J. W. van 1886. Description des quelques scarabaeides des Indes neerlandaises, appartenant au Musee du Leyde. Notes from the Leyden Museum 8: 131 - 137." type="journal article" year="1886">Lansberge, 1886</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="DA84656B9D439204FD90FDE2CC662A14" box="[515,1072,604,631]" pageId="95" pageNumber="96" type="description">
<paragraph id="922136E09D439204FD90FDE2CC662A14" blockId="95.[515,1072,568,631]" box="[515,1072,604,631]" pageId="95" pageNumber="96">
<figureCitation id="0AA52A659D439204FD90FDE2CA0B2A14" box="[515,605,604,630]" captionStart="Figs 70–73" captionStartId="93.[189,241,1896,1922]" captionTargetBox="[327,1247,269,1869]" captionTargetId="figure-67@93.[327,1254,265,1878]" captionTargetPageId="93" captionText="Figs 70–73. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 70. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 71. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 72. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 73. M. peregrinus Lansberge, 1886, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122158" httpUri="https://zenodo.org/record/4122158/files/figure.png" pageId="95" pageNumber="96">Figs 73</figureCitation>
,
<figureCitation id="0AA52A659D439204FDFAFDE2CACE2A14" box="[617,664,604,630]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="95" pageNumber="96">162</figureCitation>
,
<figureCitation id="0AA52A659D439204FD37FDE2CA852A15" box="[676,723,604,631]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="95" pageNumber="96">245</figureCitation>
,
<figureCitation id="0AA52A659D439204FD4CFDE2CB582A14" box="[735,782,604,630]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="95" pageNumber="96">276</figureCitation>
,
<figureCitation id="0AA52A659D439204FC8AFDE2CB1E2A15" box="[793,840,604,631]" captionStart="Figs 333–348" captionStartId="155.[189,241,1682,1708]" captionTargetBox="[394,1193,265,1641]" captionTargetId="figure-177@155.[394,1193,265,1655]" captionTargetPageId="155" captionText="Figs 333–348. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 333. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 334. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 335. M. peregrinus Lansberge, 1886, ♂ (DTC). 336. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 337. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 338. Same, enlarged (SMNS). 339. M. riedeli sp. nov., holotype, ♂ (SMNS). 340. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 341. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 342. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 343. M. similis sp. nov., paratype, ♂ (BMNH). 344. M. skalei sp. nov., paratype, ♂ (NME). 345. M. sougb sp. nov., holotype, ♂ (NME). 346. M. speciosus (Frey, 1969) comb. nov., ♀(BMNH). 347. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 348. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122212" httpUri="https://zenodo.org/record/4122212/files/figure.png" pageId="95" pageNumber="96">335</figureCitation>
, 408, 509, 691–693
</paragraph>
</subSubSection>
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<emphasis id="A0EAEAF29D439204FF2EFD23C93A2AD4" bold="true" box="[189,364,668,695]" pageId="95" pageNumber="96">Type material</emphasis>
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<typeStatus id="4D2588429D439204FF79FD74C9192A86" box="[234,335,714,740]" pageId="95" pageNumber="96" type="syntype">Syntype</typeStatus>
</emphasis>
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<materialsCitation id="22F63CBD9D439204FF2EFD50CBAC2B65" ID-GBIF-Occurrence="2898878354" box="[189,1018,749,776]" country="Indonesia" location="South Sulawesi" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Selatan" typeStatus="syntype">
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•
<specimenCount id="8498FD699D439204FEE7FD50C9DD2B6A" box="[372,395,750,776]" pageId="95" pageNumber="96" type="female">♀</specimenCount>
;
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;
<collectionCode id="F48FAE259D439204FDCCFD50CAEF2B6A" box="[607,697,750,776]" collectionName="NMNL" pageId="95" pageNumber="96">NMNL</collectionCode>
. Studied from photographs
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.
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<emphasis id="A0EAEAF29D439204FF2EFC91C9302B2A" bold="true" box="[189,358,814,841]" pageId="95" pageNumber="96">New material</emphasis>
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<materialsCitation id="22F63CBD9D439204FF2EFCE3CBCE2BFB" ID-GBIF-Occurrence="2898878624" collectingDate="2019-12-26" collectingDateMax="2019-12-28" collectingDateMin="2019-12-26" collectionCode="DTC" country="Indonesia" elevation="350" latitude="-2.95" location="Battang" longLatPrecision="1307" longitude="120.13333" municipality="Palopo" pageId="95" pageNumber="96" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Utara">
<collectingCountry id="EA8976709D439204FF2EFCE3C90D2B15" box="[189,347,861,887]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D439204FEEFFCE3C9952B15" box="[380,451,861,887]" pageId="95" pageNumber="96" type="female">2 ♀♀</specimenCount>
; “
<collectingCountry id="EA8976709D439204FE4CFCE3CAD32B15" box="[479,645,861,887]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FD23FCE2CB4B2B14" box="[688,797,860,886]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
,
<collectingMunicipality id="7245AC9A9D439204FCBEFCE2CB282B14" box="[813,894,860,886]" pageId="95" pageNumber="96">Palopo</collectingMunicipality>
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NW,
<location id="9741603B9D439204FB86FCE3CC262B15" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FB86FCE3CC262B15" box="[1045,1136,861,887]" country="Indonesia" latitude="-2.95" longLatPrecision="1307" longitude="120.13333" municipality="Palopo" name="Battang" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Battang</location>
vill.,
<quantity id="55669B059D439204FB2EFCE2CD5B2B15" box="[1213,1293,860,887]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="95" pageNumber="96" unit="m" value="350.0">
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,
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,
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,
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, at light, leg. local collector”;
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•
<materialsCitation id="22F63CBD9D439204FC22FCC1CD212BDE" ID-GBIF-Occurrence="2898878579" collectingDate="2019-12-28" collectionCode="DTC" country="Indonesia" elevation="985" latitude="-2.9816668" location="Sarambu" longLatPrecision="21" longitude="119.83833" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
<specimenCount id="8498FD699D439204FC22FCC1CB892BF8" box="[945,991,895,922]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="EA8976709D439204FC6BFCC1CCC82BFB" box="[1016,1182,895,921]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FB51FCC1CD792BFB" box="[1218,1327,895,921]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
, W of
<locationDeviation id="83932FC29D439204FF2EFC1CC9472BDE" box="[189,273,930,956]" pageId="95" pageNumber="96">Palopo</locationDeviation>
,
<locationDeviation id="83932FC29D439204FE8EFC1DC9DA2BDF" box="[285,396,931,957]" pageId="95" pageNumber="96">Rantepao</locationDeviation>
<quantity id="55669B059D439204FE01FC1DC99B2BDE" box="[402,461,930,957]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="95" pageNumber="96" unit="km" value="7.0">7 km</quantity>
W,
<location id="9741603B9D439204FE68FC1CCA352BDE" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FE68FC1CCA352BDE" box="[507,611,930,956]" country="Indonesia" latitude="-2.9816668" longLatPrecision="21" longitude="119.83833" name="Sarambu" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Sarambu</location>
vill.,
<quantity id="55669B059D439204FD35FC1CCAA42BDF" box="[678,754,930,957]" metricMagnitude="2" metricUnit="m" metricValue="9.85" pageId="95" pageNumber="96" unit="m" value="985.0">
<elevation id="19B3D1D39D439204FD35FC1CCAA42BDF" box="[678,754,930,957]" metricMagnitude="2" metricUnit="m" metricValue="9.85" pageId="95" pageNumber="96" unit="m" value="985.0">985 m</elevation>
</quantity>
,
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,
<geoCoordinate id="F7AA50279D439204FC1BFC1CCC732BDF" box="[904,1061,930,957]" degrees="119" direction="east" minutes="50" orientation="longitude" pageId="95" pageNumber="96" precision="15" seconds="18" value="119.83833">119°50’18”E</geoCoordinate>
, at light,
<date id="E62010209D439204FB03FC1CCD732BDE" box="[1168,1317,930,956]" pageId="95" pageNumber="96" value="2019-12-28">
<collectingDate id="F664E9C89D439204FB03FC1CCD732BDE" box="[1168,1317,930,956]" pageId="95" pageNumber="96" value="2019-12-28">28.XII.2019</collectingDate>
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”;
<collectionCode id="F48FAE259D439204FAAFFC1CCD212BDE" box="[1340,1399,930,956]" pageId="95" pageNumber="96">DTC</collectionCode>
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•
<materialsCitation id="22F63CBD9D439204FF42FC7BCA702C60" ID-GBIF-Occurrence="2898878644" collectingDate="2020-01-01" collectionCode="DTC" country="Indonesia" elevation="1490" latitude="-3.1422224" location="Palopo" longLatPrecision="21" longitude="119.824165" pageId="95" pageNumber="96" specimenCount="2" specimenCount-female="2" stateProvince="Sulawesi Utara">
1 ♂,
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; “
<collectingCountry id="EA8976709D439204FEFCFC7BCA432BBD" box="[367,533,965,991]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FDAEFC7BCAFC2BBD" box="[573,682,965,991]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
,
<location id="9741603B9D439204FD24FC7BCB5E2BBD" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FD24FC7BCB5E2BBD" box="[695,776,965,991]" country="Indonesia" latitude="-3.1422224" longLatPrecision="21" longitude="119.824165" name="Palopo" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Palopo</location>
<quantity id="55669B059D439204FC81FC7BCB082BBD" box="[786,862,965,992]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="95" pageNumber="96" unit="km" value="45.0">45 km</quantity>
SW,
<locationDeviation id="83932FC29D439204FC31FC7BCC2F2BBD" box="[930,1145,965,992]" location="Makale" pageId="95" pageNumber="96">
<quantity id="55669B059D439204FC31FC7BCB892BBD" box="[930,991,965,992]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="95" pageNumber="96" unit="km" value="5.0">5 km</quantity>
SW Makale
</locationDeviation>
,
<quantity id="55669B059D439204FB14FC7BCCB22BBD" box="[1159,1252,965,991]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="95" pageNumber="96" unit="m" value="1490.0">
<elevation id="19B3D1D39D439204FB14FC7BCCB22BBD" box="[1159,1252,965,991]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="95" pageNumber="96" unit="m" value="1490.0">1490 m</elevation>
</quantity>
,
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,
<geoCoordinate id="F7AA50279D439204FF2EFC56C90F2C60" box="[189,345,1000,1026]" degrees="119" direction="east" minutes="49" orientation="longitude" pageId="95" pageNumber="96" precision="15" seconds="27" value="119.824165">119°49’27”E</geoCoordinate>
,
<date id="E62010209D439204FEFAFC56C9992C60" box="[361,463,1000,1026]" pageId="95" pageNumber="96" value="2020-01-01">
<collectingDate id="F664E9C89D439204FEFAFC56C9992C60" box="[361,463,1000,1026]" pageId="95" pageNumber="96" value="2020-01-01">1.I.2020</collectingDate>
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”;
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•
<materialsCitation id="22F63CBD9D439204FDD6FC56CB9B2C46" ID-GBIF-Occurrence="2898878596" collectingDate="2020-01-04" collectionCode="DTC" country="Indonesia" elevation="765" latitude="-2.9586112" location="Puri Rimba" longLatPrecision="21" longitude="120.086945" municipality="Palopo" pageId="95" pageNumber="96" specimenCount="1" stateProvince="Sulawesi Utara">
1 ♂; “
<collectingCountry id="EA8976709D439204FD02FC56CB612C60" box="[657,823,1000,1026]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FCF2FC59CB982C63" box="[865,974,999,1025]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
,
<collectingMunicipality id="7245AC9A9D439204FC4EFC59CC782C63" box="[989,1070,999,1025]" pageId="95" pageNumber="96">Palopo</collectingMunicipality>
<quantity id="55669B059D439204FBAAFC56CCD02C63" box="[1081,1158,999,1026]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="95" pageNumber="96" unit="km" value="12.0">12 km</quantity>
NWW,
<location id="9741603B9D439204FB7EFC59CD202C63" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FB7EFC59CD202C63" box="[1261,1398,999,1025]" country="Indonesia" latitude="-2.9586112" longLatPrecision="21" longitude="120.086945" municipality="Palopo" name="Puri Rimba" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Puri Rimba</location>
resort,
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,
<geoCoordinate id="F7AA50279D439204FE0CFBB4CA6B2C47" box="[415,573,1034,1061]" degrees="120" direction="east" minutes="05" orientation="longitude" pageId="95" pageNumber="96" precision="15" seconds="13" value="120.086945">120°05’13”E</geoCoordinate>
,
<quantity id="55669B059D439204FDDFFBB4CACD2C47" box="[588,667,1034,1061]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="95" pageNumber="96" unit="m" value="765.0">
<elevation id="19B3D1D39D439204FDDFFBB4CACD2C47" box="[588,667,1034,1061]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="95" pageNumber="96" unit="m" value="765.0">765 m</elevation>
</quantity>
,
<date id="E62010209D439204FD39FBB4CB592C46" box="[682,783,1034,1060]" pageId="95" pageNumber="96" value="2020-01-04">
<collectingDate id="F664E9C89D439204FD39FBB4CB592C46" box="[682,783,1034,1060]" pageId="95" pageNumber="96" value="2020-01-04">4.I.2020</collectingDate>
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, at light”;
<collectionCode id="F48FAE259D439204FC01FBB4CB9B2C46" box="[914,973,1034,1060]" pageId="95" pageNumber="96">DTC</collectionCode>
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•
<materialsCitation id="22F63CBD9D439204FC78FBB4CD212C25" ID-GBIF-Occurrence="2898878330" collectingDate="2020-01-04" collectionCode="DTC" country="Indonesia" elevation="765" latitude="-2.9586112" location="Puri Rimba" longLatPrecision="21" longitude="120.086945" municipality="Palopo" pageId="95" pageNumber="96" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara">
<specimenCount id="8498FD699D439204FC78FBB4CC4D2C47" box="[1003,1051,1034,1061]" pageId="95" pageNumber="96" type="female">1 ♀</specimenCount>
; “
<collectingCountry id="EA8976709D439204FBA5FBB4CC8A2C46" box="[1078,1244,1034,1060]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FA96FBB4CD242C46" box="[1285,1394,1034,1060]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
,
<collectingMunicipality id="7245AC9A9D439204FF2EFB93C9582C25" box="[189,270,1069,1095]" pageId="95" pageNumber="96">Palopo</collectingMunicipality>
<quantity id="55669B059D439204FE8AFB93C9302C25" box="[281,358,1069,1095]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="95" pageNumber="96" unit="km" value="12.0">12 km</quantity>
NWW,
<location id="9741603B9D439204FE5DFB93CA012C25" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FE5DFB93CA012C25" box="[462,599,1069,1095]" country="Indonesia" latitude="-2.9586112" longLatPrecision="21" longitude="120.086945" municipality="Palopo" name="Puri Rimba" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Puri Rimba</location>
resort,
<geoCoordinate id="F7AA50279D439204FD24FB93CB602C2A" box="[695,822,1069,1096]" degrees="2" direction="south" minutes="57" orientation="latitude" pageId="95" pageNumber="96" precision="15" seconds="31" value="-2.9586112">2°57’31”S</geoCoordinate>
,
<geoCoordinate id="F7AA50279D439204FCD6FB93CBB52C2A" box="[837,995,1069,1096]" degrees="120" direction="east" minutes="05" orientation="longitude" pageId="95" pageNumber="96" precision="15" seconds="13" value="120.086945">120°05’13”E</geoCoordinate>
,
<quantity id="55669B059D439204FC60FB93CC142C25" box="[1011,1090,1069,1096]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="95" pageNumber="96" unit="m" value="765.0">
<elevation id="19B3D1D39D439204FC60FB93CC142C25" box="[1011,1090,1069,1096]" metricMagnitude="2" metricUnit="m" metricValue="7.65" pageId="95" pageNumber="96" unit="m" value="765.0">765 m</elevation>
</quantity>
,
<date id="E62010209D439204FBC1FB93CCE12C25" box="[1106,1207,1069,1095]" pageId="95" pageNumber="96" value="2020-01-04">
<collectingDate id="F664E9C89D439204FBC1FB93CCE12C25" box="[1106,1207,1069,1095]" pageId="95" pageNumber="96" value="2020-01-04">4.I.2020</collectingDate>
</date>
, at light”;
<collectionCode id="F48FAE259D439204FAAFFB93CD212C25" box="[1340,1399,1069,1095]" pageId="95" pageNumber="96">DTC</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D439204FF41FBEECBD22CEF" ID-GBIF-Occurrence="2898878522" collectingDate="2020-01-14" collectionCode="DTC" country="Indonesia" elevation="1430" latitude="-2.894722" location="Landorundun" longLatPrecision="21" longitude="119.849724" pageId="95" pageNumber="96" specimenCount="3" specimenCount-female="3" stateProvince="Sulawesi Utara">
2 ♂♂,
<specimenCount id="8498FD699D439204FEB9FBEEC9262C08" box="[298,368,1104,1130]" pageId="95" pageNumber="96" type="female">3 ♀♀</specimenCount>
; “
<collectingCountry id="EA8976709D439204FE1FFBEECA642C08" box="[396,562,1104,1130]" name="Indonesia" pageId="95" pageNumber="96">INDONESIA</collectingCountry>
, S
<collectingRegion id="505AF8029D439204FDCEFBEECA9C2C08" box="[605,714,1104,1130]" country="Indonesia" name="Sulawesi Utara" pageId="95" pageNumber="96">Sulawesi</collectingRegion>
, W of
<locationDeviation id="83932FC29D439204FCB2FBEECB232C08" box="[801,885,1104,1130]" pageId="95" pageNumber="96">Palopo</locationDeviation>
,
<locationDeviation id="83932FC29D439204FC16FBEECBA22C08" box="[901,1012,1104,1130]" pageId="95" pageNumber="96">Rantepao</locationDeviation>
<quantity id="55669B059D439204FC6CFBEECC1A2C08" box="[1023,1100,1104,1130]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="95" pageNumber="96" unit="km" value="10.0">10 km</quantity>
NW,
<location id="9741603B9D439204FB0BFBEECD602C08" LSID="urn:lsid:plazi:treatment:1A3787F69D43923BFDBAFD86CA372A91:9741603B9D439204FB0BFBEECD602C08" box="[1176,1334,1104,1130]" country="Indonesia" latitude="-2.894722" longLatPrecision="21" longitude="119.849724" name="Landorundun" pageId="95" pageNumber="96" stateProvince="Sulawesi Utara">Landorundun</location>
vill.,
<geoCoordinate id="F7AA50279D439204FF2EFBCCC96A2CEF" box="[189,316,1138,1165]" degrees="2" direction="south" minutes="53" orientation="latitude" pageId="95" pageNumber="96" precision="15" seconds="41" value="-2.894722">2°53’41”S</geoCoordinate>
,
<geoCoordinate id="F7AA50279D439204FEDBFBCCC9B22CEF" box="[328,484,1138,1165]" degrees="119" direction="east" minutes="50" orientation="longitude" pageId="95" pageNumber="96" precision="15" seconds="59" value="119.849724">119°50’59”E</geoCoordinate>
,
<quantity id="55669B059D439204FE62FBCDCA1A2CEF" box="[497,588,1139,1165]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="95" pageNumber="96" unit="m" value="1430.0">
<elevation id="19B3D1D39D439204FE62FBCDCA1A2CEF" box="[497,588,1139,1165]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="95" pageNumber="96" unit="m" value="1430.0">1430 m</elevation>
</quantity>
, at light,
<date id="E62010209D439204FD2AFBCDCB792CEF" box="[697,815,1139,1165]" pageId="95" pageNumber="96" value="2020-01-14">
<collectingDate id="F664E9C89D439204FD2AFBCDCB792CEF" box="[697,815,1139,1165]" pageId="95" pageNumber="96" value="2020-01-14">14.I.2020</collectingDate>
</date>
”;
<collectionCode id="F48FAE259D439204FCD5FBCDCBD22CEF" box="[838,900,1139,1165]" pageId="95" pageNumber="96">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D43923BFF2EFB0DCA372A91" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="description">
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<emphasis id="A0EAEAF29D439204FF2EFB0DC9182CAF" bold="true" box="[189,334,1203,1229]" pageId="95" pageNumber="96">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D439204FF2EFB5CCB552C9E" blockId="95.[189,771,1249,1276]" box="[189,771,1249,1276]" pageId="95" pageNumber="96">
MEASUREMENTS. Total body length
<quantity id="55669B059D439204FDC2FB5FCAA82C9E" box="[593,766,1249,1276]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.4" metricValueMin="7.6" pageId="95" pageNumber="96" unit="mm" value="8.5" valueMax="9.4" valueMin="7.6">7.60–9.40 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D43923BFF2EFA9ACB852A5D" blockId="95.[189,1399,1315,2025]" lastBlockId="96.[189,1399,265,576]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
Dorsum and venter uniformly black-brown, labroclypeus, antennae and legs brown. Male (
<figureCitation id="0AA52A659D439204FB49FA9ACD6B2D5C" box="[1242,1341,1315,1342]" captionStart="Figs 157–168" captionStartId="142.[189,241,1752,1778]" captionTargetBox="[338,1248,270,1715]" captionTargetId="figure-140@142.[338,1249,265,1724]" captionTargetPageId="142" captionText="Figs 157–168. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal (157–159, 161–167) and frontal (160, 168) views. 157. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). 158. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 159. Same, ♂ (paratype of M. arrowi Frey, 1969) (BMNH). 160. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 161. Same, ♀ (SNSD). 162. M. peregrinus Lansberge, 1886, ♂ (DTC). 163. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 164. Same, syntype, ♂ (BMNH). 165. M. riedeli sp. nov., holotype, ♂ (SMNS). 166. M. perlatus (Frey, 1969) comb. nov., holotype, ♂ (NHMB). 167. Same, ♀ (NHMB). 168. M. simplex Frey, 1969, holotype, ♂ (photograph courtesy BPBM) (BPBM). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122183" httpUri="https://zenodo.org/record/4122183/files/figure.png" pageId="95" pageNumber="96">Fig. 162</figureCitation>
) and female labroclypeus shallowly emarginate to subtruncate on anterior margin, its lateral margins nearly straight to slightly sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus obtuse, rounded, not protruding anteriad. Frons convex dorsally. Head punctures ovoid to irregularly hexagonal, very deep and dense, filled with microscopical velvety pubescence. Intervening spaces glossy, generally much smaller than punctures. Head setae suberect to erect, moderately dense, much longer on frons than on anterior part of head; each seta rises from anterior margin of corresponding puncture. Pronotum strongly transverse, sinuous on anterior margin, with slightly protruding anterolateral angles. Basal margin of pronotum very broadly rounded to subtruncate. Lateral margin of pronotum broadly rounded, crenulate all along (
<figureCitation id="0AA52A659D439204FE0EF9D9CA572EE0" box="[413,513,1639,1666]" captionStart="Figs 242–258" captionStartId="148.[189,241,1646,1672]" captionTargetBox="[394,1193,265,1617]" captionTargetId="figure-193@148.[394,1193,265,1626]" captionTargetPageId="148" captionText="Figs 242–258. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 242. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 243. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 244. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 245. M. peregrinus Lansberge, 1886, ♂ (DTC). 246. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 247. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 248. M. riedeli sp. nov., holotype, ♂ (SMNS). 249. M. rugicollis Moser, 1920, lectotype, ♀ (ZMHB). 250. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 251. M. seriepunctatus Moser, 1920, lectotype, ♂ (ZMHB). 252. M. similis sp. nov., paratype, ♂ (BMNH). 253. M. skalei sp. nov., paratype, ♂ (NME). 254. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 255. M. sougb sp. nov., holotype, ♂ (NME). 256. M. speciosus (Frey, 1969) comb. nov., ♀ (BMNH). 257. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 258. M. suwawa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122196" httpUri="https://zenodo.org/record/4122196/files/figure.png" pageId="95" pageNumber="96">Fig. 245</figureCitation>
). Long erect apically slightly curved seta present between every two crenulae. Maximum width of pronotum postmedially. Pronotal punctures ovoid, very deep and dense, filled with microscopical velvety pubescence. Intervening spaces glossy, much smaller than punctures, in part wrinkled. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Setae of pronotum generally shorter than those on head, erect, apically curved; wide median prebasal area glabrous. Hypomeron broadly emarginate and very long setose on anterior margin which is flange-like (
<figureCitation id="0AA52A659D439204FD81F881CA2D2F3B" box="[530,635,1854,1881]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="95" pageNumber="96">Fig. 276</figureCitation>
). Scutellar shield narrowly rounded apically. Elytron with tracks of three flat glabrous longitudinal carinae; sutural carina not indicated. Elytral punctures elongate ovoid and narrow, deep and dense. Intervening spaces glossy, generally larger than punctures, in part covered with microscopical velvety pubescence. Elytral setae suberect to erect, variably long. All setae rise from anterior margin of corresponding punctures, shorter ones about their length, longer ones much longer and arranged in longitudinal rows along tracks of elytral carinae. Male pygidium slightly convex dorsally, partly covered with microscopical velvety pubescence, with moderately large dense circular punctures and variably large intervening spaces (
<figureCitation id="0AA52A659D7C923BFC96FEEECB382909" box="[773,878,336,363]" captionStart="Figs 504–521" captionStartId="166.[189,241,1681,1707]" captionTargetBox="[433,1155,265,1647]" captionTargetId="figure-177@166.[433,1155,265,1658]" captionTargetPageId="166" captionText="Figs 504–521. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 504. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 505. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 506. Same, ♀ (BMNH). 507. M. pedarioides Arrow, 1941, syntype, ♀ (BMNH). 508. Same, syntype, ♂ (BMNH). 509. M. peregrinus Lansberge, 1886, ♂ (DTC). 510. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 511. M. popei (Frey, 1969) comb. nov., holotype, ♀ (BMNH). 512. Same, paratype, ♂ (BMNH). 513. M. riedeli sp. nov., holotype, ♂ (SMNS). 514. M. rugicollis Moser, 1920, ♂ (BMNH). 515. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 516. M. similis sp. nov., paratype, ♂ (BMNH). 517. M. skalei sp. nov., paratype, ♂ (NME). 518. M. sougb sp. nov., holotype, ♂ (NME). 519. M. speciosus (Frey, 1969) comb. nov., ♂ (BMNH). 520. Same, different ♂ (BMNH). 521. Same, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122228" httpUri="https://zenodo.org/record/4122228/files/figure.png" pageId="96" pageNumber="97">Fig. 509</figureCitation>
). Female pygidium with rather large obtuse median impression at posterior margin. Setae of pygidium suberect to erect, moderately long. Male and female protibia widened distally, with two obtuse distal teeth on external margin (Fig. 408). Distal margin of metatibia in both sexes with projection over basal metatarsomere. Male protibial terminal spur strongly curved, female one straight and pointed. Male lower meso- and metatibial terminal spur strongly curved, not rounded apically. Male basal metatarsomere widened and dorso-ventrally flattened. All claws with pulvilli in both sexes. Aedeagus as in
<figureCitation id="0AA52A659D7C923BFCB8FD98CB992A5D" box="[811,975,549,576]" captionStart="Figs 682–696" captionStartId="177.[189,241,1892,1918]" captionTargetBox="[436,1163,269,1860]" captionTargetId="figure-72@177.[423,1174,265,1866]" captionTargetPageId="177" captionText="Figs 682–696. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 682– 684. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 685–687. M. pauxillus Heller, 1910 comb. rest., ♂ (BMNH). 688–690. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 691–693. M. peregrinus Lansberge, 1886, ♂ (DTC). 694-696. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122248" httpUri="https://zenodo.org/record/4122248/files/figure.png" pageId="96" pageNumber="97">Figs 691–693</figureCitation>
.
</paragraph>
<paragraph id="922136E09D7C923BFF2EFDD8C9E62AE2" blockId="96.[189,432,614,640]" box="[189,432,614,640]" pageId="96" pageNumber="97">
<heading id="C969818C9D7C923BFF2EFDD8C9E62AE2" bold="true" box="[189,432,614,640]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="A0EAEAF29D7C923BFF2EFDD8C9E62AE2" bold="true" box="[189,432,614,640]" pageId="96" pageNumber="97">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D7C923BFF2EFD2ACA372A91" blockId="96.[189,1398,660,755]" pageId="96" pageNumber="97">Lamellae of female antennal club shorter; terminal spur of protibia straight and pointed; basal metatarsomere moderately widened and flattened; pygidium humped medially; lower meso- and metatibial terminal spur not curved.</paragraph>
</subSubSection>
</treatment>
</document>
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