353 lines
37 KiB
XML
353 lines
37 KiB
XML
<document id="328F71B6A735E26F73E2ADCD642B54F9" ID-DOI="10.1080/00222933.2010.535915" ID-ISSN="1464-5262" ID-Zenodo-Dep="5203373" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629048099209" checkinUser="felipe" docAuthor="Zacharda, Miloslav, Grafitti, Giuseppe & Piva, Erminio" docDate="2011" docId="0382502C900AFFEAFEDCFEF9E6EE231F" docLanguage="en" docName="JNatHist.45.11-12.667-683.pdf" docOrigin="Journal of Natural History 45 (11 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2010.535915" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Rhagidia (Deharvengiella) paralleloseta Zacharda & Grafitti & Piva 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="672" masterDocId="FFBB2854900EFFEDFFB3FFF6E7592115" masterDocTitle="New taxa of Rhagidia and Foveacheles (Acari: Prostigmata: Rhagidiidae) from Italian and French caves, with keys to adults of subgenera Deharvengiella and Mediostella" masterLastPageNumber="683" masterPageNumber="667" pageNumber="669" updateTime="1699057466393" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="439F5AD9F2EE384BD06BAB4ACECBEB4E">New taxa of Rhagidia and Foveacheles (Acari: Prostigmata: Rhagidiidae) from Italian and French caves, with keys to adults of subgenera Deharvengiella and Mediostella</mods:title>
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</mods:titleInfo>
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<mods:name id="A92BCE1D5413BA84BA0524E992A636CB" type="personal">
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<mods:namePart id="C76BC6CC72027841F3F6BEFE975C0F5F">Zacharda, Miloslav</mods:namePart>
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<mods:roleTerm id="68D4A690BBF449E9DE06F4EA83E83B4E">Author</mods:roleTerm>
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<mods:namePart id="9931A1ABB8AAF11C09FA08C4F6A85920">Grafitti, Giuseppe</mods:namePart>
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<mods:name id="1C86215D9547AB730C3C2C40DDA1488D" type="personal">
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<mods:roleTerm id="2AE564709389D7F71BF497588D8A1F06">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AD9E26FFC5FB56A6E649BECEEA2A1270">Piva, Erminio</mods:namePart>
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<mods:title id="618BE7098560D02E57A854A72E4AB9C1">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="4A9FDA0A15AC2734D3E1B79A376A468F">
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<mods:date id="53C8CCB2BE2961A018B099DBEA576596">2011</mods:date>
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<mods:number id="58C51EC4137C2560E9321158B6DAEA0D">2011-03-07</mods:number>
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<mods:number id="022242A5803D51B6815BE1A7982C4692">45</mods:number>
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<mods:detail id="ADA928D5C2D976652528A6D82CAB7A54" type="issue">
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<mods:number id="89DC38E77801BC17A0C55E89EC9223C6">11 - 12</mods:number>
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<mods:start id="3023108BAE49DD83D3387092FFEE3714">667</mods:start>
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<mods:end id="F9C55D6D118C61BF0FA06420ABA8DF41">683</mods:end>
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<mods:url id="1DE1BE4ACFD37EFEE02A41285963C2B9">http://dx.doi.org/10.1080/00222933.2010.535915</mods:url>
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<mods:identifier id="E0B5AA3AC973DF4A49837B0A97B07944" type="DOI">10.1080/00222933.2010.535915</mods:identifier>
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<mods:identifier id="934BC903086A063E60CDE7EA1EA30D57" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="261F2AE4ED61330D9F79E6EB4DA2B4F4" type="Zenodo-Dep">5203373</mods:identifier>
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<treatment id="0382502C900AFFEAFEDCFEF9E6EE231F" LSID="urn:lsid:plazi:treatment:0382502C900AFFEAFEDCFEF9E6EE231F" httpUri="http://treatment.plazi.org/id/0382502C900AFFEAFEDCFEF9E6EE231F" lastPageId="7" lastPageNumber="672" pageId="4" pageNumber="669">
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<subSubSection id="C331B2B1900AFFE9FEDCFEF9E4A12032" box="[367,1016,271,296]" pageId="4" pageNumber="669" type="nomenclature">
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<paragraph id="8B94E13A900AFFE9FEDCFEF9E4A12032" blockId="4.[367,1016,271,327]" box="[367,1016,271,296]" pageId="4" pageNumber="669">
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<heading id="D0DC5656900AFFE9FEDCFEF9E4A12032" box="[367,1016,271,296]" centered="true" fontSize="36" level="2" pageId="4" pageNumber="669" reason="7">
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<taxonomicName id="4C2B9AB9900AFFE9FEDCFEF9E4C0203D" authority="Zacharda" authorityName="Zacharda & Grafitti & Piva" authorityYear="2011" box="[367,921,271,296]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="669" phylum="Arthropoda" rank="species" species="paralleloseta" status="sp. nov." subGenus="Deharvengiella">
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<emphasis id="B95F3D28900AFFE9FEDCFEF9E47A203D" bold="true" box="[367,803,271,296]" pageId="4" pageNumber="669">
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<emphasis id="B95F3D28900AFFE9FEDCFEF9E68A203D" bold="true" box="[367,467,271,296]" italics="true" pageId="4" pageNumber="669">Rhagidia</emphasis>
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(
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<emphasis id="B95F3D28900AFFE9FE50FEF9E5D1203D" bold="true" box="[483,648,271,296]" italics="true" pageId="4" pageNumber="669">Deharvengiella</emphasis>
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)
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<emphasis id="B95F3D28900AFFE9FD24FEF9E47A203D" bold="true" box="[663,803,271,296]" italics="true" pageId="4" pageNumber="669">paralleloseta</emphasis>
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</emphasis>
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Zacharda
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26C8053900AFFE9FC17FEE6E4A12032" box="[932,1016,272,295]" pageId="4" pageNumber="669" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C331B2B1900AFFEAFDDAFED9E6EE231F" lastPageId="7" lastPageNumber="672" pageId="4" pageNumber="669" type="description">
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<paragraph id="8B94E13A900AFFE9FDDAFED9E5A72052" blockId="4.[367,1016,271,327]" box="[617,766,303,327]" pageId="4" pageNumber="669">
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||
(
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<figureCitation id="1310FDBF900AFFE9FDC3FED9E5822052" box="[624,731,303,327]" captionStart="Figure 1" captionStartId="4.[213,280,1637,1659]" captionTargetBox="[211,1171,599,1589]" captionTargetId="figure-78@4.[211,1171,599,1589]" captionTargetPageId="4" captionText="Figure 1. Rhagidia (D.) paralleloseta sp. nov., adult male: (A) prodorsum; (B) ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203375" httpUri="https://zenodo.org/record/5203375/files/figure.png" pageId="4" pageNumber="669">Figures 1</figureCitation>
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,
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<figureCitation id="1310FDBF900AFFE9FD5BFED9E5AF2052" box="[744,758,303,327]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="4" pageNumber="669">2</figureCitation>
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)
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</paragraph>
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<paragraph id="8B94E13A900AFFE9FF66FE84E6FE209F" blockId="4.[213,423,370,394]" box="[213,423,370,394]" pageId="4" pageNumber="669">
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<emphasis id="B95F3D28900AFFE9FF66FE84E6FE209F" box="[213,423,370,394]" italics="true" pageId="4" pageNumber="669">Material examined</emphasis>
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</paragraph>
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||
<paragraph id="8B94E13A900AFFE9FF66FE6FE6BB2304" blockId="4.[213,1170,409,529]" pageId="4" pageNumber="669">
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<materialsCitation id="3B43EB67900AFFE9FF66FE6FE6BB2304" collectingDate="1984-04-15" collectionCode="OSAL" collectorName="de l'Oule & L. Deharveng & Ohio State University & Type No." country="France" county="Museum of Biological Diversity" elevation="240" location="Adult" municipality="Gouffre de l'Oule" pageId="4" pageNumber="669" specimenCode="OSAL0007414" specimenCount="1" typeStatus="holotype">
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<emphasis id="B95F3D28900AFFE9FF66FE6FE61A20A4" box="[213,323,409,433]" italics="true" pageId="4" pageNumber="669">
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||
<typeStatus id="54905F98900AFFE9FF66FE6FE66720A4" box="[213,318,409,433]" pageId="4" pageNumber="669" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
<location id="8EF4B7E1900AFFE9FEF9FE6FE6D520A4" LSID="urn:lsid:plazi:treatment:0382502C900AFFEAFEDCFEF9E6EE231F:8EF4B7E1900AFFE9FEF9FE6FE6D520A4" box="[330,396,409,433]" country="France" county="Museum of Biological Diversity" municipality="Gouffre de l'Oule" name="Adult" pageId="4" pageNumber="669">Adult</location>
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female,
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<collectingCountry id="F33CA1AA900AFFE9FE5FFE6CE56520A4" box="[492,572,410,433]" name="France" pageId="4" pageNumber="669">France</collectingCountry>
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||
,
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<location id="8EF4B7E1900AFFE9FDFBFE6CE52820A4" LSID="urn:lsid:plazi:treatment:0382502C900AFFEAFEDCFEF9E6EE231F:8EF4B7E1900AFFE9FDFBFE6CE52820A4" box="[584,625,410,433]" country="France" county="Museum of Biological Diversity" municipality="Gouffre de l'Oule" name="Lot" pageId="4" pageNumber="669">Lot</location>
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and
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<location id="8EF4B7E1900AFFE9FD1FFE6FE47920A4" LSID="urn:lsid:plazi:treatment:0382502C900AFFEAFEDCFEF9E6EE231F:8EF4B7E1900AFFE9FD1FFE6FE47920A4" box="[684,800,409,433]" country="France" county="Museum of Biological Diversity" municipality="Gouffre de l'Oule" name="Dordogne" pageId="4" pageNumber="669">Dordogne</location>
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||
,
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||
<collectingMunicipality id="6BF07B40900AFFE9FC9EFE6FE4AE20A4" box="[813,1015,409,433]" pageId="4" pageNumber="669">
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||
Gouffre
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||
<collectorName id="26DE84EC900AFFE9FC3DFE6FE4AE20A4" box="[910,1015,409,433]" pageId="4" pageNumber="669">de l’Oule</collectorName>
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||
</collectingMunicipality>
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||
,
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||
<elevation id="00060609900AFFE9FBB0FE6FE66F20C4" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="669" unit="m" value="240.0">
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||
<quantity id="4CD34CDF900AFFE9FBB0FE6FE31020A4" box="[1027,1097,409,433]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="669" unit="m" value="240.0">240 m</quantity>
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above sea level
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</elevation>
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, coll.
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<collectorName id="26DE84EC900AFFE9FECFFE4CE54520C4" box="[380,540,441,465]" pageId="4" pageNumber="669">L. Deharveng</collectorName>
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||
,
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<date id="FF95C7FA900AFFE9FD9AFE4FE59120C4" box="[553,712,441,465]" pageId="4" pageNumber="669" value="1984-04-15">
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||
<collectingDate id="EFD13E12900AFFE9FD9AFE4FE59120C4" box="[553,712,441,465]" pageId="4" pageNumber="669" value="1984-04-15">15 April 1984</collectingDate>
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</date>
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||
; deposited in the
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<collectingCounty id="62F599B6900AFFE9FC27FE4CE66420E4" pageId="4" pageNumber="669">Museum of Biological Diversity</collectingCounty>
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||
,
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||
<collectorName id="26DE84EC900AFFE9FEFBFE2FE56720E4" box="[328,574,473,497]" pageId="4" pageNumber="669">Ohio State University</collectorName>
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||
,
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||
<collectorName id="26DE84EC900AFFE9FDFBFE2CE5E820E4" box="[584,689,474,498]" pageId="4" pageNumber="669">Type No.</collectorName>
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||
<specimenCode id="DB8D4941900AFFE9FD04FE2FE43820E4" box="[695,865,473,497]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="4" pageNumber="669">OSAL0007414</specimenCode>
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; adult male, the same data, lost during preparation.
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</materialsCitation>
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</paragraph>
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||
<paragraph id="8B94E13A900AFFE9FCE2F9DEE4DB2751" blockId="4.[849,898,1576,1604]" box="[849,898,1576,1604]" pageId="4" pageNumber="669">ad1</paragraph>
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||
<caption id="DF54B1B2900AFFE9FF66F993E645278C" ID-DOI="http://doi.org/10.5281/zenodo.5203375" ID-Zenodo-Dep="5203375" httpUri="https://zenodo.org/record/5203375/files/figure.png" pageId="4" pageNumber="669" startId="4.[213,280,1637,1659]" targetBox="[211,1171,599,1589]" targetPageId="4">
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||
<paragraph id="8B94E13A900AFFE9FF66F993E645278C" blockId="4.[213,1170,1637,1689]" pageId="4" pageNumber="669">
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Figure 1.
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<taxonomicName id="4C2B9AB9900AFFE9FEF5F990E53A2769" authority="Zacharda & Grafitti & Piva, 2011" authorityName="Zacharda & Grafitti & Piva" authorityYear="2011" box="[326,611,1637,1660]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="669" phylum="Arthropoda" rank="species" species="paralleloseta" status="sp. nov." subGenus="Deharvengiella">
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||
<emphasis id="B95F3D28900AFFE9FEF5F990E6F92769" box="[326,416,1638,1660]" italics="true" pageId="4" pageNumber="669">Rhagidia</emphasis>
|
||
(
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||
<emphasis id="B95F3D28900AFFE9FE05F991E6962769" box="[438,463,1639,1660]" italics="true" pageId="4" pageNumber="669">D.</emphasis>
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||
)
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||
<emphasis id="B95F3D28900AFFE9FE56F990E53A2769" box="[485,611,1638,1660]" italics="true" pageId="4" pageNumber="669">paralleloseta</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A26C8053900AFFE9FDC1F990E590276E" box="[626,713,1638,1659]" pageId="4" pageNumber="669" rank="species">sp. nov.</taxonomicNameLabel>
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, adult male: (A) prodorsum; (B) ventral aspect.
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</paragraph>
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||
</caption>
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||
<paragraph id="8B94E13A900BFFE8FE90FBA7E4AB257F" blockId="5.[291,1010,1105,1131]" box="[291,1010,1105,1131]" pageId="5" pageNumber="670">A B C,D,F,G E</paragraph>
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||
<caption id="DF54B1B2900BFFE8FF23FB7AE5B025EC" ID-DOI="http://doi.org/10.5281/zenodo.5203377" ID-Zenodo-Dep="5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="5" pageNumber="670" startId="5.[144,211,1164,1186]" targetBox="[150,1093,161,1094]" targetPageId="5">
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||
<paragraph id="8B94E13A900BFFE8FF23FB7AE5B025EC" blockId="5.[144,1101,1164,1273]" pageId="5" pageNumber="670">
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||
Figure 2.
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||
<taxonomicName id="4C2B9AB9900BFFE8FF4DFB7AE54E25B7" authority="Zacharda & Grafitti & Piva, 2011" authorityName="Zacharda & Grafitti & Piva" authorityYear="2011" box="[254,535,1164,1186]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="670" phylum="Arthropoda" rank="species" species="paralleloseta" status="sp. nov." subGenus="Deharvengiella">
|
||
<emphasis id="B95F3D28900BFFE8FF4DFB7AE60125B7" box="[254,344,1164,1186]" italics="true" pageId="5" pageNumber="670">Rhagidia</emphasis>
|
||
(
|
||
<emphasis id="B95F3D28900BFFE8FEDEFB7BE6DC25B7" box="[365,389,1165,1186]" italics="true" pageId="5" pageNumber="670">D.</emphasis>
|
||
)
|
||
<emphasis id="B95F3D28900BFFE8FE2AFB7AE54E25B7" box="[409,535,1164,1186]" italics="true" pageId="5" pageNumber="670">paralleloseta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C8053900BFFE8FD90FB7AE52125B4" box="[547,632,1164,1185]" pageId="5" pageNumber="670" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion
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||
<emphasis id="B95F3D28900BFFE8FC92FB31E47625C8" box="[801,815,1223,1245]" italics="true" pageId="5" pageNumber="670">Φ</emphasis>
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||
on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect.
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</paragraph>
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</caption>
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<paragraph id="8B94E13A900BFFE8FF23FAC7E7A5245D" blockId="5.[144,252,1329,1352]" box="[144,252,1329,1352]" pageId="5" pageNumber="670">
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||
<emphasis id="B95F3D28900BFFE8FF23FAC7E7A5245D" box="[144,252,1329,1352]" italics="true" pageId="5" pageNumber="670">Diagnosis</emphasis>
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||
</paragraph>
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<paragraph id="8B94E13A900BFFE8FF23FAAEE668271A" blockId="5.[144,1101,1368,1551]" pageId="5" pageNumber="670">
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||
Rhagidial organ I comprises four strikingly long rhagidial solenidia lying parallel in separate depressions, stellate famulus (
|
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<emphasis id="B95F3D28900BFFE8FDDAFA8EE52C2485" box="[617,629,1400,1424]" italics="true" pageId="5" pageNumber="670">ε</emphasis>
|
||
) subtending second rhagidial solenidion; rhagidial organ II with three rhagidial solenidia lying parallel in separate depressions, spiniform famulus (
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||
<emphasis id="B95F3D28900BFFE8FEDCFA4EE62224C5" box="[367,379,1464,1488]" italics="true" pageId="5" pageNumber="670">ε</emphasis>
|
||
) not observed. Coxae I, II, III, IV with 3-1-5-3 finely pubescent setae, respectively. Adoral setae on subcapitulum short, not overlapping apex of subcapitulum.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A900BFFE8FF23F9BBE53E2771" blockId="5.[144,615,1612,1636]" box="[144,615,1612,1636]" pageId="5" pageNumber="670">
|
||
<emphasis id="B95F3D28900BFFE8FF23F9BBE53E2771" box="[144,615,1612,1636]" italics="true" pageId="5" pageNumber="670">Description – adult female (one examined)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B94E13A900BFFEBFF23F982E60D23A9" blockId="5.[144,984,1652,1676]" lastBlockId="6.[213,1170,166,700]" lastPageId="6" lastPageNumber="671" pageId="5" pageNumber="670">
|
||
Length of idiosoma 768 µm. Ratio of leg I length to idiosomal length 1.04.
|
||
<emphasis id="B95F3D289008FFEBFF66FF50E63121AB" box="[213,360,166,190]" italics="true" pageId="6" pageNumber="671">
|
||
<taxonomicName id="4C2B9AB99008FFEBFF66FF50E63A21AB" authorityName="Krefft" authorityYear="1871" box="[213,355,166,190]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="6" pageNumber="671" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Subcapitulum broadly oval, subtriangular; ratio of length to breadth 1.33; distal hypostomal lips with spiniform internal malar and serrate external malar processes; adoral setae nude, short, not overlapping apex of subcapitulum (
|
||
<figureCitation id="1310FDBF9008FFEBFBA1FF10E3DD21EB" box="[1042,1156,230,254]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2F</figureCitation>
|
||
); proximal subcapitular setae pubescent, external pair slightly longer than internal pair. Dorsal surface of chelicera with saddle-shaped depression at level of bases of digits; cheliceral digits slender; fixed digit smooth along masticatory surface (
|
||
<figureCitation id="1310FDBF9008FFEBFC47FEB3E3DD2048" box="[1012,1156,325,349]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2A,B</figureCitation>
|
||
); movable digit smooth along masticatory surface. Chelicera with two setae, proximal seta inserted above articulation of movable digit; tip of proximal seta not reaching insertion of distal seta; tip of distal seta slightly overlaps apex of fixed digit. Length of chelicera 158 µm, dorsoventral width 59 µm, length of movable digit 69 µm, length of proximal and distal cheliceral setae 20 and 26 µm, respectively, distance between their insertions 26 µm. Ratios: cheliceral length to dorsoventral width 2.67; length of movable digit to length of chelicera 0.44; length of movable digit to dorsoventral width of chelicera 1.17. Palpal tarsus much shorter than femorogenu (
|
||
<figureCitation id="1310FDBF9008FFEBFC1EFDB2E37E2349" box="[941,1063,580,604]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2G</figureCitation>
|
||
); ratio of length to width of tarsus 2.0. Lengths of palpal trochanter, femorogenu, tibia and tarsus 33, 66, 23 and 59 µm, respectively. Number of setae on palpal segments was not discernible.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66FD18E3252250" blockId="6.[213,1170,750,840]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66FD18E60F2213" box="[213,342,750,774]" italics="true" pageId="6" pageNumber="671">Prodorsum.</emphasis>
|
||
Naso indistinct, only slightly developed (
|
||
<figureCitation id="1310FDBF9008FFEBFC91FD18E4C72213" box="[802,926,750,774]" captionStart="Figure 1" captionStartId="4.[213,280,1637,1659]" captionTargetBox="[211,1171,599,1589]" captionTargetId="figure-78@4.[211,1171,599,1589]" captionTargetPageId="4" captionText="Figure 1. Rhagidia (D.) paralleloseta sp. nov., adult male: (A) prodorsum; (B) ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203375" httpUri="https://zenodo.org/record/5203375/files/figure.png" pageId="6" pageNumber="671">Figure 1 A</figureCitation>
|
||
), with pair of internal vertical setae
|
||
<emphasis id="B95F3D289008FFEBFEDFFCF9E6D8223D" box="[364,385,783,808]" italics="true" pageId="6" pageNumber="671">
|
||
v
|
||
<subScript id="17AFE37F9008FFEBFECBFCE1E6D8223D" attach="left" box="[376,385,791,808]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
. Bothridial setae
|
||
<emphasis id="B95F3D289008FFEBFDFBFCF9E53F223D" box="[584,614,783,808]" italics="true" pageId="6" pageNumber="671">
|
||
sc
|
||
<subScript id="17AFE37F9008FFEBFDEEFCE1E53F223D" attach="left" box="[605,614,791,808]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
filiform, finely pubescent, clearly shorter than
|
||
<emphasis id="B95F3D289008FFEBFBC2FCF9E3C9223D" box="[1137,1168,783,808]" italics="true" pageId="6" pageNumber="671">
|
||
sc
|
||
<subScript id="17AFE37F9008FFEBFB34FCE1E3C9223D" attach="left" box="[1159,1168,791,808]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
and not overlapping disjugal suture. Length of setae:
|
||
<emphasis id="B95F3D289008FFEBFC98FCD8E466225D" box="[811,831,814,840]" italics="true" pageId="6" pageNumber="671">
|
||
v
|
||
<subScript id="17AFE37F9008FFEBFC85FCC1E466225D" attach="left" box="[822,831,823,840]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
33,
|
||
<emphasis id="B95F3D289008FFEBFCC3FCD8E4DD225D" box="[880,900,814,840]" italics="true" pageId="6" pageNumber="671">
|
||
v
|
||
<subScript id="17AFE37F9008FFEBFCC8FCC1E4DD225D" attach="left" box="[891,900,823,840]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
33,
|
||
<emphasis id="B95F3D289008FFEBFC06FCD8E48D225D" box="[949,980,814,840]" italics="true" pageId="6" pageNumber="671">
|
||
sc
|
||
<subScript id="17AFE37F9008FFEBFC78FCC1E48D225D" attach="left" box="[971,980,823,840]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
76,
|
||
<emphasis id="B95F3D289008FFEBFBB6FCD8E37D225D" box="[1029,1060,814,840]" italics="true" pageId="6" pageNumber="671">
|
||
sc
|
||
<subScript id="17AFE37F9008FFEBFBA8FCC1E37D225D" attach="left" box="[1051,1060,823,840]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
99 µm.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66FC81E3CB22DB" blockId="6.[213,1170,887,977]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66FC81E5D9229A" box="[213,640,887,911]" italics="true" pageId="6" pageNumber="671">Opisthosomal dorsum and anal region.</emphasis>
|
||
Complement and arrangement of dorsal setae typical of
|
||
<taxonomicName id="4C2B9AB99008FFEBFEF8FC61E68E22BA" authorityName="Oudemans" authorityYear="1922" box="[331,471,919,943]" class="Arachnida" family="Rhagidiidae" kingdom="Animalia" order="Trombidiformes" pageId="6" pageNumber="671" phylum="Arthropoda" rank="family">Rhagidiidae</taxonomicName>
|
||
; cupules were indiscernible. Length of setae:
|
||
<emphasis id="B95F3D289008FFEBFC55FC6EE4A222A4" box="[998,1019,920,945]" italics="true" pageId="6" pageNumber="671">
|
||
c
|
||
<subScript id="17AFE37F9008FFEBFC41FC56E4A222A4" attach="left" box="[1010,1019,928,945]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
23,
|
||
<emphasis id="B95F3D289008FFEBFB83FC6EE31C22A4" box="[1072,1093,920,945]" italics="true" pageId="6" pageNumber="671">
|
||
c
|
||
<subScript id="17AFE37F9008FFEBFB8FFC56E31C22A4" attach="left" box="[1084,1093,928,945]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
76,
|
||
<emphasis id="B95F3D289008FFEBFBC9FC61E3C922A4" box="[1146,1168,919,945]" italics="true" pageId="6" pageNumber="671">
|
||
d
|
||
<subScript id="17AFE37F9008FFEBFB34FC56E3C922A4" attach="left" box="[1159,1168,928,945]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B95F3D289008FFEBFEB5FC4EE64222C4" box="[262,283,952,977]" italics="true" pageId="6" pageNumber="671">
|
||
e
|
||
<subScript id="17AFE37F9008FFEBFEA1FC36E64222C4" attach="left" box="[274,283,960,977]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
not known (torn off),
|
||
<emphasis id="B95F3D289008FFEBFDA4FC41E57022C4" box="[535,553,951,977]" italics="true" pageId="6" pageNumber="671">
|
||
f
|
||
<subScript id="17AFE37F9008FFEBFD93FC36E57022C4" attach="left" box="[544,553,960,977]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
36,
|
||
<emphasis id="B95F3D289008FFEBFDEAFC41E53322C4" box="[601,618,951,977]" italics="true" pageId="6" pageNumber="671">
|
||
f
|
||
<subScript id="17AFE37F9008FFEBFDD2FC36E53322C4" attach="left" box="[609,618,960,977]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
30,
|
||
<emphasis id="B95F3D289008FFEBFD29FC41E5E922C4" box="[666,688,951,977]" italics="true" pageId="6" pageNumber="671">
|
||
h
|
||
<subScript id="17AFE37F9008FFEBFD14FC36E5E922C4" attach="left" box="[679,688,960,977]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
69,
|
||
<emphasis id="B95F3D289008FFEBFD53FC41E5AF22C4" box="[736,758,951,977]" italics="true" pageId="6" pageNumber="671">
|
||
h
|
||
<subScript id="17AFE37F9008FFEBFD5EFC36E5AF22C4" attach="left" box="[749,758,960,977]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
30,
|
||
<emphasis id="B95F3D289008FFEBFC95FC4EE41F22C4" box="[806,838,952,977]" italics="true" pageId="6" pageNumber="671">
|
||
ps
|
||
<subScript id="17AFE37F9008FFEBFC8EFC36E41F22C4" attach="left" box="[829,838,960,977]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
53,
|
||
<emphasis id="B95F3D289008FFEBFCC5FC4EE4CE22C4" box="[886,919,952,977]" italics="true" pageId="6" pageNumber="671">
|
||
ps
|
||
<subScript id="17AFE37F9008FFEBFC3DFC36E4CE22C4" attach="left" box="[910,919,960,977]" fontSize="7" pageId="6" pageNumber="671">2</subScript>
|
||
</emphasis>
|
||
30,
|
||
<emphasis id="B95F3D289008FFEBFC75FC4EE4BE22C4" box="[966,999,952,977]" italics="true" pageId="6" pageNumber="671">
|
||
ps
|
||
<subScript id="17AFE37F9008FFEBFC6DFC36E4BE22C4" attach="left" box="[990,999,960,977]" fontSize="7" pageId="6" pageNumber="671">3</subScript>
|
||
</emphasis>
|
||
20,
|
||
<emphasis id="B95F3D289008FFEBFBA5FC41E36322C4" box="[1046,1082,951,977]" italics="true" pageId="6" pageNumber="671">
|
||
ad
|
||
<subScript id="17AFE37F9008FFEBFB82FC36E36322C4" attach="left" box="[1073,1082,960,977]" fontSize="7" pageId="6" pageNumber="671">1</subScript>
|
||
</emphasis>
|
||
25 µm.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66FBF6E31A250D" blockId="6.[213,1091,1024,1048]" box="[213,1091,1024,1048]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66FBF6E616250D" box="[213,335,1024,1048]" italics="true" pageId="6" pageNumber="671">Podosoma.</emphasis>
|
||
Coxae I, II, III, IV with 3-1-5-3 finely pubescent setae, respectively.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66FBBCE32525B4" blockId="6.[213,1170,1098,1186]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66FBBCE6202577" box="[213,377,1098,1122]" italics="true" pageId="6" pageNumber="671">Genital region.</emphasis>
|
||
Genital valves each with five finely pubescent genital setae (
|
||
<emphasis id="B95F3D289008FFEBFB97FBBDE3682577" box="[1060,1073,1099,1122]" italics="true" pageId="6" pageNumber="671">g</emphasis>
|
||
) of similar length, about 16 µm, arranged evenly along medial edge of valve. Five pairs of aggenital setae (
|
||
<emphasis id="B95F3D289008FFEBFE3AFB7CE6FD25B4" box="[393,420,1162,1185]" italics="true" pageId="6" pageNumber="671">ag</emphasis>
|
||
) of similar length, about 30 µm. Length of genital valves 96 µm.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66FB22E37F24FF" blockId="6.[213,1170,1235,1515]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66FB22E64825FE" box="[213,273,1236,1259]" italics="true" pageId="6" pageNumber="671">Legs.</emphasis>
|
||
Leg I 800 µm long, about 1.04–1.15 times as long as idiosoma. Empodia of all legs setulose, slightly longer than claws; claws each with small basal tubercle ventrobasally. Tarsus I slender, its tip slightly tapers in lateral view, ratio length to width 4.6, with four strikingly long rhagidial solenidia lying parallel in separate depressions dorsodistally (
|
||
<figureCitation id="1310FDBF9008FFEBFEC6FAA4E6B0247F" box="[373,489,1362,1386]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2C</figureCitation>
|
||
), stellate famulus (
|
||
<emphasis id="B95F3D289008FFEBFD09FAA5E59F247E" box="[698,710,1363,1387]" italics="true" pageId="6" pageNumber="671">ε</emphasis>
|
||
) subtending second rhagidial solenidion; tarsus II with three rhagidial solenidia lying parallel in separate depressions, spiniform famulus (
|
||
<emphasis id="B95F3D289008FFEBFEF2FA65E61424BE" box="[321,333,1427,1451]" italics="true" pageId="6" pageNumber="671">ε</emphasis>
|
||
) not observed (
|
||
<figureCitation id="1310FDBF9008FFEBFDB7FA64E52624BF" box="[516,639,1426,1450]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2D</figureCitation>
|
||
). Dorsodistal lanceolate solenidion on tibia II recessed in deep insertion pit with broadly open surface pore (
|
||
<figureCitation id="1310FDBF9008FFEBFC25FA44E35524DF" box="[918,1036,1458,1482]" captionStart="Figure 2" captionStartId="5.[144,211,1164,1186]" captionTargetBox="[150,1093,161,1094]" captionTargetId="figure-7@5.[150,1094,161,1095]" captionTargetPageId="5" captionText="Figure 2. Rhagidia (D.) paralleloseta sp. nov., adult female: (A) chelicera, lateral aspect; (B) cheliceral digits in detail, lateral aspect; (C) rhagidial organ I, dorsal aspect; (D) rhagidial organ II, lateral aspect; (E) dorsodistal lanceolate solenidion Φ on tibia II, dorsal aspect; (F) subcapitulum, ventral aspect; (G) palpus, lateral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203377" httpUri="https://zenodo.org/record/5203377/files/figure.png" pageId="6" pageNumber="671">Figure 2E</figureCitation>
|
||
). Spiniform solenidia on leg segments were not discernible in badly damaged specimens.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66F9CCE6BA2744" blockId="6.[213,483,1593,1617]" box="[213,483,1593,1617]" pageId="6" pageNumber="671">
|
||
<emphasis id="B95F3D289008FFEBFF66F9CCE6BA2744" box="[213,483,1593,1617]" italics="true" pageId="6" pageNumber="671">Description – adult male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9008FFEBFF66F996E607278D" blockId="6.[213,1169,1632,1688]" pageId="6" pageNumber="671">
|
||
One examined but lost in preparation. Length of idiosoma 640 µm, similar to female (
|
||
<figureCitation id="1310FDBF9008FFEBFF6EF976E609278D" box="[221,336,1664,1688]" captionStart="Figure 1" captionStartId="4.[213,280,1637,1659]" captionTargetBox="[211,1171,599,1589]" captionTargetId="figure-78@4.[211,1171,599,1589]" captionTargetPageId="4" captionText="Figure 1. Rhagidia (D.) paralleloseta sp. nov., adult male: (A) prodorsum; (B) ventral aspect." figureDoi="http://doi.org/10.5281/zenodo.5203375" httpUri="https://zenodo.org/record/5203375/files/figure.png" pageId="6" pageNumber="671">Figure 1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9009FFEAFF23FF50E7AC21AB" blockId="7.[144,245,166,190]" box="[144,245,166,190]" pageId="7" pageNumber="672">
|
||
<emphasis id="B95F3D289009FFEAFF23FF50E7AC21AB" box="[144,245,166,190]" italics="true" pageId="7" pageNumber="672">Affinities</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9009FFEAFF23FF38E5662070" blockId="7.[144,1101,206,358]" pageId="7" pageNumber="672">
|
||
<taxonomicName id="4C2B9AB99009FFEAFF23FF38E6E121F3" authority="Zacharda & Grafitti & Piva, 2011" authorityName="Zacharda & Grafitti & Piva" authorityYear="2011" box="[144,440,206,230]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="672" phylum="Arthropoda" rank="species" species="paralleloseta" status="sp. nov." subGenus="Deharvengiella">
|
||
<emphasis id="B95F3D289009FFEAFF23FF38E7AD21F3" box="[144,244,206,230]" italics="true" pageId="7" pageNumber="672">Rhagidia</emphasis>
|
||
(
|
||
<emphasis id="B95F3D289009FFEAFEB0FF39E64721F3" box="[259,286,207,230]" italics="true" pageId="7" pageNumber="672">D.</emphasis>
|
||
)
|
||
<emphasis id="B95F3D289009FFEAFE9FFF38E6E121F3" box="[300,440,206,230]" italics="true" pageId="7" pageNumber="672">paralleloseta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C80539009FFEAFE0DFF39E54B21F3" box="[446,530,207,230]" pageId="7" pageNumber="672" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from the other known representatives of the genus by having a parallel position of the enlarged rhagidial solenidia in the rhagidial organs I and II. The extremely long, slender rhagidial solenidia are a troglomorphism, a specific morphological adaptation to life in the subterranean habitat. See also the key to adults of
|
||
<taxonomicName id="4C2B9AB99009FFEAFEF6FEB8E6B02073" authorityName="Zacharda, 1987:" authorityYear="2051" box="[325,489,334,358]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="668" phylum="Arthropoda" rank="subGenus" subGenus="Deharvengiella">
|
||
<emphasis id="B95F3D289009FFEAFEF6FEB8E6B02073" box="[325,489,334,358]" italics="true" pageId="7" pageNumber="672">Deharvengiella</emphasis>
|
||
</taxonomicName>
|
||
, below.
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9009FFEAFF23FE5DE65120D6" blockId="7.[144,264,427,451]" box="[144,264,427,451]" pageId="7" pageNumber="672">
|
||
<emphasis id="B95F3D289009FFEAFF23FE5DE65120D6" box="[144,264,427,451]" italics="true" pageId="7" pageNumber="672">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B94E13A9009FFEAFF23FE24E6EE231F" blockId="7.[144,1101,466,522]" pageId="7" pageNumber="672">
|
||
The epithet
|
||
<taxonomicName id="4C2B9AB99009FFEAFEA6FE25E6F820FE" authorityName="Zacharda & Grafitti & Piva" authorityYear="2011" box="[277,417,467,491]" class="Arachnida" family="Rhagidiidae" genus="Rhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="7" pageNumber="672" phylum="Arthropoda" rank="species" species="paralleloseta">
|
||
<emphasis id="B95F3D289009FFEAFEA6FE25E6F820FE" box="[277,417,467,491]" italics="true" pageId="7" pageNumber="672">paralleloseta</emphasis>
|
||
</taxonomicName>
|
||
reflects the parallel position of the rhagidial solenidia on tarsi I and II. Feminine gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |