treatments-xml/data/DE/A3/34/DEA3346520AD5089A5330148CDBDED5E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.83916" ID-GBIF-Dataset="e036b25d-83e3-4247-9c3b-41ad2a84b72a" ID-PMC="PMC9043006" ID-Pensoft-Pub="1313-2970-1097-1" ID-Pensoft-UUID="6593D2730A295F46B63A738B12E43A22" ID-PubMed="35837583" ID-ZooBank="66F17436DF364FFDB2D3021F14D40D62" ModsDocID="1313-2970-1097-1" checkinTime="1650371395093" checkinUser="pensoft" docAuthor="Antic, Dragan, Vagalinski, Boyan, Stoev, Pavel &amp; Akkari, Nesrine" docDate="2022" docId="DEA3346520AD5089A5330148CDBDED5E" docLanguage="en" docName="ZooKeys 1097: 1-46" docOrigin="ZooKeys 1097" docPubDate="2022-04-19" docSource="http://dx.doi.org/10.3897/zookeys.1097.83916" docTitle="Balkanodesminus dentatoides gen. nov. et 2022, sp. nov." docType="treatment" docUuid="BEF9EB03-1DC8-4C13-BE0F-DE8A2AD0C257" docUuidSource="ZooBank" docVersion="4" id="6593D2730A295F46B63A738B12E43A22" lastPageNumber="1" masterDocId="6593D2730A295F46B63A738B12E43A22" masterDocTitle="A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="1" updateTime="1668151688527" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Antic, Dragan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1231-4213</mods:nameIdentifier>
<mods:affiliation>University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski trg 16, 11000 Belgrade, Serbia &amp; Serbian Biospeleological Society, Trg Dositeja Obradovica 2, 21000 Novi Sad, Serbia</mods:affiliation>
<mods:nameIdentifier type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vagalinski, Boyan</mods:namePart>
<mods:affiliation>Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stoev, Pavel</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5702-5677</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd. 1, Sofia 1000, Bulgaria &amp; Pensoft Publishers, Sofia, Bulgaria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Akkari, Nesrine</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5019-4833</mods:nameIdentifier>
<mods:affiliation>3</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-04-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1097</mods:number>
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<mods:start>1</mods:start>
<mods:end>46</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1097.83916</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1097.83916</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1097-1</mods:identifier>
<mods:identifier type="ZooBank">66F17436DF364FFDB2D3021F14D40D62</mods:identifier>
<mods:identifier type="Pensoft-UUID">6593D2730A295F46B63A738B12E43A22</mods:identifier>
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<treatment ID-GBIF-Taxon="194660820" LSID="urn:lsid:zoobank.org:act:BEF9EB03-1DC8-4C13-BE0F-DE8A2AD0C257" httpUri="http://treatment.plazi.org/id/DEA3346520AD5089A5330148CDBDED5E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/BEF9EB03-1DC8-4C13-BE0F-DE8A2AD0C257" authority="gen. nov. et" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides" status="sp. nov.">Balkanodesminus dentatoides gen. nov. et</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">Figs 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Balkanodesminus dentatoides gen. nov. et sp. nov., right gonopods A-E paratype ♂ (NHMW MY 10258) A mesal view B lateral view C meso-distal view D antero-distal view E tip of solenomere, distal view F, G ♂ from Sopotska peshtera Cave (NHMW MY 10267), mesal and antero-disto-lateral views, respectively. Abbreviations: a acropodite, ap acuminate process of solenophore, bl basal lamellar lobe of solenophore, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, mt mesal thickening of solenophore, pf prefemorite, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore, t mesal tooth of distal solenomeral process. Scale bars: 0.02 mm (A-D, F, G), 0.005 mm (E)." figureDoi="10.3897/zookeys.1097.83916.figure9" httpUri="https://binary.pensoft.net/fig/674287" pageId="0" pageNumber="1">, 9</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">, 17M, N</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">, 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari, 2022" authorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" authorityYear="2022" baseAuthorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" baseAuthorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus bulgaricus subsp. dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="bulgaricus" status="sp. nov." subSpecies="dentatus">Bacillidesmus bulgaricus dentatus</taxonomicName>
- 
<bibRefCitation author="Stoev, P" journalOrPublisher="Acta Zoologica Bulgarica" pageId="0" pageNumber="1" pagination="145 - 154" refId="B50" refString="Stoev, P, 2004. New distributional records of millipedes from Bulgarian caves (Myriapoda: Diplopoda). Acta Zoologica Bulgarica 56 (2): 145 - 154" title="New distributional records of millipedes from Bulgarian caves (Myriapoda: Diplopoda)." volume="56" year="2004">Stoev 2004</bibRefCitation>
: 149.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authority="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari, 2022" authorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" authorityYear="2022" baseAuthorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" baseAuthorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus bulgaricus subsp. dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="bulgaricus" status="sp. nov." subSpecies="dentatus">Bacillidesmus bulgaricus dentatus</taxonomicName>
in part.- 
<bibRefCitation DOI="https://doi.org/10.1007/978-1-4020-5781-6_11" author="Stoev, P" editor="Fet, V" journalOrPublisher="Vol. 82. Biogeography and Ecology of Bulgaria. Springer, Dordrecht" pageId="0" pageNumber="1" pagination="379 - 404" refId="B51" refString="Stoev, P, 2007. Fauna and zoogeography of Myriapoda in Bulgaria. In: Fet, V, Popov, A, Eds., Monographiae Biologicae. Vol. 82. Biogeography and Ecology of Bulgaria. Springer, Dordrecht: 379 - 404, DOI: https://doi.org/10.1007/978-1-4020-5781-6_11" title="Fauna and zoogeography of Myriapoda in Bulgaria." url="https://doi.org/10.1007/978-1-4020-5781-6_11" volumeTitle="Monographiae Biologicae." year="2007">Stoev 2007</bibRefCitation>
: 384; 
<bibRefCitation author="Beron, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" refId="B10" refString="Beron, P, 2015. Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." title="Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." year="2015">Beron 2015</bibRefCitation>
: 80, 411; 
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4263.3.4" author="Bachvarova, D" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="507 - 526" refId="B9" refString="Bachvarova, D, Vagalinski, B, Doichinov, A, Stoev, P, 2017. New records of millipedes and centipedes from Bulgaria, with an annotated checklist of the Bulgarian myriapods. Zootaxa 4263 (3): 507 - 526, DOI: https://doi.org/10.11646/zootaxa.4263.3.4" title="New records of millipedes and centipedes from Bulgaria, with an annotated checklist of the Bulgarian myriapods." url="https://doi.org/10.11646/zootaxa.4263.3.4" volume="4263" year="2017">Bachvarova et al. 2017</bibRefCitation>
: 521; 
<bibRefCitation author="Kime, RD" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="0" pageNumber="1" refId="B35" refString="Kime, RD, Enghoff, H, 2011. Atlas of European millipedes (Class Diplopoda). Vol. 1. Pensoft Publishers, Sofia-Moscow" title="Atlas of European millipedes (Class Diplopoda). Vol. 1." year="2011">Kime and Enghoff 2011</bibRefCitation>
: 71, 262.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Differs from 
<taxonomicName authorityName="gen. nov., comb. nov. ex Bacillidesmus" baseAuthorityName="Strasser" baseAuthorityYear="1962" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus bulgaricus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bulgaricus">
<emphasis italics="true" pageId="0" pageNumber="1">Balkanodesminus bulgaricus</emphasis>
</taxonomicName>
gen. nov., comb. nov. and 
<taxonomicName lsidName="B. serbicus" pageId="0" pageNumber="1" rank="species" species="serbicus">
<emphasis italics="true" pageId="0" pageNumber="1">B. serbicus</emphasis>
</taxonomicName>
gen. nov. et sp. nov. by the presence of shorter (vs. longer) metatergal setae and their greater (vs. smaller) number of rows, as well as by the presence of more complicated (vs. more simplified) gonopods, with biramous (vs. uniramous) distal solenomeral process and larger and denticulated (vs. smaller and smooth) lamella of solenophore. From 
<taxonomicName lsidName="B. dentatus" pageId="0" pageNumber="1" rank="species" species="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">B. dentatus</emphasis>
</taxonomicName>
gen. nov., comb. nov., stat. nov., with which it shares similar habitus and similar gonopods, it differs by the presence of slenderer (vs. more robust) solenomere, by small triangular tooth (vs. short process) on distal solenomeral process, and by the absence (vs. presence) of additional short subdistal process at distal projection of solenophore. In addition, lateral lamella and basal lobe more robust than in 
<taxonomicName lsidName="B. dentatus" pageId="0" pageNumber="1" rank="species" species="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">B. dentatus</emphasis>
</taxonomicName>
gen. nov., comb. nov., stat. nov.
</paragraph>
<caption doi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 7.</emphasis>
<taxonomicName authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
<emphasis italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
</taxonomicName>
gen. nov. et sp. nov., holotype ♂ (
<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
, NMNHS-10815) and paratype ♂ (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
, NHMW MY10258), habitus 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
anterior part of body, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior part of body, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
midbody rings, dorsal view. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="name">
<paragraph pageId="0" pageNumber="1">Name.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species name is a combination of the name 
<taxonomicName authorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" authorityYear="2022" baseAuthorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" baseAuthorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">Bacillidesmus dentatus</emphasis>
</taxonomicName>
and the Ancient Greek suffix - 
<emphasis italics="true" pageId="0" pageNumber="1">oides</emphasis>
, referring to the 
<normalizedToken originalValue="species’">species'</normalizedToken>
particularly strong resemblance to 
<taxonomicName authorityName="gen. nov., comb. nov." authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1966" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatus</emphasis>
</taxonomicName>
gen. nov., comb. nov., stat. nov. Adjective.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1997-10-13" collectorName="B. Petrov, P. Stoev." country="Bulgaria" county="Yablanitsa Municipality" location="Saeva" municipality="Brestnitsa" specimenCount="♂" stateProvince="Lovech District" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
<specimenCount>♂</specimenCount>
(NMNHS-10815), 
<collectingCountry name="Bulgaria">Bulgaria</collectingCountry>
, 
<collectingRegion country="Bulgaria" name="Lovech">Lovech District</collectingRegion>
, 
<collectingCounty>Yablanitsa Municipality</collectingCounty>
, 
<collectingMunicipality>Brestnitsa</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:DEA3346520AD5089A5330148CDBDED5E:B600A078872833C511EBD1215C38C6AF" country="Bulgaria" county="Yablanitsa Municipality" municipality="Brestnitsa" name="Saeva" stateProvince="Lovech District">Saeva</location>
dupka 
<location LSID="urn:lsid:plazi:treatment:DEA3346520AD5089A5330148CDBDED5E:43F37FFD986FACDF0F8E9CAB7A50F383" country="Bulgaria" county="Yablanitsa Municipality" municipality="Brestnitsa" name="Cave" stateProvince="Lovech District">Cave</location>
, clay, 
<collectingDate value="1997-10-13">13.X.1997</collectingDate>
, leg. 
<collectorName>B. Petrov</collectorName>
&amp; 
<collectorName>P. Stoev.</collectorName>
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount type="male">2 ♂♂</specimenCount>
(NMNHS-10816, 10817), 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="B. Petrov &amp; P. Stoev." country="Bulgaria" county="Yablanitsa Municipality" location="Saeva" municipality="Brestnitsa" specimenCount="1" specimenCount-female="1" stateProvince="Lovech" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
, (NMNHS-10818), same data as for holotype
</materialsCitation>
; 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="B. Petrov &amp; P. Stoev." country="Bulgaria" county="Yablanitsa Municipality" location="Saeva" municipality="Brestnitsa" specimenCount="1" specimenCount-male="1" stateProvince="Lovech" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
(used for SEM, NHMW MY10258) same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<specimenCount type="male">1 ♂</specimenCount>
(right gonopod used for SEM, NHMW MY10267), 
<specimenCount type="male">2 ♂♂</specimenCount>
(NMNHS-10819, 10820), 
<materialsCitation collectingDate="2004-05-08" collectorName="P. Beron." county="Cave" location="Lovech District" municipality="Sopotska" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2 ♀♀</specimenCount>
(NMNHS-10821, 10822), 
<location LSID="urn:lsid:plazi:treatment:DEA3346520AD5089A5330148CDBDED5E:E6FE2838079785F130743CE4ACFE96BC" county="Cave" municipality="Sopotska" name="Lovech District">Lovech District</location>
, village of 
<location LSID="urn:lsid:plazi:treatment:DEA3346520AD5089A5330148CDBDED5E:C07255CCBD106DBE3BAA975D167415F0" county="Cave" municipality="Sopotska" name="Sopot">Sopot</location>
, 
<collectingMunicipality>Sopotska</collectingMunicipality>
peshtera 
<collectingCounty>Cave</collectingCounty>
, 
<collectingDate value="2004-05-08">8.V.2004</collectingDate>
, leg. 
<collectorName>P. Beron.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Number of body rings and measurements</emphasis>
: Body with 19 rings (including telson) in adults, moniliform. Holotype male 3.8 mm long, width of midbody pro- and metazonae 0.25 mm and 0.30 mm, respectively. Paratype males 3.4-3.9 mm long, width of midbody pro- and metazonae 0.23-0.25 mm and 0.29-0.31 mm, respectively. Paratype female 4.3 mm long, width of midbody pro- and metazona 0.30 and 0.35 mm, respectively.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
: Entirely pallid, slightly translucent (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: Broader than collum, setose (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7B</figureCitation>
); epicranial suture poorly developed; isthmus between antennae ≈ 1.3 
<normalizedToken originalValue="×">x</normalizedToken>
broader than diameter of antennal socket. Labrum with three labral teeth, and with 3+3 labral and five supralabral setae (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8B</figureCitation>
). Gnathochilarium without peculiarities. Antennae rather short, clavate (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7B</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8C, D</figureCitation>
). Antennomere length 6&gt; 2 = 3 = 4&gt; 5&gt; 7 = 1. Antennae 0.6 mm long in the holotype male; length/breadth ratios of antennomeres 1-7: 1 (1), 2 (2), 3 (2), 4 (2), 5 (1), 6 (1) and 7 (1). Antennomere 6 with four sensilla trichodea and with strongly developed disto-dorsal pit with numerous long sensilla basiconica partially exposed outside the pit (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8D</figureCitation>
). Antennomere 7 with one sensillum trichodeum and a small bulge with three sensilla basiconica spiniformia (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8C, D</figureCitation>
). Four apical cones (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8D</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 8.</emphasis>
<taxonomicName authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
<emphasis italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
</taxonomicName>
gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY10258) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior part of body, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
labrum, anterior view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) 
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
body rings 8-10, lateral and dorsal views, respectively 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
right ozopore 10, lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">H, I</emphasis>
body rings 17-19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, C, E, F, H, I</emphasis>
), 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B, D</emphasis>
), 0.01 mm (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Collum</emphasis>
: Semi-circular, with one or two lateral incisions and ≈ 7 irregular rows of medium sized and trichoid setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Body rings</emphasis>
: Tegument shining, texture alveolate, reticulate and scaly. Rings densely setose. Setae rather short and trichoid, originating from small tubercules (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8A, E-G, I</figureCitation>
). Posteriormost tubercules mostly with a small thorn. Rings 2-4 with ≈ 4 rows of setae (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8A</figureCitation>
). Rings 5-18 with ≈ 6-8 irregular rows of setae (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8E-G, I</figureCitation>
). Paraterga serrated, with 5-7 teeth (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7D</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8E-G, I</figureCitation>
). Pore formula normal: 5, 7, 9, 10, 12, 13, 15-18. Poriferous metazonae with an enlarged postero-lateral cone bearing an ozopore and three medium-sized setae (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8H</figureCitation>
). Epiproct blunt, directed slightly ventrad (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7C</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8F, I</figureCitation>
). Paraprocts semi-spherical, each with two long setae originating from small tubercules and ≈ 10 shorter setae without tubercules (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8I</figureCitation>
). Hypoproct trapeziform with 2 long distal setae and numerous shorter setae throughout (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Balkanodesminus dentatoides gen. nov. et sp. nov., holotype ♂ (A-C, NMNHS- 10815) and paratype ♂ (D, NHMW MY 10258), habitus A lateral view B anterior part of body, lateral view C posterior part of body, lateral view D midbody rings, dorsal view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure7" httpUri="https://binary.pensoft.net/fig/674285" pageId="0" pageNumber="1">7C</figureCitation>
). Sterna unmodified, poorly setose. Pleurosternal carinae absent, only a few small teeth sometimes present on rings 2 and 3 (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Balkanodesminus dentatoides gen. nov. et sp. nov., paratype ♂ habitus (NHMW MY 10258) A anterior part of body, lateral view B labrum, anterior view C right antenna, anterior view (arrow indicates small bulge with sensilla basiconica spiniformia) D distal antennomeres of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) E, F body rings 8 - 10, lateral and dorsal views, respectively G right ozopore 10, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively. Scale bars: 0.1 mm (A, C, E, F, H, I), 0.05 mm (B, D), 0.01 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure8" httpUri="https://binary.pensoft.net/fig/674286" pageId="0" pageNumber="1">8A</figureCitation>
). Gonopod aperture large, subsemi-circular.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Walking legs</emphasis>
: Legs 1-3 in males with swollen femur; coxa 2 with a short mesal apophysis (cf. 
<bibRefCitation author="Strasser, K" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="1" pagination="325 - 385" refId="B54" refString="Strasser, K, 1966a. Ueber Diplopoden Bulgariens. Annales Zoologici 23 (12): 325 - 385" title="Ueber Diplopoden Bulgariens." volume="23" year="1966 a">Strasser 1966a</bibRefCitation>
: 341, fig. 13). No other peculiarities.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName genus="Gonopods" lsidName="Gonopods" pageId="0" pageNumber="1" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
</taxonomicName>
(Figs 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Balkanodesminus dentatoides gen. nov. et sp. nov., right gonopods A-E paratype ♂ (NHMW MY 10258) A mesal view B lateral view C meso-distal view D antero-distal view E tip of solenomere, distal view F, G ♂ from Sopotska peshtera Cave (NHMW MY 10267), mesal and antero-disto-lateral views, respectively. Abbreviations: a acropodite, ap acuminate process of solenophore, bl basal lamellar lobe of solenophore, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, mt mesal thickening of solenophore, pf prefemorite, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore, t mesal tooth of distal solenomeral process. Scale bars: 0.02 mm (A-D, F, G), 0.005 mm (E)." figureDoi="10.3897/zookeys.1097.83916.figure9" httpUri="https://binary.pensoft.net/fig/674287" pageId="0" pageNumber="1">9</figureCitation>
, 
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">17M, N</figureCitation>
): Coxa (cx) large, semi-circular in ventral and lateral views, with differentiated gonocoel mesally; lateral part swollen, alveolate, with three long setae near mesal ridge. Cannula (ca) long, C-shaped. Telopodite relatively long compared to coxa, consisting of a transverse, setose prefemorite (pf) and a somewhat C-shaped (in lateral and mesal views) acropodite (a) longitudinally divided in the distal half into two branches, solenomeral branch (sb) and solenophore (sph). Solenomeral branch positioned mesally, with a narrow 
<normalizedToken originalValue="“neck”">&quot;neck&quot;</normalizedToken>
, then abruptly expands and transversely divides into two processes, solenomere (s) and distal solenomeral process (dsp). Extended part of solenomeral branch with spiculiform outgrowths. Solenomere (s) very long, slender, subdistally with a small bifurcation. Distal solenomeral process (dsp) extends in the same direction as solenomere and is half as long as solenomere; bifurcated=with small additional mesal tooth (t). Solenophore (sph) longer and more robust than solenomeral branch, characterized by a robust, lateral, ear-shaped lamella (ll) and a distal projection (dp). Lateral lamella (ll) begins at bifurcation of solenomeral branch and solenophore, surrounding laterally solenophore up to beginning of distal ending; lateral margins of lamella denticulated. Distal projection (dp) with strongly developed, basal lamellar lobe (bl), with mesal thickening (mt) and with relativelly short and acuminate process (ap). Seminal groove (sg) starts from seminal fossa (sf) mesally on prefemorite, extends along mesal side of acropodite up to bifurcation of solenomeral branch and solenophore, then passes on lateral side of solenomeral branch, further on solenomere, ending distally.
</paragraph>
<caption doi="10.3897/zookeys.1097.83916.figure9" httpUri="https://binary.pensoft.net/fig/674287" pageId="0" pageNumber="1" start="Figure 9" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 9.</emphasis>
<taxonomicName authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
<emphasis italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
</taxonomicName>
gen. nov. et sp. nov., right gonopods 
<emphasis bold="true" pageId="0" pageNumber="1">A-E</emphasis>
paratype ♂ (NHMW MY10258) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
mesal view 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
lateral view 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
meso-distal view 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
antero-distal view 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
tip of solenomere, distal view 
<emphasis bold="true" pageId="0" pageNumber="1">F, G</emphasis>
♂ from Sopotska peshtera Cave (NHMW MY10267), mesal and antero-disto-lateral views, respectively. Abbreviations: 
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
acropodite, 
<emphasis bold="true" pageId="0" pageNumber="1">ap</emphasis>
acuminate process of solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">bl</emphasis>
basal lamellar lobe of solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">ca</emphasis>
cannula, 
<emphasis bold="true" pageId="0" pageNumber="1">cx</emphasis>
coxa, 
<emphasis bold="true" pageId="0" pageNumber="1">dp</emphasis>
distal projection of solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">dsp</emphasis>
distal solenomeral process, 
<emphasis bold="true" pageId="0" pageNumber="1">ll</emphasis>
lamella of solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">mt</emphasis>
mesal thickening of solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">pf</emphasis>
prefemorite, 
<emphasis bold="true" pageId="0" pageNumber="1">s</emphasis>
solenomere, 
<emphasis bold="true" pageId="0" pageNumber="1">sb</emphasis>
solenomeral branch, 
<emphasis bold="true" pageId="0" pageNumber="1">sf</emphasis>
seminal fossa, 
<emphasis bold="true" pageId="0" pageNumber="1">sg</emphasis>
seminal groove, 
<emphasis bold="true" pageId="0" pageNumber="1">sph</emphasis>
solenophore, 
<emphasis bold="true" pageId="0" pageNumber="1">t</emphasis>
mesal tooth of distal solenomeral process. Scale bars: 0.02 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A-D, F, G</emphasis>
), 0.005 mm (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
Saeva dupka Cave is a show cave which is now heavily impacted by electrification and continuous touristic flow. The cave has naturally formed 400 meters of corridors and halls. The samples from the cave were taken in 1997, under stones in clay, when the cave was temporarily closed for visitors due to the change of its governance during the democratic changes in Bulgaria. After more than 20 years of active exploration of the cave, new material needs to be collected to assess whether the species was influenced by the human activities. Saeva dupka Cave is inhabited by numerous and diverse invertebrate taxa, but the only troglobiont currently on record is the local endemic 
<taxonomicName authorityName="Karaman" authorityYear="1958" class="Insecta" family="Leiodidae" genus="Bulgariella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulgariella tranteevi" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tranteevi">
<emphasis italics="true" pageId="0" pageNumber="1">Bulgariella tranteevi</emphasis>
</taxonomicName>
Z. Karaman, 1958 (
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, 
<taxonomicName authorityName="Fleming" authorityYear="1821" class="Insecta" family="Leiodidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
) (
<bibRefCitation author="Beron, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" refId="B10" refString="Beron, P, 2015. Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." title="Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." year="2015">Beron 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
So far known only from two caves in Lovech District (Fig. 
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on material from Saeva dupka Cave, 
<bibRefCitation author="Stoev, P" journalOrPublisher="Acta Zoologica Bulgarica" pageId="0" pageNumber="1" pagination="145 - 154" refId="B50" refString="Stoev, P, 2004. New distributional records of millipedes from Bulgarian caves (Myriapoda: Diplopoda). Acta Zoologica Bulgarica 56 (2): 145 - 154" title="New distributional records of millipedes from Bulgarian caves (Myriapoda: Diplopoda)." volume="56" year="2004">Stoev (2004)</bibRefCitation>
already noticed that there were certain differences in the structure of gonopods of that sample and 
<taxonomicName authorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" authorityYear="2022" baseAuthorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" baseAuthorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">Bacillidesmus dentatus</emphasis>
</taxonomicName>
, and he did not exclude the possibility that it belonged to a new taxon. However, he still treated this as 
<taxonomicName authorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" authorityYear="2022" baseAuthorityName="Antić &amp; Vagalinski &amp; Stoev &amp; Akkari" baseAuthorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus bulgaricus subsp. dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="bulgaricus" subSpecies="dentatus">
<emphasis italics="true" pageId="0" pageNumber="1">Bacillidesmus bulgaricus dentatus</emphasis>
</taxonomicName>
. After reviewing the material that was available to him, as well as based on the newly studied material, we describe it above as a new species.
</paragraph>
</subSubSection>
</treatment>
</document>