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<document id="B6FD6484003FACC09F9103DFF07DCBBD" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D2B9262FD93FAF6C9C72EDD" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius interruptocarinulatus Heller 1914" docType="treatment" docVersion="10" lastPageNumber="58" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="56" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="DBA3DF43DB8355F52E32FCC0E917A2C7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5C8636C5F7759C9FF54AEDF5D21D7A7E">
<mods:title id="33224FBF988446D86C2C563CE3333513">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="E50FEE9803FD3AC16681A299B7312491">Telnov, Dmitry</mods:namePart>
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<mods:title id="86EF88D3F4D33AD076ED748077C482D0">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
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<mods:date id="2C734491890D412252607B29FA29F262">2020</mods:date>
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<mods:number id="A13E0FFB730A8D0DC88F43E7276D3A2C">2020-10-19</mods:number>
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<mods:number id="592A34A8DEF7869EC973165CAB655AD4">721</mods:number>
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<mods:classification id="8FD53FD018BA868C0C8C4C8131996021">journal article</mods:classification>
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<mods:identifier id="9FBE8F990CE3303AB4EAA08FF3FF7F30" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D2B9262FD93FAF6C9C72EDD" ID-DOI="http://doi.org/10.5281/zenodo.4344419" ID-GBIF-Taxon="168648136" ID-Zenodo-Dep="4344419" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD" httpUri="http://treatment.plazi.org/id/1A3787F69D2B9262FD93FAF6C9C72EDD" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56">
<subSubSection id="DA84656B9D2B926CFD93FAF6CC652D01" box="[512,1075,1352,1379]" pageId="55" pageNumber="56" type="nomenclature">
<paragraph id="922136E09D2B926CFD93FAF6CC652D01" blockId="55.[512,1075,1352,1414]" box="[512,1075,1352,1379]" pageId="55" pageNumber="56">
<heading id="C969818C9D2B926CFD93FAF6CC652D01" box="[512,1075,1352,1379]" centered="true" fontSize="11" level="2" pageId="55" pageNumber="56" reason="2">
<taxonomicName id="559E4D639D2B926CFD93FAF6CC652D01" ID-CoL="3XFJN" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[512,1075,1352,1379]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D2B926CFD93FAF6CBCC2D01" bold="true" box="[512,922,1352,1379]" italics="true" pageId="55" pageNumber="56">Maechidius interruptocarinulatus</emphasis>
<bibRefCitation id="F60F4B119D2B926CFC31FAF7CC652D01" author="Heller K. M." box="[930,1075,1353,1379]" pageId="55" pageNumber="56" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2B926CFDC8FAD2CB8E2DE4" box="[603,984,1388,1414]" pageId="55" pageNumber="56" type="description">
<paragraph id="922136E09D2B926CFDC8FAD2CB8E2DE4" blockId="55.[512,1075,1352,1414]" box="[603,984,1388,1414]" pageId="55" pageNumber="56">
<figureCitation id="0AA52A659D2B926CFDC8FAD2CAE32DE4" box="[603,693,1388,1414]" captionStart="Figs 46–49" captionStartId="56.[189,241,1892,1918]" captionTargetBox="[364,1221,267,1852]" captionTargetId="figure-64@56.[361,1224,254,1856]" captionTargetPageId="56" captionText="Figs 46–49. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 46. M.interruptocarinulatusHeller, 1914,holotype, ♀ (SNSD).47. M.jobiensis Moser, 1920,lectotype, ♂ (ZMHB). 48. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 49. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122146" httpUri="https://zenodo.org/record/4122146/files/figure.png" pageId="55" pageNumber="56">Figs 46</figureCitation>
,
<figureCitation id="0AA52A659D2B926CFD52FAD2CAA62DE4" box="[705,752,1388,1414]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="55" pageNumber="56">126</figureCitation>
,
<figureCitation id="0AA52A659D2B926CFD6FFAD2CB7D2DE4" box="[764,811,1388,1414]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="55" pageNumber="56">219</figureCitation>
,
<figureCitation id="0AA52A659D2B926CFCA4FAD2CB302DE4" box="[823,870,1388,1414]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="55" pageNumber="56">309</figureCitation>
, 383, 477
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2B926CFF2EFA10C94B2F8B" pageId="55" pageNumber="56" type="materials_examined">
<paragraph id="922136E09D2B926CFF2EFA10C93A2DAA" blockId="55.[189,364,1454,1480]" box="[189,364,1454,1480]" pageId="55" pageNumber="56">
<heading id="C969818C9D2B926CFF2EFA10C93A2DAA" bold="true" box="[189,364,1454,1480]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
<emphasis id="A0EAEAF29D2B926CFF2EFA10C93A2DAA" bold="true" box="[189,364,1454,1480]" pageId="55" pageNumber="56">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2B926CFF79FA63C90A2D95" blockId="55.[189,1398,1501,1598]" box="[234,348,1501,1527]" pageId="55" pageNumber="56">
<emphasis id="A0EAEAF29D2B926CFF79FA63C90A2D95" bold="true" box="[234,348,1501,1527]" pageId="55" pageNumber="56">
<typeStatus id="4D2588429D2B926CFF79FA63C90A2D95" box="[234,348,1501,1527]" pageId="55" pageNumber="56" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D2B926CFF2EF9BECB0F2E5C" blockId="55.[189,1398,1501,1598]" pageId="55" pageNumber="56">
<materialsCitation id="22F63CBD9D2B926CFF2EF9BECB0F2E5C" ID-GBIF-Occurrence="2898878335" collectionCode="SNSD" country="Indonesia" location="INDONESIA" pageId="55" pageNumber="56" specimenCount="2" specimenCount-female="1" typeStatus="typus">
<collectingCountry id="EA8976709D2B926CFF2EF9BEC90D2E78" box="[189,347,1536,1562]" name="Indonesia" pageId="55" pageNumber="56">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D2B926CFEE5F9BFC9DB2E79" box="[374,397,1537,1563]" pageId="55" pageNumber="56" type="female">♀</specimenCount>
; “Nieuw.Guinea-Expeditie 190[p] 4 Etna-bai [h] //
<typeStatus id="4D2588429D2B926CFC7AF9BECC792E78" box="[1001,1071,1536,1562]" pageId="55" pageNumber="56">typus</typeStatus>
<specimenCount id="8498FD699D2B926CFBBDF9BECC642E78" box="[1070,1074,1536,1562]" pageId="55" pageNumber="56" type="male">♂</specimenCount>
[p, red label]// 1913 [p] 18 [h] // Staatl. Museum für Tierkunde.Dresden [p]”;
<collectionCode id="F48FAE259D2B926CFC95F99ACB032E5C" box="[774,853,1572,1598]" collectionName="SNSD" pageId="55" pageNumber="56">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D2B926CFF2EF9D8C9302E1D" blockId="55.[189,358,1637,1664]" box="[189,358,1637,1664]" pageId="55" pageNumber="56">
<heading id="C969818C9D2B926CFF2EF9D8C9302E1D" bold="true" box="[189,358,1637,1664]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
<emphasis id="A0EAEAF29D2B926CFF2EF9D8C9302E1D" bold="true" box="[189,358,1637,1664]" pageId="55" pageNumber="56">New material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2B926CFF2EF92AC9BB2E94" blockId="55.[189,1399,1684,1782]" pageId="55" pageNumber="56">
<materialsCitation id="22F63CBD9D2B926CFF2EF92AC9432EB0" ID-GBIF-Occurrence="2898878384" collectingDate="1933-07" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Papua New Guinea" elevation="914" location="PAPUA" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1">
<collectingCountry id="EA8976709D2B926CFF2EF92AC9B62ECC" box="[189,480,1684,1711]" name="Papua New Guinea" pageId="55" pageNumber="56">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D2B926CFD93F92ACA672ECD" box="[512,561,1684,1711]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
; “
<location id="9741603B9D2B926CFDDEF92ACAE42ECC" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFDDEF92ACAE42ECC" box="[589,690,1684,1710]" country="Papua New Guinea" name="PAPUA" pageId="55" pageNumber="56">PAPUA</location>
:
<location id="9741603B9D2B926CFD53F92ACB5F2ECC" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFD53F92ACB5F2ECC" box="[704,777,1684,1710]" country="Papua New Guinea" name="Orrori" pageId="55" pageNumber="56">Orrori</location>
<quantity id="55669B059D2B926CFC86F92ACB272ECC" box="[789,881,1684,1710]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="55" pageNumber="56" unit="ft" value="3000.0">
<elevation id="19B3D1D39D2B926CFC86F92ACB272ECC" box="[789,881,1684,1710]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="55" pageNumber="56" unit="ft" value="3000.0">3,000ft.</elevation>
</quantity>
<date id="E62010209D2B926CFCE8F92ACBB62ECC" box="[891,992,1684,1710]" pageId="55" pageNumber="56" value="1933-07">
<collectingDate id="F664E9C89D2B926CFCE8F92ACBB62ECC" box="[891,992,1684,1710]" pageId="55" pageNumber="56" value="1933-07">vii.1933</collectingDate>
</date>
.
<collectorName id="3F6B53369D2B926CFC7DF92ACCF52ECC" box="[1006,1187,1684,1710]" pageId="55" pageNumber="56">L.E.Cheesman.</collectorName>
B.M.1933-492.”;
<collectionCode id="F48FAE259D2B926CFF2EF906C9432EB0" box="[189,277,1720,1746]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D2B926CFEA1F906C9B72EB0" ID-GBIF-Occurrence="2898878519" box="[306,481,1720,1746]" collectionCode="NEW" country="Papua New Guinea" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1">
<specimenCount id="8498FD699D2B926CFEA1F906C9362EB0" box="[306,352,1720,1746]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
; “W.NEW
</materialsCitation>
<materialsCitation id="22F63CBD9D2B926CFE79F906C9BC2E94" ID-GBIF-Occurrence="2898878338" collectingDate="1936-02" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="213" location="Mt. Nomo." municipality="Mt. Bougainville." pageId="55" pageNumber="56" specimenCount="1">
<collectingCountry id="EA8976709D2B926CFE79F906CA092EB0" box="[490,607,1720,1746]" name="Guinea-Bissau" pageId="55" pageNumber="56">GUINEA</collectingCountry>
:
<location id="9741603B9D2B926CFDF8F906CAB02EB0" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFDF8F906CAB02EB0" box="[619,742,1720,1746]" country="Guinea-Bissau" municipality="Mt. Bougainville." name="Mt. Nomo." pageId="55" pageNumber="56">Mt.Nomo.</location>
S.of
<collectingMunicipality id="7245AC9A9D2B926CFCBBF906CBA72EB0" box="[808,1009,1720,1746]" pageId="55" pageNumber="56">Mt.Bougainville.</collectingMunicipality>
<quantity id="55669B059D2B926CFC69F906CC1E2EB0" box="[1018,1096,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="55" pageNumber="56" unit="ft" value="700.0">
<elevation id="19B3D1D39D2B926CFC69F906CC1E2EB0" box="[1018,1096,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="55" pageNumber="56" unit="ft" value="700.0">700 ft.</elevation>
</quantity>
<date id="E62010209D2B926CFBC2F906CCF12EB0" box="[1105,1191,1720,1746]" pageId="55" pageNumber="56" value="1936-02">
<collectingDate id="F664E9C89D2B926CFBC2F906CCF12EB0" box="[1105,1191,1720,1746]" pageId="55" pageNumber="56" value="1936-02">ii.1936</collectingDate>
</date>
.//
<collectorName id="3F6B53369D2B926CFB51F906CD212EB0" box="[1218,1399,1720,1746]" pageId="55" pageNumber="56">L.E.Cheesman.</collectorName>
B.M.1936-271.”;
<collectionCode id="F48FAE259D2B926CFE1DF962C9BC2E94" box="[398,490,1756,1782]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="922136E09D2B926CFF2EF8A3C94B2F8B" blockId="55.[189,1399,1821,2025]" pageId="55" pageNumber="56">
<materialsCitation id="22F63CBD9D2B926CFF2EF8A3CAF42F39" ID-GBIF-Occurrence="2898878420" collectingDate="1936-06" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Indonesia" elevation="610" location="DUTCH NEW GUINEA" municipality="Sabron." pageId="55" pageNumber="56" specimenCount="1">
<collectingCountry id="EA8976709D2B926CFF2EF8A3C90D2F55" box="[189,347,1821,1847]" name="Indonesia" pageId="55" pageNumber="56">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D2B926CFE16F8A3CA402F55" box="[389,534,1821,1847]" pageId="55" pageNumber="56" type="generic">1 specimen</specimenCount>
; “
<location id="9741603B9D2B926CFDA0F8A3CB242F55" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFDA0F8A3CB242F55" box="[563,882,1821,1847]" country="Indonesia" municipality="Sabron." name="DUTCH NEW GUINEA" pageId="55" pageNumber="56">DUTCH NEW GUINEA</location>
:
<location id="9741603B9D2B926CFC16F8A3CC7B2F55" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFC16F8A3CC7B2F55" box="[901,1069,1821,1847]" country="Indonesia" municipality="Sabron." name="Cyclops Mts." pageId="55" pageNumber="56">Cyclops Mts.</location>
,
<collectingMunicipality id="7245AC9A9D2B926CFBD2F8A3CCCA2F55" box="[1089,1180,1821,1847]" pageId="55" pageNumber="56">Sabron.</collectingMunicipality>
<quantity id="55669B059D2B926CFB38F8A3CD412F55" box="[1195,1303,1821,1847]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="55" pageNumber="56" unit="ft" value="2000.0">
<elevation id="19B3D1D39D2B926CFB38F8A3CD412F55" box="[1195,1303,1821,1847]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="55" pageNumber="56" unit="ft" value="2000.0">2,000 ft.</elevation>
</quantity>
<date id="E62010209D2B926CFA84F8A3CD242F55" box="[1303,1394,1821,1847]" pageId="55" pageNumber="56" value="1936-06">
<collectingDate id="F664E9C89D2B926CFA84F8A3CD242F55" box="[1303,1394,1821,1847]" pageId="55" pageNumber="56" value="1936-06">vi.1936</collectingDate>
</date>
.
<collectorName id="3F6B53369D2B926CFF2EF8FEC9272F38" box="[189,369,1856,1882]" pageId="55" pageNumber="56">L.E.Cheesman.</collectorName>
B.M.1936-271.”;
<collectionCode id="F48FAE259D2B926CFDD9F8FFCAF42F39" box="[586,674,1857,1883]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D2B926CFD2EF8FFCC6E2F1C" ID-GBIF-Occurrence="2898878453" collectingDate="1936-06" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="610" location="DUTCH NEW GUINEA" municipality="Sabron." pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1">
<specimenCount id="8498FD699D2B926CFD2EF8FFCABD2F39" box="[701,747,1857,1883]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
; “Sabron Camp 2. //
<location id="9741603B9D2B926CFC4DF8FFCD5D2F39" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFC4DF8FFCD5D2F39" box="[990,1291,1857,1883]" country="Guinea-Bissau" municipality="Sabron." name="DUTCH NEW GUINEA" pageId="55" pageNumber="56">DUTCH NEW GUINEA</location>
:
<location id="9741603B9D2B926CFA85F8FEC8A42F1C" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFA85F8FEC8A42F1C" country="Guinea-Bissau" municipality="Sabron." name="Cyclops Mts." pageId="55" pageNumber="56">Cyclops Mts.</location>
,
<collectingMunicipality id="7245AC9A9D2B926CFF65F8DAC9022F1C" box="[246,340,1892,1918]" pageId="55" pageNumber="56">Sabron.</collectingMunicipality>
<quantity id="55669B059D2B926CFEF7F8DAC9872F1C" box="[356,465,1892,1918]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="55" pageNumber="56" unit="ft" value="2000.0">
<elevation id="19B3D1D39D2B926CFEF7F8DAC9872F1C" box="[356,465,1892,1918]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="55" pageNumber="56" unit="ft" value="2000.0">2,000 ft.</elevation>
</quantity>
<date id="E62010209D2B926CFE42F8DACA7A2F1C" box="[465,556,1892,1918]" pageId="55" pageNumber="56" value="1936-06">
<collectingDate id="F664E9C89D2B926CFE42F8DACA7A2F1C" box="[465,556,1892,1918]" pageId="55" pageNumber="56" value="1936-06">vi.1936</collectingDate>
</date>
.
<collectorName id="3F6B53369D2B926CFDD2F8DACAA02F1C" box="[577,758,1892,1918]" pageId="55" pageNumber="56">L.E.Cheesman.</collectorName>
B.M.1936-271.”;
<collectionCode id="F48FAE259D2B926CFC73F8DACC6E2F1C" box="[992,1080,1892,1918]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D2B926CFBF0F8DACD702FC0" ID-GBIF-Occurrence="2898878628" collectingDate="1937-09" collectionCode="BMNH" collectorName="W. Stuber." country="Guinea-Bissau" location="DUTCH NEW GUINEA" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1">
<specimenCount id="8498FD699D2B926CFBF0F8DACCCC2F1C" box="[1123,1178,1892,1918]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
; “
<location id="9741603B9D2B926CFB28F8DAC9642FC3" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFB28F8DAC9642FC3" country="Guinea-Bissau" name="DUTCH NEW GUINEA" pageId="55" pageNumber="56">DUTCH NEW GUINEA</location>
:
<location id="9741603B9D2B926CFEADF839CA982FC0" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFEADF839CA982FC0" box="[318,718,1927,1954]" country="Guinea-Bissau" name="Humboldt Bay Dist. Bewani Mts." pageId="55" pageNumber="56">Humboldt Bay Dist. Bewani Mts.</location>
<date id="E62010209D2B926CFD45F839CB642FC3" box="[726,818,1927,1953]" pageId="55" pageNumber="56" value="1937-09">
<collectingDate id="F664E9C89D2B926CFD45F839CB642FC3" box="[726,818,1927,1953]" pageId="55" pageNumber="56" value="1937-09">ix.1937</collectingDate>
</date>
. [p] //
<collectorName id="3F6B53369D2B926CFCEBF839CBBD2FC3" box="[888,1003,1927,1953]" pageId="55" pageNumber="56">W.Stüber.</collectorName>
B.M. 1938-177.”;
<collectionCode id="F48FAE259D2B926CFB5DF836CD702FC0" box="[1230,1318,1928,1954]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D2B926CFAD2F839C94F2F8B" ID-GBIF-Occurrence="2898878424" collectingDate="1938-04" collectionCode="BMNH" collectorName="L. E. Cheesman." country="Guinea-Bissau" elevation="762" location="DUTCH NEW GUINEA" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1">
<specimenCount id="8498FD699D2B926CFAD2F839CD392FC0" box="[1345,1391,1927,1954]" pageId="55" pageNumber="56" type="female">1 ♀</specimenCount>
; “N.
<location id="9741603B9D2B926CFF73F815CA432FA7" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFF73F815CA432FA7" box="[224,533,1963,1989]" country="Guinea-Bissau" name="DUTCH NEW GUINEA" pageId="55" pageNumber="56">DUTCH NEW GUINEA</location>
:
<location id="9741603B9D2B926CFDB2F815CB452FA7" LSID="urn:lsid:plazi:treatment:1A3787F69D2B9262FD93FAF6C9C72EDD:9741603B9D2B926CFDB2F815CB452FA7" box="[545,787,1963,1989]" country="Guinea-Bissau" name="Waigeu. Camp Nok." pageId="55" pageNumber="56">Waigeu. Camp Nok.</location>
<quantity id="55669B059D2B926CFC8FF815CBD62FA7" box="[796,896,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="55" pageNumber="56" unit="ft" value="2500.0">
<elevation id="19B3D1D39D2B926CFC8FF815CBD62FA7" box="[796,896,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="55" pageNumber="56" unit="ft" value="2500.0">2,500 ft.</elevation>
</quantity>
<date id="E62010209D2B926CFC1AF815CBB52FA7" box="[905,995,1963,1989]" pageId="55" pageNumber="56" value="1938-04">
<collectingDate id="F664E9C89D2B926CFC1AF815CBB52FA7" box="[905,995,1963,1989]" pageId="55" pageNumber="56" value="1938-04">iv.1938</collectingDate>
</date>
.
<collectorName id="3F6B53369D2B926CFC63F815CCF32FA7" box="[1008,1189,1963,1989]" pageId="55" pageNumber="56">L.E.Cheesman.</collectorName>
B.M.1938-593.”;
<collectionCode id="F48FAE259D2B926CFF2EF871C94F2F8B" box="[189,281,1999,2025]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="55" pageNumber="56">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="C6E166689D249263FF2EF8DAC9072F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122146" ID-Zenodo-Dep="4122146" httpUri="https://zenodo.org/record/4122146/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,241,1892,1918]" targetBox="[364,1221,267,1852]" targetPageId="56">
<paragraph id="922136E09D249263FF2EF8DAC9072F8A" blockId="56.[189,1399,1892,2024]" pageId="56" pageNumber="57">
<emphasis id="A0EAEAF29D249263FF2EF8DAC9092F1C" bold="true" box="[189,351,1892,1918]" pageId="56" pageNumber="57">Figs 46–49.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D249263FD5EF8DACC6E2F1C" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[717,1080,1892,1918]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D249263FD5EF8DACB002F1C" box="[717,854,1892,1918]" italics="true" pageId="56" pageNumber="57">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D249263FCE0F8DACC6E2F1C" author="Macleay W. S." box="[883,1080,1892,1918]" pageId="56" pageNumber="57" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D249263FF2EF839C88C2FC3" bold="true" box="[189,218,1927,1953]" pageId="56" pageNumber="57">46</emphasis>
.
<taxonomicName id="559E4D639D249263FF76F836CACF2FC3" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[229,665,1927,1953]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D249263FF76F836CA502FC3" box="[229,518,1927,1953]" italics="true" pageId="56" pageNumber="57">M.interruptocarinulatus</emphasis>
<bibRefCitation id="F60F4B119D249263FD9AF839CACF2FC3" author="Heller K. M." box="[521,665,1927,1953]" pageId="56" pageNumber="57" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
,holotype, ♀ (SNSD).
<emphasis id="A0EAEAF29D249263FC04F839CBE22FC3" bold="true" box="[919,948,1927,1953]" pageId="56" pageNumber="57">47</emphasis>
.
<taxonomicName id="559E4D639D249263FC53F836CCB72FC3" authority="Moser, 1920" authorityName="Moser" authorityYear="1920" box="[960,1249,1927,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="jobiensis">
<emphasis id="A0EAEAF29D249263FC53F836CC1D2FC3" box="[960,1099,1927,1953]" italics="true" pageId="56" pageNumber="57">M.jobiensis</emphasis>
<bibRefCitation id="F60F4B119D249263FBDCF836CCB72FC3" author="Moser J." box="[1103,1249,1927,1954]" pageId="56" pageNumber="57" pagination="3 - 28" refId="ref98398" refString="Moser J. 1920. Beitrag zur Kenntnis der Melolonthiden XI. Stettiner Entomologische Zeitung 81: 3 - 28." type="journal article" year="1920">Moser, 1920</bibRefCitation>
</taxonomicName>
,lectotype, ♂ (ZMHB).
<emphasis id="A0EAEAF29D249263FEA0F814C9062FA6" bold="true" box="[307,336,1962,1988]" pageId="56" pageNumber="57">48</emphasis>
.
<taxonomicName id="559E4D639D249263FECDF815C9A92FA6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[350,511,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="kazantsevi" status="sp. nov.">
<emphasis id="A0EAEAF29D249263FECDF815C9A92FA6" box="[350,511,1962,1988]" italics="true" pageId="56" pageNumber="57">M. kazantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D249263FD96F815CA342FA7" box="[517,610,1963,1989]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH).
<emphasis id="A0EAEAF29D249263FCE4F814CBC22FA6" bold="true" box="[887,916,1962,1988]" pageId="56" pageNumber="57">49</emphasis>
.
<taxonomicName id="559E4D639D249263FC31F815CC5F2FA6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[930,1033,1962,1988]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="konjo" status="sp. nov.">
<emphasis id="A0EAEAF29D249263FC31F815CC5F2FA6" box="[930,1033,1962,1988]" italics="true" pageId="56" pageNumber="57">M. konjo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D249263FB83F815CC3A2FA7" box="[1040,1132,1963,1989]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NMPC). Not to scale.
</paragraph>
</caption>
<subSubSection id="DA84656B9D259262FF2EFEB7C9D329D7" pageId="57" pageNumber="58" type="discussion">
<paragraph id="922136E09D259262FF2EFEB7C9792941" blockId="57.[189,1399,265,437]" box="[189,303,265,291]" pageId="57" pageNumber="58">
<heading id="C969818C9D259262FF2EFEB7C9792941" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="A0EAEAF29D259262FF2EFEB7C9792941" bold="true" box="[189,303,265,291]" pageId="57" pageNumber="58">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D259262FF2EFE8BC9D329D7" blockId="57.[189,1399,265,437]" pageId="57" pageNumber="58">
<bibRefCitation id="F60F4B119D259262FF2EFE8BC9CE2932" author="Heller K. M." box="[189,408,309,336]" pageId="57" pageNumber="58" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 624)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D259262FD2CFE88CBB2292D" authorityName="Heller" authorityYear="1914" box="[703,996,309,335]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="interruptocarinulatus">
<emphasis id="A0EAEAF29D259262FD2CFE88CBB2292D" box="[703,996,309,335]" italics="true" pageId="57" pageNumber="58">M. interruptocarinulatus</emphasis>
</taxonomicName>
on a single specimen, mentioning that the
<typeStatus id="4D2588429D259262FE8CFEE9C9D02913" box="[287,390,343,369]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
was deposited at ZMHB. However, the only type specimen with label data as in the original description was allocated in SNSD and has never been deposited in ZMHB (B. Jaeger, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D259262FF2EFE64C9C72EDD" pageId="57" pageNumber="58" type="description">
<paragraph id="922136E09D259262FF2EFE64C9182996" blockId="57.[189,334,474,500]" box="[189,334,474,500]" pageId="57" pageNumber="58">
<heading id="C969818C9D259262FF2EFE64C9182996" bold="true" box="[189,334,474,500]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="A0EAEAF29D259262FF2EFE64C9182996" bold="true" box="[189,334,474,500]" pageId="57" pageNumber="58">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D259262FF2EFDB6CD382A40" blockId="57.[189,1390,519,546]" box="[189,1390,519,546]" pageId="57" pageNumber="58">
MEASUREMENTS. Total body length 7.40 (
<typeStatus id="4D2588429D259262FD00FDB9CAAB2A40" box="[659,765,519,546]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
) to 8.60 (selected specimen from Cyclops Mts.) mm.
</paragraph>
<paragraph id="922136E09D259262FF2EFDF6CCEC2E31" blockId="57.[189,1399,583,1619]" pageId="57" pageNumber="58">
Dorsum and venter uniformly black-brown except for brown appendages and labroclypeus. Head slightly convex dorsally on frons between eyes, glossy dorsally and ventrally. Female labroclypeus (
<figureCitation id="0AA52A659D259262FF55FD34C97D2AC7" box="[198,299,650,677]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="57" pageNumber="58">Fig. 126</figureCitation>
) broadly emarginate anteriorly, its lateral margins nearly straight in dorsal, sinuous in lateral view. Anterolateral angles almost right-angled, not protruding, raised up at angle of nearly 80° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Upper- and underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus nearly straight in dorsal view. Punctures of frons ovoid, deep and dense; inner margin with thin membrane covered with dense microscopical velvety pubescence. Intervening spaces glossy, generally smaller than punctures. Setae inconspicuous, suberect; each seta rises from anterior margin of corresponding puncture, not or hardly surpassing its length. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum broadly sinuous with protruding anterolateral angles (stronger) and mesal portion (slightly), basal margin broadly rounded. Lateral margin of pronotum broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D259262FDB6FC65CAD82B97" box="[549,654,987,1013]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="57" pageNumber="58">Fig. 219</figureCitation>
). An appressed short seta present between every two crenulae. Lateral and basal margins, antero- and posterolateral angles partly covered with dense microscopical velvety pubescence. Lateral margin of pronotum slightly sinuous in lateral view. Hypomeron emarginate and long setose on anterior margin which is flange-like protruding. Antennal pocket deep. Punctures of pronotal disc generally larger and stronger oblong than those on head, inner margin with thin membrane covered with dense microscopical velvety pubescence. Intervening spaces glossy, in part wrinkled. Setation scale-like, appressed to suberect, short. Seta generally not surpassing length of corresponding puncture. Scutellar shield narrowly rounded apically. Elytron glossy, with five poorly defined glabrous variously broadly interrupted longitudinal carinae, including sutural one. Punctures of elytral disc ovoid, generally smaller than on forebody, deep (
<figureCitation id="0AA52A659D259262FD3EFAB4CB432D46" box="[685,789,1290,1316]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="57" pageNumber="58">Fig. 309</figureCitation>
). Each puncture encircled with dense microscopical velvety pubescence. Intervening spaces glossy, in part glabrous, generally smaller than punctures. Setae inconspicuous, similar to those on pronotum, becoming longer and less appressed along longitudinal carinae. Each elytron with long lateral subhumeral seta on lateral margin. Female pygidium flattened dorsally, with vague median longitudinal impression, covered with dense microscopical fur-like velvety pubescence (
<figureCitation id="0AA52A659D259262FEC1FA0CC9EE2DAE" box="[338,440,1458,1484]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="57" pageNumber="58">Fig. 477</figureCitation>
). Punctures of pygidium dense, setae long, dense, erect, intervening spaces opaque, microreticulate. Punctures of venter narrowly surrounded by dense fur-like microscopical pubescence. Female protibia with upper longitudinal carina, slightly widened distally, with two distal teeth of which basal one is broader and stronger obtuse (Fig. 383). Female protibial terminal spur straight. Female metatibial terminal spurs rounded apically. Tarsal claws with pulvilli (in female only?).
</paragraph>
<paragraph id="922136E09D259262FF2EF9C6C9E62EF0" blockId="57.[189,432,1656,1682]" box="[189,432,1656,1682]" pageId="57" pageNumber="58">
<heading id="C969818C9D259262FF2EF9C6C9E62EF0" bold="true" box="[189,432,1656,1682]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="A0EAEAF29D259262FF2EF9C6C9E62EF0" bold="true" box="[189,432,1656,1682]" pageId="57" pageNumber="58">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D259262FF2EF91BC9C72EDD" blockId="57.[189,401,1701,1727]" box="[189,401,1701,1727]" pageId="57" pageNumber="58">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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