treatments-xml/data/2A/1C/03/2A1C034EFFF4562360C8FF158662FA6C.xml

<document id="56E4E54832F4BAB323AEBAACFBF7C061" ID-DOI="10.11606/0031-1049.2017.57.02" ID-ISSN="1807-0205" ID-Zenodo-Dep="4656696" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617279465628" checkinUser="carolina" docAuthor="Díaz, Florentina, Felippe-Bauer, Maria Luiza &amp; Spinelli, Gustavo Ricardo" docDate="2017" docId="2A1C034EFFF4562360C8FF158662FA6C" docLanguage="en" docName="PapAvZool.57.2.17-22.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 57 (2)" docSource="http://dx.doi.org/10.11606/0031-1049.2017.57.02" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli 2017" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="D6257B36FFF65621605BFFE28524FF9C" masterDocTitle="Two new Neotropical species of the Dasyhelea mutabilis group (Culicomorpha: Ceratopogonidae)" masterLastPageNumber="22" masterPageNumber="17" pageNumber="19" updateTime="1698899678169" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5D66CF96C8AAA16C090FD331E8AB7CE9">Two new Neotropical species of the Dasyhelea mutabilis group (Culicomorpha: Ceratopogonidae)</mods:title>
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<mods:namePart id="42946AB9B6B0FE3FB94018EE66007991">Díaz, Florentina</mods:namePart>
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<mods:namePart id="1DBCDC2BB53A0BED3BE93847C63701E9">Felippe-Bauer, Maria Luiza</mods:namePart>
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<mods:title id="66E697A351BEA381971BA0985F923E87">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="0F67B0E620622A935F6A0C6CF13F6663">2017</mods:date>
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<paragraph id="A20AB258FFF4562360C8FF1587BEFE8C" blockId="2.[147,666,247,336]" box="[147,666,247,272]" pageId="2" pageNumber="19">
<heading id="F9420534FFF4562360C8FF1587BEFE8C" bold="true" box="[147,666,247,272]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="19" reason="2">
<taxonomicName id="65B5C9DBFFF4562360C8FF1587BEFE8C" authority="Diaz, Felippe-Bauer &amp; Spinelli" authorityName="Diaz, Felippe-Bauer &amp; Spinelli" authorityYear="2017" box="[147,666,247,272]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="jorgei" status="sp. nov.">
<emphasis id="90C16E4AFFF4562360C8FF1587BEFE8C" bold="true" box="[147,666,247,272]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF4562360C8FF158460FE8C" bold="true" box="[147,324,247,272]" italics="true" pageId="2" pageNumber="19">Dasyhelea jorgei</emphasis>
Díaz, Felippe-Bauer &amp; Spinelli
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<emphasis id="90C16E4AFFF456236136FEFA84E4FEAC" bold="true" box="[365,448,280,304]" pageId="2" pageNumber="19">sp. nov.</emphasis>
</paragraph>
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<subSubSection id="EAAFE1D3FFF456236178FEDA8662FA6C" pageId="2" pageNumber="19" type="description">
<paragraph id="A20AB258FFF456236178FEDA872EFECC" blockId="2.[147,666,247,336]" box="[291,522,312,336]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236178FEDA872EFECC" bold="true" box="[291,522,312,336]" pageId="2" pageNumber="19">
(
<figureCitation id="3A8EAEDDFFF456236172FEDA8490FECC" box="[297,436,312,336]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Figures 1-10</figureCitation>
, 
<figureCitation id="3A8EAEDDFFF45623619AFEDA8726FECC" box="[449,514,312,336]" captionStart="FIGURES 20‑27" captionStartId="4.[133,223,1800,1819]" captionTargetBox="[133,1264,548,1773]" captionTargetId="figure-179@4.[132,1266,548,1776]" captionTargetPageId="4" captionText="FIGURES 20‑27: (20-23) Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male (24-27) Dasyhelea mineira Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, (20, 24) genitalia; (21, 25) sternite 9; (22, 26) paramere and gonocoxal apodemes; (23, 27) aedeagus. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656706" httpUri="https://zenodo.org/record/4656706/files/figure.png" pageId="2" pageNumber="19">20-23</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="A20AB258FFF4562360DFFE9A84ABFE4C" blockId="2.[132,680,373,464]" pageId="2" pageNumber="19">
<materialsCitation id="12DDB805FFF4562360DFFE9A84B9FE2C" collectingDate="2004-07" collectionCode="MPLA" collectorName="J. Williams" country="Peru" location="Pagoreni" pageId="2" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="holotype">
<emphasis id="90C16E4AFFF4562360DFFE9A85C5FE0C" box="[132,225,376,400]" italics="true" pageId="2" pageNumber="19">
<typeStatus id="7D0E0CFAFFF4562360DFFE9A85F8FE0C" box="[132,220,376,400]" pageId="2" pageNumber="19" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="B4B379D1FFF4562360B3FE9785DFFE0C" box="[232,251,373,400]" pageId="2" pageNumber="19" type="male">♂</specimenCount>
, 
<collectingCountry id="DAA2F2C8FFF45623615CFE9A841CFE13" box="[263,312,376,399]" name="Peru" pageId="2" pageNumber="19">Peru</collectingCountry>
, 
<collectingRegion id="60717CBAFFF456236118FE9A84ACFE0C" box="[323,392,376,400]" country="Peru" name="Cusco" pageId="2" pageNumber="19">Cuzco</collectingRegion>
: 
<location id="A76AE483FFF4562361CAFE9A84C9FE0C" LSID="urn:lsid:plazi:treatment:2A1C034EFFF4562360C8FF158662FA6C:A76AE483FFF4562361CAFE9A84C9FE0C" box="[401,493,376,400]" country="Peru" name="Pagoreni" pageId="2" pageNumber="19" stateProvince="Cusco">Pagoreni</location>
, 
<date id="D60B9498FFF4562361ADFE9A877DFE0C" box="[502,601,376,400]" pageId="2" pageNumber="19" value="2004-07">
<collectingDate id="C64F6D70FFF4562361ADFE9A877DFE0C" box="[502,601,376,400]" pageId="2" pageNumber="19" value="2004-07">VII.2004</collectingDate>
</date>
, 
<collectorName id="0F40D78EFFF456236238FE9A859EFE2C" pageId="2" pageNumber="19">J. Williams</collectorName>
col, at light (
<collectionCode id="C4A42A9DFFF456236112FE7A84B2FE2C" box="[329,406,408,432]" pageId="2" pageNumber="19">MPLA</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="12DDB805FFF4562361F2FE7B84AFFE4C" collectingDate="2004-07" collectionCode="MLPA" collectorName="J. Williams" country="Peru" location="Pagoreni" pageId="2" pageNumber="19" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="90C16E4AFFF4562361F2FE7B872EFE2C" box="[425,522,409,432]" italics="true" pageId="2" pageNumber="19">
<typeStatus id="7D0E0CFAFFF4562361F2FE7B8721FE2C" box="[425,517,409,432]" pageId="2" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="B4B379D1FFF456236249FE7A871FFE2C" box="[530,571,405,432]" pageId="2" pageNumber="19" type="male">3 ♂</specimenCount>
, 
<specimenCount id="B4B379D1FFF456236212FE7A874AFE2C" box="[585,622,405,432]" pageId="2" pageNumber="19" type="female">1 ♀</specimenCount>
same data as holotype (
<collectionCode id="C4A42A9DFFF456236163FE5A84A0FE4C" box="[312,388,440,464]" pageId="2" pageNumber="19">MLPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A20AB258FFF4562360DFFE1B8434FD2C" blockId="2.[132,681,504,688]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF4562360DFFE1B85CDFD8C" box="[132,233,505,528]" italics="true" pageId="2" pageNumber="19">Diagnosis:</emphasis>
male with posterolateral arms single and subparallel, and posteromedian projection of aedeagus stout and rounded with notched tip. Female with palpal segment 5 very slender and slightly shorter than segment 3, and spermatheca pyriform with stout and straight neck.
</paragraph>
<paragraph id="A20AB258FFF4562360DFFD3B8599F973" blockId="2.[132,681,728,1775]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF4562360DFFD3B85DFFD6C" box="[132,251,729,752]" italics="true" pageId="2" pageNumber="19">Description:</emphasis>
Male adult. Antennal flagellum brown; with flagellomeres 2-4 globular, 5-9 rhomboid, 10-13 elongates (
<figureCitation id="3A8EAEDDFFF456236162FCFA8451FCAC" box="[313,373,792,816]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 1</figureCitation>
). Clypeus with 4 pairs of setae (
<figureCitation id="3A8EAEDDFFF4562360EFFCDA85D2FCCC" box="[180,246,824,848]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 2</figureCitation>
). Palpus (
<figureCitation id="3A8EAEDDFFF456236139FCDA8485FCCC" box="[354,417,824,848]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 3</figureCitation>
) pale brown; segment 5 0.60 length of segment 3; 
<collectingRegion id="60717CBAFFF4562361D4FCBA848BFCF3" box="[399,431,856,879]" country="Brazil" name="Para" pageId="2" pageNumber="19">PR</collectingRegion>
5.33-6.16 (5.70, n = 4). Scutellum with 6 large, 2 thinner setae. Wing (
<figureCitation id="3A8EAEDDFFF45623623FFC9A8785FC0C" box="[612,673,888,912]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 4</figureCitation>
) length 0.80-0.88 (0.84, n = 4) mm, width 0.26-0.30 (0.27, n = 4) mm; CR 0.44-0.47 (0.46, n = 4). Genitalia (
<figureCitation id="3A8EAEDDFFF45623609AFC3A8428FC6C" box="[193,268,984,1008]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Figs. 5</figureCitation>
, 
<figureCitation id="3A8EAEDDFFF456236147FC3A841EFC6C" box="[284,314,984,1008]" captionStart="FIGURES 20‑27" captionStartId="4.[133,223,1800,1819]" captionTargetBox="[133,1264,548,1773]" captionTargetId="figure-179@4.[132,1266,548,1776]" captionTargetPageId="4" captionText="FIGURES 20‑27: (20-23) Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male (24-27) Dasyhelea mineira Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, (20, 24) genitalia; (21, 25) sternite 9; (22, 26) paramere and gonocoxal apodemes; (23, 27) aedeagus. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656706" httpUri="https://zenodo.org/record/4656706/files/figure.png" pageId="2" pageNumber="19">20</figureCitation>
): tergite 9 broad on anterior half, distal half tapering to rounded tip, reaching to level of apex of gonocoxites, apicolateral process slender with apical seta; cercus short with 3-4 setae; sternite 9 (
<figureCitation id="3A8EAEDDFFF4562360D7FBBA85F2FBEC" box="[140,214,1112,1136]" captionStart="FIGURES 20‑27" captionStartId="4.[133,223,1800,1819]" captionTargetBox="[133,1264,548,1773]" captionTargetId="figure-179@4.[132,1266,548,1776]" captionTargetPageId="4" captionText="FIGURES 20‑27: (20-23) Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male (24-27) Dasyhelea mineira Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, (20, 24) genitalia; (21, 25) sternite 9; (22, 26) paramere and gonocoxal apodemes; (23, 27) aedeagus. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656706" httpUri="https://zenodo.org/record/4656706/files/figure.png" pageId="2" pageNumber="19">Fig. 21</figureCitation>
) 0.4 length of greatest width, posteromedian projection rounded, reaching ½ of total aedeagus length. Gonocoxite stout, 1.8 X longer than greatest, mesal width, with short mesal process; gonostylus 0.9 length of gonocoxite, base wide, slightly curved with pointed tip. Paramere and gonocoxal apodemes (
<figureCitation id="3A8EAEDDFFF4562360D0FAFA85F4FAAC" box="[139,208,1304,1328]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Figs. 5</figureCitation>
, 
<figureCitation id="3A8EAEDDFFF456236081FAFA85D2FAAC" box="[218,246,1304,1328]" captionStart="FIGURES 20‑27" captionStartId="4.[133,223,1800,1819]" captionTargetBox="[133,1264,548,1773]" captionTargetId="figure-179@4.[132,1266,548,1776]" captionTargetPageId="4" captionText="FIGURES 20‑27: (20-23) Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male (24-27) Dasyhelea mineira Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, (20, 24) genitalia; (21, 25) sternite 9; (22, 26) paramere and gonocoxal apodemes; (23, 27) aedeagus. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656706" httpUri="https://zenodo.org/record/4656706/files/figure.png" pageId="2" pageNumber="19">22</figureCitation>
) forming an asymmetrical structure; gonocoxal apodemes stout, proximal ⅔ nearly straight, distal ⅓ directed mesally, with short, pointed, subbasal tooth; right apodeme broadly fused with paramere, left apodeme not contacting paramere, latter slender, simple, tapering distally, tip rounded, produced beyond apex of sternite 9 by short distance. Aedeagus (
<figureCitation id="3A8EAEDDFFF4562360D0FA1A85F0F98C" box="[139,212,1528,1552]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Figs. 5</figureCitation>
, 
<figureCitation id="3A8EAEDDFFF4562360B8FA1A85DAF98C" box="[227,254,1528,1552]" captionStart="FIGURES 20‑27" captionStartId="4.[133,223,1800,1819]" captionTargetBox="[133,1264,548,1773]" captionTargetId="figure-179@4.[132,1266,548,1776]" captionTargetPageId="4" captionText="FIGURES 20‑27: (20-23) Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male (24-27) Dasyhelea mineira Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, (20, 24) genitalia; (21, 25) sternite 9; (22, 26) paramere and gonocoxal apodemes; (23, 27) aedeagus. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656706" httpUri="https://zenodo.org/record/4656706/files/figure.png" pageId="2" pageNumber="19">23</figureCitation>
) 1.1 X longer than greatest width, W/L ratio 0.38-0.45 (0.41, n = 4); anterior margin slightly convex, basal arms slender, well sclerotized, directed anterolaterally; posterolateral arms single, subparallel, each with apex abruptly recurved ventrolaterally; posteromedian projection stout, rounded with notched tip, produced beyond recurved apices of posterolateral arms.
</paragraph>
<paragraph id="A20AB258FFF4562360DFF8FB811DFECC" blockId="2.[132,681,1816,1872]" lastBlockId="2.[718,1266,184,336]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF4562360DFF8FB85DFF8AC" box="[132,251,1817,1840]" italics="true" pageId="2" pageNumber="19">Description:</emphasis>
Female adult. Similar to male, except: Head brown. Eyes contiguous for distance of width of 2-3 ommatidia. Antennal flagellum brown; with flagellomeres 2-8 vasiform, 9-13 more elongate vasiform, 13 longest (
<figureCitation id="3A8EAEDDFFF4562363CAFF1A86F5FE8C" box="[913,977,248,272]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 6</figureCitation>
) AR 0.74. Clypeus with 5 pairs of setae. Palpus (
<figureCitation id="3A8EAEDDFFF4562363EDFEFA86D0FEAC" box="[950,1012,280,304]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 7</figureCitation>
) pale brown; segment 5 0.83 length of segment 3; 
<collectingRegion id="60717CBAFFF456236386FEDA86D9FED3" box="[989,1021,312,335]" country="Brazil" name="Para" pageId="2" pageNumber="19">PR</collectingRegion>
4.33.
</paragraph>
<paragraph id="A20AB258FFF456236295FE9A87DBFDEC" blockId="2.[718,1266,376,624]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236295FE9A8638FE0C" box="[718,796,376,400]" italics="true" pageId="2" pageNumber="19">Thorax:</emphasis>
Uniformly brown; scutellum with 6 large, 1 thinner setae. Legs brown; apex of hind tibia with 8 spines; foreleg TR 1.60, midleg TR 2.03, hind leg TR 2.14. Wing (
<figureCitation id="3A8EAEDDFFF4562363F2FE3A86C3FE6C" box="[937,999,472,496]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 8</figureCitation>
) length 
<quantity id="654D1FBDFFF45623641FFE3A818DFE73" box="[1092,1193,472,496]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="19" unit="mm" value="0.56">0.56 mm</quantity>
, width 
<quantity id="654D1FBDFFF456236295FE1A8615FD93" box="[718,817,504,528]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="19" unit="mm" value="0.28">0.28 mm</quantity>
; CR 0.46; membrane hyaline, densely covered with macrotrichia; radial cells obliterated; cubital fork at level of beginning of second radial cell. Halter pale.
</paragraph>
<paragraph id="A20AB258FFF456236295FD7A8178FCD3" blockId="2.[718,1267,664,848]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236295FD7A8616FD2C" box="[718,818,664,688]" italics="true" pageId="2" pageNumber="19">Abdomen:</emphasis>
Dark brown. Subgenital plate (
<figureCitation id="3A8EAEDDFFF45623642AFD7A8189FD2C" box="[1137,1197,664,688]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Fig. 9</figureCitation>
) semicircular, with broad, lumen, posterior margin straight; posteromedian projection slender, pointed; posterolateral arms stout, curved. Spermatheca (
<figureCitation id="3A8EAEDDFFF456236424FD1A81CEFC8C" box="[1151,1258,760,784]" captionStart="FIGURES 1‑10" captionStartId="1.[95,185,1826,1845]" captionTargetBox="[94,1228,404,1802]" captionTargetId="figure-88@1.[94,1228,404,1802]" captionTargetPageId="1" captionText="FIGURES 1‑10: Dasyhelea jorgei Diaz, Felippe-Bauer &amp; Spinelli sp. nov. Holotype male, adult. (1-5) male; (6-10) female. (1, 6) antennal flagellum; (2) clypeus; (3, 7) palpus; (4, 8) wing; (5) genitalia; (9) subgenital plate; (10) spermathecae. Scale bars: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4656700" httpUri="https://zenodo.org/record/4656700/files/figure.png" pageId="2" pageNumber="19">Figs. 9-10</figureCitation>
) pyriform, well sclerotized, measuring 26 × 34 um, neck stout, straight, measuring 16 um.
</paragraph>
<paragraph id="A20AB258FFF456236295FC9A811BFC6C" blockId="2.[718,1267,888,1008]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236295FC9A861DFC0C" box="[718,825,888,912]" italics="true" pageId="2" pageNumber="19">Etymology:</emphasis>
This species is named after Prof. Jorge D. Williams, herpetologist at the Museo de La Plata, 
<collectingCountry id="DAA2F2C8FFF45623648BFC7A8639FC4C" name="Argentina" pageId="2" pageNumber="19">Argentina</collectingCountry>
, in recognition of his friendship and important help collecting ceratopogonids.
</paragraph>
<paragraph id="A20AB258FFF456236295FBFA86C1FBAC" blockId="2.[718,997,1048,1072]" box="[718,997,1048,1072]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236295FBFA8674FBAC" box="[718,848,1048,1072]" italics="true" pageId="2" pageNumber="19">Distribution:</emphasis>
<collectingCountry id="DAA2F2C8FFF45623630DFBFA86A0FBB3" box="[854,900,1048,1071]" name="Peru" pageId="2" pageNumber="19">Peru</collectingCountry>
(
<collectingRegion id="60717CBAFFF4562363C8FBFA86FEFBAC" box="[915,986,1048,1072]" country="Peru" name="Cusco" pageId="2" pageNumber="19">Cuzco</collectingRegion>
).
</paragraph>
<paragraph id="A20AB258FFF456236295FBBB86DEFAAC" blockId="2.[718,1267,1112,1520]" pageId="2" pageNumber="19">
<emphasis id="90C16E4AFFF456236295FBBB86C7FBEC" box="[718,995,1112,1136]" italics="true" pageId="2" pageNumber="19">
Discussion: 
<taxonomicName id="65B5C9DBFFF456236318FBBA86C7FBEC" authority="Díaz &amp; Felippe-Bauer &amp; Spinelli, 2017" authorityName="Díaz &amp; Felippe-Bauer &amp; Spinelli" authorityYear="2017" box="[835,995,1112,1136]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="jorgei" status="sp. nov.">Dasyhelea jorgei</taxonomicName>
</emphasis>
<taxonomicNameLabel id="8BF2D331FFF4562363B7FBBA8118FBF3" box="[1004,1084,1112,1135]" pageId="2" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
is very similar to the other species herein described as new, 
<taxonomicName id="65B5C9DBFFF4562364D8FB9B81D6FB0C" authority="Díaz &amp; Felippe-Bauer &amp; Spinelli, 2017" authorityName="Díaz &amp; Felippe-Bauer &amp; Spinelli" authorityYear="2017" box="[1155,1266,1145,1168]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="mineira" status="sp. nov.">
<emphasis id="90C16E4AFFF4562364D8FB9B81D6FB0C" box="[1155,1266,1145,1168]" italics="true" pageId="2" pageNumber="19">D. mineira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BF2D331FFF456236295FB7A8607FB33" box="[718,803,1176,1199]" pageId="2" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, by virtue of the very elongate and slender palpal segment 5. However, in males of 
<taxonomicName id="65B5C9DBFFF4562364DAFB5B81D6FB4C" authority="Díaz &amp; Felippe-Bauer &amp; Spinelli, 2017" authorityName="Díaz &amp; Felippe-Bauer &amp; Spinelli" authorityYear="2017" box="[1153,1266,1209,1232]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="mineira" status="sp. nov.">
<emphasis id="90C16E4AFFF4562364DAFB5B81D6FB4C" box="[1153,1266,1209,1232]" italics="true" pageId="2" pageNumber="19">D. mineira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8BF2D331FFF456236295FB3A863AFB73" box="[718,798,1240,1263]" pageId="2" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
the paramere and gonocoxal apodemes form a symmetrical structure, and the female spermatheca has a stout and oblique neck.
</paragraph>
<paragraph id="A20AB258FFF45623635CFADA8662FA6C" blockId="2.[718,1267,1112,1520]" pageId="2" pageNumber="19">
This new species is also similar to the Patagonian species 
<taxonomicName id="65B5C9DBFFF456236346FABB81CBFAF3" authority="Diaz &amp; Spinelli. However" authorityName="Diaz &amp; Spinelli. However" box="[797,1263,1368,1392]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="pseudolacustris">
<emphasis id="90C16E4AFFF456236346FABB86F5FAEC" box="[797,977,1368,1392]" italics="true" pageId="2" pageNumber="19">D. pseudolacustris</emphasis>
Díaz &amp; Spinelli. However
</taxonomicName>
, the palpal segment 5 of the latter species is shorter and stouter, the paramere and gonocoxal apodemes form a symmetrical structure, and the female spermatheca is spherical.
</paragraph>
</subSubSection>
</treatment>
</document>