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<document id="DDCDC779252F55B0566F4F4CD4AF267A" ID-CLB-Dataset="24935" ID-DOI="10.11646/zootaxa.4695.3.1" ID-GBIF-Dataset="256591b2-6d7b-465d-a818-9a158017ca09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3532515" ID-ZooBank="32E17629-F29A-47C8-ACCB-69A32C17B147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573197328991" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03ED0167FFB5FF8CFF1DF234FEE2FEEB" docLanguage="en" docName="zootaxa.4695.3.1.pdf" docOrigin="Zootaxa 4695 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus reticulatus Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="715F686B-32B6-49D6-8376-9ADD10996BD2" docUuidSource="ZooBank" docVersion="8" lastPageNumber="249" masterDocId="FFD4791FFF85FFBDFF8AF379FF9DFFA8" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups" masterLastPageNumber="252" masterPageNumber="201" pageNumber="249" updateTime="1698764404900" updateUser="ExternalLinkService">
<mods:mods id="70F6705DCA4F1B083010FCA1BE1FE3F9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9C279C86FCA61C51CA48EE6F2A0DEC40">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups</mods:title>
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<mods:name id="70E56A5C2A08F02BEF75944D550D2766" type="personal">
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<mods:namePart id="C217F092FDE8D07D2A39D870A4392B73">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="DAF778AF98EBD128D1CA21A6B3F59BD4">Weir, Thomas A.</mods:namePart>
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<mods:title id="08C1909EFE1FEC7FBE47211B04A7B4FC">Zootaxa</mods:title>
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<mods:part id="5ED45722A21F3BA3315105D7FBE037F1">
<mods:date id="9F40AB28F1C77560653283772C283344">2019</mods:date>
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<mods:number id="34D37A107F282AAC2F6F9358B322F38F">2019-11-08</mods:number>
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<mods:number id="B221533C61ABE7C106A5F7208D3E620B">4695</mods:number>
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<mods:classification id="FC4C1BE3DEDF4817DAFE8666CD2AF33B">journal article</mods:classification>
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<mods:identifier id="9974FB9CAA8D0F185EC68534A55D8688" type="DOI">10.11646/zootaxa.4695.3.1</mods:identifier>
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<mods:identifier id="C804F1BD28A2D1EEFA89BC3CECDF5019" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="89175AE2A43E601B7FF1134BF9478DCD" type="Zenodo-Dep">3532515</mods:identifier>
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<treatment id="03ED0167FFB5FF8CFF1DF234FEE2FEEB" ID-GBIF-Taxon="160217993" LSID="urn:lsid:zoobank.org:act:715F686B-32B6-49D6-8376-9ADD10996BD2" httpUri="http://treatment.plazi.org/id/03ED0167FFB5FF8CFF1DF234FEE2FEEB" lastPageId="49" lastPageNumber="249" pageId="48" pageNumber="249">
<subSubSection id="C35EE3FAFFB5FF8DFF1DF234FD77FECF" box="[151,746,333,360]" pageId="48" pageNumber="249" type="nomenclature">
<paragraph id="8BFBB071FFB5FF8DFF1DF234FD77FECF" blockId="48.[151,746,333,395]" box="[151,746,333,360]" pageId="48" pageNumber="249">
<heading id="D0B3071DFFB5FF8DFF1DF234FD77FECF" bold="true" box="[151,746,333,360]" fontSize="11" level="1" pageId="48" pageNumber="249" reason="1">
<emphasis id="B9306C63FFB5FF8DFF1DF234FD77FECF" bold="true" box="[151,746,333,360]" pageId="48" pageNumber="249">
<taxonomicName id="4C44CBF2FFB5FF8DFF1DF234FDD0FECF" ID-CoL="3T8VM" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,589,333,360]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="reticulatus" status="sp. nov.">
<emphasis id="B9306C63FFB5FF8DFF1DF234FE15FEC0" bold="true" box="[151,392,333,360]" italics="true" pageId="48" pageNumber="249">Lepanus reticulatus</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FFB5FF8DFDD0F234FD77FECF" box="[602,746,333,359]" pageId="48" pageNumber="249" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8DFF1DF208FE5CFE23" box="[151,449,369,395]" pageId="48" pageNumber="249" type="description">
<paragraph id="8BFBB071FFB5FF8DFF1DF208FE5CFE23" blockId="48.[151,746,333,395]" box="[151,449,369,395]" pageId="48" pageNumber="249">
(
<figureCitation id="137FACF4FFB5FF8DFF15F208FE99FE23" box="[159,260,369,395]" captionStart="FIGURE 16" captionStartId="39.[151,250,1781,1806]" captionTargetBox="[177,1410,166,1762]" captionTargetId="figure@39.[177,1410,166,1762]" captionTargetPageId="39" captionText="FIGURE 16. Some diagnostic characters of the Lepanus villosus species group. A, lateral view of elytron of Lepanus villosus, white arrow points to deep pit at the base of stria 8; B, lateral view of elytron of Lepanus vestitus, note the absence of any deep pit; C, elytron of Lepanus villosus dotted line showing humeral callus between striae 6 and 7 and white arrow showing depression at base of striae 4 and 5; D, male metatibiae of Lepanus vestitus, notch on inner edge; E, male protibia of Lepanus pseudovillosus, showing typical configuration for the species group including the small comb of setae (grey arrow); F, close up of elytral interval surface of Lepanus reticulatus, showing reticulation of short longitudinal striations; G, close up of elytral interval surface of Lepanus vestitus showing reticulations of fine meshes." figureDoi="http://doi.org/10.5281/zenodo.3532547" httpUri="https://zenodo.org/record/3532547/files/figure.png" pageId="48" pageNumber="249">Figs 16F</figureCitation>
,
<figureCitation id="137FACF4FFB5FF8DFE85F208FEDDFE23" box="[271,320,369,395]" captionStart="FIGURE 17" captionStartId="40.[151,250,1855,1880]" captionTargetBox="[155,1434,184,1811]" captionTargetId="figure@40.[151,1434,174,1836]" captionTargetPageId="40" captionText="FIGURE 17. Habitus of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532549" httpUri="https://zenodo.org/record/3532549/files/figure.png" pageId="48" pageNumber="249">17B</figureCitation>
,
<figureCitation id="137FACF4FFB5FF8DFEC6F208FEE0FE23" box="[332,381,369,395]" captionStart="FIGURE 18" captionStartId="41.[151,250,1897,1922]" captionTargetBox="[153,1430,180,1864]" captionTargetId="figure@41.[151,1436,166,1876]" captionTargetPageId="41" captionText="FIGURE 18. Left, dorsal, and right view of aedeagi of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532551" httpUri="https://zenodo.org/record/3532551/files/figure.png" pageId="48" pageNumber="249">18B</figureCitation>
,
<figureCitation id="137FACF4FFB5FF8DFE02F208FE25FE23" box="[392,440,369,395]" captionStart="FIGURE 19" captionStartId="42.[151,250,859,884]" captionTargetBox="[159,1427,184,840]" captionTargetId="figure@42.[151,1436,174,840]" captionTargetPageId="42" captionText="FIGURE 19. Pygidial configuration of species within the Lepanus villosus species group. A, Lepanus pseudovillosus; B, Lepanus reticulatus; C, Lepanus vestitus; D, Lepanus villosus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532553" httpUri="https://zenodo.org/record/3532553/files/figure.png" pageId="48" pageNumber="249">19B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8DFF1DF2C0FF6CFD03" pageId="48" pageNumber="249" type="materials_examined">
<paragraph id="8BFBB071FFB5FF8DFF1DF2C0FF6CFD03" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF1DF2C0FEB8FE7B" bold="true" box="[151,293,440,467]" pageId="48" pageNumber="249">Type series.</emphasis>
<materialsCitation id="3B2CBA2CFFB5FF8DFEA5F2C0FEB0FDB3" ID-GBIF-Occurrence="2448757379" collectingDate="1976-04-29" collectingDateMax="1976-05-02" collectingDateMin="1976-04-29" collectingMethod="Hu- man Dung Trap" collectionCode="ANIC" collectorName="R. I. Storey" latitude="-16.583334" location="Mt. Lewis Rd." municipality="Julatten" pageId="48" pageNumber="249" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9306C63FFB5FF8DFEA5F2C0FE38FE7B" bold="true" box="[303,421,441,467]" pageId="48" pageNumber="249">
<typeStatus id="54FF0ED3FFB5FF8DFEA5F2C0FE3DFE7B" box="[303,416,441,467]" pageId="48" pageNumber="249" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="9D427BF8FFB5FF8DFE25F2C0FE59FE7B" box="[431,452,441,467]" pageId="48" pageNumber="249" type="male">♂</specimenCount>
“
<quantity id="4CBC1D94FFB5FF8DFE53F2C0FDB9FE7B" box="[473,548,441,467]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="48" pageNumber="249" unit="km" value="13.0">
<locationDeviation id="9A49A953FFB5FF8DFE53F2C0FDB9FE7B" box="[473,548,441,467]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="48" pageNumber="249" unit="km" value="13.0">13 km</locationDeviation>
</quantity>
up
<location id="8E9BE6AAFFB5FF8DFDD9F2C0FD66FE7B" LSID="urn:lsid:plazi:treatment:03ED0167FFB5FF8CFF1DF234FEE2FEEB:8E9BE6AAFFB5FF8DFDD9F2C0FD66FE7B" box="[595,763,441,467]" latitude="-16.583334" municipality="Julatten" name="Mt. Lewis Rd." pageId="48" pageNumber="249">Mt. Lewis Rd.</location>
via
<collectingMunicipality id="6B9F2A0BFFB5FF8DFCBAF2C0FC17FE7B" box="[816,906,441,467]" pageId="48" pageNumber="249">Julatten</collectingMunicipality>
, QLD.
<date id="FFFA96B1FFB5FF8DFC6AF2C0FB4AFE7B" box="[992,1239,441,467]" pageId="48" pageNumber="249" value="1976-04-29" valueMax="1976-05-02" valueMin="1976-04-29">
<collectingDate id="EFBE6F59FFB5FF8DFC6AF2C0FB4AFE7B" box="[992,1239,441,467]" pageId="48" pageNumber="249" value="1976-04-29" valueMax="1976-05-02" valueMin="1976-04-29">Apr.29- May 2, 1976</collectingDate>
</date>
,
<collectorName id="26B1D5A7FFB5FF8DFB6FF2C0FAFEFE7B" box="[1253,1379,441,467]" pageId="48" pageNumber="249">R.I. Storey</collectorName>
,
<collectingMethod id="5205C866FFB5FF8DFAFBF2C0FED3FE5F" pageId="48" pageNumber="249">Hu- man Dung Trap</collectingMethod>
” / “
<collectionCode id="ED5528B4FFB5FF8DFEF1F2A4FE5FFE5F" box="[379,450,477,503]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="48" pageNumber="249">ANIC</collectionCode>
Database No. 25-054120” / “
<taxonomicName id="4C44CBF2FFB5FF8DFC9DF2A4FCE4FE5F" authorityName="Balthasar" authorityYear="1966" box="[791,889,477,503]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NQ7, det TA
<bibRefCitation id="EFD5CD80FFB5FF8DFB94F2A4FB05FE5F" author="Weir, T. A. &amp; Monteith, G. B." box="[1054,1176,477,503]" pageId="48" pageNumber="249" pagination="235 - 241" refId="ref56456" refString="Weir, T. A. &amp; Monteith, G. B. (2010) Lepanus storeyi, a new species of dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) from Southeast Queensland, The Australian Entomologist, 36, 235 - 241." type="journal article" year="2010">Weir 2010</bibRefCitation>
” in
<collectionCode id="ED5528B4FFB5FF8DFB42F2A4FA91FE5F" box="[1224,1292,477,503]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="48" pageNumber="249">ANIC</collectionCode>
(
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,
<geoCoordinate id="EE70D6B6FFB5FF8DFF1DF179FEB9FDB3" box="[151,292,512,539]" pageId="48" pageNumber="249">145°17”0”E</geoCoordinate>
)
</materialsCitation>
.
<emphasis id="B9306C63FFB5FF8DFEB3F178FE62FDB3" bold="true" box="[313,511,512,539]" pageId="48" pageNumber="249">
<typeStatus id="54FF0ED3FFB5FF8DFEB3F178FE2CFDB3" box="[313,433,513,539]" pageId="48" pageNumber="249" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D427BF8FFB5FF8DFE4BF179FE70FDB3" box="[449,493,512,539]" pageId="48" pageNumber="249" type="female">3 ♀</specimenCount>
):
</emphasis>
<materialsCitation id="3B2CBA2CFFB5FF8DFD8DF178FC31FD97" ID-GBIF-Occurrence="2448756380" collectingDate="1976-04-29" collectingDateMax="1976-05-02" collectingDateMin="1976-04-29" collectionCode="ANIC" collectorName="R. I. Storey" latitude="-16.583334" location="Julatten" longLatPrecision="21" longitude="145.28334" municipality="Mount Lewis Road" pageId="48" pageNumber="249" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<quantity id="4CBC1D94FFB5FF8DFD8DF178FDD2FDB3" box="[519,591,513,539]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="48" pageNumber="249" unit="km" value="13.0">13 km</quantity>
up
<collectingMunicipality id="6B9F2A0BFFB5FF8DFDF0F178FCC9FDB3" box="[634,852,513,539]" pageId="48" pageNumber="249">Mount Lewis Road</collectingMunicipality>
via
<location id="8E9BE6AAFFB5FF8DFC0CF178FC7DFDB3" LSID="urn:lsid:plazi:treatment:03ED0167FFB5FF8CFF1DF234FEE2FEEB:8E9BE6AAFFB5FF8DFC0CF178FC7DFDB3" box="[902,992,513,539]" latitude="-16.583334" longLatPrecision="21" longitude="145.28334" municipality="Mount Lewis Road" name="Julatten" pageId="48" pageNumber="249" stateProvince="Queensland">Julatten</location>
,
<collectingRegion id="49807E93FFB5FF8DFC66F178FBE9FDB3" box="[1004,1140,513,539]" country="Australia" name="Queensland" pageId="48" pageNumber="249">Queensland</collectingRegion>
. [
<geoCoordinate id="EE70D6B6FFB5FF8DFB0CF179FB62FDB3" box="[1158,1279,512,539]" degrees="16" direction="south" minutes="35" orientation="latitude" pageId="48" pageNumber="249" precision="15" seconds="0" value="-16.583334">16°35’0”S</geoCoordinate>
,
<geoCoordinate id="EE70D6B6FFB5FF8DFA8EF179FA0DFDB3" box="[1284,1424,512,539]" degrees="145" direction="east" minutes="17" orientation="longitude" pageId="48" pageNumber="249" precision="15" seconds="0" value="145.28334">145°17’0”E</geoCoordinate>
],
<date id="FFFA96B1FFB5FF8DFF1DF15CFEDBFD97" box="[151,326,549,575]" pageId="48" pageNumber="249" value="1976-04-29" valueMax="1976-05-02" valueMin="1976-04-29">
<collectingDate id="EFBE6F59FFB5FF8DFF1DF15CFEDBFD97" box="[151,326,549,575]" pageId="48" pageNumber="249" value="1976-04-29" valueMax="1976-05-02" valueMin="1976-04-29">29.iv– 2.v.1976</collectingDate>
</date>
,
<collectorName id="26B1D5A7FFB5FF8DFED8F15CFE51FD97" box="[338,460,549,575]" pageId="48" pageNumber="249">R.I. Storey</collectorName>
,
<collectingMethod id="5205C866FFB5FF8DFE5DF15CFD09FD97" box="[471,660,549,575]" pageId="48" pageNumber="249">human dung trap</collectingMethod>
(25-065535 in
<collectionCode id="ED5528B4FFB5FF8DFCB7F15CFC1CFD97" box="[829,897,549,575]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="48" pageNumber="249">ANIC</collectionCode>
<specimenCount id="9D427BF8FFB5FF8DFC02F15CFC00FD97" box="[904,925,549,575]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B2CBA2CFFB5FF8DFC36F15CFB29FDCB" ID-GBIF-Occurrence="2448756379" collectingDate="2008-02-14" collectingDateMax="2008-03-17" collectingDateMin="2008-02-14" collectionCode="QM" collectorName="M. Aristophanous" elevation="1000" latitude="-16.555832" location="Pit" longLatPrecision="21" longitude="145.27834" municipality="Mount Lewis Road" pageId="48" pageNumber="249" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<collectingMunicipality id="6B9F2A0BFFB5FF8DFC36F15CFB0BFD97" box="[956,1174,549,575]" pageId="48" pageNumber="249">Mount Lewis Road</collectingMunicipality>
,
<quantity id="4CBC1D94FFB5FF8DFB2AF15CFB6AFD97" box="[1184,1271,549,575]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="249" unit="m" value="1000.0">
<elevation id="00695742FFB5FF8DFB2AF15CFB6AFD97" box="[1184,1271,549,575]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="249" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, CUDAZ
<location id="8E9BE6AAFFB5FF8DFAE3F15CFA15FD97" LSID="urn:lsid:plazi:treatment:03ED0167FFB5FF8CFF1DF234FEE2FEEB:8E9BE6AAFFB5FF8DFAE3F15CFA15FD97" box="[1385,1416,549,575]" latitude="-16.555832" longLatPrecision="21" longitude="145.27834" municipality="Mount Lewis Road" name="Pit" pageId="48" pageNumber="249" stateProvince="Queensland">Pit</location>
3 [
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,
<geoCoordinate id="EE70D6B6FFB5FF8DFEA2F131FE59FDCB" box="[296,452,584,611]" degrees="145" direction="east" minutes="16" orientation="longitude" pageId="48" pageNumber="249" precision="15" seconds="42" value="145.27834">145°16’42”E</geoCoordinate>
],
<date id="FFFA96B1FFB5FF8DFE5CF130FD0FFDCB" box="[470,658,585,611]" pageId="48" pageNumber="249" value="2008-02-14" valueMax="2008-03-17" valueMin="2008-02-14">
<collectingDate id="EFBE6F59FFB5FF8DFE5CF130FD0FFDCB" box="[470,658,585,611]" pageId="48" pageNumber="249" value="2008-02-14" valueMax="2008-03-17" valueMin="2008-02-14">14.ii–17.iii.2008</collectingDate>
</date>
,
<collectorName id="26B1D5A7FFB5FF8DFD17F130FCFAFDCB" box="[669,871,585,611]" pageId="48" pageNumber="249">M. Aristophanous</collectorName>
, ID 2637 (T
<quantity id="4CBC1D94FFB5FF8DFC61F130FBC1FDCB" box="[1003,1116,585,611]" metricMagnitude="3" metricUnit="m" metricValue="4.2935906" pageId="48" pageNumber="249" unit="in" value="169039.0">169039 in</quantity>
<collectionCode id="ED5528B4FFB5FF8DFBEBF130FB13FDCB" box="[1121,1166,585,611]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="48" pageNumber="249" type="Museum">QM</collectionCode>
<specimenCount id="9D427BF8FFB5FF8DFB1EF130FB35FDCB" box="[1172,1192,585,611]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B2CBA2CFFB5FF8DFB4AF130FF76FD03" ID-GBIF-Occurrence="2448756378" collectingDate="2009-01-03" collectingDateMax="2009-01-30" collectingDateMin="2009-01-03" collectionCode="QM" collectorName="M. Aristophanous &amp; Trip" elevation="1000" latitude="-16.555832" location="Pit" longLatPrecision="21" longitude="145.27834" municipality="Mount Lewis Road" pageId="48" pageNumber="249" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
<collectingMunicipality id="6B9F2A0BFFB5FF8DFB4AF130FA05FDCB" box="[1216,1432,585,611]" pageId="48" pageNumber="249">Mount Lewis Road</collectingMunicipality>
,
<quantity id="4CBC1D94FFB5FF8DFF1DF114FF72FD2F" box="[151,239,621,647]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="249" unit="m" value="1000.0">
<elevation id="00695742FFB5FF8DFF1DF114FF72FD2F" box="[151,239,621,647]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="249" unit="m" value="1000.0">1000 m</elevation>
</quantity>
, CUDAZ
<location id="8E9BE6AAFFB5FF8DFEE9F114FE1FFD2F" LSID="urn:lsid:plazi:treatment:03ED0167FFB5FF8CFF1DF234FEE2FEEB:8E9BE6AAFFB5FF8DFEE9F114FE1FFD2F" box="[355,386,621,647]" latitude="-16.555832" longLatPrecision="21" longitude="145.27834" municipality="Mount Lewis Road" name="Pit" pageId="48" pageNumber="249" stateProvince="Queensland">Pit</location>
4 [
<geoCoordinate id="EE70D6B6FFB5FF8DFE2CF115FDB6FD2F" box="[422,555,620,647]" degrees="16" direction="south" minutes="33" orientation="latitude" pageId="48" pageNumber="249" precision="15" seconds="21" value="-16.555832">16°33’21”S</geoCoordinate>
,
<geoCoordinate id="EE70D6B6FFB5FF8DFDBAF115FD51FD2F" box="[560,716,620,647]" degrees="145" direction="east" minutes="16" orientation="longitude" pageId="48" pageNumber="249" precision="15" seconds="42" value="145.27834">145°16’42”E</geoCoordinate>
],
<date id="FFFA96B1FFB5FF8DFD6AF114FCFAFD2F" box="[736,871,621,647]" pageId="48" pageNumber="249" value="2009-01-03" valueMax="2009-01-30" valueMin="2009-01-03">
<collectingDate id="EFBE6F59FFB5FF8DFD6AF114FCFAFD2F" box="[736,871,621,647]" pageId="48" pageNumber="249" value="2009-01-03" valueMax="2009-01-30" valueMin="2009-01-03">3–30.i.2009</collectingDate>
</date>
,
<collectorName id="26B1D5A7FFB5FF8DFCFEF114FBDDFD2F" box="[884,1088,621,647]" pageId="48" pageNumber="249">M. Aristophanous</collectorName>
, ID 8284,
<collectorName id="26B1D5A7FFB5FF8DFB3CF114FB78FD2F" box="[1206,1253,621,647]" pageId="48" pageNumber="249">Trip</collectorName>
20 (T
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<collectionCode id="ED5528B4FFB5FF8DFF1DF1E8FF59FD03" box="[151,196,657,683]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="48" pageNumber="249" type="Museum">QM</collectionCode>
<specimenCount id="9D427BF8FFB5FF8DFF41F1E8FF42FD03" box="[203,223,657,683]" pageId="48" pageNumber="249" type="female">♀</specimenCount>
)
</materialsCitation>
;
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8DFF4DF1CDFDC2F987" pageId="48" pageNumber="249" type="description">
<paragraph id="8BFBB071FFB5FF8DFF4DF1CDFB5BFD67" blockId="48.[151,1437,440,2015]" box="[199,1222,692,719]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF1CDFEC4FD67" bold="true" box="[199,345,692,719]" pageId="48" pageNumber="249">Description.</emphasis>
Ranging in colour from light brown to black. Antennal clubs yellowish white.
</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF1A0FB27FD5B" blockId="48.[151,1437,440,2015]" box="[199,1210,728,755]" pageId="48" pageNumber="249">
Total length:
<quantity id="4CBC1D94FFB5FF8DFED6F1A1FE28FD5B" box="[348,437,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="48" pageNumber="249" unit="mm" value="3.5">3.5 mm</quantity>
.
<typeStatus id="54FF0ED3FFB5FF8DFE4AF1A0FDB5FD5B" box="[448,552,729,755]" pageId="48" pageNumber="249" type="holotype">Holotype</typeStatus>
measurements: total length
<quantity id="4CBC1D94FFB5FF8DFCEEF1A1FC21FD5B" box="[868,956,728,755]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="48" pageNumber="249" unit="mm" value="3.5">3.5 mm</quantity>
, elytral width
<quantity id="4CBC1D94FFB5FF8DFBD7F1A0FB2BFD5B" box="[1117,1206,729,755]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="48" pageNumber="249" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF184FD6AFC2B" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF184FE9CFCBF" box="[199,257,765,791]" italics="true" pageId="48" pageNumber="249">Male</emphasis>
. Head. Clypeal surface anteriorly smooth, nitid and punctate, with rest of the head reticulate with dense ocellate punctures each bearing a seta. Broad U-shaped between clypeal teeth that are long and upturned. Genal angle weakly defined. Dorsal part of the eyes narrow, separated by an interocular space approximately 12 times the eye width (48: 4). Head 1.4 x wider than long (68:50).
</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF0F4FDEAFB90" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">Prothorax: Pronotal surface with moderate ocellate punctation anteriorly to heavy ocellate punctation posteriorly, each puncture bearing a posteriorly directed seta. Punctures along the basal margin similar to punctures on hind part of disc. Anterior angles about 90°. Basal margin rounded. Hypomeral surface punctured and reticulate. Hypomeral stria about half the length of hypomeron, with a few punctures between hypomeral stria and lateral edge. Pronotum 1.65 x wider than long (110:66).</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF738FD55FB0B" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">Elytra: Intervals heavily reticulate with dense, short, longitudinally aligned striae and a double row of recurved setae. Striae distinctly geminate. Stria 7 extending or almost extending to elytral base. Stria 8 not ending basally in a deep pit. Elytra 1.1 x wider than long (123:112).</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF7D4FAA0FB43" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">Legs: Protibiae with the 3 teeth not equidistant, apical 2 slightly closer together. Apical digit of protibiae short, with a relatively long spur and small brush of setae at inner apex. Metatibiae normal, not bent or indented.</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF78CFA9BFA9B" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">Abdomen: Pygidium somewhat convex, reticulate, heavily punctate, setate and with an indication of a raised longitudinal midline and with an overhanging basal edge. Abdominal ventrites reticulate right across.</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF644FF63FA6B" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">Pterothorax: Medial lobe of metaventrite punctate and narrowly margined between the mesocoxae. Lateral lobe of metaventrite with ocellate punctures. Meso-metaventral suture slightly arched, with a slight angulation medially. Mesoventrite surface smooth and punctate. Mesepimeron reticulate and punctate. Metanepisternum reticulate and punctate.</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF6B4FE23FA4F" blockId="48.[151,1437,440,2015]" box="[199,446,1485,1511]" pageId="48" pageNumber="249">Hind wings: vestigial.</paragraph>
<paragraph id="8BFBB071FFB5FF8DFF4DF688FDC2F987" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF688FE86F9A3" box="[199,283,1521,1547]" italics="true" pageId="48" pageNumber="249">Female</emphasis>
. Clypeal surface wholly reticulate and protibiae with a small comb of stout setae at inner apex, otherwise as per the
<taxonomicName id="4C44CBF2FFB5FF8DFECEF56FFE2AF987" box="[324,439,1557,1583]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B9306C63FFB5FF8DFECEF56FFE2AF987" box="[324,439,1557,1583]" italics="true" pageId="48" pageNumber="249">L. villosus</emphasis>
</taxonomicName>
species group.
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8DFF4DF540FD7BF9DF" pageId="48" pageNumber="249" type="etymology">
<paragraph id="8BFBB071FFB5FF8DFF4DF540FD7BF9DF" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF540FED3F9FB" bold="true" box="[199,334,1593,1619]" pageId="48" pageNumber="249">Etymology.</emphasis>
Refers to the distinct reticulations covering the body surface, from
<taxonomicName id="4C44CBF2FFB5FF8DFBD5F540FAB2F9FC" authority="(Latin)" authorityName="Latin" baseAuthorityName="Latin" box="[1119,1327,1593,1620]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9306C63FFB5FF8DFBD5F540FB4BF9FB" box="[1119,1238,1593,1619]" italics="true" pageId="48" pageNumber="249">reticulatus</emphasis>
(Latin)
</taxonomicName>
- net-like. This name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8DFF4DF5F9FF63F94B" pageId="48" pageNumber="249" type="distribution">
<paragraph id="8BFBB071FFB5FF8DFF4DF5F9FF63F94B" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF5F9FEFCF933" bold="true" box="[199,353,1664,1691]" pageId="48" pageNumber="249">Distribution.</emphasis>
Known only from four specimens taken on Mount Lewis in the Australian Wet Tropics bioregion (
<figureCitation id="137FACF4FFB5FF8DFF50F5DCFED9F917" box="[218,324,1701,1727]" captionStart="FIGURE 21" captionStartId="50.[151,250,1013,1038]" captionTargetBox="[151,1436,301,994]" captionTargetId="figure@50.[151,1436,301,994]" captionTargetPageId="50" captionText="FIGURE 21. Map of known records for A, Lepanus reticulatus; B, Lepanus vestitus including predicted distribution estimated with BIOCLIM models. QLD, Queensland." figureDoi="http://doi.org/10.5281/zenodo.3532557" httpUri="https://zenodo.org/record/3532557/files/figure.png" pageId="48" pageNumber="249">Fig. 21A</figureCitation>
). These specimens were collected at high elevation (
<quantity id="4CBC1D94FFB5FF8DFC17F5DCFC6BF917" box="[925,1014,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="249" unit="m" value="1000.0">1000 m</quantity>
) and this appears to be a short range endemic.
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFFB5FF8CFF4DF595FEE2FEEB" lastPageId="49" lastPageNumber="250" pageId="48" pageNumber="249" type="discussion">
<paragraph id="8BFBB071FFB5FF8DFF4DF595FB07F883" blockId="48.[151,1437,440,2015]" pageId="48" pageNumber="249">
<emphasis id="B9306C63FFB5FF8DFF4DF595FED2F8AF" bold="true" box="[199,335,1772,1799]" pageId="48" pageNumber="249">Comments.</emphasis>
This species has been collected in pitfall traps including those baited with human faeces. It has been referred to under the informal nomenclature
<emphasis id="B9306C63FFB5FF8DFD4CF46BFC99F883" box="[710,772,1810,1835]" italics="true" pageId="48" pageNumber="249">sensu</emphasis>
<bibRefCitation id="EFD5CD80FFB5FF8DFC80F468FC4CF883" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[778,977,1809,1835]" pageId="48" pageNumber="249" pagination="92 - 99" refId="ref56502" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modeling. In: West, J. (Ed.), 21 st Century Taxonomy: Accelerating Research and Discovery of Australia's Biodiversity, Taxonomy Research and Identification Network Final Report, March, 2011, pp. 92 - 99. Available from: www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 6 September 2019)" type="journal article" year="2011">
Yeates
<emphasis id="B9306C63FFB5FF8DFCD1F46BFC0EF883" box="[859,915,1809,1835]" italics="true" pageId="48" pageNumber="249">et al.</emphasis>
2011
</bibRefCitation>
as
<taxonomicName id="4C44CBF2FFB5FF8DFC7CF46BFBC8F883" authorityName="Balthasar" authorityYear="1966" box="[1014,1109,1810,1835]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B9306C63FFB5FF8DFC7CF46BFBC8F883" box="[1014,1109,1810,1835]" italics="true" pageId="48" pageNumber="249">Lepanus</emphasis>
</taxonomicName>
NQ7.
</paragraph>
<paragraph id="8BFBB071FFB5FF8CFF4DF44CFEE2FEEB" blockId="48.[151,1437,440,2015]" lastBlockId="49.[151,1436,152,323]" lastPageId="49" lastPageNumber="250" pageId="48" pageNumber="249">
The labels of the two
<typeStatus id="54FF0ED3FFB5FF8DFE32F44CFDBEF8E7" box="[440,547,1845,1871]" pageId="48" pageNumber="249" type="paratype">paratypes</typeStatus>
collected at Mount Lewis Road (
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) by M. Aristophanous do not list coordinates, however we report putative geocoded coordinates extropolated from
<bibRefCitation id="EFD5CD80FFB5FF8DFC72F420FB6DF8DB" author="Aristophanous, M." box="[1016,1264,1881,1907]" pageId="48" pageNumber="249" refId="ref55830" refString="Aristophanous, M. (2014) Understanding Patterns Of Endemic Dung Beetle (Coleoptera: Scarabaeidae: Scarabaeinae) Biodiversity In The Australian Wet Tropics Rainforest: Implications Of Climate Change. PhD Thesis, James Cook University, Townsville, Australia. Available from: https: // researchonline. jcu. edu. au / 41143 (accessed 6 September 2019)" type="url" year="2014">Aristophanous (2014)</bibRefCitation>
.
<bibRefCitation id="EFD5CD80FFB5FF8DFB73F420FF7FF83F" author="Aristophanous, M." pageId="48" pageNumber="249" refId="ref55830" refString="Aristophanous, M. (2014) Understanding Patterns Of Endemic Dung Beetle (Coleoptera: Scarabaeidae: Scarabaeinae) Biodiversity In The Australian Wet Tropics Rainforest: Implications Of Climate Change. PhD Thesis, James Cook University, Townsville, Australia. Available from: https: // researchonline. jcu. edu. au / 41143 (accessed 6 September 2019)" type="url" year="2014">Aristophanous (2014)</bibRefCitation>
surveyed dung beetles of elevational transects in the Australian Wet Tropics including seven elevational gradients in Mount Lewis National Park, Carbine Tableland. Specimens from his study were collected between
<date id="FFFA96B1FFB5FF8DFF1DF4BCFE88F877" box="[151,277,1989,2015]" pageId="48" pageNumber="249" value="2007" valueMax="2009">2007–2009</date>
(
<bibRefCitation id="EFD5CD80FFB5FF8DFEAEF4BCFD90F877" author="Aristophanous, M." box="[292,525,1989,2015]" pageId="48" pageNumber="249" refId="ref55830" refString="Aristophanous, M. (2014) Understanding Patterns Of Endemic Dung Beetle (Coleoptera: Scarabaeidae: Scarabaeinae) Biodiversity In The Australian Wet Tropics Rainforest: Implications Of Climate Change. PhD Thesis, James Cook University, Townsville, Australia. Available from: https: // researchonline. jcu. edu. au / 41143 (accessed 6 September 2019)" type="url" year="2014">Aristophanous 2014</bibRefCitation>
). Eight specimens identified as
<taxonomicName id="4C44CBF2FFB5FF8DFCE5F4BFFBB6F877" box="[879,1067,1989,2015]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="249" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B9306C63FFB5FF8DFCE5F4BFFBB6F877" box="[879,1067,1989,2015]" italics="true" pageId="48" pageNumber="249">Lepanus villosus</emphasis>
</taxonomicName>
were collected at Carbine Table- land and the species was only collected at site CU10, 1016 m,
<geoCoordinate id="EE70D6B6FFB4FF8CFCECF3E1FC50FF1B" box="[870,973,152,179]" degrees="16.5558" direction="south" orientation="latitude" pageId="49" pageNumber="250" precision="5" value="-16.5558">-16.5558</geoCoordinate>
,
<geoCoordinate id="EE70D6B6FFB4FF8CFC50F3E1FBDAFF1B" box="[986,1095,152,179]" degrees="145.2784" direction="east" orientation="longitude" pageId="49" pageNumber="250" precision="5" value="145.2784">145.2784</geoCoordinate>
. These coordinates fall on the main northern sector of the Mount Lewis Road. Given the likeness of
<taxonomicName id="4C44CBF2FFB4FF8CFC2DF3C7FB87FF7F" box="[935,1050,189,215]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="villosus">
<emphasis id="B9306C63FFB4FF8CFC2DF3C7FB87FF7F" box="[935,1050,189,215]" italics="true" pageId="49" pageNumber="250">L. villosus</emphasis>
</taxonomicName>
and
<emphasis id="B9306C63FFB4FF8CFBDAF3C7FB76FF7F" box="[1104,1259,189,215]" italics="true" pageId="49" pageNumber="250">
<taxonomicName id="4C44CBF2FFB4FF8CFBDAF3C7FB7BFF7F" box="[1104,1254,189,215]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="250" phylum="Arthropoda" rank="species" species="reticulatus">L. reticulatus</taxonomicName>
,
</emphasis>
the congruence of locality data, collector and collection year, we believe these reported specimens of
<bibRefCitation id="EFD5CD80FFB4FF8CFBC5F398FAD9FF53" author="Aristophanous, M." box="[1103,1348,225,251]" pageId="49" pageNumber="250" refId="ref55830" refString="Aristophanous, M. (2014) Understanding Patterns Of Endemic Dung Beetle (Coleoptera: Scarabaeidae: Scarabaeinae) Biodiversity In The Australian Wet Tropics Rainforest: Implications Of Climate Change. PhD Thesis, James Cook University, Townsville, Australia. Available from: https: // researchonline. jcu. edu. au / 41143 (accessed 6 September 2019)" type="url" year="2014">Aristophanous (2014)</bibRefCitation>
include the
<typeStatus id="54FF0ED3FFB4FF8CFF48F27CFEB0FEB7" box="[194,301,261,287]" pageId="49" pageNumber="250" type="paratype">paratypes</typeStatus>
listed here and as such we list a more detailed geocoded locality than would otherwise be available from the label alone.
</paragraph>
</subSubSection>
</treatment>
</document>
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