257 lines
36 KiB
XML
257 lines
36 KiB
XML
<document id="A0D71B1162C2EBF09D0508E9D9716BC0" ID-CLB-Dataset="2419" ID-DOI="10.21248/contrib.entomol.66.1.13-111" ID-GBIF-Dataset="d2933fad-1cac-4a5a-a558-2633f8314541" ID-ISSN="0005-805X" ID-Zenodo-Dep="6421171" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630270776760" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="766F7C36FF9FFFC4FF687734D9A4FEC2" docLanguage="en" docName="ContribEntomol.2016.66.1.13-111.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (1)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Orphnebius biformis Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="60" masterDocId="8A56044EFFB1FFEBFF8B7527D878FF83" masterDocTitle="On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="111" masterPageNumber="13" pageNumber="59" updateTime="1699235685730" updateUser="ExternalLinkService">
|
||
<mods:mods id="0297E69FAB2040714897AD93939043A3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="25585C48BF8446AA745795C20457BD43">
|
||
<mods:title id="3216209BA83CCCA36DB22861675E5CD5">On some Lomechusini of the Palaearctic and Oriental regions (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="B4D3362762B06DDC08E606C03DEE8299" type="personal">
|
||
<mods:role id="12E54E5AA3C1CBFD6752080234F2AF59">
|
||
<mods:roleTerm id="FCD0A4E21ABBCD51DF55A02CD8A94BC0">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EE2112F2E0487EFB7E409BA0E4994510">Assing, Volker</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DC2198F714E34C0A4F617D699A5DD913">text</mods:typeOfResource>
|
||
<mods:relatedItem id="CA4E16A9D2AA75DB4D229BB66E549A36" type="host">
|
||
<mods:titleInfo id="19EC4A877E951D70A3C04B4B855E0C38">
|
||
<mods:title id="101E7B16356CB8A5B72A07DBF31138D4">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="3C605D89891281762FE68C8A64224351">
|
||
<mods:date id="1F4EE69E02E648AC2D303D12E4F150E8">2016</mods:date>
|
||
<mods:detail id="B00AB3D1E86018B79CA6153EC0651359" type="pubDate">
|
||
<mods:number id="24B6A4438F314080A22791BC0791E237">2016-12-20</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B0D009E42676B17EECC6AA7A2ED296B1" type="volume">
|
||
<mods:number id="C26C94C53ADC1756CFFC041351544CE0">66</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A58DBBA763B5B2CCFA68F2F31834C4EF" type="issue">
|
||
<mods:number id="5B1EEE1C7764CCE1ADD6C841F6B25D97">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="DF564DE25D9E8A4A6E5D25BC692288CF" unit="page">
|
||
<mods:start id="117D0640166902A838BCC307288E2A9D">13</mods:start>
|
||
<mods:end id="ED5E9991D4CEC234725F241FD13C896B">111</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="CCDCA6481CB147C0EF9E3DF84E8C5664">
|
||
<mods:url id="F67EB610AB40A4369F2BEB797ADDDB95">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="D3E5A2DF91E91C0BA3C1624AC66B8CF2">journal article</mods:classification>
|
||
<mods:identifier id="C96AFF1653D18E3EE77F8C3B645C1073" type="CLB-Dataset">2419</mods:identifier>
|
||
<mods:identifier id="CE81B708062E6FE07E904738716750E9" type="DOI">10.21248/contrib.entomol.66.1.13-111</mods:identifier>
|
||
<mods:identifier id="947CD3856C0A5E3F80FE12F7C3D05643" type="GBIF-Dataset">d2933fad-1cac-4a5a-a558-2633f8314541</mods:identifier>
|
||
<mods:identifier id="3FD7DE3CC652A52A5DD4A0FC989D6D53" type="ISSN">0005-805X</mods:identifier>
|
||
<mods:identifier id="762DEA13992999ED0BF6107F1E4FA062" type="Zenodo-Dep">6421171</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="766F7C36FF9FFFC4FF687734D9A4FEC2" ID-DOI="http://doi.org/10.5281/zenodo.5903428" ID-GBIF-Taxon="192602870" ID-Zenodo-Dep="5903428" LSID="urn:lsid:plazi:treatment:766F7C36FF9FFFC4FF687734D9A4FEC2" httpUri="http://treatment.plazi.org/id/766F7C36FF9FFFC4FF687734D9A4FEC2" lastPageId="47" lastPageNumber="60" pageId="46" pageNumber="59">
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF687734DA5AFDAD" box="[227,546,531,558]" pageId="46" pageNumber="59" type="nomenclature">
|
||
<paragraph id="FE79CD20FF9FFFC5FF687734DA5AFDAD" blockId="46.[227,546,531,591]" box="[227,546,531,558]" pageId="46" pageNumber="59">
|
||
<heading id="A5317A4CFF9FFFC5FF687734DA5AFDAD" box="[227,546,531,558]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="59" reason="3">
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FF687734D9CAFDAD" ID-CoL="8Y79B" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[227,434,531,558]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="biformis" status="sp. nov.">
|
||
<emphasis id="CCB21132FF9FFFC5FF687734D9CAFDAD" box="[227,434,531,558]" italics="true" pageId="46" pageNumber="59">Orphnebius biformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D781AC49FF9FFFC5FE337734DA5AFDAD" box="[440,546,531,558]" pageId="46" pageNumber="59" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF68771ED999FDCC" box="[227,481,569,591]" pageId="46" pageNumber="59" type="description">
|
||
<paragraph id="FE79CD20FF9FFFC5FF68771ED999FDCC" blockId="46.[227,546,531,591]" box="[227,481,569,591]" pageId="46" pageNumber="59">
|
||
(
|
||
<figureCitation id="66FDD1A5FF9FFFC5FF6C771ED930FDCC" box="[231,328,569,591]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="46" pageNumber="59">Figs 19–20</figureCitation>
|
||
,
|
||
<figureCitation id="66FDD1A5FF9FFFC5FEDB771ED9F0FDCC" box="[336,392,569,591]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="46" pageNumber="59">42–43</figureCitation>
|
||
,
|
||
<figureCitation id="66FDD1A5FF9FFFC5FE1B771ED9A4FDCC" box="[400,476,569,591]" captionStart-0="Figs 258–276" captionStart-1="Figs 277–295" captionStartId-0="22.[189,229,1970,1992]" captionStartId-1="23.[151,191,1971,1993]" captionTargetBox-0="[192,1432,240,1938]" captionTargetBox-1="[154,1395,232,1955]" captionTargetId-0="figure-135@22.[189,1436,232,1954]" captionTargetId-1="figure-127@23.[151,1398,232,1963]" captionTargetPageId-0="22" captionText-0="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." captionText-1="Figs 277–295. Orphnebius biformis (277–278), O. retunsus (279–282), O. fodens (283–286), O. effeminatus (287–292), and O. multimpressus (293–295): median lobe of aedeagus in lateral and in ventral view (277–280, 283–284, 287–288, 293–294); paramere (281, 285, 289, 295); tergite VIII (282, 286, 290); median portion of tergite VIII (291); male tergites IX-X (292). Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6421204" figureDoi-1="http://doi.org/10.5281/zenodo.6421206" httpUri-0="https://zenodo.org/record/6421204/files/figure.png" httpUri-1="https://zenodo.org/record/6421206/files/figure.png" pageId="46" pageNumber="59">274–278</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF36774EDB72FCA2" pageId="46" pageNumber="59" type="materials_examined">
|
||
<paragraph id="FE79CD20FF9FFFC5FF36774ED950FC82" blockId="46.[189,794,617,802]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FF36774ED92CFD02" bold="true" box="[189,340,617,641]" pageId="46" pageNumber="59">Type material</emphasis>
|
||
:
|
||
<materialsCitation id="4EAEC77DFF9FFFC5FEEB774ED950FC82" ID-GBIF-Occurrence="3455118372" collectingDate="2003-02-25" collectingDateMax="2003-03-02" collectingDateMin="2003-02-25" collectionCode="NHMB" collectorName="Dembicky" determinerName="V. Assing" elevation="250" latitude="-1.75" location="Seblat N. P." longLatPrecision="1308" longitude="101.416664" municipality="Kerinci" pageId="46" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Sumatra" typeStatus="holotype">
|
||
<typeStatus id="217D7382FF9FFFC5FEEB774ED9BBFD02" box="[352,451,617,641]" pageId="46" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="E8C006A9FF9FFFC5FE42774DD9A6FD02" box="[457,478,618,641]" pageId="46" pageNumber="59" type="male">♂</specimenCount>
|
||
: “
|
||
<collectingRegion id="3C0203C2FF9FFFC5FE7F774EDA28FD02" box="[500,592,617,641]" country="Indonesia" name="Sumatera Utara" pageId="46" pageNumber="59">Sumatra</collectingRegion>
|
||
:
|
||
<collectingRegion id="3C0203C2FF9FFFC5FDD1774EDAC4FD02" box="[602,700,617,641]" pageId="46" pageNumber="59">Jambi pr.</collectingRegion>
|
||
:
|
||
<collectingMunicipality id="1E1D575AFF9FFFC5FD4E774EDB6EFD02" box="[709,790,617,641]" pageId="46" pageNumber="59">Kerinci</collectingMunicipality>
|
||
,
|
||
<location id="FB199BFBFF9FFFC5FF3677AED94DFD22" LSID="urn:lsid:plazi:treatment:766F7C36FF9FFFC4FF687734D9A4FEC2:FB199BFBFF9FFFC5FF3677AED94DFD22" box="[189,309,649,673]" latitude="-1.75" longLatPrecision="1308" longitude="101.416664" municipality="Kerinci" name="Seblat N. P." pageId="46" pageNumber="59" stateProvince="Sumatra">Seblat N.P.</location>
|
||
:
|
||
<locationDeviation id="EFCBD402FF9FFFC5FEC977ADD981FD21" box="[322,505,649,674]" location="Kayuaro" pageId="46" pageNumber="59">
|
||
<quantity id="393E60C5FF9FFFC5FEC977ADD900FD22" box="[322,376,649,673]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="46" pageNumber="59" unit="km" value="7.0">7 km</quantity>
|
||
E Kayuaro
|
||
</locationDeviation>
|
||
:
|
||
<location id="FB199BFBFF9FFFC5FD8D77ADDA0EFD22" LSID="urn:lsid:plazi:treatment:766F7C36FF9FFFC4FF687734D9A4FEC2:FB199BFBFF9FFFC5FD8D77ADDA0EFD22" box="[518,630,649,673]" latitude="-1.75" longLatPrecision="1308" longitude="101.416664" municipality="Kerinci" name="Mt. Tujuh" pageId="46" pageNumber="59" stateProvince="Sumatra">Mt. Tujuh</location>
|
||
, 1750±
|
||
<quantity id="393E60C5FF9FFFC5FD5A77AEDB6EFD22" box="[721,790,649,673]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="59" unit="m" value="250.0">
|
||
<elevation id="75EB2A13FF9FFFC5FD5A77AEDB6EFD22" box="[721,790,649,673]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="46" pageNumber="59" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="9BF2ABE7FF9FFFC5FF36778ED978FD42" box="[189,256,681,705]" degrees="1" direction="south" minutes="45" orientation="latitude" pageId="46" pageNumber="59" precision="925" value="-1.75">1°45'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9BF2ABE7FF9FFFC5FE87778ED913FD42" box="[268,363,681,705]" degrees="101" direction="east" minutes="25" orientation="longitude" pageId="46" pageNumber="59" precision="925" value="101.416664">101°25'E</geoCoordinate>
|
||
,
|
||
<collectorName id="5333A8F6FF9FFFC5FEFC778ED99CFD42" box="[375,484,681,705]" pageId="46" pageNumber="59">Dembický</collectorName>
|
||
leg.,
|
||
<date id="8A78EBE0FF9FFFC5FD94778EDABAFD42" box="[543,706,681,705]" pageId="46" pageNumber="59" value="2003-02-25" valueMax="2003-03-02" valueMin="2003-02-25">
|
||
<collectingDate id="9A3C1208FF9FFFC5FD94778EDABAFD42" box="[543,706,681,705]" pageId="46" pageNumber="59" value="2003-02-25" valueMax="2003-03-02" valueMin="2003-02-25">25.ii.–2.iii.2003</collectingDate>
|
||
</date>
|
||
/ Holotypus ♂
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FE9777EED98CFD62" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[284,500,713,737]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="biformis" status="sp. nov.">
|
||
<emphasis id="CCB21132FF9FFFC5FE9777EED98CFD62" box="[284,500,713,737]" italics="true" pageId="46" pageNumber="59">Orphnebius biformis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D781AC49FF9FFFC5FE7777EDDA4CFD62" box="[508,564,714,737]" pageId="46" pageNumber="59" rank="species">sp. n.</taxonomicNameLabel>
|
||
, det.
|
||
<determinerName id="0BA521E4FF9FFFC5FDFA77EDDAABFD62" box="[625,723,713,737]" pageId="46" pageNumber="59">V. Assing</determinerName>
|
||
2015” (
|
||
<collectionCode id="98D755E5FF9FFFC5FF4C77CED965FC82" box="[199,285,745,769]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="46" pageNumber="59">NHMB</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FF36762DDB72FCA2" blockId="46.[189,794,617,802]" box="[189,778,777,802]" pageId="46" pageNumber="59">
|
||
<materialsCitation id="4EAEC77DFF9FFFC5FF36762DDB7DFCA2" ID-GBIF-Occurrence="3455118403" box="[189,773,777,802]" collectingDate="2003-02-25" collectingDateMax="2003-03-02" collectingDateMin="2003-02-25" collectionCode="NHMB" collectorName="Dembicky" determinerName="V. Assing" elevation="250" latitude="-1.75" location="Mt. Tujuh" longLatPrecision="1308" longitude="101.416664" municipality="Kerinci" pageId="46" pageNumber="59" specimenCount="2" specimenCount-female="2" stateProvince="Sumatera Utara" typeStatus="paratype">
|
||
<typeStatus id="217D7382FF9FFFC5FF36762DD95FFCA1" box="[189,295,778,802]" pageId="46" pageNumber="59" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="E8C006A9FF9FFFC5FEBB762ED911FCA2" box="[304,361,777,801]" pageId="46" pageNumber="59" type="female">2 ♀♀</specimenCount>
|
||
: same data as holotype (
|
||
<collectionCode id="98D755E5FF9FFFC5FDE7762EDAC7FCA2" box="[620,703,777,801]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="46" pageNumber="59">NHMB</collectionCode>
|
||
, cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF36766ED9ADFC22" pageId="46" pageNumber="59" type="etymology">
|
||
<paragraph id="FE79CD20FF9FFFC5FF36766ED9ADFC22" blockId="46.[189,794,841,930]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FF36766ED94CFCE2" bold="true" box="[189,308,841,865]" pageId="46" pageNumber="59">Etymology</emphasis>
|
||
: The specific epithet (Latin, adjective) alludes to the hypothesized pronounced sexual dimorphism of the eyes and the antennae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FF3676EEDD2FFBA2" pageId="46" pageNumber="59" type="description">
|
||
<paragraph id="FE79CD20FF9FFFC5FF3676EEDA78FB42" blockId="46.[188,796,969,2081]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FF3676EED947FC62" bold="true" box="[189,319,969,993]" pageId="46" pageNumber="59">Description</emphasis>
|
||
: Body length
|
||
<quantity id="393E60C5FF9FFFC5FE6E76EEDA1EFC62" box="[485,614,969,993]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="46" pageNumber="59" unit="mm" value="4.55" valueMax="4.8" valueMin="4.3">4.3–4.8 mm</quantity>
|
||
; length of forebody 1.9–2.0 mm. Coloration: head dark-brown with the clypeus yellowish-red; pronotum dark-brown; elytra dark-brown with the anterior and sutural portions more or less extensively paler; abdomen reddish-yellow; legs yellowish; antennae dark-brown with antennomeres I–IV reddish-yellow; maxillary palpi reddish-yellow with yellowish terminal palpomere.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FF3671EED919F8C2" blockId="46.[188,796,969,2081]" pageId="46" pageNumber="59">
|
||
Head (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FE8271EED9F8FB62" box="[265,384,1225,1249]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="46" pageNumber="59">Figs 42–43</figureCitation>
|
||
) sexually dimorphic, strongly transverse, approximately 1.3 (female) and 1.4 (male) times as broad as long, respectively; laterally with few scattered and very fine punctures; remainder of dorsal portion extensively impunctate; interstices without microsculpture. Eye size sexually dimorphic, in male enormous and strongly bulging, covering all of lateral margins, in female less large and less convex. Antenna (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FF4F70EED943FA62" box="[196,315,1481,1505]" captionStart="Figs 1–26" captionStartId="9.[151,191,1971,1993]" captionTargetBox="[151,1398,238,1937]" captionTargetId="figure-172@9.[151,1398,232,1948]" captionTargetPageId="9" captionText="Figs 1–26. Antenna (1–21) and forebody (22–26) of Orphnebius spp. of the O. hauseri group: spinans (1, 23), cernens (2, 24), lunatus (3, 25), bakeri (4, 26), breviceps (5), extensus (6), serratus (7), integer (8), bicuspis, Laos (9), bicuspis, NE India (10), fusicollis (11), retunsus (12), fodens (13), opticus (14), dilatatus (15), fuscapicalis (16), grandicollis (17, 22), reductus (18), biformis, male (19), biformis, female (20), effeminatus (21). Scale bars: 22–26: 1.0 mm; 1–21: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6421173" httpUri="https://zenodo.org/record/6421173/files/figure.png" pageId="46" pageNumber="59">Figs 19–20</figureCitation>
|
||
) sexually dimorphic, approximately 1.6 and
|
||
<quantity id="393E60C5FF9FFFC5FF3670CED977F982" box="[189,271,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="46" pageNumber="59" unit="mm" value="1.3">1.3 mm</quantity>
|
||
long in male and in female respectively, distinctly incrassate, and distinctly asymmetric; antennomere IV small and moderately transverse; antennomere V moderately transverse and weakly transverse in female, strongly transverse and strongly asymmetric in male; antennomeres VI–X of gradually and distinctly increasing width, increasingly asymmetric, and increasingly transverse, more transverse in male than in female; X more than twice as wide as long; XI elongate, approximately three times as long as broad (male) or twice as long as broad (female).
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FF36726DD9ACF7A2" blockId="46.[188,796,969,2081]" pageId="46" pageNumber="59">
|
||
Pronotum (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FEC8726ED9C2F8E2" box="[323,442,1865,1889]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="46" pageNumber="59">Figs 42–43</figureCitation>
|
||
) moderately transverse, 1.21– 1.26 times as broad as long and approximately 0.95 (male) or 1.1 times (female) as broad as head, moderately strongly convex in cross-section; posterior angles nearly obsolete; disc with a median pair of punctures, otherwise practically impunctate; lateral margins each with three long and erect black setae.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB475CEDBD0FE22" blockId="46.[831,1437,233,1058]" pageId="46" pageNumber="59">
|
||
Elytra (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FC1975CEDC71FE82" box="[914,1033,233,257]" captionStart="Figs 40–53" captionStartId="11.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1971]" captionTargetId="figure-131@11.[151,1398,232,1971]" captionTargetPageId="11" captionText="Figs 40–53. Forebody (40–50), abdomen (51), and antenna (52–53) of Orphnebius spp. of the O. hauseri group (40–49) and of the subgenus Deroleptus (50–53): opticus (40), effeminatus (41), biformis, male (42), biformis, female (43), bicuspis, NE India (44), retunsus (45), fusicollis (46–47), fodens (48), reductus (49), niger (50–52), siamensis (53). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6421179" httpUri="https://zenodo.org/record/6421179/files/figure.png" pageId="46" pageNumber="59">Figs 42–43</figureCitation>
|
||
) approximately 0.7 times as long as pronotum; suture strongly gaping posteriorly; punctation sparse and very fine; pubescence long and pale. Hind wings fully developed. Metatarsomere I approximately as long as, or slightly longer than, the combined length of II and III.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB4748EDD59FCA2" blockId="46.[831,1437,233,1058]" pageId="46" pageNumber="59">
|
||
Abdomen: paratergites moderately elevated and moderately sharply edged; tergites III–VI each with two lateral setiferous punctures on either side, tergite VI additionally with median pair of punctures at posterior margin; tergites IV–V with or without median pair of setiferous punctures at posterior margin; tergite VII with non-setiferous punctation in posterior two-thirds and with eight setiferous punctures near posterior margin, posterior margin with palisade fringe; tergite VIII (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FA8F778EDD24FD42" box="[1284,1372,681,705]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="46" pageNumber="59">Fig. 275</figureCitation>
|
||
) with two transverse rows of black setae posteriorly and with broadly truncate posterior margin; sternite VIII with broadly and weakly convex posterior margin.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB4760DDC5CFC62" blockId="46.[831,1437,233,1058]" pageId="46" pageNumber="59">
|
||
<specimenCount id="E8C006A9FF9FFFC5FCB4760DDB2CFCC2" box="[831,852,810,833]" pageId="46" pageNumber="59" type="male">♂</specimenCount>
|
||
: hemi-tergites IX and tergite X with extremely dense and long pubescence; median lobe of aedeagus (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FCCC764EDBAFFC02" box="[839,983,873,897]" captionStart="Figs 277–295" captionStartId="23.[151,191,1971,1993]" captionTargetBox="[154,1395,232,1955]" captionTargetId="figure-127@23.[151,1398,232,1963]" captionText="Figs 277–295. Orphnebius biformis (277–278), O. retunsus (279–282), O. fodens (283–286), O. effeminatus (287–292), and O. multimpressus (293–295): median lobe of aedeagus in lateral and in ventral view (277–280, 283–284, 287–288, 293–294); paramere (281, 285, 289, 295); tergite VIII (282, 286, 290); median portion of tergite VIII (291); male tergites IX-X (292). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421206" httpUri="https://zenodo.org/record/6421206/files/figure.png" pageId="46" pageNumber="59">Figs 277–278</figureCitation>
|
||
)
|
||
<quantity id="393E60C5FF9FFFC5FC62764EDC30FC02" box="[1001,1096,873,897]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="46" pageNumber="59" unit="mm" value="0.51">0.51 mm</quantity>
|
||
long and of distinctive shape; internal sac without strongly sclerotized structures; paramere (
|
||
<figureCitation id="66FDD1A5FF9FFFC5FC01768EDB99FC42" box="[906,993,937,961]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="46" pageNumber="59">Fig. 274</figureCitation>
|
||
) approximately
|
||
<quantity id="393E60C5FF9FFFC5FB11768EDC94FC42" box="[1178,1260,937,961]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="59" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long, condylite very short and broad.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB476CDDD2FFBA2" blockId="46.[831,1437,233,1058]" pageId="46" pageNumber="59">
|
||
<specimenCount id="E8C006A9FF9FFFC5FCB476CDDB2CFB82" box="[831,852,1002,1025]" pageId="46" pageNumber="59" type="female">♀</specimenCount>
|
||
: abdominal segments IX–X of similar shape and chaetotaxy as in
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FC34712EDC52FBA2" authorityName="Eppelsheim" authorityYear="1895" box="[959,1066,1033,1057]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="hauseri">
|
||
<emphasis id="CCB21132FF9FFFC5FC34712EDC52FBA2" box="[959,1066,1033,1057]" italics="true" pageId="46" pageNumber="59">O. hauseri</emphasis>
|
||
</taxonomicName>
|
||
; spermatheca as in
|
||
<figureCitation id="66FDD1A5FF9FFFC5FB70712EDD2AFBA2" box="[1275,1362,1033,1057]" captionStart="Figs 258–276" captionStartId="22.[189,229,1970,1992]" captionTargetBox="[192,1432,240,1938]" captionTargetId="figure-135@22.[189,1436,232,1954]" captionText="Figs 258–276. Orphnebius bicuspis (258–260; 258–259: India), O. opticus, syntype (261–262), O. fusicollis (263–268), O. reductus (269–273), and O. biformis (274–276): median lobe of aedeagus in lateral and in ventral view (258–259, 263–264, 269–270); tergite VIII (260, 261, 266, 272, 275); spermatheca (262, 268, 273, 276); paramere (265, 271, 274); anteromedian portion of sternite VIII (267). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6421204" httpUri="https://zenodo.org/record/6421204/files/figure.png" pageId="46" pageNumber="59">Fig. 276</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9FFFC5FCB4716EDC5EF862" pageId="46" pageNumber="59" type="discussion">
|
||
<paragraph id="FE79CD20FF9FFFC5FCB4716EDBC6FA22" blockId="46.[831,1438,1097,1441]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FCB4716EDBD7FBE2" bold="true" box="[831,943,1097,1121]" pageId="46" pageNumber="59">Comment</emphasis>
|
||
: The
|
||
<typeStatus id="217D7382FF9FFFC5FC70716EDC22FBE2" box="[1019,1114,1097,1121]" pageId="46" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
male is distinguished from the
|
||
<specimenCount id="E8C006A9FF9FFFC5FCE0714DDB9CFB02" box="[875,996,1129,1153]" pageId="46" pageNumber="59" type="female">two female</specimenCount>
|
||
<typeStatus id="217D7382FF9FFFC5FC65714DDC2EFB01" box="[1006,1110,1130,1154]" pageId="46" pageNumber="59" type="paratype">paratypes</typeStatus>
|
||
by the different shape of the antennae, the more transverse head (broader than the pronotum), and the distinctly larger and more convex eyes. These differences, however, are interpreted as sexual dimorphism, not only because the
|
||
<specimenCount id="E8C006A9FF9FFFC5FA9171CEDB01FAA2" pageId="46" pageNumber="59" type="generic">three specimens</specimenCount>
|
||
were collected in the same locality on the same date, but also because they are extremely similar in other respects and because a pronounced sexual dimorphism (of the antennae) was also observed in the closely related
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FCB470AEDBC0FA22" authorityName="Assing" authorityYear="2016" box="[831,952,1417,1441]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="reductus">
|
||
<emphasis id="CCB21132FF9FFFC5FCB470AEDBC0FA22" box="[831,952,1417,1441]" italics="true" pageId="46" pageNumber="59">O. reductus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB470EEDC27F8E2" blockId="46.[830,1438,1481,2017]" pageId="46" pageNumber="59">
|
||
<emphasis id="CCB21132FF9FFFC5FCB470EEDC62FA62" bold="true" box="[831,1050,1481,1505]" pageId="46" pageNumber="59">Comparative notes</emphasis>
|
||
: Based on the modifications of the abdominal segments IX–X, the shape and chaetotaxy of tergite and sternite VIII, the pronounced sexual dimorphism of the antennae, the absence of strongly sclerotized structures in the internal sac of the aedeagus, and particularly on the very similar morphology of the spermatheca, this species is closely allied to
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FB6F73AEDD24F922" authorityName="Assing" authorityYear="2016" box="[1252,1372,1673,1697]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="reductus">
|
||
<emphasis id="CCB21132FF9FFFC5FB6F73AEDD24F922" box="[1252,1372,1673,1697]" italics="true" pageId="46" pageNumber="59">O. reductus</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs by different coloration, a pronounced sexual dimorphism of the head (including the eyes), by the shape of the male and female antennae, the coloration, and the primary sexual characters, particularly the completely different shape of the median lobe and the parameres of the aedeagus.
|
||
</paragraph>
|
||
<paragraph id="FE79CD20FF9FFFC5FCB4724DDC5EF862" blockId="46.[830,1438,1481,2017]" pageId="46" pageNumber="59">
|
||
None of the three
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FC71724EDCBEF802" box="[1018,1222,1897,1921]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="CCB21132FF9FFFC5FC71724EDC0CF802" box="[1018,1140,1897,1921]" italics="true" pageId="46" pageNumber="59">Orphnebius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
previously recorded from
|
||
<collectingRegion id="3C0203C2FF9FFFC5FCF572AEDBAFF822" box="[894,983,1929,1953]" country="Indonesia" name="Sumatera Utara" pageId="46" pageNumber="59">Sumatra</collectingRegion>
|
||
(
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FC6672AEDD6FF822" authority="CAMERON, 1925" authorityName="CAMERON" authorityYear="1925" box="[1005,1303,1929,1953]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="laticeps">
|
||
<emphasis id="CCB21132FF9FFFC5FC6672AEDC2FF822" box="[1005,1111,1929,1953]" italics="true" pageId="46" pageNumber="59">O. laticeps</emphasis>
|
||
<bibRefCitation id="9A57B0D1FF9FFFC5FBE872AEDD6FF822" author="CAMERON, M." box="[1123,1303,1929,1953]" pageId="46" pageNumber="59" pagination="34 - 49" refId="ref77074" refString="CAMERON, M. 1925: Descriptions of new species of oriental Staphylinidae. - Annali del Museo Civico di Storia Naturale di Genova 52: 34 - 49." type="journal article" year="1925">CAMERON, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FAAD72AEDC44F842" authority="CAMERON, 1936" authorityName="CAMERON" authorityYear="1936" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="ophthalmicus">
|
||
<emphasis id="CCB21132FF9FFFC5FAAD72AEDB04F842" italics="true" pageId="46" pageNumber="59">O. ophthalmicus</emphasis>
|
||
CAMERON, 1936
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FBC7728EDD00F842" authority="PACE, 2010" authorityName="PACE" authorityYear="2010" box="[1100,1400,1961,1985]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="harpagonum">
|
||
<emphasis id="CCB21132FF9FFFC5FBC7728EDC88F842" box="[1100,1264,1961,1985]" italics="true" pageId="46" pageNumber="59">O. harpagonum</emphasis>
|
||
<bibRefCitation id="9A57B0D1FF9FFFC5FB77728EDD00F842" author="PACE, R." box="[1276,1400,1961,1985]" pageId="46" pageNumber="59" pagination="19 - 54" refId="ref77979" refString="PACE, R. 2010: Thamiaraeini, Lomechusini, Oxypodini, Hoplandriini e Aleocharini di Taiwan (Coleoptera, Staphylinidae). - Bollettino del Museo Civico di Storia Naturale di Verona 34: 19 - 54." type="journal article" year="2010">PACE, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
) is similar to
|
||
<taxonomicName id="39C6B6A3FF9FFFC5FC2072EEDC58F862" authorityName="Assing" authorityYear="2016" box="[939,1056,1993,2017]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="59" phylum="Arthropoda" rank="species" species="biformis">
|
||
<emphasis id="CCB21132FF9FFFC5FC2072EEDC58F862" box="[939,1056,1993,2017]" italics="true" pageId="46" pageNumber="59">O. biformis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6DC9EABFF9EFFC4FF1C75CED9A4FEC2" pageId="47" pageNumber="60" type="distribution">
|
||
<paragraph id="FE79CD20FF9EFFC4FF1C75CED9A4FEC2" blockId="47.[151,756,233,321]" pageId="47" pageNumber="60">
|
||
<emphasis id="CCB21132FF9EFFC4FF1C75CEDA71FE82" bold="true" box="[151,521,233,257]" pageId="47" pageNumber="60">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="217D7382FF9EFFC4FDDB75CDDA06FE81" box="[592,638,234,258]" pageId="47" pageNumber="60">type</typeStatus>
|
||
locality is situated in
|
||
<collectingRegion id="3C0203C2FF9EFFC4FE80742ED9D7FEA2" box="[267,431,265,289]" country="Indonesia" name="Sumatera Barat" pageId="47" pageNumber="60">Sumatera Barat</collectingRegion>
|
||
,
|
||
<collectingRegion id="3C0203C2FF9EFFC4FE33742EDA6BFEA2" box="[440,531,265,289]" country="Indonesia" name="Sumatera Utara" pageId="47" pageNumber="60">Sumatra</collectingRegion>
|
||
,
|
||
<collectingCountry id="86D18DB0FF9EFFC4FD96742EDAF0FEA2" box="[541,648,265,289]" name="Indonesia" pageId="47" pageNumber="60">Indonesia</collectingCountry>
|
||
, at an altitude of approximately
|
||
<quantity id="393E60C5FF9EFFC4FE0D740ED9AFFEC2" box="[390,471,297,321]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="47" pageNumber="60" unit="m" value="1750.0">1750 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |