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<document id="FBBF1C4AE63B81A6B3E9D4865263D08E" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D0F924FFDC9FB8DCCE22EF0" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius alesbezdeki Telnov 2020, sp. nov." docType="treatment" docUuid="A7930BCD-9290-497E-ABAF-137253AB5D2E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="21" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="20" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B5BD89033C080EE1D11DD6AA9C45468F">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="5A3D78AD538DE1DCA96767B0D9AFEB16" type="personal">
<mods:role id="7E9742FECB4225BB0F85A0E8DD498ACE">
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<mods:namePart id="6178F6AD73D05D99EBDA7B1AAC63ACB0">Telnov, Dmitry</mods:namePart>
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<mods:title id="8373C908F31E3B9DB2AF52F1E1F278FE">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="CB56BF3834AF900E6F7380B1E6915F14">
<mods:date id="C6FD1BD78AC7B2BF95AF260C95B1764B">2020</mods:date>
<mods:detail id="741F1B1174D0B0F690C8C0324D6383FC" type="pubDate">
<mods:number id="7857CF1DB6E350A950B6A66A4EA6367E">2020-10-19</mods:number>
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<mods:number id="5C27911A1513F8BEEBF1C51B6B09745E">721</mods:number>
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<mods:classification id="5AE9438DA87FFD49D30D2605A87AB941">journal article</mods:classification>
<mods:identifier id="DCDD71026E95EE059E84DE7FC21EBD7B" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="57552D74DCBDA2A03A324FFD2820D0FD" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="AB60A5E3380A6989EA564DA8E00D694F" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="1E1C0CDE27A9FF5823B3BE0E1EB65181" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="D8A7DCD69818F40E94211A4E576D8926" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D0F924FFDC9FB8DCCE22EF0" ID-DOI="http://doi.org/10.5281/zenodo.4344549" ID-GBIF-Taxon="168648180" ID-Zenodo-Dep="4344549" LSID="urn:lsid:zoobank.org:act:A7930BCD-9290-497E-ABAF-137253AB5D2E" httpUri="http://treatment.plazi.org/id/1A3787F69D0F924FFDC9FB8DCCE22EF0" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="DA84656B9D0F9248FDC9FB8DCB8C2C2C" box="[602,986,1075,1102]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="922136E09D0F9248FDC9FB8DCB8C2C2C" blockId="19.[359,1228,1075,1171]" box="[602,986,1075,1102]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FDC9FB8DCB8C2C2C" box="[602,986,1075,1102]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName id="559E4D639D0F9248FDC9FB8DCB2E2C2C" ID-CoL="7X3W5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[602,888,1075,1102]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alesbezdeki" status="sp. nov.">
<emphasis id="A0EAEAF29D0F9248FDC9FB8DCB2E2C2C" bold="true" box="[602,888,1075,1102]" italics="true" pageId="19" pageNumber="20">Maechidius alesbezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0F9248FCECFB8ACB8C2C2C" box="[895,986,1076,1102]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FEF4FBE8CC692CF1" pageId="19" pageNumber="20" type="description">
<paragraph id="922136E09D0F9248FEF4FBE8CC9A2C13" blockId="19.[359,1228,1075,1171]" box="[359,1228,1110,1137]" pageId="19" pageNumber="20">
<uri id="E60F3AE29D0F9248FEF4FBE8CC9A2C13" box="[359,1228,1110,1137]" pageId="19" pageNumber="20">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D0F9248FD12FBE8CC9A2C13" box="[641,1228,1110,1137]" pageId="19" pageNumber="20">A7930BCD-9290-497E-ABAF-137253AB5D2E</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D0F9248FE66FBC6CC692CF1" blockId="19.[359,1228,1075,1171]" box="[501,1087,1144,1171]" pageId="19" pageNumber="20">
<figureCitation id="0AA52A659D0F9248FE66FBC6CA2C2CF0" box="[501,634,1144,1170]" captionStart="Figs 18–21" captionStartId="17.[188,240,1928,1954]" captionTargetBox="[353,1214,268,1904]" captionTargetId="figure-12@17.[346,1240,250,1910]" captionTargetPageId="17" captionText="Figs 18–21. Papuan Maechidius Macleay, 1819, habitus, dorsal and lateral views. 18. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 19. M. aiyura sp. nov., holotype, ♂ (BMNH). 20–21. M. alesbezdeki sp. nov. ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122130" httpUri="https://zenodo.org/record/4122130/files/figure.png" pageId="19" pageNumber="20">Figs 20–21</figureCitation>
,
<figureCitation id="0AA52A659D0F9248FD14FBC6CAF12CF0" box="[647,679,1144,1170]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="19" pageNumber="20">99</figureCitation>
,
<figureCitation id="0AA52A659D0F9248FD20FBC6CAB42CF0" box="[691,738,1144,1170]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="19" pageNumber="20">199</figureCitation>
,
<figureCitation id="0AA52A659D0F9248FD7EFBC6CB4A2CF0" box="[749,796,1144,1170]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="19" pageNumber="20">288</figureCitation>
, 361, 454, 567, 573–575
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FF2EFB07CA062DE9" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="922136E09D0F9248FF2EFB07C99F2CB1" blockId="19.[189,457,1209,1235]" box="[189,457,1209,1235]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EFB07C99F2CB1" bold="true" box="[189,457,1209,1235]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EFB07C99F2CB1" bold="true" box="[189,457,1209,1235]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EFB59CA062DE9" blockId="19.[189,1399,1255,1419]" pageId="19" pageNumber="20">
Among Papuan congeners,
<taxonomicName id="559E4D639D0F9248FD82FB56CA932D63" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[529,709,1255,1281]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="alesbezdeki" status="sp. nov.">
<emphasis id="A0EAEAF29D0F9248FD82FB56CA932D63" box="[529,709,1255,1281]" italics="true" pageId="19" pageNumber="20">M. alesbezdeki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0F9248FD41FB59CB642D63" box="[722,818,1255,1281]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
is readily distinguishable primarily due to the combination of the following characters: labroclypeus deeply U-shaped emarginate anteriorly with protruding obtuse angulate anterolateral angles and sinuous lateral margins, dense punctured frons, denticulate lateral margin of pronotum (crenulate in most congeners), linear (incision-shaped) punctured elytra and shape of male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FF2EFA0CCA802E7F" pageId="19" pageNumber="20" type="etymology">
<paragraph id="922136E09D0F9248FF2EFA0CC9122DAE" blockId="19.[189,324,1458,1484]" box="[189,324,1458,1484]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EFA0CC9122DAE" bold="true" box="[189,324,1458,1484]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EFA0CC9122DAE" bold="true" box="[189,324,1458,1484]" pageId="19" pageNumber="20">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EFA61CA802E7F" blockId="19.[189,1399,1503,1566]" pageId="19" pageNumber="20">
Patronymic. This species is named after the famous
<taxonomicName id="559E4D639D0F9248FCB9FA61CB932D9B" authorityName="Latreille" authorityYear="1802" box="[810,965,1503,1529]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
expert and respected colleague Aleš Bezděk (České Budějovice,
<collectingCountry id="EA8976709D0F9248FD9AF9BCCA9F2E7F" box="[521,713,1538,1565]" name="Czech Republic" pageId="19" pageNumber="20">Czech Republic</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F9248FF2EF9FDCA762E90" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="922136E09D0F9248FF2EF9FDC93A2E3E" blockId="19.[189,364,1602,1629]" box="[189,364,1602,1629]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EF9FDC93A2E3E" bold="true" box="[189,364,1602,1629]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EF9FDC93A2E3E" bold="true" box="[189,364,1602,1629]" pageId="19" pageNumber="20">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF79F9CFC90A2EE9" blockId="19.[189,1399,1649,1778]" box="[234,348,1649,1675]" pageId="19" pageNumber="20">
<emphasis id="A0EAEAF29D0F9248FF79F9CFC90A2EE9" bold="true" box="[234,348,1649,1675]" pageId="19" pageNumber="20">
<typeStatus id="4D2588429D0F9248FF79F9CFC90A2EE9" box="[234,348,1649,1675]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D0F9248FF2EF92DCA762E90" blockId="19.[189,1399,1649,1778]" pageId="19" pageNumber="20">
<materialsCitation id="22F63CBD9D0F9248FF2EF92DCA762E90" ID-GBIF-Occurrence="2898878635" collectedFrom="edge of primary mid montane rainforest" collectingDate="2015-09-10" collectingDateMax="2015-09-11" collectingDateMin="2015-09-10" collectingMethod="white light" collectionCode="NME" country="Indonesia" elevation="2200" latitude="-1.3013889" location="Arfak mts" longLatPrecision="21" longitude="133.90666" municipality="Anggi Gigi Lake" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<collectingCountry id="EA8976709D0F9248FF2EF92DC90D2ECF" box="[189,347,1683,1709]" name="Indonesia" pageId="19" pageNumber="20">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D0F9248FEE4F92DC9D92ECF" box="[375,399,1683,1709]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D0F9248FE3BF92DCA1F2ECF" box="[424,585,1683,1709]" name="Indonesia" pageId="19" pageNumber="20">INDONESIA</collectingCountry>
E,
<locationDeviation id="83932FC29D0F9248FDE7F92DCB672ECF" box="[628,817,1683,1709]" pageId="19" pageNumber="20">W New Guinea</locationDeviation>
, Doberai Peninsula,
<location id="9741603B9D0F9248FBB5F92DCCF62ECF" LSID="urn:lsid:plazi:treatment:1A3787F69D0F924FFDC9FB8DCCE22EF0:9741603B9D0F9248FBB5F92DCCF62ECF" box="[1062,1184,1683,1709]" country="Indonesia" latitude="-1.3013889" longLatPrecision="21" longitude="133.90666" municipality="Anggi Gigi Lake" name="Arfak mts" pageId="19" pageNumber="20">Arfak mts</location>
,
<collectingMunicipality id="7245AC9A9D0F9248FB3FF92DCD212ECF" box="[1196,1399,1683,1709]" pageId="19" pageNumber="20">Anggi Gigi Lake</collectingMunicipality>
S env., Uper vill.,
<geoCoordinate id="F7AA50279D0F9248FE32F90BCA762EB2" box="[417,544,1717,1744]" degrees="1" direction="south" minutes="18" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="05" value="-1.3013889">1°18’05”S</geoCoordinate>
,
<geoCoordinate id="F7AA50279D0F9248FDA3F90BCA9B2EB2" box="[560,717,1717,1744]" degrees="133" direction="east" minutes="54" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="24" value="133.90666">133°54’24”E</geoCoordinate>
,
<date id="E62010209D0F9248FD4EF90BCBDA2EB2" box="[733,908,1717,1744]" pageId="19" pageNumber="20" value="2015-09-10" valueMax="2015-09-11" valueMin="2015-09-10">
<collectingDate id="F664E9C89D0F9248FD4EF90BCBDA2EB2" box="[733,908,1717,1744]" pageId="19" pageNumber="20" value="2015-09-10" valueMax="2015-09-11" valueMin="2015-09-10">10-11.IX.2015</collectingDate>
</date>
,
<quantity id="55669B059D0F9248FC0FF908CBAC2EB2" box="[924,1018,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="m" value="2200.0">
<elevation id="19B3D1D39D0F9248FC0FF908CBAC2EB2" box="[924,1018,1718,1744]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="20" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<collectedFrom id="2182751D9D0F9248FB98F90BC9672E90" pageId="19" pageNumber="20">edge of primary mid montane rainforest</collectedFrom>
,
<collectingMethod id="4BDF4EF79D0F9248FEAFF966C9972E90" box="[316,449,1752,1778]" pageId="19" pageNumber="20">white light</collectingMethod>
”;
<collectionCode id="F48FAE259D0F9248FE44F966CA4A2E90" box="[471,540,1752,1778]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="19" pageNumber="20">NME</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0F924FFF2EF8A6C9FD2DD3" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="922136E09D0F9248FF2EF8A6C9182F50" blockId="19.[189,334,1816,1842]" box="[189,334,1816,1842]" pageId="19" pageNumber="20">
<heading id="C969818C9D0F9248FF2EF8A6C9182F50" bold="true" box="[189,334,1816,1842]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="A0EAEAF29D0F9248FF2EF8A6C9182F50" bold="true" box="[189,334,1816,1842]" pageId="19" pageNumber="20">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0F9248FF2EF8F9CD212FE1" blockId="19.[189,1399,1862,1924]" pageId="19" pageNumber="20">
MEASUREMENTS. Total body length
<quantity id="55669B059D0F9248FDC2F8F8CAEA2F03" box="[593,700,1862,1889]" metricMagnitude="-3" metricUnit="m" metricValue="8.67" pageId="19" pageNumber="20" unit="mm" value="8.67">8.67 mm</quantity>
. Head
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long, across eyes
<quantity id="55669B059D0F9248FBD8F8F8CCE22F03" box="[1099,1204,1862,1889]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="19" pageNumber="20" unit="mm" value="1.86">1.86 mm</quantity>
wide. Pronotum
<quantity id="55669B059D0F9248FF2EF8D7C9732FE1" box="[189,293,1897,1923]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="20" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D0F9248FDA7F8D7CAC92FE1" box="[564,671,1897,1923]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="19" pageNumber="20" unit="mm" value="2.6">2.60 mm</quantity>
. Elytral length
<quantity id="55669B059D0F9248FCC2F8D7CBED2FE1" box="[849,955,1897,1924]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="19" pageNumber="20" unit="mm" value="5.3">5.30 mm</quantity>
, maximum combined width
<quantity id="55669B059D0F9248FA9BF8D7CD252FE1" box="[1288,1395,1897,1924]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="20" unit="mm" value="3.5">3.50 mm</quantity>
.
</paragraph>
<paragraph id="922136E09D0F924FFF2EF815CD212D20" blockId="19.[189,1398,1963,2025]" lastBlockId="20.[189,1399,265,1346]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Dorsum black-brown with castaneous labroclypeus, anterior and lateral margins of pronotum, mouthparts and legs. Venter uniformly dark castaneous brown. Head flattened between eyes, subopaque dorsally and ventrally, with large slightly prominent eyes occupying about half side of head. Male labroclypeus very deeply and rather narrowly U-shaped emarginate (
<figureCitation id="0AA52A659D08924FFC96FE93CB0A2925" box="[773,860,301,327]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="20" pageNumber="21">Fig. 99</figureCitation>
), its lateral margins sinuous in both dorsal and lateral views, anterolateral angles large, obtuse angulate and raised up at angle of nearly 80° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Punctures of frons annular, large and shallow. Intervening spaces microreticulate, generally much smaller than punctures. Pubescence dirty yellowish, sparse, erect to suberect with longest setae near eyes and on underside of labroclypeus. Each seta rises from anterior margin of corresponding puncture. Male antenna 8-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 transverse. Pronotum flattened dorsally, glossy dorsally and laterally. Anterior margin broadly emarginate, slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin broadly rounded. Lateral margin broadly rounded, constricted towards base in basal third. Along lateral margin with 12–13 acute denticles (two anterior less conspicuous, median ones stronger) (
<figureCitation id="0AA52A659D08924FFD32FD03CB5E2AB5" box="[673,776,701,727]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="20" pageNumber="21">Fig. 199</figureCitation>
). Long erect seta present between each two denticles. Lateral margin of pronotum nearly straight in lateral view. Punctures on disc ovoid, annular, generally larger and deeper than those on head, becoming flatter and regularly circular along lateral margins. Intervening spaces generally smaller than punctures, glossy. Setation like on head. Hypomeron nearly straight, separated from prosternum by low carina, with long setae on anterolateral margin. Antennal pocket shallow. Scutellar shield narrowly rounded apically. Elytra cylindrical, maximum width across median third, opaque and flattened dorsally, with rounded humeri. Obtuse hump on each elytron near apex. No tracks of longitudinal carinae, sutural carinae also not present. Disc and lateral sides of elytra in irregular delicate wrinkles between dense, narrow and rather deep linear incision-shaped punctures (
<figureCitation id="0AA52A659D08924FFF55FBBAC97B2C7D" box="[198,301,1028,1055]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="20" pageNumber="21">Fig. 288</figureCitation>
). Inconspicuous suberect seta rises from anterior margin of each puncture; setae variably long. Epipleuron with row of cilia-like erect setae. Male pygidium and abdominal ventrites densely covered with large and shallow annular punctures, each provided with short seta (
<figureCitation id="0AA52A659D08924FFB9CFBF3CC202C0A" box="[1039,1142,1101,1128]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="20" pageNumber="21">Fig. 454</figureCitation>
). Abdominal ventrites with no track of lateral longitudinal ridge. Legs long and slender, femora and tibiae covered with long erect setae arranged in 4–5 lines. Protibiae straight, metatibiae stouter than protibiae and thickened distally. Protibia with track of inconspicuous carina on dorsal surface. Three teeth on external margin of male protibia: two large pointed distal ones and broad obtuse inconspicuous basal one same distance from median as median one from distal (Fig. 361). Male protibial terminal spur straight. Tarsal claws with large pulvilli (in males only?). Spiculum gastrale as in
<figureCitation id="0AA52A659D08924FFCE2FA99CB802D20" box="[881,982,1319,1346]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="20" pageNumber="21">Fig. 567</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D08924FFB43FA99CD252D20" box="[1232,1395,1319,1346]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="20" pageNumber="21">Figs 573–575</figureCitation>
.
</paragraph>
<paragraph id="922136E09D08924FFF2EFAD6C9E62DE1" blockId="20.[189,432,1384,1411]" box="[189,432,1384,1411]" pageId="20" pageNumber="21">
<heading id="C969818C9D08924FFF2EFAD6C9E62DE1" bold="true" box="[189,432,1384,1411]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="A0EAEAF29D08924FFF2EFAD6C9E62DE1" bold="true" box="[189,432,1384,1411]" pageId="20" pageNumber="21">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D08924FFF2EFA29C9FD2DD3" blockId="20.[189,427,1431,1457]" box="[189,427,1431,1457]" pageId="20" pageNumber="21">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D08924FFF2EFA67CCC32E43" box="[189,1173,1497,1570]" pageId="20" pageNumber="21" type="biology_ecology">
<paragraph id="922136E09D08924FFF2EFA67C9762D91" blockId="20.[189,288,1497,1523]" box="[189,288,1497,1523]" pageId="20" pageNumber="21">
<heading id="C969818C9D08924FFF2EFA67C9762D91" bold="true" box="[189,288,1497,1523]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="A0EAEAF29D08924FFF2EFA67C9762D91" bold="true" box="[189,288,1497,1523]" pageId="20" pageNumber="21">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D08924FFF2EF9B6CCC32E43" blockId="20.[189,1173,1543,1570]" box="[189,1173,1543,1570]" pageId="20" pageNumber="21">
Occurs in primary mid-montane rainforests at ~
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altitude. Possibly nocturnal.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D08924FFF2EF9F7CCE22EF0" box="[189,1204,1609,1682]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="922136E09D08924FFF2EF9F7C9002E01" blockId="20.[189,342,1609,1635]" box="[189,342,1609,1635]" pageId="20" pageNumber="21">
<heading id="C969818C9D08924FFF2EF9F7C9002E01" bold="true" box="[189,342,1609,1635]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="A0EAEAF29D08924FFF2EF9F7C9002E01" bold="true" box="[189,342,1609,1635]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D08924FFF2EF9C6CCE22EF0" blockId="20.[189,1204,1656,1682]" box="[189,1204,1656,1682]" pageId="20" pageNumber="21">
Hitherto only known from Arfak Mountains, Doberai Peninsula, western New
<collectingCountry id="EA8976709D08924FFBC4F9C6CCE62EF0" box="[1111,1200,1656,1682]" name="Guinea-Bissau" pageId="20" pageNumber="21">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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