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<document id="09D64C33761657EFFF025A79F8AD024D" ID-CLB-Dataset="300289" ID-DOI="10.6620/ZS.2022.61-73" ID-GBIF-Dataset="2a11fc58-cec5-4063-8891-f3c66ad521c9" ID-ISSN="1810-522X" ID-PMC="PMC10061213" ID-PubMed="37007805" ID-Zenodo-Dep="12827273" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721933110989" checkinUser="felipe" docAuthor="Hidaka, Chiho, Yang, Chien-Hui &amp; Wakabayashi, Kaori" docDate="2022" docId="145E87EB562F3972FF2F1009C7044246" docLanguage="en" docName="ZoolStud.61.73.1-12.pdf" docOrigin="Zoological Studies 61 (73)" docSource="http://dx.doi.org/10.5281/zenodo.8074244" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Scyllarides squammosus Nisto" docType="treatment" docVersion="2" lastPageNumber="8" masterDocId="E867FF93562B3975FF811114C26C4333" masterDocTitle="Parascorpaena poseidon Chou and Liao 2022" masterLastPageNumber="12" masterPageNumber="1" pageNumber="5" updateTime="1721937811008" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3C0E51DF066C95238249A91A4DD0421B">Parascorpaena poseidon Chou and Liao 2022</mods:title>
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<mods:namePart id="E4566C10D2641A0441775C701741B675">Hidaka, Chiho</mods:namePart>
<mods:affiliation id="056A4F5688FD35FC96D580513E1C75E6">School of Applied Biological Science, Hiroshima University, Kagamiyama 1 - 4 - 4, Higashi-Hiroshima, Hiroshima 739 - 8528, Japan.</mods:affiliation>
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<mods:namePart id="6069363CC93752AC046108B327FE9EBF">Yang, Chien-Hui</mods:namePart>
<mods:affiliation id="14106EF8093A108D3428B318AACBABB0">Institute of Marine Biology and Centre of Excellence for the Oceans, National Taiwan Ocean University, 2 Pei-Ning Road, Keelung 202301, Taiwan. E-mail: chyang @ mail. ntou. edu. tw (Yang)</mods:affiliation>
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<mods:namePart id="784EC184B032750E43999277A94ECB36">Wakabayashi, Kaori</mods:namePart>
<mods:affiliation id="00240225306C73B5E0145EFD950798C7">Graduate School of Integrated Sciences for Life, Hiroshima University, Kagamiyama 1 - 4 - 4, Higashi-Hiroshima, Hiroshima 739 - 8528, Japan. * Correspondence: E-mail: kaoriw @ hiroshima-u. ac. jp (Wakabayashi)</mods:affiliation>
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<mods:title id="570861A8FCD1ED4025B23D0DD9A6ACA8">Zoological Studies</mods:title>
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<mods:date id="24382C43A83FD377020D03198B3EB706">2022</mods:date>
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<treatment id="145E87EB562F3972FF2F1009C7044246" ID-DOI="http://doi.org/10.5281/zenodo.12827277" ID-Zenodo-Dep="12827277" LSID="urn:lsid:plazi:treatment:145E87EB562F3972FF2F1009C7044246" httpUri="http://treatment.plazi.org/id/145E87EB562F3972FF2F1009C7044246" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5">
<subSubSection id="D4ED6576562F3971FF2F1009C3904265" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="9C4836FD562F3971FF2F1009C3904265" blockId="4.[174,775,252,342]" pageId="4" pageNumber="5">
<heading id="C7008191562F3971FF2F1009C3904265" centered="true" fontSize="36" level="2" pageId="4" pageNumber="5" reason="4">
<taxonomicName id="5BF74D7E562F3971FF2F1009C3904265" ID-CoL="854SS" authority="(H. Milne Edwards, 1837) Nisto" authorityName="Nisto" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="AE83EAEF562F3971FF2F1009C3904265" bold="true" pageId="4" pageNumber="5">
<emphasis id="AE83EAEF562F3971FF2F1009C3B04205" bold="true" box="[174,476,285,311]" italics="true" pageId="4" pageNumber="5">Scyllarides squammosus</emphasis>
(H. Milne Edwards, 1837) Nisto
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4ED6576562F3972FF6B1069C7044246" lastPageId="7" lastPageNumber="8" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="9C4836FD562F3971FF6B1069C0A34286" blockId="4.[174,775,381,886]" pageId="4" pageNumber="5">
<emphasis id="AE83EAEF562F3971FF6B1069C3D942A6" box="[234,437,381,405]" italics="true" pageId="4" pageNumber="5">Material examined</emphasis>
: Nisto, NSMT-Cr 30950, BL =
<quantity id="5B0F9B18562F3971FF2F108AC2974286" box="[174,251,414,438]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="5" unit="mm" value="39.0">39 mm</quantity>
, CL =
<quantity id="5B0F9B18562F3971FEC21089C3C94286" box="[323,421,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="5" unit="mm" value="14.5">14.5 mm</quantity>
, CW =
<quantity id="5B0F9B18562F3971FE77108AC0284286" box="[502,580,414,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="5" unit="mm" value="17.0">17 mm</quantity>
, TL =
<quantity id="5B0F9B18562F3971FD0B108AC0A74286" box="[650,715,414,438]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="5" unit="mm" value="9.0">9 mm</quantity>
.
</paragraph>
<paragraph id="9C4836FD562F3971FF6B10AAC74E4266" blockId="4.[174,775,381,886]" lastBlockId="4.[850,1452,253,886]" pageId="4" pageNumber="5">
<emphasis id="AE83EAEF562F3971FF6B10AAC30542E6" box="[234,361,446,469]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
: Carapace (
<figureCitation id="04CC2A78562F3971FE6E10A9C01442E5" box="[495,632,445,470]" captionStart="Fig" captionStartId="5.[136,167,1925,1944]" captionTargetBox="[392,1151,1193,1899]" captionTargetId="figure-6@5.[359,1191,1180,1917]" captionTargetPageId="5" captionText="Fig. 3. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Body, dorsal, a pair of antennules omitted; B: Body, lateral (left), ventral side of telson in gray. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12827281" httpUri="https://zenodo.org/record/12827281/files/figure.png" pageId="4" pageNumber="5">Fig. 3A, 3B</figureCitation>
): Wider than longer (CW/CL 1.17) with maximum width in the middle; surface with few tubercules, entirely furnished with fine setae; anterior margin smooth, a prominent horn present at both angles; lateral margin possessing a distinct post-cervical incision, with 8 (right) to 9 (left) anterolateral and 14 (right) to 12 (left) post-lateral teeth; rostral short with blunt end; 2 prominent tubercules and 1 small tubercule on the inner and posterior edges of orbits; gastric tooth possessing 2 prominent tubercles anteriorly and 4 small setae posteriorly; cardiac tooth bearing 2 rows of prominent tubercules anteriorly and 2 rows of small setae posteriorly, cardiac ridge with small setae; 10 small setae (8 each on the right and left sides,
<quantity id="5B0F9B18562F3971FCD311EAC1114226" box="[850,893,253,278]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="5" unit="in" value="2.0">2 in</quantity>
the middle) present along the post-cervical groove; 10 small setae in a row along post-lateral teeth; 5 small setae present at middle of posterior margin.
</paragraph>
<paragraph id="9C4836FD562F3971FC0E1049C6344166" blockId="4.[850,1452,253,886]" pageId="4" pageNumber="5">
Antennule (
<figureCitation id="04CC2A78562F3971FB9E1049C6EF4245" box="[1055,1155,349,374]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="4" pageNumber="5">Fig. 4A</figureCitation>
): Peduncle 3-segmented with flagellated distal end; proximal segment bearing dense long setae on ventral side; second segment possessing small setae scattered; third segment almost naked; accessory flagellum 18-articulated as long as the primary flagellum; primary flagellum thicker than accessory flagellum, 15-articulated, group of aesthetascs present on articles 5–12.
</paragraph>
<paragraph id="9C4836FD562F3971FC0E134AC6224066" blockId="4.[850,1452,253,886]" pageId="4" pageNumber="5">
Antenna (
<figureCitation id="04CC2A78562F3971FC791349C6224145" box="[1016,1102,605,630]" captionStart="Fig" captionStartId="5.[136,167,1925,1944]" captionTargetBox="[392,1151,1193,1899]" captionTargetId="figure-6@5.[359,1191,1180,1917]" captionTargetPageId="5" captionText="Fig. 3. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Body, dorsal, a pair of antennules omitted; B: Body, lateral (left), ventral side of telson in gray. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12827281" httpUri="https://zenodo.org/record/12827281/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
): 4-segmented, biramous, broad, flat; proximal segment inverted triangle, anterior margin with a ridge at middle; second segment with serrated lateral margin, prominent horn at anterolateral angle, a small notch on anteromedial margin, 2 small teeth on medial margin; third segment bearing a prominent tooth on anteromedial angle; distal segment surrounded with finely serrated margins.
</paragraph>
<paragraph id="9C4836FD562F3970FC0E1249C3C34226" blockId="4.[850,1452,253,886]" lastBlockId="5.[136,737,253,1142]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Mandible (
<figureCitation id="04CC2A78562F3971FB8D1249C60B4045" box="[1036,1127,861,886]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="4" pageNumber="5">Fig. 4B</figureCitation>
): Poorly developed, slightly asymmetrical; palp present.
</paragraph>
<caption id="C8886675562F3971FF2F165AC0124486" ID-DOI="http://doi.org/10.5281/zenodo.12827279" ID-Zenodo-Dep="12827279" httpUri="https://zenodo.org/record/12827279/files/figure.png" pageId="4" pageNumber="5" startId="4.[174,205,1870,1889]" targetBox="[174,1451,953,1837]" targetPageId="4" targetType="figure">
<paragraph id="9C4836FD562F3971FF2F165AC0124486" blockId="4.[174,1452,1870,1973]" pageId="4" pageNumber="5">
<emphasis id="AE83EAEF562F3971FF2F165AC28B4452" bold="true" box="[174,231,1870,1889]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
Bayesian phylogenetic tree of the subfamily
<taxonomicName id="5BF74D7E562F3971FDF9165AC0BE4452" authorityName="Holthuis" authorityYear="1985" box="[632,722,1870,1889]" class="Malacostraca" family="Scyllaridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Arctidinae">Arctidinae</taxonomicName>
with partial sequences of
<emphasis id="AE83EAEF562F3971FC36165AC1B04452" box="[951,988,1870,1889]" italics="true" pageId="4" pageNumber="5">COI</emphasis>
and
<emphasis id="AE83EAEF562F3971FB86165AC6444452" box="[1031,1064,1870,1889]" italics="true" pageId="4" pageNumber="5">16S</emphasis>
. Posterior probability (Pp) values are shown when above 0.8. The tree was constructed basically with both the
<emphasis id="AE83EAEF562F3971FD64167EC166444E" box="[741,778,1898,1917]" italics="true" pageId="4" pageNumber="5">COI</emphasis>
and
<emphasis id="AE83EAEF562F3971FCB5167EC139444E" box="[820,853,1898,1917]" italics="true" pageId="4" pageNumber="5">16S</emphasis>
sequences, but the operational taxonomic units (OTUs) with hash (#) were inferred only from the
<emphasis id="AE83EAEF562F3971FE261692C3A044AA" box="[423,460,1926,1945]" italics="true" pageId="4" pageNumber="5">COI</emphasis>
and those with delta (Δ) were inferred only from the
<emphasis id="AE83EAEF562F3971FC251692C1A944AA" box="[932,965,1926,1945]" italics="true" pageId="4" pageNumber="5">16S</emphasis>
, due to a lack of information. Asterisks (*) indicate the OTUs with sequences resulting from the mitogenome.
</paragraph>
</caption>
<paragraph id="9C4836FD562E3970FF44100AC03B42A6" blockId="5.[136,737,253,1142]" pageId="5" pageNumber="6">
Maxillule (
<figureCitation id="04CC2A78562E3970FEDF100AC3AE4205" box="[350,450,285,310]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="5" pageNumber="6">Fig. 4 C</figureCitation>
): Uniramous; poorly developed; basal endite with 5 small tubercules on anterior margin; coxal endite unarmed on anterior margin, bearing 3 small setae on basal part.
</paragraph>
<paragraph id="9C4836FD562E3970FF441089C0244146" blockId="5.[136,737,253,1142]" pageId="5" pageNumber="6">
Maxilla (
<figureCitation id="04CC2A78562E3970FEAE1089C3E24285" box="[303,398,413,438]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="5" pageNumber="6">Fig. 4D</figureCitation>
): Flattened; endopod slightly differentiated with 8 setae and 3 small spines on the medial margin; endites differentiated, 4 (2+2) and 7 (5+2) setae on anterior ends of coxal and basal endites, 1 and 2 setae present on the inner proximal parts of coxal and basal endites, respectively; hair-like setae lining the outer margin of scaphognathite.
</paragraph>
<paragraph id="9C4836FD562E3970FF441369C0064006" blockId="5.[136,737,253,1142]" pageId="5" pageNumber="6">
First maxilliped (
<figureCitation id="04CC2A78562E3970FE0D1369C38B41A5" box="[396,487,637,662]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="5" pageNumber="6">Fig. 4E</figureCitation>
): Biramous, coxa and basis undifferentiated, flattened; endopod unsegmented, smooth; exopod with 15 marginal setae on proximal part, small setae on distal end; epipod membranous, expanding posteriorly, bearing 5 long setae; 2 small setae present at the distal end of basal endite.
</paragraph>
<paragraph id="9C4836FD562E3970FF441229C0524086" blockId="5.[136,737,253,1142]" pageId="5" pageNumber="6">
Second maxilliped (
<figureCitation id="04CC2A78562E3970FE351229C07D4065" box="[436,529,829,854]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="5" pageNumber="6">Fig. 4F</figureCitation>
): Biramous, coxa and basis undifferentiated; endopod 4-segmented with 0,1,13,0 setae; exopod incompletely segmented, longer than endopod, 2 small setae at distal end.
</paragraph>
<paragraph id="9C4836FD562E3970FF4412A9C03E4766" blockId="5.[136,737,253,1142]" pageId="5" pageNumber="6">
Third maxilliped (
<figureCitation id="04CC2A78562E3970FE1012A9C38040E5" box="[401,492,957,982]" captionStart="Fig" captionStartId="6.[174,205,1897,1916]" captionTargetBox="[225,1424,250,1863]" captionTargetId="figure-5@6.[189,1437,232,1870]" captionTargetPageId="6" captionText="Fig. 4. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="5" pageNumber="6">Fig. 4G</figureCitation>
): Biramous, coxa and basis differentiated; endopod 5-segmented, carpus with dense setae (~30), propodus with 14 setae, dactylus with 4 long setae on inner margin; exopod segmented, poorly developed with 7 small setae on distal end.
</paragraph>
<paragraph id="9C4836FD562E3970FF441549C6A342E6" blockId="5.[136,737,253,1142]" lastBlockId="5.[813,1414,253,1142]" pageId="5" pageNumber="6">
Pereiopods (
<figureCitation id="04CC2A78562E3970FECE1549C3C64746" box="[335,426,1117,1141]" captionStart="Fig" captionStartId="7.[136,167,1898,1917]" captionTargetBox="[137,1415,451,1863]" captionTargetId="figure-5@7.[122,1425,441,1882]" captionTargetPageId="7" captionText="Fig. 5. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Pereiopods, left, ventral, degenerated exopods shown in shaded; B: Pleopod 3, right, anterior, with enlarged distal ends of pleopods 1 and 3 (a) and pleopods 2 and 4 (b); C: Telson, dorsal; D: Uropod, right, dorsal. Scale bars: A, D = 3 mm; C = 2 mm, B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827285" httpUri="https://zenodo.org/record/12827285/files/figure.png" pageId="5" pageNumber="6">Fig. 5A</figureCitation>
): Pereiopods 1–4 biramous, coxa and basis differentiated, endopods 5-segmented with almost equal length, exopods degenerated, merus and carpus bearing a prominent horn at distal end, ischium of pereiopod 2–4 with 4 small spines; Pereiopod 5 uniramous, shortest, merus possessing a prominent horn at the distal end, ischium bearing a conspicuous spine pointing posteriorly.
</paragraph>
<paragraph id="9C4836FD562E3970FCE810C9C65341E6" blockId="5.[813,1414,253,1142]" pageId="5" pageNumber="6">
Pleon (
<figureCitation id="04CC2A78562E3970FC3510C9C65B42C5" box="[948,1079,477,502]" captionStart="Fig" captionStartId="5.[136,167,1925,1944]" captionTargetBox="[392,1151,1193,1899]" captionTargetId="figure-6@5.[359,1191,1180,1917]" captionTargetPageId="5" captionText="Fig. 3. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Body, dorsal, a pair of antennules omitted; B: Body, lateral (left), ventral side of telson in gray. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12827281" httpUri="https://zenodo.org/record/12827281/files/figure.png" pageId="5" pageNumber="6">Fig. 3A, 3B</figureCitation>
): Tergum of pleonite 1 smooth; median carina distinct on tergites of pleonites 2–5; terga of pleonites 2–3 with a few spicules, small spines on median carina, a small tooth at center of posterior margin; terga of pleonites 4–5 smooth, a large tooth present at center of posterior margin; pleura of pleonites 2–5 serrated with large tooth, 2 small spines on the medial margins of pleura.
</paragraph>
<paragraph id="9C4836FD562E3970FCE813C9C6CB4765" blockId="5.[813,1414,253,1142]" pageId="5" pageNumber="6">
Pleopods (
<figureCitation id="04CC2A78562E3970FC6413C9C62E41C6" box="[997,1090,733,757]" captionStart="Fig" captionStartId="7.[136,167,1898,1917]" captionTargetBox="[137,1415,451,1863]" captionTargetId="figure-5@7.[122,1425,441,1882]" captionTargetPageId="7" captionText="Fig. 5. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Pereiopods, left, ventral, degenerated exopods shown in shaded; B: Pleopod 3, right, anterior, with enlarged distal ends of pleopods 1 and 3 (a) and pleopods 2 and 4 (b); C: Telson, dorsal; D: Uropod, right, dorsal. Scale bars: A, D = 3 mm; C = 2 mm, B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827285" httpUri="https://zenodo.org/record/12827285/files/figure.png" pageId="5" pageNumber="6">Fig. 5B</figureCitation>
): Present on pleonites 2–5; biramous; 2 setae at medial distal angle of protopod; exopod surrounded with 35 (pleopod 1), 37 (2), 38 (3), 33 (4) long plumose natatory setae, each possessing 6 (pleopod 1), 12 (2), 14 (3), 12 (4) small plumose setae at distal end; endopod surrounded with 48 (pleopod 1), 52 (2), 48 (3), 48 (4) long plumose natatory setae, each possessing 0 (pleopod 1), 2 (2), 5 (3), 4 (4) small plumose setae at distal end, appendix interna not reaching distal end of endopod, with approximately 12 cincinnuli at the tip, a plumose seta present just beside the cincinnuli for pleopods 2 and 4.
</paragraph>
<paragraph id="9C4836FD562E3972FCE81549C32B4206" blockId="5.[813,1414,253,1142]" lastBlockId="7.[136,737,253,373]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
Uropod (
<figureCitation id="04CC2A78562E3970FC561549C6554746" box="[983,1081,1117,1141]" captionStart="Fig" captionStartId="7.[136,167,1898,1917]" captionTargetBox="[137,1415,451,1863]" captionTargetId="figure-5@7.[122,1425,441,1882]" captionTargetPageId="7" captionText="Fig. 5. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Pereiopods, left, ventral, degenerated exopods shown in shaded; B: Pleopod 3, right, anterior, with enlarged distal ends of pleopods 1 and 3 (a) and pleopods 2 and 4 (b); C: Telson, dorsal; D: Uropod, right, dorsal. Scale bars: A, D = 3 mm; C = 2 mm, B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827285" httpUri="https://zenodo.org/record/12827285/files/figure.png" pageId="5" pageNumber="6">Fig. 5C</figureCitation>
): A tooth present at lateral margins of exopod and endopod; medial margin of endopod serrated.
</paragraph>
<caption id="C8886675562E3970FF091691C3CF4487" ID-DOI="http://doi.org/10.5281/zenodo.12827281" ID-Zenodo-Dep="12827281" httpUri="https://zenodo.org/record/12827281/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,167,1925,1944]" targetBox="[392,1151,1193,1899]" targetPageId="5" targetType="figure">
<paragraph id="9C4836FD562E3970FF091691C3CF4487" blockId="5.[136,1414,1925,1972]" pageId="5" pageNumber="6">
<emphasis id="AE83EAEF562E3970FF091691C2AC44AB" bold="true" box="[136,192,1925,1944]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="5BF74D7E562E3970FF4A1691C0E844AB" authority="(H. Milne Edwards, 1837)" authorityName="Nisto" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[203,644,1925,1944]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="AE83EAEF562E3970FF4A1691C3F044AB" box="[203,412,1925,1944]" italics="true" pageId="5" pageNumber="6">Scyllarides squammosus</emphasis>
(H. Milne Edwards, 1837)
</taxonomicName>
, nisto. A: Body, dorsal, a pair of antennules omitted; B: Body, lateral (left), ventral side of telson in gray. Scale bar = 5 mm.
</paragraph>
</caption>
<caption id="C8886675562D3973FF2F167DC0134486" ID-DOI="http://doi.org/10.5281/zenodo.12827283" ID-Zenodo-Dep="12827283" httpUri="https://zenodo.org/record/12827283/files/figure.png" pageId="6" pageNumber="7" startId="6.[174,205,1897,1916]" targetBox="[225,1424,250,1863]" targetPageId="6" targetType="figure">
<paragraph id="9C4836FD562D3973FF2F167DC0134486" blockId="6.[174,1452,1897,1973]" pageId="6" pageNumber="7">
<emphasis id="AE83EAEF562D3973FF2F167DC284444E" bold="true" box="[174,232,1897,1917]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="5BF74D7E562D3973FF76167EC0D3444E" authority="(H. Milne Edwards, 1837)" authorityName="Nisto" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[247,703,1898,1917]" class="Malacostraca" family="Scyllaridae" genus="Scyllarides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="squammosus">
<emphasis id="AE83EAEF562D3973FF76167EC3A1444F" box="[247,461,1898,1917]" italics="true" pageId="6" pageNumber="7">Scyllarides squammosus</emphasis>
(H. Milne Edwards, 1837)
</taxonomicName>
, nisto. A: Antennule, right, ventrolateral; B: Mandibles, ventral; C: Maxillule, right, ventral; D: Maxilla, right, ventral; E: First maxilliped, right, dorsal; F: Second maxilliped, right, ventral; G: Third maxilliped, right, dorsal. Scale bars:A, E, G = 1 mm; B, F = 0.5 mm; C, D = 0.1 mm.
</paragraph>
</caption>
<paragraph id="9C4836FD562C3972FF441029C3424246" blockId="7.[136,737,253,373]" pageId="7" pageNumber="8">
Telson (
<figureCitation id="04CC2A78562C3972FE9D1029C3184266" box="[284,372,317,341]" captionStart="Fig" captionStartId="7.[136,167,1898,1917]" captionTargetBox="[137,1415,451,1863]" captionTargetId="figure-5@7.[122,1425,441,1882]" captionTargetPageId="7" captionText="Fig. 5. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: Pereiopods, left, ventral, degenerated exopods shown in shaded; B: Pleopod 3, right, anterior, with enlarged distal ends of pleopods 1 and 3 (a) and pleopods 2 and 4 (b); C: Telson, dorsal; D: Uropod, right, dorsal. Scale bars: A, D = 3 mm; C = 2 mm, B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12827285" httpUri="https://zenodo.org/record/12827285/files/figure.png" pageId="7" pageNumber="8">Fig. 5D</figureCitation>
): A tooth present at the proximal, lateral margins.
</paragraph>
<paragraph id="9C4836FD562C3972FCE811E9C7044246" blockId="7.[813,1413,253,373]" pageId="7" pageNumber="8">
Color (
<figureCitation id="04CC2A78562C3972FC3811E9C1904225" box="[953,1020,253,278]" captionStart="Fig" captionStartId="2.[174,205,1898,1917]" captionTargetBox="[348,1277,1354,1871]" captionTargetId="figure-550@2.[348,1277,1354,1871]" captionTargetPageId="2" captionText="Fig. 1. Scyllarides squammosus (H. Milne Edwards, 1837), nisto. A: in situ, live (photographed by Hideki Abe, reproduced with permission from Wakabayashi et al. 2017a, p. 59 in which the nisto was named as Scyllarides sp.); B: fixed specimen (NSMT-Cr 30950) in 99% ethanol. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.12827275" httpUri="https://zenodo.org/record/12827275/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
): Body entirely transparent in live specimen except a yellow line along the anterior margin of carapace and gills in pink. Specimen preserved in ethanol entirely white with transparent body margins.
</paragraph>
</subSubSection>
</treatment>
</document>
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