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<document id="715AC88455A0700734F549B7F86F5BFB" ID-CLB-Dataset="6148" ID-DOI="10.11646/zootaxa.4975.3.2" ID-GBIF-Dataset="330b4589-7947-450b-857f-812cb72beb6d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4808307" ID-ZooBank="222B81FA-E0F5-43A9-960D-5C9ACC2ADD0A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622026211124" checkinUser="plazi" docAuthor="Martynov, Alexander V., Selvakumar, C., Subramanian, K. A., Sivaramakrishnan, K. G., Vasanth, M., Sinha, Bikramjit &amp; Jacobus, Luke M." docDate="2021" docId="1D7287CDFFDDD663FF090CD70ED4FBD4" docLanguage="en" docName="zootaxa.4975.3.2.pdf" docOrigin="Zootaxa 4975 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torleya coheri" docType="treatment" docVersion="8" lastPageNumber="467" masterDocId="E14BFFB5FFD2D673FF9E0C080A62FFC5" masterDocTitle="Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions" masterLastPageNumber="482" masterPageNumber="451" pageNumber="466" updateTime="1698991974093" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="386BADA4C6FBD98DDBE0F463CE2C84B7">Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions</mods:title>
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<mods:namePart id="72AA868AD260F7F6B40DF4FF680CD35B">Martynov, Alexander V.</mods:namePart>
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<mods:affiliation id="69F8E1290BA6ECE28450C27EEA8DC8E5">National Museum of Natural History, National Academy of Sciences of Ukraine, Bohdan Khmelnytsky Str., 15, 01030, Kyiv, Ukraine. martynov _ av @ ukr. net; centroptilum @ gmail. com; https: // orcid. org / 0000 - 0002 - 6506 - 5134</mods:affiliation>
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<mods:namePart id="3EC41FF8BE2C28E67699A3074D20DFF5">Selvakumar, C.</mods:namePart>
<mods:affiliation id="D5381DED1610D9D5717980F855562494">Department of Zoology, The Madura College (Autonomous), Madurai- 625011, Tamil Nadu, India.</mods:affiliation>
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<mods:affiliation id="35734CAB61B0F167D5D0797F803744C8">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India. &amp; subbuka. zsi @ gmail. com; https: // orcid. org / 0000 - 0003 - 0872 - 9771</mods:affiliation>
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<mods:namePart id="BC16F6F22F9F9CD1AFBDE3D4B21E363D">Vasanth, M.</mods:namePart>
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<mods:affiliation id="5FC6DF822E30912BD1A67DDC46048D57">Zoological Survey of India, Southern Regional Centre, Santhome High Road, Chennai- 600028, India. &amp; vasan 071994 @ gmail. com; https: // orcid. org / 0000 - 0003 - 4446 - 4889</mods:affiliation>
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<mods:namePart id="C1CC8E6B741E27AE8D9C62321957D4D3">Sinha, Bikramjit</mods:namePart>
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<mods:namePart id="80472289B56AF36CF5063ABDD0C5B70E">Jacobus, Luke M.</mods:namePart>
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<mods:affiliation id="2C7B69093A376844298898556DE69DEC">Division of Science, Indiana University Purdue University Columbus, Columbus, Indiana, 47203, USA. lukemjacobus @ alumni. purdue. edu; https: // orcid. org / 0000 - 0002 - 7201 - 3922</mods:affiliation>
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<taxonomicName id="52DB4D58FFDDD67CFF090CD708EEFF3C" ID-CoL="57FRW" authority="(Allen &amp; Edmunds, 1963)" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" box="[151,652,223,250]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="coheri">
<emphasis id="A7AFEAC9FFDDD67CFF090CD708EEFF3C" bold="true" box="[151,652,223,250]" pageId="15" pageNumber="466">
<emphasis id="A7AFEAC9FFDDD67CFF090CD70B24FF3F" bold="true" box="[151,326,223,250]" italics="true" pageId="15" pageNumber="466">Torleya coheri</emphasis>
(
<bibRefCitation id="F14A4B2AFFDDD67CFECB0CD708E6FF3C" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[341,644,223,249]" pageId="15" pageNumber="466" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
)
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(
<figureCitation id="0DE02A5EFFDDD67CFF3E0D0B0A93FED8" box="[160,241,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1854,1878]" captionTargetBox="[151,1436,766,1811]" captionTargetId="figure-176@15.[151,1436,766,1830]" captionTargetPageId="15" captionText="FIGURE 10. Larva of Torleya coheri (Allen &amp; Edmunds, 1963). A.—total view of larva; B.—tarsus and tarsal claw, black arrow shows palisade of denticles; C.—abdomen, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808351" httpUri="https://zenodo.org/record/4808351/files/figure.png" pageId="15" pageNumber="466">Fig. 10</figureCitation>
)
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<taxonomicName id="52DB4D58FFDDD67CFF090D40083FFE9A" authority="Allen &amp; Edmunds, 1963" authorityName="Allen &amp; Edmunds" authorityYear="1963" box="[151,605,327,352]" class="Insecta" family="Ephemerellidae" genus="Ephemerella" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="coheri">
<emphasis id="A7AFEAC9FFDDD67CFF090D400B3DFEA5" box="[151,351,328,352]" italics="true" pageId="15" pageNumber="466">Ephemerella coheri</emphasis>
<bibRefCitation id="F14A4B2AFFDDD67CFEFB0D40083FFE9A" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[357,605,327,352]" pageId="15" pageNumber="466" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
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<taxonomicName id="52DB4D58FFDDD67CFF090D6E0855FEBB" authority="Wang &amp; Sites, 1999" authorityName="Wang &amp; Sites" authorityYear="1999" box="[151,567,358,383]" class="Insecta" family="Ephemerellidae" genus="Crinitella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="permkami">
<emphasis id="A7AFEAC9FFDDD67CFF090D6E0B07FEBB" box="[151,357,358,382]" italics="true" pageId="15" pageNumber="466">Crinitella permkami</emphasis>
<bibRefCitation id="F14A4B2AFFDDD67CFEF20D6F0855FEBB" author="Wang, T. &amp; Sites, R. W." box="[364,567,358,383]" pageId="15" pageNumber="466" pagination="73 - 77" refId="ref16266" refString="Wang, T. &amp; Sites, R. W. (1999) Description of a new species of Crinitella (Ephemeroptera: Ephemerellidae) from Thailand. Journal of the New York Entomological Society, 107 (1), 73 - 77." type="journal article" year="1999">Wang &amp; Sites, 1999</bibRefCitation>
</taxonomicName>
(junior synonym,
<bibRefCitation id="F14A4B2AFFDDD67CFD6C0D6E0E71FEBB" author="Jacobus, L. M. &amp; McCafferty, W. P." box="[754,1043,358,382]" pageId="15" pageNumber="466" pagination="48 - 50" refId="ref14824" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2003) Revisions to the genus Crinitella (Ephemeroptera: Ephmerellidae). Journal of the New York Entomological Society, 111 (1), 48 - 50." type="journal article" year="2003">Jacobus &amp; McCafferty 2003</bibRefCitation>
)
</paragraph>
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<emphasis id="A7AFEAC9FFDDD67CFF090DA40BE0FE03" bold="true" box="[151,386,428,454]" pageId="15" pageNumber="466">Material examined.</emphasis>
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<collectingCountry id="EDCC764BFFDDD67CFE150DA40BBFFE03" box="[395,477,428,454]" name="India" pageId="15" pageNumber="466">INDIA</collectingCountry>
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:
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a.s.l.
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, h ~
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a.s.l.
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&amp;
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<collectorName id="382E530DFFDDD67CFDBA0E1008E5FDF6" box="[548,647,536,563]" pageId="15" pageNumber="466">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFDDD67CFD110E110962FDF6" box="[655,768,537,563]" italics="true" pageId="15" pageNumber="466">5347/H13</emphasis>
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;
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<elevation id="1EF6D1E8FFDDD67CFD190E35096AFD93" box="[647,776,572,599]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="15" pageNumber="466" unit="m" value="128.0">
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a.s.l.
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,
<date id="E165101BFFDDD67CFC8D0E3409EEFD92" box="[787,908,572,599]" pageId="15" pageNumber="466" value="2015-04-22">
<collectingDate id="F121E9F3FFDDD67CFC8D0E3409EEFD92" box="[787,908,572,599]" pageId="15" pageNumber="466" value="2015-04-22">22.iv.2015</collectingDate>
</date>
,
<collectorName id="382E530DFFDDD67CFC080E340ECCFD93" box="[918,1198,572,599]" pageId="15" pageNumber="466">Colls. K.A. Subramanian</collectorName>
&amp;
<collectorName id="382E530DFFDDD67CFB510E340FE2FD93" box="[1231,1408,572,598]" pageId="15" pageNumber="466">Bikramjit Sinha</collectorName>
<collectorName id="382E530DFFDDD67CFF090E690A99FDBE" box="[151,251,609,635]" pageId="15" pageNumber="466">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFDDD67CFE9E0E680B12FDBF" box="[256,368,608,634]" italics="true" pageId="15" pageNumber="466">5609/H13</emphasis>
[
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]
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;
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<specimenCount id="83DDFD52FFDDD67CFE5E0E69086DFDBF" box="[448,527,608,635]" pageId="15" pageNumber="466" type="larva">1 larva</specimenCount>
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<collectingRegion id="571FF839FFDDD67CFD870E6908CAFDBF" box="[537,680,608,635]" country="India" name="West Bengal" pageId="15" pageNumber="466">West Bengal</collectingRegion>
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(
<collectingRegion id="571FF839FFDDD67CFCA80E6809EBFDBF" box="[822,905,608,634]" country="India" name="Sikkim" pageId="15" pageNumber="466">Sikkim</collectingRegion>
border),
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,
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, h ~
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a.s.l.
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,
<date id="E165101BFFDDD67CFD9C0E8C081CFD5B" box="[514,638,644,670]" pageId="15" pageNumber="466" value="2013-03-23">
<collectingDate id="F121E9F3FFDDD67CFD9C0E8C081CFD5B" box="[514,638,644,670]" pageId="15" pageNumber="466" value="2013-03-23">23.iii.2013</collectingDate>
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<collectorName id="382E530DFFDDD67CFC090E8D099EFD5A" box="[919,1020,645,671]" pageId="15" pageNumber="466">Reg. No.</collectorName>
<emphasis id="A7AFEAC9FFDDD67CFB9B0E8D0E17FD5A" box="[1029,1141,645,671]" italics="true" pageId="15" pageNumber="466">5349/H13</emphasis>
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.
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<emphasis id="A7AFEAC9FFDDD67CFB550E8C0F4AFD5B" bold="true" box="[1227,1320,644,670]" pageId="15" pageNumber="466">
<collectingCountry id="EDCC764BFFDDD67CFB550E8C0F4AFD5B" box="[1227,1320,644,670]" name="Nepal" pageId="15" pageNumber="466">NEPAL</collectingCountry>
</emphasis>
:
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<collectingRegion id="571FF839FFDDD67CFF090EA00B5FFD06" box="[151,317,680,707]" country="Nepal" name="Bagmati" pageId="15" pageNumber="466">Bagmati Zone</collectingRegion>
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<collectingMunicipality id="7500ACA1FFDDD67CFED20EA10B1EFD06" box="[332,380,681,707]" pageId="15" pageNumber="466">East</collectingMunicipality>
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<location id="90046000FFDDD67CFE500EA0086CFD07" LSID="urn:lsid:plazi:treatment:1D7287CDFFDDD663FF090CD70ED4FBD4:90046000FFDDD67CFE500EA0086CFD07" box="[462,526,680,706]" country="Nepal" latitude="27.57163" longLatPrecision="1" longitude="84.668755" municipality="East" name="River" pageId="15" pageNumber="466" stateProvince="Bagmati">River</location>
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, h—
<elevation id="1EF6D1E8FFDDD67CFCC60EA00980FD07" box="[856,994,680,707]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="466" unit="m" value="230.0">
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a.s.l.
</elevation>
,
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<collectingDate id="F121E9F3FFDDD67CFC6C0EA00E3CFD07" box="[1010,1118,680,706]" pageId="15" pageNumber="466" value="2014-01-27">27.i.2014</collectingDate>
</date>
,
<collectorName id="382E530DFFDDD67CFBF30EA00F74FD06" box="[1133,1302,680,707]" pageId="15" pageNumber="466">Chertoprud M.</collectorName>
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. leg.—
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<materialsCitation id="25B33C86FFDDD67CFF090EC40860FD22" ID-GBIF-Occurrence="3128883304" box="[151,514,716,743]" collectionCode="NMNH, NASU" country="Nepal" location="Torsp" pageId="15" pageNumber="466" specimenCount="1">
<emphasis id="A7AFEAC9FFDDD67CFF090EC40B59FD23" box="[151,315,716,742]" italics="true" pageId="15" pageNumber="466">
<taxonomicName id="52DB4D58FFDDD67CFF090EC40AB1FD23" box="[151,211,716,742]" class="Insecta" family="Nepidae" genus="Nepa" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="genus">Nepa</taxonomicName>
16
<location id="90046000FFDDD67CFF6E0EC40B52FD23" LSID="urn:lsid:plazi:treatment:1D7287CDFFDDD663FF090CD70ED4FBD4:90046000FFDDD67CFF6E0EC40B52FD23" box="[240,304,716,742]" country="Nepal" name="Torsp" pageId="15" pageNumber="466">Torsp</location>
1
</emphasis>
[
<collectionCode id="F3CAAE1EFFDDD67CFED60EC50BC3FD22" box="[328,417,717,743]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="466" type="Museum">NMNH</collectionCode>
<collectionCode id="F3CAAE1EFFDDD67CFE360EC40B99FD22" box="[424,507,716,743]" pageId="15" pageNumber="466">NASU</collectionCode>
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.
</paragraph>
</subSubSection>
<caption id="C1A46653FFDDD67CFF090B3609DEF8BF" ID-DOI="http://doi.org/10.5281/zenodo.4808351" ID-Zenodo-Dep="4808351" httpUri="https://zenodo.org/record/4808351/files/figure.png" pageId="15" pageNumber="466" startId="15.[151,250,1854,1878]" targetBox="[151,1436,766,1811]" targetPageId="15">
<paragraph id="956436DBFFDDD67CFF090B3609DEF8BF" blockId="15.[151,1436,1854,1914]" pageId="15" pageNumber="466">
<emphasis id="A7AFEAC9FFDDD67CFF090B360B79F893" bold="true" box="[151,283,1854,1878]" pageId="15" pageNumber="466">FIGURE 10</emphasis>
. Larva of
<taxonomicName id="52DB4D58FFDDD67CFE150B36095EF893" authority="(Allen &amp; Edmunds, 1963)" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" box="[395,828,1854,1878]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="coheri">
<emphasis id="A7AFEAC9FFDDD67CFE150B360843F893" box="[395,545,1854,1878]" italics="true" pageId="15" pageNumber="466">Torleya coheri</emphasis>
(
<bibRefCitation id="F14A4B2AFFDDD67CFDAE0B360951F893" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[560,819,1854,1878]" pageId="15" pageNumber="466" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="A7AFEAC9FFDDD67CFCD90B360902F893" bold="true" box="[839,864,1854,1878]" pageId="15" pageNumber="466">A.</emphasis>
—total view of larva;
<emphasis id="A7AFEAC9FFDDD67CFBD60B360E3DF893" bold="true" box="[1096,1119,1854,1878]" pageId="15" pageNumber="466">B.</emphasis>
—tarsus and tarsal claw, black arrow shows palisade of denticles;
<emphasis id="A7AFEAC9FFDDD67CFE640B6A0871F8BF" bold="true" box="[506,531,1890,1914]" pageId="15" pageNumber="466">C.</emphasis>
—abdomen, dorsal view. Scale bar 1 mm.
</paragraph>
</caption>
<subSubSection id="DDC16550FFDDD663FF590BA00EBCFDA4" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466" type="diagnosis">
<paragraph id="956436DBFFDDD663FF590BA00EBCFDA4" blockId="15.[151,1436,1960,2023]" lastBlockId="16.[151,1437,151,1042]" lastPageId="16" lastPageNumber="467" pageId="15" pageNumber="466">
<emphasis id="A7AFEAC9FFDDD67CFF590BA00B20F807" bold="true" box="[199,322,1960,1986]" pageId="15" pageNumber="466">Diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="52DB4D58FFDDD67CFCFB0BA00E74F807" box="[869,1046,1960,1986]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A7AFEAC9FFDDD67CFCFB0BA009D8F807" box="[869,954,1960,1986]" italics="true" pageId="15" pageNumber="466">Torleya</emphasis>
species
</taxonomicName>
by the following combination of characters:
<emphasis id="A7AFEAC9FFDDD67CFE880BC50B35F823" box="[278,343,1997,2022]" italics="true" pageId="15" pageNumber="466">Larva</emphasis>
: (i) dorsal surface of body covered by long hair-like setae (
<figureCitation id="0DE02A5EFFDDD67CFC670BC40E02F822" box="[1017,1120,1996,2023]" captionStart="FIGURE 10" captionStartId="15.[151,250,1854,1878]" captionTargetBox="[151,1436,766,1811]" captionTargetId="figure-176@15.[151,1436,766,1830]" captionTargetPageId="15" captionText="FIGURE 10. Larva of Torleya coheri (Allen &amp; Edmunds, 1963). A.—total view of larva; B.—tarsus and tarsal claw, black arrow shows palisade of denticles; C.—abdomen, dorsal view. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4808351" httpUri="https://zenodo.org/record/4808351/files/figure.png" pageId="15" pageNumber="466">Fig. 10A</figureCitation>
; fig.
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); (ii) maxilla with rudimentary palp (fig.
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<bibRefCitation id="F14A4B2AFFC2D663FD7F0C9F098DFF77" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[737,1007,151,178]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (iii) anteromedian emargination very shallow, almost absent (fig.
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<bibRefCitation id="F14A4B2AFFC2D663FD930CB3097BFF13" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[525,793,187,214]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (iv) labium highly reduced (fig.
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); (v) abdominal segment VIII with a unique shape and setation (
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; fig.
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<bibRefCitation id="F14A4B2AFFC2D663FBC70CD70F38FF3C" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[1113,1370,223,249]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
; fig.
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FE8E0D0B0B25FED8" box="[272,327,259,285]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
2007
</bibRefCitation>
); (vi) posterolateral projections of segment IX extremely elongated (
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; fig.
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<bibRefCitation id="F14A4B2AFFC2D663FAC10D0B0B02FE87" author="Allen, R. K. &amp; Edmunds Jr., G. F." pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (vii) tarsal claw with distinct distal palisade of denticles (
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; fig.
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<bibRefCitation id="F14A4B2AFFC2D663FB4F0D2F0AB0FEA3" author="Allen, R. K. &amp; Edmunds Jr., G. F." pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (viii) lateral margins of abdominal segments IVVIII with short stout setae; stout setae on segment VIII pass over acute posterior projection (fig.
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<bibRefCitation id="F14A4B2AFFC2D663FDD60D67092CFE4F" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[584,846,367,394]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (ix) paired submedial projections on abdominal terga are absent (
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; fig.
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<bibRefCitation id="F14A4B2AFFC2D663FE420D9B0886FE6B" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[476,740,403,430]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds 1963</bibRefCitation>
); (x) gills IIIVI with outer posterolateral protuberances, gill III not covering following gills entirely (figs 1, 16 in
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).
<emphasis id="A7AFEAC9FFC2D663FC590DBF0E28FE15" box="[967,1098,439,465]" italics="true" pageId="16" pageNumber="467">Male imago</emphasis>
: (xi) penis short, lobes apically rounded (fig.
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FE230DD30B97FE30" box="[445,501,475,501]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
2007
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); (xii) dorsal projection of penis small, rounded, with broad apical cleft (figs 2, 4 in
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FEB60DF70B02FDDC" box="[296,352,511,537]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
2007
</bibRefCitation>
) (xiii) genital forceps segment II robust, with relatively straight lateral profile, not expanded apically and segment III subovoid (fig.
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Jacobus
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2007
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); (xiv) abdomen with light general coloration and purple shading, lacking a prominent, middorsal, longitudinal stripe (figs 3, 4 in
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FBC70E4F0EF2FDA4" box="[1113,1168,583,609]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
2007
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).
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC2D663FF590E620994FD08" pageId="16" pageNumber="467" type="distribution">
<paragraph id="956436DBFFC2D663FF590E620994FD08" blockId="16.[151,1437,151,1042]" pageId="16" pageNumber="467">
<emphasis id="A7AFEAC9FFC2D663FF590E620B03FD41" bold="true" box="[199,353,618,644]" pageId="16" pageNumber="467">Distribution.</emphasis>
<collectingCountry id="EDCC764BFFC2D663FEF50E630BC7FD40" box="[363,421,619,645]" name="India" pageId="16" pageNumber="467">India</collectingCountry>
(
<bibRefCitation id="F14A4B2AFFC2D663FE280E630859FD40" author="Allen, R. K." box="[438,571,619,645]" pageId="16" pageNumber="467" pagination="71 - 91" refId="ref14005" refString="Allen, R. K. (1980) Geographic distribution and reclassification of the subfamily Ephemerellinae (Ephemeroptera: Ephemerellidae). In: Flannagan, J. F. &amp; Marshall, K. E. (Eds.), Advances in Ephemeroptera Biology. Plenum Press, New York, pp. 71 - 91. https: // doi. org / 10.1007 / 978 - 1 - 4613 - 3066 - 0 _ 6" type="book chapter" year="1980">Allen 1980</bibRefCitation>
,
<bibRefCitation id="F14A4B2AFFC2D663FDD40E6309F6FD40" author="Jacobus, L. M. &amp; McCafferty, W. P." box="[586,916,619,645]" pageId="16" pageNumber="467" pagination="48 - 50" refId="ref14824" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2003) Revisions to the genus Crinitella (Ephemeroptera: Ephmerellidae). Journal of the New York Entomological Society, 111 (1), 48 - 50." type="journal article" year="2003">Jacobus &amp; McCafferty 2003</bibRefCitation>
,
<bibRefCitation id="F14A4B2AFFC2D663FC3C0E630EE4FD40" author="Jacobus, L. M. &amp; McCafferty, W. P. &amp; Sites, R. W." box="[930,1158,619,645]" pageId="16" pageNumber="467" pagination="45 - 53" refId="ref15040" refString="Jacobus, L. M., McCafferty, W. P. &amp; Sites, R. W. (2007) A new species and first stage associations in Crinitella (Ephemeroptera: Ephemerellidae: Ephemerellinae). Zootaxa, 1611 (1), 45 - 53. https: // doi. org / 10.11646 / zootaxa. 1611.1.3" type="journal article" year="2007">
Jacobus
<emphasis id="A7AFEAC9FFC2D663FB9B0E630E58FD40" box="[1029,1082,619,645]" italics="true" pageId="16" pageNumber="467">et al</emphasis>
. 2007
</bibRefCitation>
, new data), India-China border region (new data),
<collectingCountry id="EDCC764BFFC2D663FE590E87084CFD6C" box="[455,558,655,681]" name="Malaysia" pageId="16" pageNumber="467">Malaysia</collectingCountry>
(
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FD3A0E8708BAFD6C" box="[676,728,655,681]" italics="true" pageId="16" pageNumber="467">et al</emphasis>
. 2007
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),
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(
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, new data),
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(
<bibRefCitation id="F14A4B2AFFC2D663FF010EBB0B1FFD08" author="Wang, T. &amp; Sites, R. W." box="[159,381,691,717]" pageId="16" pageNumber="467" pagination="73 - 77" refId="ref16266" refString="Wang, T. &amp; Sites, R. W. (1999) Description of a new species of Crinitella (Ephemeroptera: Ephemerellidae) from Thailand. Journal of the New York Entomological Society, 107 (1), 73 - 77." type="journal article" year="1999">Wang &amp; Sites 1999</bibRefCitation>
;
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FE770EBB0878FD08" box="[489,538,691,717]" italics="true" pageId="16" pageNumber="467">et al</emphasis>
. 2007
</bibRefCitation>
) and
<collectingCountry id="EDCC764BFFC2D663FD010EBB089CFD08" box="[671,766,691,717]" name="Vietnam" pageId="16" pageNumber="467">Vietnam</collectingCountry>
(
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Jacobus
<emphasis id="A7AFEAC9FFC2D663FCF00EBB09FDFD08" box="[878,927,691,717]" italics="true" pageId="16" pageNumber="467">et al</emphasis>
. 2007
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).
</paragraph>
</subSubSection>
<subSubSection id="DDC16550FFC2D663FF590EDF0ED4FBD4" pageId="16" pageNumber="467" type="discussion">
<paragraph id="956436DBFFC2D663FF590EDF0E90FC60" blockId="16.[151,1437,151,1042]" pageId="16" pageNumber="467">
<emphasis id="A7AFEAC9FFC2D663FF590EDF0B59FD34" bold="true" box="[199,315,727,753]" pageId="16" pageNumber="467">Remarks.</emphasis>
The larva of this species was adequately described and illustrated from
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by
<bibRefCitation id="F14A4B2AFFC2D663FB4E0EDF0A80FCD3" author="Allen, R. K. &amp; Edmunds Jr., G. F." pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds (1963)</bibRefCitation>
and from
<collectingCountry id="EDCC764BFFC2D663FEC80EF30BD8FCD0" box="[342,442,763,789]" name="Thailand" pageId="16" pageNumber="467">Thailand</collectingCountry>
by
<bibRefCitation id="F14A4B2AFFC2D663FE7D0EF308B3FCD0" author="Wang, T. &amp; Sites, R. W." box="[483,721,763,789]" pageId="16" pageNumber="467" pagination="73 - 77" refId="ref16266" refString="Wang, T. &amp; Sites, R. W. (1999) Description of a new species of Crinitella (Ephemeroptera: Ephemerellidae) from Thailand. Journal of the New York Entomological Society, 107 (1), 73 - 77." type="journal article" year="1999">Wang &amp; Sites (1999)</bibRefCitation>
(as
<taxonomicName id="52DB4D58FFC2D663FC9E0EF30EB2FCD0" authority="Wang &amp; Sites, 1999" authorityName="Wang &amp; Sites" authorityYear="1999" box="[768,1232,763,789]" class="Insecta" family="Ephemerellidae" genus="Crinitella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="permkami">
<emphasis id="A7AFEAC9FFC2D663FC9E0EF30986FCD0" box="[768,996,763,789]" italics="true" pageId="16" pageNumber="467">Crinitella permkami</emphasis>
<bibRefCitation id="F14A4B2AFFC2D663FC750EF30EB2FCD0" author="Wang, T. &amp; Sites, R. W." box="[1003,1232,763,789]" pageId="16" pageNumber="467" pagination="73 - 77" refId="ref16266" refString="Wang, T. &amp; Sites, R. W. (1999) Description of a new species of Crinitella (Ephemeroptera: Ephemerellidae) from Thailand. Journal of the New York Entomological Society, 107 (1), 73 - 77." type="journal article" year="1999">Wang &amp; Sites, 1999</bibRefCitation>
</taxonomicName>
, junior synonym). Larvae of this species already were reported from Kashmir, and the adult description and details of species variability were provided by
<bibRefCitation id="F14A4B2AFFC2D663FEFB0F4B0836FC98" author="Jacobus, L. M. &amp; McCafferty, W. P. &amp; Sites, R. W." box="[357,596,835,861]" pageId="16" pageNumber="467" pagination="45 - 53" refId="ref15040" refString="Jacobus, L. M., McCafferty, W. P. &amp; Sites, R. W. (2007) A new species and first stage associations in Crinitella (Ephemeroptera: Ephemerellidae: Ephemerellinae). Zootaxa, 1611 (1), 45 - 53. https: // doi. org / 10.11646 / zootaxa. 1611.1.3" type="journal article" year="2007">
Jacobus
<emphasis id="A7AFEAC9FFC2D663FE580F4B0B9AFC98" box="[454,504,835,861]" italics="true" pageId="16" pageNumber="467">et al</emphasis>
. (2007)
</bibRefCitation>
. Following
<bibRefCitation id="F14A4B2AFFC2D663FD450F4B0E53FC98" author="Jacobus, L. M. &amp; McCafferty, W. P." box="[731,1073,835,861]" pageId="16" pageNumber="467" pagination="185 - 274" refId="ref14939" refString="Jacobus, L. M. &amp; McCafferty, W. P. (2008) Revision of Ephemerellidae genera (Ephemeroptera). Transactions of the American Entomological Society, 134 (1 - 2), 185 - 274. https: // doi. org / 10.3157 / 0002 - 8320 (2008) 134 [185: ROEGE] 2.0. CO; 2" type="journal article" year="2008">Jacobus &amp; McCafferty (2008)</bibRefCitation>
, we consider
<taxonomicName id="52DB4D58FFC2D663FB530F4B0B2BFC44" authority="Allen &amp; Edmunds, 1963" authorityName="Allen &amp; Edmunds" authorityYear="1963" class="Insecta" family="Ephemerellidae" genus="Crinitella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC2D663FB530F4B0F58FC98" box="[1229,1338,835,861]" italics="true" pageId="16" pageNumber="467">Crinitella</emphasis>
<bibRefCitation id="F14A4B2AFFC2D663FADE0F4B0B2BFC44" author="Allen, R. K. &amp; Edmunds Jr., G. F." pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
</taxonomicName>
as a junior synonym of
<taxonomicName id="52DB4D58FFC2D663FDF80F6F0904FC44" authority="Lestage, 1917" authorityName="Lestage" authorityYear="1917" box="[614,870,871,897]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC2D663FDF80F6F08D9FC44" box="[614,699,871,897]" italics="true" pageId="16" pageNumber="467">Torleya</emphasis>
Lestage, 1917
</taxonomicName>
.
<bibRefCitation id="F14A4B2AFFC2D663FCED0F6F0E32FC44" author="Ogden, T. H. &amp; Osborne, J. T. &amp; Jacobus, L. M. &amp; Whiting, M. F." box="[883,1104,871,897]" pageId="16" pageNumber="467" pagination="28 - 42" refId="ref15591" refString="Ogden, T. H., Osborne, J. T., Jacobus, L. M. &amp; Whiting, M. F. (2009) Combined molecular and morphological phylogeny of Ephemerellinae (Ephemerellidae: Ephemeroptera), with remarks about classification. Zootaxa, 1991 (1), 28 - 42. https: // doi. org / 10.11646 / zootaxa. 1991.1.2" type="journal article" year="2009">
Ogden
<emphasis id="A7AFEAC9FFC2D663FC590F6F0E60FC44" box="[967,1026,871,897]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
(2009
</bibRefCitation>
: figs 1, 2) could not confirm monophyly of
<taxonomicName id="52DB4D58FFC2D663FEA20F830BCBFC60" authorityName="Allen &amp; Edmunds" authorityYear="1963" box="[316,425,907,933]" class="Insecta" family="Ephemerellidae" genus="Crinitella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC2D663FEA20F830BCBFC60" box="[316,425,907,933]" italics="true" pageId="16" pageNumber="467">Crinitella</emphasis>
</taxonomicName>
+
<taxonomicName id="52DB4D58FFC2D663FE590F83087EFC60" authorityName="Lestage" authorityYear="1917" box="[455,540,907,933]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC2D663FE590F83087EFC60" box="[455,540,907,933]" italics="true" pageId="16" pageNumber="467">Torleya</emphasis>
</taxonomicName>
, however, based on their partial sampling of these genus groups.
</paragraph>
<paragraph id="956436DBFFC2D663FF590FA70ED4FBD4" blockId="16.[151,1437,151,1042]" pageId="16" pageNumber="467">
The other Indomalayan species of
<taxonomicName id="52DB4D58FFC2D663FDCB0FA708C8FC0C" authorityName="Lestage" authorityYear="1917" box="[597,682,943,969]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="A7AFEAC9FFC2D663FDCB0FA708C8FC0C" box="[597,682,943,969]" italics="true" pageId="16" pageNumber="467">Torleya</emphasis>
</taxonomicName>
include
<taxonomicName id="52DB4D58FFC2D663FC900FA709C3FC0C" authority="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus, 2021" authorityName="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus" authorityYear="2021" box="[782,929,943,969]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="dibruensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC2D663FC900FA709C3FC0C" box="[782,929,943,969]" italics="true" pageId="16" pageNumber="467">T. dibruensis</emphasis>
</taxonomicName>
<emphasis id="A7AFEAC9FFC2D663FC340FA70E64FC0C" bold="true" box="[938,1030,943,969]" pageId="16" pageNumber="467">
<taxonomicNameLabel id="BC9C57B2FFC2D663FC340FA70E64FC0C" box="[938,1030,943,969]" pageId="16" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="52DB4D58FFC2D663FB880FA70B42FC28" authority="(Jacobus, McCafferty &amp; Sites, 2007)" baseAuthorityName="Jacobus, McCafferty &amp; Sites" baseAuthorityYear="2007" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="lacuna">
<emphasis id="A7AFEAC9FFC2D663FB880FA70E1DFC0C" box="[1046,1151,943,969]" italics="true" pageId="16" pageNumber="467">T. lacuna</emphasis>
(
<bibRefCitation id="F14A4B2AFFC2D663FB0F0FA70B7AFC28" author="Jacobus, L. M. &amp; McCafferty, W. P. &amp; Sites, R. W." pageId="16" pageNumber="467" pagination="45 - 53" refId="ref15040" refString="Jacobus, L. M., McCafferty, W. P. &amp; Sites, R. W. (2007) A new species and first stage associations in Crinitella (Ephemeroptera: Ephemerellidae: Ephemerellinae). Zootaxa, 1611 (1), 45 - 53. https: // doi. org / 10.11646 / zootaxa. 1611.1.3" type="journal article" year="2007">Jacobus, McCafferty &amp; Sites, 2007</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="52DB4D58FFC2D663FEB50FDB0883FC28" authority="(Gui, Zhou &amp; Su, 1999)" baseAuthorityName="Gui, Zhou &amp; Su" baseAuthorityYear="1999" box="[299,737,979,1005]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="longforceps">
<emphasis id="A7AFEAC9FFC2D663FEB50FDB0BA8FC28" box="[299,458,979,1005]" italics="true" pageId="16" pageNumber="467">T. longforceps</emphasis>
(
<bibRefCitation id="F14A4B2AFFC2D663FE470FDB08B8FC28" author="Gui, H. &amp; Zhou, C. - F. &amp; Su, C. - R." box="[473,730,979,1005]" pageId="16" pageNumber="467" refId="ref14572" refString="Gui, H., Zhou, C. - F. &amp; Su, C. - R. (1999) Ephemeroptera. In: Huang, B. - K. (Ed.), Fujian Insect Fauna. Vol. 1. Fujian Science &amp; Technology Press, Fuzhou, pp. 324 - 346." type="book" year="1999">Gui, Zhou &amp; Su, 1999</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="52DB4D58FFC2D663FD730FDB0E55FC28" authority="Kang &amp; Yang, 1995" authorityName="Kang &amp; Yang" authorityYear="1995" box="[749,1079,978,1005]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="lutosa">
<emphasis id="A7AFEAC9FFC2D663FD730FDB092FFC28" box="[749,845,979,1005]" italics="true" pageId="16" pageNumber="467">T. lutosa</emphasis>
<bibRefCitation id="F14A4B2AFFC2D663FCCB0FDB0E55FC28" author="Kang, S. C. &amp; Yang, C. T." box="[853,1079,978,1005]" pageId="16" pageNumber="467" pagination="95 - 116" refId="ref15298" refString="Kang, S. C. &amp; Yang, C. T. (1995) Ephemerellidae of Taiwan (Insecta, Ephemeroptera). Bulletin of National Museum of Natural Science, 5, 95 - 116." type="journal article" year="1995">Kang &amp; Yang, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="52DB4D58FFC2D663FBDD0FDB0FFEFC28" authority="Jacobus &amp; McCafferty" authorityName="Jacobus &amp; McCafferty" authorityYear="2004" box="[1091,1436,979,1005]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="naga">
<emphasis id="A7AFEAC9FFC2D663FBDD0FDB0EF4FC29" box="[1091,1174,979,1004]" italics="true" pageId="16" pageNumber="467">T. naga</emphasis>
Jacobus &amp; McCafferty
</taxonomicName>
(in Jacobus
<emphasis id="A7AFEAC9FFC2D663FE830FFF0B37FBD4" box="[285,341,1015,1041]" italics="true" pageId="16" pageNumber="467">et al.</emphasis>
), 2004,
<taxonomicName id="52DB4D58FFC2D663FE2C0FFF0938FBD7" authority="(Allen &amp; Edmunds, 1963)" authorityName="" baseAuthorityName="Allen &amp; Edmunds" baseAuthorityYear="1963" box="[434,858,1015,1042]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="nepalica">
<emphasis id="A7AFEAC9FFC2D663FE2C0FFF084CFBD4" box="[434,558,1015,1041]" italics="true" pageId="16" pageNumber="467">T. nepalica</emphasis>
(
<bibRefCitation id="F14A4B2AFFC2D663FDA30FFF0933FBD7" author="Allen, R. K. &amp; Edmunds Jr., G. F." box="[573,849,1015,1042]" pageId="16" pageNumber="467" pagination="11 - 22" refId="ref14173" refString="Allen, R. K. &amp; Edmunds Jr., G. F. (1963) New and little known Ephemerellidae from southern Asia, Africa and Madagascar. Pacific Insects, 5 (1), 11 - 22." type="journal article" year="1963">Allen &amp; Edmunds, 1963</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="52DB4D58FFC2D663FC0E0FFF0E36FBD4" authority="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus, 2021" authorityName="Martynov &amp; Selvakumar &amp; Subramanian &amp; Sivaramakrishnan &amp; Vasanth &amp; Sinha &amp; Jacobus" authorityYear="2021" box="[912,1108,1015,1041]" class="Insecta" family="Ephemerellidae" genus="Torleya" kingdom="Animalia" order="Ephemeroptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="simbalbarensis" status="sp. nov.">
<emphasis id="A7AFEAC9FFC2D663FC0E0FFF0E36FBD4" box="[912,1108,1015,1041]" italics="true" pageId="16" pageNumber="467">T. simbalbarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BC9C57B2FFC2D663FBC50FFF0ED4FBD4" box="[1115,1206,1015,1041]" pageId="16" pageNumber="467" rank="species">
<emphasis id="A7AFEAC9FFC2D663FBC50FFF0ECDFBD4" bold="true" box="[1115,1199,1015,1041]" pageId="16" pageNumber="467">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>