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<document id="0DEF12A8756DED97BDEF8A2EC99241B6" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D0B9242FD16FBDEC9A62ACF" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius awu Telnov 2020, sp. nov." docType="treatment" docUuid="86803286-3157-4DB2-9A03-6040AD2D2226" docUuidSource="ZooBank" docVersion="10" lastPageNumber="26" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="24" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="52AE320AF8D6717FE16265CD9FA154E4">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="CE3437D71CB6EE9A5BF1390244BED444">Telnov, Dmitry</mods:namePart>
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<mods:title id="511D78CF9318B17E4A4A40F88CDD4A92">European Journal of Taxonomy</mods:title>
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<mods:date id="FD4DF129BBC8BC54BBA2B67B901A7925">2020</mods:date>
<mods:detail id="3BD360A8E06C9212580B45E67003213C" type="pubDate">
<mods:number id="13A86A084426225F8A9A60A0C39BD760">2020-10-19</mods:number>
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<mods:identifier id="C0066FEEED505EBD1936847AEFC275A4" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="9CA432F7B1C716A8C2775F0847E0A58D" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D0B9242FD16FBDEC9A62ACF" ID-DOI="http://doi.org/10.5281/zenodo.4344485" ID-GBIF-Taxon="168648182" ID-Zenodo-Dep="4344485" LSID="urn:lsid:zoobank.org:act:86803286-3157-4DB2-9A03-6040AD2D2226" httpUri="http://treatment.plazi.org/id/1A3787F69D0B9242FD16FBDEC9A62ACF" lastPageId="25" lastPageNumber="26" pageId="23" pageNumber="24">
<subSubSection id="DA84656B9D0B924CFD16FBDECBF92C19" box="[645,943,1120,1147]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="922136E09D0B924CFD16FBDECBF92C19" blockId="23.[374,1214,1120,1223]" box="[645,943,1120,1147]" pageId="23" pageNumber="24">
<heading id="C969818C9D0B924CFD16FBDECBF92C19" box="[645,943,1120,1147]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="24" reason="2">
<taxonomicName id="559E4D639D0B924CFD16FBDECB1B2C19" ID-CoL="7X3W8" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[645,845,1120,1147]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="awu" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFD16FBDECB1B2C19" bold="true" box="[645,845,1120,1147]" italics="true" pageId="23" pageNumber="24">Maechidius awu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFCC7FBDFCBF92C19" box="[852,943,1121,1147]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0B924CFEE5FB38CC1B2CA5" pageId="23" pageNumber="24" type="description">
<paragraph id="922136E09D0B924CFEE5FB38CCE82CC3" blockId="23.[374,1214,1120,1223]" box="[374,1214,1158,1185]" pageId="23" pageNumber="24">
<uri id="E60F3AE29D0B924CFEE5FB38CCE82CC3" box="[374,1214,1158,1185]" pageId="23" pageNumber="24">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D0B924CFD00FB38CCE82CC3" box="[659,1214,1158,1185]" pageId="23" pageNumber="24">86803286-3157-4DB2-9A03-6040AD2D2226</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D0B924CFE75FB12CC1B2CA5" blockId="23.[374,1214,1120,1223]" box="[486,1101,1196,1223]" pageId="23" pageNumber="24">
<figureCitation id="0AA52A659D0B924CFE75FB12CA162CA5" box="[486,576,1196,1223]" captionStart="Figs 22–24" captionStartId="22.[189,241,1928,1954]" captionTargetBox="[354,1240,272,1893]" captionTargetId="figure-55@22.[344,1244,264,1901]" captionTargetPageId="22" captionText="Figs 22–24. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 22. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 23. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 24. M. awu sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122132" httpUri="https://zenodo.org/record/4122132/files/figure.png" pageId="23" pageNumber="24">Figs 24</figureCitation>
,
<figureCitation id="0AA52A659D0B924CFDDFFB13CAE32CA5" box="[588,693,1197,1223]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="23" pageNumber="24">103–104</figureCitation>
,
<figureCitation id="0AA52A659D0B924CFD52FB13CAA62CA5" box="[705,752,1197,1223]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="23" pageNumber="24">202</figureCitation>
,
<figureCitation id="0AA52A659D0B924CFD6FFB13CB7D2CA5" box="[764,811,1197,1223]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="23" pageNumber="24">292</figureCitation>
, 364, 457, 537, 579–581
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0B924CFF2EFB4FCCF12E43" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="922136E09D0B924CFF2EFB4FC99F2D69" blockId="23.[189,457,1265,1291]" box="[189,457,1265,1291]" pageId="23" pageNumber="24">
<heading id="C969818C9D0B924CFF2EFB4FC99F2D69" bold="true" box="[189,457,1265,1291]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A0EAEAF29D0B924CFF2EFB4FC99F2D69" bold="true" box="[189,457,1265,1291]" pageId="23" pageNumber="24">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0B924CFF2EFA9CCCF12E43" blockId="23.[189,1399,1314,1569]" pageId="23" pageNumber="24">
Undoubtedly very close to
<collectingRegion id="505AF8029D0B924CFE60FA9CCA0B2D5E" box="[499,605,1314,1340]" country="Indonesia" name="Sulawesi Utara" pageId="23" pageNumber="24">Sulawesi</collectingRegion>
congeners, differs primarily in shape of male genitalia (cf.
<figureCitation id="0AA52A659D0B924CFA97FA9CC8BA2D01" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="23" pageNumber="24">Figs 579– 581</figureCitation>
), comparatively deeper emarginate anterior margin of male labroclypeus and somewhat medially angulate lateral margin of female pronotum (seen in dorsal view). In the shape of the aedeagus,
<taxonomicName id="559E4D639D0B924CFA8CFAD1CD212DEA" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1311,1399,1391,1416]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="awu" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFA8CFAD1CD212DEA" box="[1311,1399,1391,1416]" italics="true" pageId="23" pageNumber="24">M. awu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFF2EFA2BC94F2DCD" box="[189,281,1429,1455]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
strongly resembles
<taxonomicName id="559E4D639D0B924CFD94FA2BCADA2DCC" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[519,652,1428,1454]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFD94FA2BCADA2DCC" box="[519,652,1428,1454]" italics="true" pageId="23" pageNumber="24">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFD07FA2BCAA62DCD" box="[660,752,1429,1455]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
from
<collectingRegion id="505AF8029D0B924CFCA9FA2ACBA62DCC" box="[826,1008,1428,1454]" country="Indonesia" name="Sulawesi Utara" pageId="23" pageNumber="24">North Sulawesi</collectingRegion>
(see description below), but it is readily distinguishable in male pygidium evenly convex (humped posteriorly in
<taxonomicName id="559E4D639D0B924CFBE8FA05CD572DB6" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1147,1281,1466,1492]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFBE8FA05CD572DB6" box="[1147,1281,1466,1492]" italics="true" pageId="23" pageNumber="24">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFA99FA05CD3F2DB7" box="[1290,1385,1467,1493]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
), dorsally evenly convex pronotal disc (with circular impression on either side of the midline in
<taxonomicName id="559E4D639D0B924CFB67FA5FCD212D98" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1268,1399,1504,1530]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFB67FA5FCD212D98" box="[1268,1399,1504,1530]" italics="true" pageId="23" pageNumber="24">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFF2EF9B9C94F2E43" box="[189,281,1543,1569]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
) and different shape of parameres (cf.
<figureCitation id="0AA52A659D0B924CFD4CF9B8CB292E42" box="[735,895,1542,1568]" captionStart="Figs 601–616" captionStartId="172.[189,241,1892,1918]" captionTargetBox="[458,1119,274,1854]" captionTargetId="figure-73@172.[450,1138,265,1868]" captionTargetPageId="172" captionText="Figs 601–616. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 601– 603. M. brocki sp. nov., holotype, ♂ (BMNH). 604–606. M. dani sp. nov., holotype, ♂ (IECA). 607–610. M. deltouri sp. nov., holotype, ♂ (BMNH). 611–613. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 614–616. M. esau Heller, 1914, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122238" httpUri="https://zenodo.org/record/4122238/files/figure.png" pageId="23" pageNumber="24">Figs 607–610</figureCitation>
for
<taxonomicName id="559E4D639D0B924CFC23F9B9CC622E42" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[944,1076,1542,1568]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="deltouri" status="sp. nov.">
<emphasis id="A0EAEAF29D0B924CFC23F9B9CC622E42" box="[944,1076,1542,1568]" italics="true" pageId="23" pageNumber="24">M. deltouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D0B924CFBA8F9B9CCCC2E43" box="[1083,1178,1543,1569]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0B924CFF2EF9F5CD212EDE" pageId="23" pageNumber="24" type="etymology">
<paragraph id="922136E09D0B924CFF2EF9F5C9122E07" blockId="23.[189,324,1611,1637]" box="[189,324,1611,1637]" pageId="23" pageNumber="24">
<heading id="C969818C9D0B924CFF2EF9F5C9122E07" bold="true" box="[189,324,1611,1637]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A0EAEAF29D0B924CFF2EF9F5C9122E07" bold="true" box="[189,324,1611,1637]" pageId="23" pageNumber="24">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0B924CFF2EF9C2CD212EDE" blockId="23.[189,1399,1660,1725]" pageId="23" pageNumber="24">
Named after Awu, the largest volcano in the Sangihe chain, which caused its last powerful eruption in 1966 and developed a
<quantity id="55669B059D0B924CFE56F91CCA402EDE" box="[453,534,1698,1725]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="23" pageNumber="24" unit="km" value="4.5">4.5 km</quantity>
wide crater at the summit as a result of this eruption. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D0B924CFF2EF959CD212F8A" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="922136E09D0B924CFF2EF959C93A2F62" blockId="23.[189,364,1766,1793]" box="[189,364,1766,1793]" pageId="23" pageNumber="24">
<heading id="C969818C9D0B924CFF2EF959C93A2F62" bold="true" box="[189,364,1766,1793]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="A0EAEAF29D0B924CFF2EF959C93A2F62" bold="true" box="[189,364,1766,1793]" pageId="23" pageNumber="24">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D0B924CFF79F8A6C90A2F50" blockId="23.[189,1399,1816,1921]" box="[234,348,1816,1842]" pageId="23" pageNumber="24">
<emphasis id="A0EAEAF29D0B924CFF79F8A6C90A2F50" bold="true" box="[234,348,1816,1842]" pageId="23" pageNumber="24">
<typeStatus id="4D2588429D0B924CFF79F8A6C90A2F50" box="[234,348,1816,1842]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D0B924CFF2EF880CCF22F1C" blockId="23.[189,1399,1816,1921]" pageId="23" pageNumber="24">
<materialsCitation id="22F63CBD9D0B924CFF2EF880CCF22F1C" ID-GBIF-Occurrence="2898878561" collectingDate="2009-11-21" collectionCode="IECA" country="Indonesia" county="St. Jakl" elevation="400" location="Indonesia" municipality="Bukit Bembaluh" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="EA8976709D0B924CFF2EF880C90D2F3A" box="[189,347,1854,1880]" name="Indonesia" pageId="23" pageNumber="24">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D0B924CFEE2F880C9DF2F3A" box="[369,393,1854,1880]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
; “
<location id="9741603B9D0B924CFE0CF880CA412F3A" LSID="urn:lsid:plazi:treatment:1A3787F69D0B9242FD16FBDEC9A62ACF:9741603B9D0B924CFE0CF880CA412F3A" box="[415,535,1854,1880]" country="Indonesia" county="St. Jakl" municipality="Bukit Bembaluh" name="Indonesia" pageId="23" pageNumber="24">Indonesia</location>
,
<quantity id="55669B059D0B924CFDB2F880CAC92F3A" box="[545,671,1854,1880]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">
<elevation id="19B3D1D39D0B924CFDB2F880CAC92F3A" box="[545,671,1854,1880]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="23" pageNumber="24" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">200-600 m</elevation>
</quantity>
alt. SANGIR Isl.,
<location id="9741603B9D0B924CFCE0F880CC5F2F3A" LSID="urn:lsid:plazi:treatment:1A3787F69D0B9242FD16FBDEC9A62ACF:9741603B9D0B924CFCE0F880CC5F2F3A" box="[883,1033,1854,1880]" country="Indonesia" county="St. Jakl" municipality="Bukit Bembaluh" name="Tahuna distr." pageId="23" pageNumber="24">Tahuna distr.</location>
<collectingMunicipality id="7245AC9A9D0B924CFB9DF880CC872F3A" box="[1038,1233,1854,1880]" pageId="23" pageNumber="24">Bukit Bembaluh</collectingMunicipality>
,
<locationDeviation id="83932FC29D0B924CFB48F880CD212F3A" box="[1243,1399,1854,1880]" location="Tahuna" pageId="23" pageNumber="24">SE of Tahuna</locationDeviation>
14.-
<date id="E62010209D0B924CFF77F8DAC9272F1C" box="[228,369,1892,1918]" pageId="23" pageNumber="24" value="2009-11-21">
<collectingDate id="F664E9C89D0B924CFF77F8DAC9272F1C" box="[228,369,1892,1918]" pageId="23" pageNumber="24" value="2009-11-21">21.11.2009</collectingDate>
</date>
,
<collectingCounty id="7B404E6C9D0B924CFEEEF8DAC9852F1C" box="[381,467,1892,1918]" pageId="23" pageNumber="24">St. Jakl</collectingCounty>
lgt. // coll.
<collectionCode id="F48FAE259D0B924CFDC0F8DACAC02F1C" box="[595,662,1892,1918]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="23" pageNumber="24" type="Other institutional type">IECA</collectionCode>
České Budějovice, Czech Republic”;
<collectionCode id="F48FAE259D0B924CFBCBF8DACCF62F1C" box="[1112,1184,1892,1918]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="23" pageNumber="24" type="Other institutional type">IECA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D0B924CFF79F816CA452FA1" blockId="23.[189,1399,1960,2025]" box="[234,531,1960,1987]" pageId="23" pageNumber="24">
<emphasis id="A0EAEAF29D0B924CFF79F816C9312FA0" bold="true" box="[234,359,1960,1986]" pageId="23" pageNumber="24">
<typeStatus id="4D2588429D0B924CFF79F816C9312FA0" box="[234,359,1960,1986]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D0B924CFEEBF816CA5D2FA1" box="[376,523,1960,1987]" pageId="23" pageNumber="24" type="generic">4 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D0B924CFF2EF870CD212F8A" blockId="23.[189,1399,1960,2025]" box="[189,1399,1998,2025]" pageId="23" pageNumber="24">
<materialsCitation id="22F63CBD9D0B924CFF2EF870CBC42F8A" ID-GBIF-Occurrence="2898878451" box="[189,914,1998,2025]" collectionCode="IECA" country="Indonesia" location="INDONESIA" pageId="23" pageNumber="24" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="EA8976709D0B924CFF2EF870C90D2F8A" box="[189,347,1998,2024]" name="Indonesia" pageId="23" pageNumber="24">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D0B924CFEE7F870C9F72F8B" box="[372,417,1998,2025]" pageId="23" pageNumber="24" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D0B924CFE3CF870C9A42F8B" box="[431,498,1998,2025]" pageId="23" pageNumber="24" type="female">2 ♀♀</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D0B924CFCDCF870CBC42F8A" box="[847,914,1998,2024]" collectionName="Czech Republic, Budejovice, Czech Academy of Science, Institute of Entomology" country="Czech Republic" httpUri="http://grbio.org/cool/06zb-203s" name="Biology Centre of the Academy of Sciences of the Czech Republic, Institute of Entomology" pageId="23" pageNumber="24" type="Other institutional type">IECA</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D0B924CFC39F870CD242F8A" ID-GBIF-Occurrence="2898878324" box="[938,1394,1998,2025]" collectionCode="DTC" country="Indonesia" location="INDONESIA" pageId="23" pageNumber="24" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="8498FD699D0B924CFC39F870CB812F8B" box="[938,983,1998,2025]" pageId="23" pageNumber="24" type="female">1 ♀</specimenCount>
; same labels as for holotype;
<collectionCode id="F48FAE259D0B924CFAA7F870CD242F8A" box="[1332,1394,1998,2024]" collectionName="Dmitrijs Telnovs" pageId="23" pageNumber="24">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D049242FF2EFEB6CBE029FD" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="922136E09D049243FF2EFEB6C9182940" blockId="24.[189,334,264,290]" box="[189,334,264,290]" pageId="24" pageNumber="25">
<heading id="C969818C9D049243FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="A0EAEAF29D049243FF2EFEB6C9182940" bold="true" box="[189,334,264,290]" pageId="24" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D049243FF2EFE82CACD29C6" blockId="24.[189,1399,315,421]" pageId="24" pageNumber="25">
MEASUREMENTS.
<typeStatus id="4D2588429D049243FEEFFE85C9BA2937" box="[380,492,315,341]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D049243FD54FE85CB652934" box="[711,819,315,342]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" pageId="24" pageNumber="25" unit="mm" value="7.05">7.05 mm</quantity>
. Head
<quantity id="55669B059D049243FC16FE85CBB82934" box="[901,1006,315,342]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="25" unit="mm" value="1.5">1.50 mm</quantity>
long, across eyes
<quantity id="55669B059D049243FB55FE85CD792934" box="[1222,1327,315,342]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="24" pageNumber="25" unit="mm" value="1.81">1.81 mm</quantity>
wide. Pronotum
<quantity id="55669B059D049243FEA4FEDDC9C9291F" box="[311,415,355,381]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="25" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D049243FD3EFEDDCB41291F" box="[685,791,355,381]" metricMagnitude="-3" metricUnit="m" metricValue="2.78" pageId="24" pageNumber="25" unit="mm" value="2.78">2.78 mm</quantity>
. Elytral length
<quantity id="55669B059D049243FC54FEDCCC64291F" box="[967,1074,354,381]" metricMagnitude="-3" metricUnit="m" metricValue="3.85" pageId="24" pageNumber="25" unit="mm" value="3.85">3.85 mm</quantity>
, maximum combined width
<quantity id="55669B059D049243FF2EFE34C97F29C6" box="[189,297,394,420]" metricMagnitude="-3" metricUnit="m" metricValue="3.12" pageId="24" pageNumber="25" unit="mm" value="3.12">3.12 mm</quantity>
.
<typeStatus id="4D2588429D049243FEA7FE34C9F029C6" box="[308,422,394,420]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
<quantity id="55669B059D049243FE3EFE34CA0E29C6" box="[429,600,394,421]" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.8" metricValueMin="5.3" pageId="24" pageNumber="25" unit="mm" value="6.05" valueMax="6.8" valueMin="5.3">5.30–6.80 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D049243FF2EFE6ECCA52F8B" blockId="24.[189,1399,463,2025]" pageId="24" pageNumber="25">
Dorsum uniformly black to black-brown with reddish brown labroclypeus, mouthparts, antennae, legs and all of venter. Lateral margins of pronotum and elytra, pronotal hypomera and venter with dense microscopical velvety pubescence in some specimens. Head transverse, glossy to subopaque dorsally and ventrally, convex between eyes. Compound eye large, occupying about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D049243FEF6FDD3C99D2AE5" box="[357,459,621,647]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="24" pageNumber="25">Fig. 103</figureCitation>
) rather deeply V-like emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views, anterolateral angles rather strongly protruding, acute in dorsal view, bent up at ~90° to frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D049243FC8FFD02CBD72AB4" box="[796,897,700,726]" captionStart="Figs 97–108" captionStartId="137.[189,241,1786,1812]" captionTargetBox="[351,1228,270,1756]" captionTargetId="figure-127@137.[351,1237,265,1772]" captionTargetPageId="137" captionText="Figs 97–108. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 97. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 98. M. aiyura sp. nov., holotype, ♂ (BMNH). 99. M. alesbezdeki sp. nov., holotype, ♂ (NME). 100. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 101. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 102. M. bintang sp. nov., holotype, ♂ (NME). 103. M. awu sp. nov., holotype, ♂ (IECA). 104. Same, paratype, ♀ (IECA). 105. M. boessnecki sp. nov., holotype, ♂ (SNSD). 106. M. babyrousa sp. nov., holotype, ♂ (BMNH). 107. Same, paratype, ♀ (BMNH). 108. M. caperatus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122173" httpUri="https://zenodo.org/record/4122173/files/figure.png" pageId="24" pageNumber="25">Fig. 104</figureCitation>
) with vague and very broad emargination, its anterolateral angles broadly rounded and not protruding, nearly straight in both dorsal and lateral views. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus slightly rounded in dorsal view. Anterior and lateral margins of labroclypeus smooth. Dorsal punctures ovoid, large and very deep; inner margin with delicate membrane. Intervening spaces glossy, glabrous, generally distinctly smaller than punctures. Very long erect seta rises from anterior margin of each puncture. Antenna in both sexes 9-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 trapezoid, transverse. Pronotum glossy dorsally and laterally. Anterior margin sinuous in male (slightly sinuous with anterolateral angles (stronger) and mesal portion protruding anteriad), broadly emarginate in female. Basal margin broadly rounded. Lateral margin of pronotum in dorsal view broadly rounded in male (
<figureCitation id="0AA52A659D049243FE95FBD2C93A2CE5" box="[262,364,1132,1159]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="24" pageNumber="25">Fig. 202</figureCitation>
), straight and widened towards obtuse postmedian angulation and afterwards constricted towards base in female. Lateral margin delicately crenulate all along, a long erect seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Punctures ovoid, very deep, variably large; inner margin with delicate membrane. Intervening spaces glossy and glabrous, generally narrower than punctures, nearly wrinkled at lateral margins. Setae curved, as on head. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Hypomeron emarginate and very long setose on anterior margin, which is flange-like protruding. Antennal pocket deep. Median anterior process of prosternum long brushy setose, short, moderately raised. Scutellar shield pointed apically. Elytra slightly widened postmedially, maximum width across midlength, glossy dorsally, with distinct humeri. Vague tracks of two flat glabrous carinae on each elytron. Punctures of elytral disc ovoid, generally smaller and much shallower than those on forebody, irregular to partly arranged in longitudinal rows; inner margin with delicate membrane covered with microscopical velvety pubescence (
<figureCitation id="0AA52A659D049243FCB2F9FBCBC62E3D" box="[801,912,1605,1631]" captionStart="Figs 286–301" captionStartId="152.[189,241,1716,1742]" captionTargetBox="[364,1224,266,1689]" captionTargetId="figure-172@152.[364,1224,266,1695]" captionTargetPageId="152" captionText="Figs 286–301. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 286. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 287. M. aiyura sp. nov., holotype, ♂ (BMNH). 288. M. alesbezdeki sp. nov., holotype, ♂ (NME). 289. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 290. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 291. M. paupianus Heller, 1910, ♂ (BMNH). 292. M. awu sp. nov., holotype, ♂ (IECA). 293. M. babyrousa sp. nov., holotype, ♂ (BMNH). 294. M. bintang sp. nov., holotype, ♂ (NME). 295. M. boessnecki sp. nov., holotype, ♂ (SNSD). 296. M. brocki sp. nov., holotype, ♂ (BMNH). 297. M. caperatus sp. nov., holotype, ♂ (NME). 298. M. ciliatus sp. nov., holotype, ♂ (IECA). 299. M. crypticus sp. nov., paratype, ♀ (BMNH). 300. M. dani sp. nov., holotype, ♂ (IECA). 301. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122206" httpUri="https://zenodo.org/record/4122206/files/figure.png" pageId="24" pageNumber="25">Fig. 292</figureCitation>
). Setae long, suberect to erect, sparse, arranged in longitudinal rows. Epipleuron all along with short suberect setae. Male pygidium slightly convex, deeply punctate (Fig. 537). Pygidium intervening spaces subopaque, microreticulate and covered with microscopical velvety pubescence, with sparse moderately long suberect setae. Female pygidium flattened, glossy on intervening spaces. Venter covered with sparse small shallow punctures, each provided with very short seta, in part covered with microscopical velvety pubescence. Legs long and slender. Male protibia straight on external margin, delicately crenulate all along, with complete dorsal fovea (
<figureCitation id="0AA52A659D049243FECDF8E6C9902F11" box="[350,454,1880,1907]" captionStart="Figs 451–468" captionStartId="163.[189,241,1681,1707]" captionTargetBox="[456,1130,262,1650]" captionTargetId="figure-178@163.[456,1131,262,1657]" captionTargetPageId="163" captionText="Figs 451–468. Papuan and Wallacean Maechidius Macleay, 1819, pygidium, dorsal view. 451. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 452. M. aiyura sp. nov., holotype, ♂ (BMNH). 453. Same, different aspect (BMNH). 454. M. alesbezdeki sp. nov., holotype, ♂ (NME). 455. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 456. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 457. M. awu sp. nov., holotype, ♂ (IECA). 458. M. babyrousa sp. nov., holotype, ♂ (BMNH). 459. Same, paratype, ♀ (BMNH). 460. M. bintang sp. nov., holotype, ♂ (NME). 461. M. boessnecki sp. nov., holotype, ♂ (SNSD). 462. M. brocki sp. nov., holotype, ♂ (BMNH). 463. M. caperatus sp. nov., holotype, ♂ (NME). 464. M. ciliatus sp. nov., holotype, ♂ (IECA). 465. M. crypticus sp. nov., paratype, ♂ (BMNH). 466. Same, paratype, ♀ (BMNH). 467. M. dani sp. nov., holotype, ♂ (IECA). 468. M. deltouri sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122222" httpUri="https://zenodo.org/record/4122222/files/figure.png" pageId="24" pageNumber="25">Fig. 457</figureCitation>
). Two acute, rather large distal teeth on protibia in both sexes, distal longer than predistal. Male protibial terminal spur large, curved, female one shorter and straight. Male metatibial terminal spurs paired, almost equally long, subacute, lower one curved; female ones shorter, nearly straight, obtuse. Tarsal claws with pulvilli in both sexes. Male aedeagus as in
<figureCitation id="0AA52A659D049243FBD8F870CCB92F8B" box="[1099,1263,1998,2025]" captionStart="Figs 570–585" captionStartId="170.[189,241,1892,1918]" captionTargetBox="[425,1155,265,1858]" captionTargetId="figure-74@170.[416,1172,265,1864]" captionTargetPageId="170" captionText="Figs 570–585. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 570– 572. M. aiyura sp. nov., holotype, ♂ (BMNH). 573–575. M. alesbezdeki sp. nov., holotype, ♂ (NME). 576–578. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 579–581. M. awu sp. nov., holotype, ♂ (IECA). 582–585. M. babyrousa sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122232" httpUri="https://zenodo.org/record/4122232/files/figure.png" pageId="24" pageNumber="25">Figs 579–581</figureCitation>
.
</paragraph>
<paragraph id="922136E09D059242FF2EFEB6C9E62940" blockId="25.[189,432,264,290]" box="[189,432,264,290]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFEB6C9E62940" bold="true" box="[189,432,264,290]" pageId="25" pageNumber="26">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EFE87CBE029FD" blockId="25.[189,1399,313,415]" pageId="25" pageNumber="26">Male with longer lamellae of antennal club, stronger emarginate anterior margin of labroclypeus, dorsally convex pygidium (flat in female), larger and curved pro- and lower metatibial terminal spurs, and evenly rounded lateral margin on pronotum (in dorsal view).</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FF2EFE76CB472A71" box="[189,785,456,532]" pageId="25" pageNumber="26" type="biology_ecology">
<paragraph id="922136E09D059242FF2EFE76C9762980" blockId="25.[189,288,456,482]" box="[189,288,456,482]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFE76C9762980" bold="true" box="[189,288,456,482]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFE76C9762980" bold="true" box="[189,288,456,482]" pageId="25" pageNumber="26">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EFE47CB472A71" blockId="25.[189,785,505,532]" box="[189,785,505,532]" pageId="25" pageNumber="26">
Studied specimens sampled at
<quantity id="55669B059D059242FDB6FE47CAFC2A76" box="[549,682,505,532]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="25" pageNumber="26" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">200–600 m</quantity>
altitude.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D059242FF2EFD83C9A62ACF" pageId="25" pageNumber="26" type="distribution">
<paragraph id="922136E09D059242FF2EFD83C9002A35" blockId="25.[189,342,573,599]" box="[189,342,573,599]" pageId="25" pageNumber="26">
<heading id="C969818C9D059242FF2EFD83C9002A35" bold="true" box="[189,342,573,599]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="A0EAEAF29D059242FF2EFD83C9002A35" bold="true" box="[189,342,573,599]" pageId="25" pageNumber="26">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D059242FF2EFDD0C9A62ACF" blockId="25.[189,1398,622,685]" pageId="25" pageNumber="26">
Hitherto only known from Sangir, the largest of the Sangihe Islands. This is hitherto the northernmost record of the
<taxonomicName id="559E4D639D059242FEC9FD2DC9BA2ACF" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[346,492,659,685]" class="Insecta" family="Scarabaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="5" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Maechidiini">Maechidiini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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