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<document id="A2B9AAB57A397433BA3810594CBDA6BC" ID-CLB-Dataset="55444" ID-DOI="10.11646/zootaxa.5138.4.4" ID-GBIF-Dataset="0a2c45b6-6635-448e-91dc-4b68c25c12e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="6571654" ID-ZooBank="505B5461-19AC-4C2C-9F9D-FC9E46EB896B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1653288933666" checkinUser="plazi" docAuthor="Das, Abhijit, Gower, David J., Narayanan, Surya, Pal, Saunak, Boruah, Bitupan, Magar, Sanjay, Das, Sanjib, Moulick, Soumabrata &amp; Deepak, V." docDate="2022" docId="18078786EC1DFFA5209CFEBEBBE1F9F2" docLanguage="en" docName="zootaxa.5138.4.4.pdf" docOrigin="Zootaxa 5138 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oligodon melaneus Wall 1909" docType="treatment" docVersion="8" lastPageNumber="421" masterDocId="E43EFFFEEC19FFA1200BFFD0BF36FFFA" masterDocTitle="Rediscovery and systematics of the rarely encountered Blue-bellied kukri snake (Oligodon melaneus Wall, 1909) from Assam, India" masterLastPageNumber="430" masterPageNumber="417" pageNumber="421" updateTime="1699337905646" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="78942561BC14D8CCD204306DFFD6534D">Rediscovery and systematics of the rarely encountered Blue-bellied kukri snake (Oligodon melaneus Wall, 1909) from Assam, India</mods:title>
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<mods:namePart id="C970ACF5B5328E38FED3CCF2E978DDD5">Gower, David J.</mods:namePart>
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<mods:namePart id="2F1072FE7B143D7D5236AD05D7A06462">Narayanan, Surya</mods:namePart>
<mods:affiliation id="8FB78DF6A6080DAF3118E2446334A004">SMS Centre for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka - 560064, India.</mods:affiliation>
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<mods:namePart id="A07B02997EC2DF2AB5E88BECDCFC6403">Pal, Saunak</mods:namePart>
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<mods:namePart id="1FC5CCC56BB560837746124CB8EB9B0F">Boruah, Bitupan</mods:namePart>
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<mods:affiliation id="A6FBA4BCD1AE84933B4217EDF9776959">Gyati gaon, Barengabari, Baksa, Assam- 781315.</mods:affiliation>
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<mods:affiliation id="F3092B3C1FB0D0C827E4C32FEBAEF730">Department of Life Sciences, Natural History Museum, London SW 7 5 BD, UK. &amp; Senckenberg Dresden, Königsbrücker Landstrasse 159, 01109 Dresden, Germany.</mods:affiliation>
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<treatment id="18078786EC1DFFA5209CFEBEBBE1F9F2" ID-DOI="http://doi.org/10.5281/zenodo.6571667" ID-GBIF-Taxon="195580672" ID-Zenodo-Dep="6571667" LSID="urn:lsid:plazi:treatment:18078786EC1DFFA5209CFEBEBBE1F9F2" httpUri="http://treatment.plazi.org/id/18078786EC1DFFA5209CFEBEBBE1F9F2" lastPageNumber="421" pageId="4" pageNumber="421">
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<taxonomicName id="57AE4D13EC1DFFA5209CFEBEBD38FE72" authority="Wall, 1909" authorityName="Wall" authorityYear="1909" box="[151,526,366,393]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="421" phylum="Chordata" rank="species" species="melaneus">
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<emphasis id="A2DAEA82EC1DFFA5209CFEBEBEB5FE73" bold="true" box="[151,387,366,393]" italics="true" pageId="4" pageNumber="421">Oligodon melaneus</emphasis>
Wall, 1909
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<figureCitation id="08952A15EC1DFFA5209CFE42BE34FE56" box="[151,258,402,428]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="5.[151,250,1942,1966]" captionStartId-1="6.[151,250,1679,1703]" captionStartId-2="7.[151,250,1943,1967]" captionTargetBox-0="[221,1367,188,1916]" captionTargetBox-1="[151,1436,181,1655]" captionTargetBox-2="[179,1408,181,1920]" captionTargetId-0="figure-17@5.[220,1367,181,1919]" captionTargetId-1="figure-17@6.[151,1436,181,1655]" captionTargetId-2="figure-17@7.[179,1408,181,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 1. Phylogenetic relationships of Oligodon melaneus.A) ML phylogeny with bootstrap support values, B) BI phylogeny with Bayesian posterior probability support values." captionText-1="FIGURE 2. Designated lectotype of Oligodon melaneus (BMNH 1910.12.31.3). Photographs by Kevin Webb. Scale bar increments in mm." captionText-2="FIGURE 3. Paralectotype of Oligodon melaneus BNHS 958. Photographs by Saunak Pal. Image not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6571656" figureDoi-1="http://doi.org/10.5281/zenodo.6571658" figureDoi-2="http://doi.org/10.5281/zenodo.6571660" httpUri-0="https://zenodo.org/record/6571656/files/figure.png" httpUri-1="https://zenodo.org/record/6571658/files/figure.png" httpUri-2="https://zenodo.org/record/6571660/files/figure.png" pageId="4" pageNumber="421">Figs. 13</figureCitation>
,
<tableCitation id="DD2C032BEC1DFFA52105FE42BEB8FE56" box="[270,398,402,428]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="2.[151,239,485,511]" captionStartId-1="8.[151,244,150,176]" captionTargetPageId-0="2" captionTargetPageId-1="8" captionText-0="TABLE 1. GenBank accession and voucher numbers for the sequences used in this study." captionText-1="TABLE 2. Morphometric (in mm) and meristic data for the two previously known (type) specimens and the newly reported specimen of Oligodon melaneus. Counts are given in right, left order. ““damaged scales. * on the right side, the 2nd and 3rd and (especially) the 6th and 7th supralabials are incompletely distinct (are partially fused). Values in square brackets are those reported by Wall (1909)." httpUri-0="http://table.plazi.org/id/C4D16618EC1BFFA3209CFE35BB63FE05" httpUri-1="http://table.plazi.org/id/C4D16618EC11FFA9209CFF46BC0FFEE7" pageId="4" pageNumber="421" tableUuid-0="C4D16618EC1BFFA3209CFE35BB63FE05" tableUuid-1="C4D16618EC11FFA9209CFF46BC0FFEE7">Tables 12</tableCitation>
.
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Wall (1909) provided relatively detailed descriptions of the
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. We note differences between Walls and our observations in the text and
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, and provide the first photographs of the
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and
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), but otherwise focus on a more detailed description of the new specimen because it was collected more than
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from the type locality.
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.
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.31.3 (
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), a female specimen from
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elevation),
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,
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, collected
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.
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specimen is here designated as the lectotype because it is the best preserved of the two syntypes.
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</paragraph>
</subSubSection>
<subSubSection id="D8B4651BEC1DFFA520CCFD06BD84FB87" pageId="4" pageNumber="421" type="description">
<paragraph id="90113690EC1DFFA520CCFD06BD44FCC3" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
Specimen in very good condition; ventral incision into coelom ca.
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to ca.
<quantity id="57569B75EC1DFFA5245BFD06BBA9FD0B" box="[1104,1183,726,753]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="421" unit="mm" value="40.0">40 mm</quantity>
from vent, midventral incision ca.
<quantity id="57569B75EC1DFFA5212BFD2ABE46FCEF" box="[288,368,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="421" unit="mm" value="14.0">14 mm</quantity>
into anterior end of tail;
<quantity id="57569B75EC1DFFA52286FD2ABDDDFCEF" box="[653,747,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="3.29" pageId="4" pageNumber="421" unit="mm" value="329.0">329 mm</quantity>
total length (snoutvent length
<quantity id="57569B75EC1DFFA5245EFD2ABB83FCEF" box="[1109,1205,762,789]" metricMagnitude="-1" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="421" unit="mm" value="280.0">280 mm</quantity>
, tail length
<quantity id="57569B75EC1DFFA52530FD2ABAA6FCEF" box="[1339,1424,762,789]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="421" unit="mm" value="49.0">49 mm</quantity>
); head length
<quantity id="57569B75EC1DFFA5212BFCCEBEB1FCC3" box="[288,391,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="4" pageNumber="421" unit="mm" value="10.4">10.4 mm</quantity>
; head width
<quantity id="57569B75EC1DFFA5221EFCCEBD58FCC3" box="[533,622,798,825]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="421" unit="mm" value="5.6">5.6 mm</quantity>
.
</paragraph>
<paragraph id="90113690EC1DFFA520CCFC92BC0CFC32" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
<taxonomicName id="57AE4D13EC1DFFA520CCFC92BE04FCA6" box="[199,306,834,860]" class="Insecta" family="Syrphidae" genus="Lepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="A2DAEA82EC1DFFA520CCFC92BE04FCA6" box="[199,306,834,860]" italics="true" pageId="4" pageNumber="421">Lepidosis</emphasis>
</taxonomicName>
. 7,7 supralabials but on the right side the 2
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and 3
<superScript id="67DB9BD8EC1DFFA52354FC92BC5BFCAA" attach="left" box="[863,877,834,848]" fontSize="6" pageId="4" pageNumber="421">rd</superScript>
and the 6
<superScript id="67DB9BD8EC1DFFA523DCFC92BCD2FCAA" attach="left" box="[983,996,834,848]" fontSize="6" pageId="4" pageNumber="421">th</superScript>
and the 7
<superScript id="67DB9BD8EC1DFFA52445FC92BB6DFCAA" attach="left" box="[1102,1115,834,848]" fontSize="6" pageId="4" pageNumber="421">th</superScript>
supralabial are incompletely distinct (partially fused). The 3rd and 4th supralabials in contact with the spectacle covering the eye. Partly subdivided nasal contacts 1
<superScript id="67DB9BD8EC1DFFA521C3FC5ABEE5FC62" attach="left" box="[456,467,906,920]" fontSize="6" pageId="4" pageNumber="421">st</superScript>
and 2
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supralabial. 1,1 preocular; 7,7 infralabials; 1+2,1+2 temporals; 2,2 postoculars; dorsal scales in 17-15-15 rows; 159 ventrals; 39 subcaudals.
</paragraph>
<paragraph id="90113690EC1DFFA520CCFC02BD84FB87" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
<emphasis id="A2DAEA82EC1DFFA520CCFC02BECEFC16" box="[199,504,978,1004]" italics="true" pageId="4" pageNumber="421">Colouration in preservative</emphasis>
. Body colour brown; top of the head without typical
<taxonomicName id="57AE4D13EC1DFFA5243FFC02BBA8FC16" authorityName="Fitzinger" authorityYear="1826" box="[1076,1182,978,1004]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="421" phylum="Chordata" rank="genus">
<emphasis id="A2DAEA82EC1DFFA5243FFC02BBA8FC16" box="[1076,1182,978,1004]" italics="true" pageId="4" pageNumber="421">Oligodon</emphasis>
</taxonomicName>
markings, same colour as dorsum; rostral cream; supralabials heavily speckled with dark brown, paler edged; postoculars and temporals (see
<figureCitation id="08952A15EC1DFFA520C0FBCABE38FBCE" box="[203,270,1050,1076]" captionStart="FIGURE 2" captionStartId="6.[151,250,1679,1703]" captionTargetBox="[151,1436,181,1655]" captionTargetId="figure-17@6.[151,1436,181,1655]" captionTargetPageId="6" captionText="FIGURE 2. Designated lectotype of Oligodon melaneus (BMNH 1910.12.31.3). Photographs by Kevin Webb. Scale bar increments in mm." figureDoi="http://doi.org/10.5281/zenodo.6571658" httpUri="https://zenodo.org/record/6571658/files/figure.png" pageId="4" pageNumber="421">Fig. 2</figureCitation>
) unpatterned; infralabials and genial region cream with four dark brown spots medially; anterior ventral scales are edged with dark brown; posterior to the thirteenth ventral, colouration uniform dark brown throughout to vent. Subcaudals dark brown with paler centres.
</paragraph>
</subSubSection>
<subSubSection id="D8B4651BEC1DFFA520CCFB56BEF5FB3E" pageId="4" pageNumber="421" type="materials_examined">
<paragraph id="90113690EC1DFFA520CCFB56BEF5FB3E" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
<materialsCitation id="20C63CCDEC1DFFA520CCFB56BE88FB3E" ID-GBIF-Occurrence="3778517302" collectionCode="BNHS" pageId="4" pageNumber="421" specimenCode="BNHS 958" specimenCount="2" typeStatus="paralectotype">
<emphasis id="A2DAEA82EC1DFFA520CCFB56BE45FB5A" bold="true" box="[199,371,1158,1184]" pageId="4" pageNumber="421">
<typeStatus id="4F158832EC1DFFA520CCFB56BE58FB5A" box="[199,366,1158,1184]" pageId="4" pageNumber="421" type="paralectotype">Paralectotype</typeStatus>
.
</emphasis>
<specimenCode id="C0089EEBEC1DFFA52175FB56BEC8FB5B" box="[382,510,1158,1185]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="4" pageNumber="421">BNHS 958</specimenCode>
(fide
<bibRefCitation id="F43F4B61EC1DFFA52242FB56BDF2FB5A" author="Wall, F." box="[585,708,1158,1184]" pageId="4" pageNumber="421" pagination="598 - 632" refId="ref7922" refString="Wall, F. (1924) A handlist of the snakes of the Indian empire. Journal of Bombay Natural History Society, 29 (3 - 4), 598 - 632." type="journal article" year="1924">Wall 1924</bibRefCitation>
;
<figureCitation id="08952A15EC1DFFA522DFFB56BC2DFB5A" box="[724,795,1158,1184]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@7.[179,1408,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Paralectotype of Oligodon melaneus BNHS 958. Photographs by Saunak Pal. Image not to scale." figureDoi="http://doi.org/10.5281/zenodo.6571660" httpUri="https://zenodo.org/record/6571660/files/figure.png" pageId="4" pageNumber="421">Fig. 3</figureCitation>
), a male specimen from the same locality and date of collection as the
<typeStatus id="4F158832EC1DFFA5215FFB7ABE88FB3E" box="[340,446,1194,1220]" pageId="4" pageNumber="421" type="lectotype">lectotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D8B4651BEC1DFFA520CCFB1EBBE1F9F2" pageId="4" pageNumber="421" type="description">
<paragraph id="90113690EC1DFFA520CCFB1EBD00FACA" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
Specimen dehydrated to some extent and possibly moulting when it was fixed; outer layer of most head shields and some body scales missing;
<quantity id="57569B75EC1DFFA521F8FB22BD79FAF7" box="[499,591,1266,1293]" metricMagnitude="-1" metricUnit="m" metricValue="3.12" pageId="4" pageNumber="421" unit="mm" value="312.0">312 mm</quantity>
total length (snoutvent length
<quantity id="57569B75EC1DFFA523A4FB22BB3BFAF7" box="[943,1037,1266,1293]" metricMagnitude="-1" metricUnit="m" metricValue="2.61" pageId="4" pageNumber="421" unit="mm" value="261.0">261 mm</quantity>
, tail length
<quantity id="57569B75EC1DFFA52486FB22BBD6FAF7" box="[1165,1248,1266,1293]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="4" pageNumber="421" unit="mm" value="51.0">51 mm</quantity>
); head length
<quantity id="57569B75EC1DFFA52572FB22BFF0FACB" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="4" pageNumber="421" unit="mm" value="9.1">9.1 mm</quantity>
; head width
<quantity id="57569B75EC1DFFA5215FFAC6BE9BFACB" box="[340,429,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="421" unit="mm" value="5.3">5.3 mm</quantity>
; adult male.
</paragraph>
<paragraph id="90113690EC1DFFA520CCFAEABD2CFA66" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
<taxonomicName id="57AE4D13EC1DFFA520CCFAEABE04FAAE" box="[199,306,1338,1364]" class="Insecta" family="Syrphidae" genus="Lepidosis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="421" phylum="Arthropoda" rank="genus">
<emphasis id="A2DAEA82EC1DFFA520CCFAEABE04FAAE" box="[199,306,1338,1364]" italics="true" pageId="4" pageNumber="421">Lepidosis</emphasis>
</taxonomicName>
. 7,7 supralabials with the 3
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and 4
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in contact with the eye. Partly subdivided nasal contacts 1
<superScript id="67DB9BD8EC1DFFA5259AFAEABAAAFAB2" attach="none" box="[1425,1436,1338,1352]" fontSize="6" pageId="4" pageNumber="421">st</superScript>
supralabial only. 1,1 small preocular; 7,7 infralabials; 1+2, 1+2 temporals; 2,2 postocular; dorsal scales in 17-15-15 rows; 152 ventrals; 43 subcaudals.
</paragraph>
<paragraph id="90113690EC1DFFA520CCFA76BBE1F9F2" blockId="4.[151,1437,474,1545]" pageId="4" pageNumber="421">
<emphasis id="A2DAEA82EC1DFFA520CCFA76BEC9FA3A" box="[199,511,1446,1472]" italics="true" pageId="4" pageNumber="421">Colouration in preservative</emphasis>
. Body colour dark grey merging to greyish on the flanks; individual scales finely speckled with black on a grey background. Venter grey and iridescent; ventral scales finely speckled with black, less densely on the anterior part of the body than on the posterior of the body and subcaudals (
<figureCitation id="08952A15EC1DFFA5248EFA3EBBFCF9F2" box="[1157,1226,1518,1544]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-17@7.[179,1408,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Paralectotype of Oligodon melaneus BNHS 958. Photographs by Saunak Pal. Image not to scale." figureDoi="http://doi.org/10.5281/zenodo.6571660" httpUri="https://zenodo.org/record/6571660/files/figure.png" pageId="4" pageNumber="421">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>