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<document id="D13657AB0FD8EC9D731EE37684B3074C" ID-CLB-Dataset="39033" ID-DOI="10.11646/zootaxa.3658.1.1" ID-GBIF-Dataset="0adf09ba-5301-4828-abf6-8dbe89f041ca" ID-ISSN="1175-5326" ID-Zenodo-Dep="246551" ID-ZooBank="C02D1C74-25C0-41DD-B098-62098EB7B62A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463647879570" checkinUser="plazi" docAuthor="Dragan P. Chobanov, Beata Grzywacz, Ionuţ Ş. Iorgu, Battal Cιplak, Maya B. Ilieva &amp; Elżbieta Warchałowska-Śliwa" docDate="2013" docId="F26F31283917FFB2B1B00D5FFF0B9B00" docLanguage="en" docName="zt03658p081.pdf.imf" docOrigin="Zootaxa 3658 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Isophya modesta subsp. modesta Frivaldszky 1867" docType="treatment" docVersion="12" lastPageNumber="33" masterDocId="0E5649503908FF92B1270A03FFD39875" masterDocTitle="Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data" masterLastPageNumber="81" masterPageNumber="1" pageNumber="32" updateTime="1698662571276" updateUser="ExternalLinkService">
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<mods:title id="B68728C76B9ECEF96216B69A0CAD861C">Review of the Balkan Isophya (Orthoptera: Phaneropteridae) with particular emphasis on the Isophya modesta group and remarks on the systematics of the genus based on morphological and acoustic data</mods:title>
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<mods:namePart id="36C1BEB2F1DF2FC8078B41425BBDB71F">Dragan P. Chobanov</mods:namePart>
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<mods:namePart id="1889265AFDC1E60C5A303081FBC3FD57">Ionuţ Ş. Iorgu</mods:namePart>
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<treatment id="F26F31283917FFB2B1B00D5FFF0B9B00" ID-DOI="http://doi.org/10.5281/zenodo.5617359" ID-GBIF-Taxon="120682299" ID-Zenodo-Dep="5617359" LSID="urn:lsid:plazi:treatment:F26F31283917FFB2B1B00D5FFF0B9B00" httpUri="http://treatment.plazi.org/id/F26F31283917FFB2B1B00D5FFF0B9B00" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
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<paragraph id="7A79803E3917FF8DB1B00D5FFD119F00" blockId="31.[151,706,1883,1942]" box="[151,706,1883,1910]" pageId="31" pageNumber="32">
<heading id="213137523917FF8DB1B00D5FFD119F00" bold="true" box="[151,706,1883,1910]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="BDC6FBBD3917FF8DB1B00D5FFD119F00" authority="Frivaldszky, 1867" authorityName="Frivaldszky" authorityYear="1867" box="[151,706,1883,1910]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3917FF8DB1B00D5FFD119F00" bold="true" box="[151,706,1883,1910]" pageId="31" pageNumber="32">
<emphasis id="48B25C2C3917FF8DB1B00D5FFE189F03" bold="true" box="[151,459,1884,1910]" italics="true" pageId="31" pageNumber="32">Isophya modesta modesta</emphasis>
(Frivaldszky, 1867)
</emphasis>
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</heading>
</paragraph>
<paragraph id="7A79803E3917FF8DB1B00D7DFDFC9FE3" blockId="31.[151,706,1883,1942]" box="[151,559,1918,1942]" pageId="31" pageNumber="32">
(
<figureCitation id="E2FD9CBB3917FF8DB1B80D7DFF269FE3" box="[159,245,1918,1942]" captionStart="FIGURES 1 18" captionStartId="5.[151,264,1882,1905]" captionTargetBox="[167,1425,179,1856]" captionTargetId="figure@5.[166,1428,177,1859]" captionTargetPageId="5" captionText="FIGURES 1 18. General appearance of some Balkan Isophya. 1, 2 — I. hospodar (1 — male, 2 — female); 3 — I. thracica (male); 4, 5 — I. rectipennis (4 — male, 5 — female); 6, 7 — I. pavelii (6 — male, 7 — female); 8, 9 — I. bureschi (8 — male, 9 — female); 10, 11 — I. andreevae (10 — male, 11 — female); 12, 13 — I. plevnensis (12 — male, 13 — female); 14, 15 — I. m. modesta (= intermedia) (14 — male, 15 — female); 16 — I. l. longicaudata (male); 17 — I. rhodopensis leonorae (male); 18 — I. rh. petkovi (male) (Figures 6 11, 17, 18 — photo M. Langourov)." httpUri="https://zenodo.org/record/246552/files/figure.png" pageId="31" pageNumber="32">Figs 16</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB0260D7DFEF39FE3" box="[257,288,1918,1942]" captionStart="FIGURES 31 55" captionStartId="12.[151,264,244,267]" captionText="FIGURES 31 55. Morphology of male head, pronotum and tegmina of the Balkan Isophya and I. yaraligozi: 31 — I. hospodar (BG: Momchilgrad, 31.05. 1965, NMNHS); 32 — I. rectipennis (BG: Ropotamo, 7.07. 1965, NMNHS); 33 — I. pavelii (holotype of I. rammei Peshev, BG: Chernogorovo, 19.06. 1973, NMNHS); 34 — I. thracica (TR: Elmali, 3.07. 2011, CC); 35 — I. bureschi (BG: “ Pirin, 1300 1450 m ”, 17.08. 1961, NMNHS); 36 — I. yaraligozi (TR: Yaraligoz Pass, 9.07. 2011, CC); 37 — I. tosevski (MK: Moklishte, 30.05. 2010, CC); 38 — I. andreevae (holotype, BG: Kresna Gorge, 2.06. 1961, NMNHS); 39 — I. clara (BA: Sarajevo, NMW); 40 — I. miksici (holotype, BG: Berkovitsa, 700 m, 21.06. 1958, NMNHS); 41 — I. plevnensis (holotype, BG: Chernelka, 30.07. 1973, NMNHS); 42 — I. longicaudata adamovici (holotype, BG: “ Sliven ”, NMNHS); 43 — I. l. longicaudata (BG: Batova Gora, 26.06. 1956, NMNHS); 44 — I. m. modesta (= intermedia) (RO: Dobrogea, Babadag, CI); 45 — I. rhodopensis leonorae (BG: Popovi Livadi, 1300 m, 4.07. 1961, NMNHS); 46 — I. rh. rhodopensis (BG: Chepelare, 16.07. 1956, NMNHS); 47 — I. rh. petkovi (BG: Mandritsa, 4.06. 1965, NMNHS); 48 — I. modestior (BG: Kopitoto, 17.07. 1958, NMNHS); 49 — I. dobrogensis (RO: Popina Island, CI); 50 — I. zubowskii (RO: “ Jasi pād, Birnari, 25.06. 1956, NMNHS); 51 — I. aff. camptoxypha (“ Montenegro ”, NMW); 52 — I. gulae (BG: Dolna Topchiya, 23.06. 2008, NMNHS); 53 — I. obtusa (BG: Vezhen Peak, 1900 m, 1.09. 1960, NMNHS); 54 — I. amplipennis (TR: “ Bursa | Gursu, | Kazanci &amp; | Eskibaglar | 14. vi. 1959 ”, “ by Dr. E. Can. | B. M. 1960 173 ”, NHM); 55 — I. speciosa (BG: Plevoun, 5.06. 1965, NMNHS)." pageId="31" pageNumber="32">44</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB00C0D7DFE9A9FE3" box="[299,329,1918,1942]" captionStart="FIGURES 80 104" captionStartId="14.[151,264,1835,1858]" captionTargetBox="[226,1360,681,1815]" captionTargetId="figure@14.[226,1362,681,1816]" captionTargetPageId="14" captionText="FIGURES 80 104. Male abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55): 80 — I. hospodar; 81 — I. rectipennis; 82 — I. pavelii; 83 — I. thracica; 84 — I. bureschi; 85 — I. yaraligozi; 86 — I. tosevski; 87 — I. andreevae; 88 — I. clara; 89 — I. miksici; 90 — I. plevnensis; 91 — I. longicaudata adamovici; 92 — I. l. longicaudata; 93 — I. m. modesta (= intermedia); 94 — I. rhodopensis leonorae; 95 — I. rh. rhodopensis; 96 — I. rh. petkovi; 97 — I. modestior; 98 — I. dobrogensis; 99 — I. zubowskii; 100 — I. aff. camptoxypha; 101 — I. gulae; 102 — I. obtusa; 103 — I. amplipennis; 104 — I. speciosa. Scale (if present) = 2 mm." httpUri="https://zenodo.org/record/246555/files/figure.png" pageId="31" pageNumber="32">93</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB0710D7DFE529FE3" box="[342,385,1918,1942]" captionStart="FIGURES 105 129" captionStartId="17.[151,264,1173,1196]" captionTargetBox="[151,1436,195,1152]" captionTargetId="figure@17.[151,1436,195,1152]" captionTargetPageId="17" captionText="FIGURES 105 129. Female abdominal terminalia of the Balkan Isophya and I. yaraligozi (localities as in Figs 31 55 unless specifically given); figures of the whole ovipositor are followed by enlarged figure of the ovipositor base: 105 — I. hospodar (BG: Dolni Glavanak, 25.04. 2005, CC); 106 — I. rectipennis (BG: Byala Vill., 27.06. 2002, CC); 107 — I. pavelii (BG: Chernogorovo, 29.05. 2006, CC); 108 — I. thracica; 109 — I. bureschi (BG: Bansko, 8.08. 2006, CC); 110 — I. yaraligozi; 111 — I. tosevski; 112 — I. andreevae (BG: Kresnensko Hanche, 19.04. 2006, CC); 113 — I. clara; 114 — I. miksici (BG: Gorski Dom Lodge, 24.06. 2006, CC); 115 — I. plevnensis (BG: Levishte, 14.07. 2009, CC); 116 — I. longicaudata adamovici (BG: Karandila, 24.06. 2008, CC); 117 — I. l. longicaudata (BG: Sofia University Botanical garden, 26.06. 1.07.2004, CC); 118 — I. m. modesta (= intermedia); 119 — I. rhodopensis leonorae (BG: Livade, 9.08. 2006, CC); 120 — I. rh. rhodopensis (BG: Bachkovo, 600 m, 23.05. 2004, CC); 121 — I. rh. petkovi (BG: Gluhite Kamani, 23.06. 2008, CC); 122 — I. modestior (BG: Bankya, 24.05. 2006, CC); 123 — I. dobrogensis; 124 — I. zubowskii; 125 — I. aff. camptoxypha (data as in Fig. 75); 126 — I. gulae; 127 — I. obtusa (BG: Ravnets Ridge, 14.07. 2009, CC); 128 — I. amplipennis (as in Fig. 78); 129 — I. speciosa (BG: Bankya, 24.05. 2006, CC). Indications on Fig. 127 show, respectively: G—gonangulum; L—lamella; P—lateral pit between lamella and gonangulum. Scale (if present) = 5 mm." httpUri="https://zenodo.org/record/246556/files/figure.png" pageId="31" pageNumber="32">118</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB0AB0D7DFE6B9FE3" box="[396,440,1918,1942]" captionStart="FIGURES 150 155" captionStartId="27.[151,264,1462,1485]" captionTargetBox="[151,1435,193,1440]" captionTargetId="figure@27.[151,1436,193,1441]" captionTargetPageId="27" captionText="FIGURES 150 155. SEM of taxonomically important structures of Isophya: 150, 151 — I. miksici (150 — BG: Gorski Dom Lodge; 151 — BG: Iskar); 152 — I. plevnensis (152 A—BG: Chernelka Canyon; 152 B, C—BG: Apriltsi); 153 — I. longicaudata adamovici (BG: Karandila Locality); 154 — I. longicaudata longicaudata (BG: Bolata Valley); 155 — I. m. modesta (= intermedia) (RO: Babadag). A—male stridulatory file (scale 1 mm); B—ventro-apical view of apex of male cerci (152 B deformed) (scale 100 μm); C—female stridulatory apparatus (scale 500 μm)." httpUri="https://zenodo.org/record/246561/files/figure.png" pageId="31" pageNumber="32">155</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB0E30D7DFE239FE3" box="[452,496,1918,1942]" captionStart="FIGURES 156 161" captionStartId="28.[151,264,1682,1705]" captionTargetBox="[151,1436,331,1661]" captionTargetId="figure@28.[151,1436,331,1662]" captionTargetPageId="28" captionText="FIGURES 156 161. Male calling song at two different speeds (air temperature at recording shown; time scale below): 156 — I. miksici (BG: Gorski Dom Lodge); 157 — I. plevnensis (BG: Apriltsi); 158, 159 — I. longicaudata adamovici (BG: Karandila Locality, two males); 160 — I. longicaudata longicaudata (BG: Bolata Valley); 161 — I. modesta modesta (RO: Babadag). Break-signs in Fig. 161 B indicate the missing silent interval of the syllable shown in Fig. 161 A." httpUri="https://zenodo.org/record/246562/files/figure.png" pageId="31" pageNumber="32">161</figureCitation>
,
<figureCitation id="E2FD9CBB3917FF8DB0DB0D7DFDF49FE3" box="[508,551,1918,1942]" captionStart="FIGURE 192" captionStartId="48.[151,250,1249,1272]" captionTargetBox="[151,1435,445,1228]" captionTargetId="figure@48.[151,1436,445,1228]" captionTargetPageId="48" captionText="FIGURE 192. Distribution of I. bureschi, I. andreevae, I. tosevski, I. clara, I. miksici, I. plevnensis, I. longicaudata adamovici, I. l. longicaudata, and I. modesta on the Balkan Peninsula." httpUri="https://zenodo.org/record/246569/files/figure.png" pageId="31" pageNumber="32">192</figureCitation>
)
</paragraph>
<paragraph id="7A79803E3917FF8DB1B00DC4FD0E9FA8" blockId="31.[151,733,1990,2013]" box="[151,733,1990,2013]" pageId="31" pageNumber="32">
<taxonomicName id="BDC6FBBD3917FF8DB1B00DC4FE069FA9" authority="Frivaldszky" authorityName="Frivaldszky" box="[151,469,1990,2013]" class="Insecta" family="Phaneropteridae" genus="Odontura" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C3917FF8DB1B00DC4FE879FA8" box="[151,340,1991,2013]" italics="true" pageId="31" pageNumber="32">Odontura modesta</emphasis>
Frivaldszky
</taxonomicName>
: Frivaldszky 1867 (sp.n.).
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53928FFB2B1B00A9BFDEA9AB4" pageId="32" pageNumber="33" type="description">
<paragraph id="7A79803E3928FFB2B1B00A9BFD4898DB" blockId="32.[151,748,152,205]" box="[151,667,152,174]" pageId="32" pageNumber="33">
<taxonomicName id="BDC6FBBD3928FFB2B1B00A9BFDFC98DB" authority="Frivaldszky" authorityName="Frivaldszky" box="[151,559,152,174]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3928FFB2B1B00A9BFE4F98DB" box="[151,412,152,174]" italics="true" pageId="32" pageNumber="33">Isophya modesta modesta</emphasis>
(Frivaldszky)
</taxonomicName>
: Kis 1960.
</paragraph>
<paragraph id="7A79803E3928FFB2B1B00AB4FD3F98B8" blockId="32.[151,748,152,205]" box="[151,748,182,205]" pageId="32" pageNumber="33">
<taxonomicName id="BDC6FBBD3928FFB2B1B00AB4FE3198B9" authority="Kis" authorityName="Kis" box="[151,482,182,205]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="intermedia">
<emphasis id="48B25C2C3928FFB2B1B00AB4FE6598B8" box="[151,438,183,205]" italics="true" pageId="32" pageNumber="33">Isophya modesta intermedia</emphasis>
Kis
</taxonomicName>
,
<emphasis id="48B25C2C3928FFB2B0CB0AB4FDF898B9" bold="true" box="[492,555,183,204]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="5381E1573928FFB2B0CB0AB4FDF898B9" box="[492,555,183,204]" pageId="32" pageNumber="33">syn.n.</taxonomicNameLabel>
</emphasis>
: Kis 1960 (ssp.n.).
</paragraph>
<paragraph id="7A79803E3928FFB2B1B00AFAFE7A9940" blockId="32.[151,1436,249,885]" pageId="32" pageNumber="33">
<emphasis id="48B25C2C3928FFB2B1B00AFAFE179964" box="[151,452,249,273]" italics="true" pageId="32" pageNumber="33">Morphological description</emphasis>
: see the references above; Harz 1969; Heller
<emphasis id="48B25C2C3928FFB2B2EF0AF9FBD29964" box="[968,1025,249,273]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2004.
<emphasis id="48B25C2C3928FFB2B5690AF9FB0E9964" box="[1102,1245,250,273]" italics="true" pageId="32" pageNumber="33">Bioacoustics</emphasis>
: Orci and Heller 2004; Heller
<emphasis id="48B25C2C3928FFB2B00C0B1DFEB09940" box="[299,355,285,309]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2004.
</paragraph>
<paragraph id="7A79803E3928FFB2B1E00B41FDEA9AB4" blockId="32.[151,1436,249,885]" pageId="32" pageNumber="33">
<emphasis id="48B25C2C3928FFB2B1E00B41FEEA992C" box="[199,313,322,345]" italics="true" pageId="32" pageNumber="33">Synonymy</emphasis>
: Kis (1960) described
<taxonomicName id="BDC6FBBD3928FFB2B3640B41FCEF992C" box="[579,828,321,345]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="intermedia">
<emphasis id="48B25C2C3928FFB2B3640B41FCEF992C" box="[579,828,321,345]" italics="true" pageId="32" pageNumber="33">I. modesta intermedia</emphasis>
</taxonomicName>
on the basis of slight morphological differences and different colouration of male tegmina recognising it from
<taxonomicName id="BDC6FBBD3928FFB2B2130B65FBC19908" box="[820,1042,357,381]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3928FFB2B2130B65FBC19908" box="[820,1042,357,381]" italics="true" pageId="32" pageNumber="33">I. modesta modesta</emphasis>
</taxonomicName>
. At the time of this publication
<taxonomicName id="BDC6FBBD3928FFB2B4AB0B65FF2799D4" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C3928FFB2B4AB0B65FF2799D4" italics="true" pageId="32" pageNumber="33">I. modesta</emphasis>
</taxonomicName>
was known from few localities in Central and Southwestern
<collectingCountry id="02D1C0AE3928FFB2B2F90B8AFB9099D4" box="[990,1091,393,417]" name="Romania" pageId="32" pageNumber="33">Romania</collectingCountry>
and little was known on its morphological variation. Investigation of the song of typical
<taxonomicName id="BDC6FBBD3928FFB2B2780BADFBCB99B0" box="[863,1048,429,453]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="intermedia">
<emphasis id="48B25C2C3928FFB2B2780BADFBCB99B0" box="[863,1048,429,453]" italics="true" pageId="32" pageNumber="33">I. m. intermedia</emphasis>
</taxonomicName>
showed it coincides with that of typical
<taxonomicName id="BDC6FBBD3928FFB2B1CC0BD1FE51999C" box="[235,386,465,489]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3928FFB2B1CC0BD1FE51999C" box="[235,386,465,489]" italics="true" pageId="32" pageNumber="33">I. m. modesta</emphasis>
</taxonomicName>
. The measurements at 23°C were as follows: main part of syllables—56170 ms with 1526 impulses and impulse interval of 38 ms; second part—6401340 ms with 2542 impulses and impulse interval of 4(at the end)127 ms. Both parts were separated by an interval of about 46 seconds and thus the total syllable duration was 53007460 ms. These characteristics fit well the measurements at higher temperatures given by Orci and Heller (2004) except the shorter main syllable part. However, regarding the great variability of the temporal song characteristics of
<taxonomicName id="BDC6FBBD3928FFB2B08F0885FD9A9AE8" box="[424,585,645,669]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3928FFB2B08F0885FD9A9AE8" box="[424,585,645,669]" italics="true" pageId="32" pageNumber="33">I. m. modesta</emphasis>
</taxonomicName>
(compare Orci and Heller 2004) we consider
<taxonomicName id="BDC6FBBD3928FFB2B5510885FAE69AE8" box="[1142,1333,645,669]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="intermedia">
<emphasis id="48B25C2C3928FFB2B5510885FAE69AE8" box="[1142,1333,645,669]" italics="true" pageId="32" pageNumber="33">I. m. intermedia</emphasis>
</taxonomicName>
a junior subjective synonym of
<taxonomicName id="BDC6FBBD3928FFB2B0BC08A9FDE19AB4" box="[411,562,681,705]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="modesta" subSpecies="modesta">
<emphasis id="48B25C2C3928FFB2B0BC08A9FDE19AB4" box="[411,562,681,705]" italics="true" pageId="32" pageNumber="33">I. m. modesta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53928FFB2B1E008CDFB729A90" box="[199,1185,717,741]" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="7A79803E3928FFB2B1E008CDFB729A90" blockId="32.[151,1436,249,885]" box="[199,1185,717,741]" pageId="32" pageNumber="33">
<emphasis id="48B25C2C3928FFB2B1E008CDFEEA9A90" box="[199,313,718,741]" italics="true" pageId="32" pageNumber="33">Diagnosis</emphasis>
: see the data for
<taxonomicName id="BDC6FBBD3928FFB2B0DF08CDFDBE9A90" box="[504,621,717,741]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="modesta">
<emphasis id="48B25C2C3928FFB2B0DF08CDFDBE9A90" box="[504,621,717,741]" italics="true" pageId="32" pageNumber="33">I. modesta</emphasis>
</taxonomicName>
. Recognition from
<emphasis id="48B25C2C3928FFB2B26F08CDFBFE9A90" box="[840,1069,717,741]" italics="true" pageId="32" pageNumber="33">
<taxonomicName id="BDC6FBBD3928FFB2B26F08CDFC6E9A90" box="[840,957,717,741]" class="Insecta" family="Phaneropteridae" genus="Isophya" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="modesta">I. modesta</taxonomicName>
rossica—
</emphasis>
see below.
</paragraph>
</subSubSection>
<subSubSection id="32DCD3B53928FFB2B1E008F2FF0B9B00" pageId="32" pageNumber="33" type="distribution">
<paragraph id="7A79803E3928FFB2B1E008F2FF0B9B00" blockId="32.[151,1436,249,885]" pageId="32" pageNumber="33">
<emphasis id="48B25C2C3928FFB2B1E008F2FD5D9B7C" box="[199,654,753,777]" italics="true" pageId="32" pageNumber="33">
Distribution (
<figureCitation id="E2FD9CBB3928FFB2B04108F2FE1E9B7C" box="[358,461,753,777]" captionStart="FIGURE 192" captionStartId="48.[151,250,1249,1272]" captionTargetBox="[151,1435,445,1228]" captionTargetId="figure@48.[151,1436,445,1228]" captionTargetPageId="48" captionText="FIGURE 192. Distribution of I. bureschi, I. andreevae, I. tosevski, I. clara, I. miksici, I. plevnensis, I. longicaudata adamovici, I. l. longicaudata, and I. modesta on the Balkan Peninsula." httpUri="https://zenodo.org/record/246569/files/figure.png" pageId="32" pageNumber="33">Fig. 192</figureCitation>
) and phenology
</emphasis>
: Known from
<collectingCountry id="02D1C0AE3928FFB2B26008F2FC7F9B7C" box="[839,940,753,777]" name="Slovakia" pageId="32" pageNumber="33">Slovakia</collectingCountry>
,
<collectingCountry id="02D1C0AE3928FFB2B29A08F1FBF39B7F" box="[957,1056,754,778]" name="Hungary" pageId="32" pageNumber="33">Hungary</collectingCountry>
,
<collectingCountry id="02D1C0AE3928FFB2B51508F2FB499B7C" box="[1074,1178,753,777]" name="Romania" pageId="32" pageNumber="33">Romania</collectingCountry>
, Western and Central
<collectingCountry id="02D1C0AE3928FFB2B1B00916FF279B58" box="[151,244,789,813]" name="Ukraine" pageId="32" pageNumber="33">Ukraine</collectingCountry>
, and possibly occurs also in
<collectingCountry id="02D1C0AE3928FFB2B36E0916FD619B58" box="[585,690,789,813]" name="Moldova" pageId="32" pageNumber="33">Moldova</collectingCountry>
. The subspecies prefers lowland and hilly terrains, rarely it was found in the middle mountain belt in mesophyte grass-bush associations. Nymphs—(III)IVV, imago—VVII( VIII).
</paragraph>
</subSubSection>
</treatment>
</document>