295 lines
38 KiB
XML
295 lines
38 KiB
XML
<document id="A527C17A75068D6B0F6DB7241C79D25F" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="A6E8E3440CF48E6EF2DE1685B01AB307" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha beaumontii Le Guillou 1841" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="6" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="9C3FE0BC10C4CA9E06D9658459D70079">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="2A2C4824044B4164B90B3879B37A8A8A">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="989327E139B8D8DAFC01C7370A0D4E5E">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="5BE4D22179BBB13F3ED73100A182A3E7">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="EB7F2A7F9D6F5C97A225AD1E4ED2CB36">text</mods:typeOfResource>
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<mods:title id="015BC853356A842F654AA8CF5052A5D6">ZooKeys</mods:title>
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<mods:date id="A9B41DEF60B6555D704A64E055CF492A">2011</mods:date>
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<mods:number id="F24D5DB383955FFA35A27DFAB87A1006">131</mods:number>
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<mods:url id="98F110B7B39A6072C89E5212539193DC">http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification id="E4E41B69603F61341927587A59DFED98">journal article</mods:classification>
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<mods:identifier id="116A9F28BFFFA522BE77A21886541C42" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier id="B71DF9CBD27A6DAD18428E402F30D01A" type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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<treatment id="A6E8E3440CF48E6EF2DE1685B01AB307" ID-GBIF-Taxon="152031107" LSID="urn:lsid:plazi:treatment:A6E8E3440CF48E6EF2DE1685B01AB307" httpUri="http://treatment.plazi.org/id/A6E8E3440CF48E6EF2DE1685B01AB307" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="E304579AFB18040B15289CCA90187740" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="55AF0432ED175148E6ACE7215A7ED412" pageId="5" pageNumber="6">
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<taxonomicName id="1E563097093FECC51C8FA35EDA6D367B" ID-CoL="758NZ" LSID="http://species-id.net/wiki/Orthomorpha_beaumontii" authority="Le Guillou, 1841" authorityName="Le Guillou" authorityYear="1841" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii (Le Guillou, 1841)</taxonomicName>
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Figs 13
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</paragraph>
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</subSubSection>
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<subSubSection id="706172D09914D6E89945C25122921015" pageId="5" pageNumber="6" type="reference_group">
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<paragraph id="01429F5E5C422D05BFD8F88D691C27AE" pageId="5" pageNumber="6">
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<taxonomicName id="54BDDD48678884FFAEA309D7AF383C8A" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus (Beaumontii)" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Beaumontii">Polydesmus Beaumontii</taxonomicName>
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<bibRefCitation id="3CBB1BBF798F9598DA5BA4EF1091DDC0" author="Le Guillou, EJF" journalOrPublisher="Socie ́ te ́ philomathique de Paris" pageId="114" pageNumber="115" pagination="279 - 280" title="[Catalogue raisonne ́ des insectes recueillis pendant le voyage de circumnavigation des corvettes l'Astrolabe et la Zelee]. L'Institut, journal universal des sciences et des societes savantes." volume="9" year="1841">Le Guillou 1841</bibRefCitation>
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: 279 (D).
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</paragraph>
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<paragraph id="2773E98257B48666CBB910F4E111C7A9" pageId="5" pageNumber="6">
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<taxonomicName id="35B3F928C8C960A9A213A6953F9C6E3E" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus (Beaumontii)" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Beaumontii">Polydesmus Beaumontii</taxonomicName>
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-
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<bibRefCitation id="9B078274050B9A6EAA6534D2107F184D" pageId="5" pageNumber="6">Gervais 1847</bibRefCitation>
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: 101 (D).
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</paragraph>
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<paragraph id="ACE6EDCA504DF90E0FD15E5CB4956877" pageId="5" pageNumber="6">
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<taxonomicName id="45C876A9B35447F7EA943C761220710F" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus (Parademus)" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Parademus">Polydesmus (Parademus) Beaumontii</taxonomicName>
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-
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<bibRefCitation id="6132B247096DA0D6326AD47DC454AFC3" author="De Saussaure, H" journalOrPublisher="Linnaea Entomologica" pageId="114" pageNumber="115" pagination="318 - 327" title="Note sur la famille des Polydesmides, principalement au point de vue des especes americaines." volume="13" year="1859">De Saussure 1859</bibRefCitation>
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: 326 (D); Humbert &
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<bibRefCitation id="15BA2738D7314207596B2CF8869A8E1E" author="De Saussure, H" journalOrPublisher="Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve" pageId="114" pageNumber="115" pagination="259 - 394" title="Essai d'une faune de Myriapodes du Mexique, avec la description de quelques especes des autres parties de l'Amerique." volume="15" year="1860">De Saussure 1860</bibRefCitation>
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: 670 (D).
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</paragraph>
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<paragraph id="51FD632D9F7F3066945D0F42307F43DE" pageId="5" pageNumber="6">
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<taxonomicName id="01843BC5ABAC21681571C2C2D763381B" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumonti" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beaumonti">Orthomorpha beaumonti</taxonomicName>
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-
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<bibRefCitation id="7EA427727BE46A93BE824EC1EAD03E44" author="Bollman, CH" journalOrPublisher="Bulletin of the United States National Museum" pageId="114" pageNumber="115" pagination="150 - 152" title=": Notes upon the North American myriapods described by C. L. Koch." volume="46" year="1893">Bollman 1893</bibRefCitation>
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: 196 (M, R).
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</paragraph>
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<paragraph id="7652DB7454C1F33F0FBA0FC7608E4BF0" pageId="5" pageNumber="6">
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<taxonomicName id="A2E39BE79F1365B957B4AD9845658485" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica subsp. spinala" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="hydrobiologica" subSpecies="spinala">Orthomorpha hydrobiologica spinala</taxonomicName>
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<bibRefCitation id="02C8EAE50EE41CA86A0FBEBD4795B7AC" author="Attems, C" journalOrPublisher="Treubia" pageId="114" pageNumber="115" pagination="29 - 41" title="Neue Polydesmiden des Museums Buitenzorg." volume="14" year="1932">Attems 1932</bibRefCitation>
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: 39 (D), syn. n.
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</paragraph>
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<paragraph id="703C83BCC6A389D96175A6A41003AFA4" pageId="5" pageNumber="6">
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<taxonomicName id="BEE5CF33EB90134E32197B567F1DEC5C" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha spinala" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinala">Orthomorpha spinala</taxonomicName>
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-
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<bibRefCitation id="C4663DF16054A3B011A1FFBC2266A86F" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
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: 45 (M);
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<bibRefCitation id="F0F035E59AD06B07F98EFF7C77C844E7" author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
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: 362 (M);
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<bibRefCitation id="5CB57FD6CA1D8CD3C910996CD38878DC" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation id="6F6361F1C1C5B86DCB55458DFDFAE1E6" pageId="5" pageNumber="6">Golovatch 1998</bibRefCitation>
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: 42 (D).
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</paragraph>
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<paragraph id="8682D4815F2FE8C6A744A3AEDDE9D25E" pageId="5" pageNumber="6">
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<taxonomicName id="3FD37DAE5E36C876FDB45F8EC4A67A7D" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
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-
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<bibRefCitation id="4C98614ED205771C3B12B7A530DEC943" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 269 (D);
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<bibRefCitation id="F2DA71D4B715AD14617B736D1D15A239" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 45 (M);
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<bibRefCitation id="A19B6F4E536D7A88D04C56552ED9450B" author="Golovatch, SI" journalOrPublisher="Miscellania Zoologica" pageId="114" pageNumber="115" pagination="73 - 80" title="Review of the Sunda weberi-group of Orthomorpha Bollman, 1893, with the description of two new species (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1997 b">Golovatch 1997b</bibRefCitation>
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: 79 (D);
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<bibRefCitation id="3D03C1745A0C1EC84E295C355BB103E7" pageId="5" pageNumber="6">1998</bibRefCitation>
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: 42 (D).
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</paragraph>
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<paragraph id="077739CF068EB500DA6E4C60D8B1A5C4" pageId="5" pageNumber="6">
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<taxonomicName id="4CB1EAF2A604B3400B1D704B69D1116D" class="Diplopoda" family="Paradoxosomatidae" genus="Prionopeltis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prionopeltis" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Prionopeltis</taxonomicName>
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sp. (
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<taxonomicName id="33C381C941F878D3FC407ABE65ED1E66" genus="Beaumontii" lsidName="Beaumontii" pageId="5" pageNumber="6" rank="genus">Beaumontii</taxonomicName>
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) (sic!) -
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<bibRefCitation id="5D0DCDFC62B368F015B9ECA70480A8A6" author="Attems, C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, mathematisch-naturwissenschaftliche Classe" pageId="114" pageNumber="115" pagination="221 - 482" title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems 1898</bibRefCitation>
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: 357 (D, R) [Non],
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</paragraph>
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<paragraph id="313066621DAA4EBF2C898475CA0F4427" pageId="5" pageNumber="6">
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<taxonomicName id="05EC968BE52507ACC490850F284A2D3E" class="Diplopoda" family="Paradoxosomatidae" genus="Prionopeltis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prionopeltis (Beaumontii)" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Beaumontii">Prionopeltis Beaumontii</taxonomicName>
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Attems (sic!) -
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<bibRefCitation id="7DF40261CE1BB11477B6E159A6DE52DB" pageId="5" pageNumber="6">Attems 1914</bibRefCitation>
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: 207 (M) [nec],
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</paragraph>
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<paragraph id="C8BBACD74141D643C79199512A039788" pageId="5" pageNumber="6">
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<taxonomicName id="A72E4EAD4C495BC7DE7FD5C02BB0A64B" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus beaumontii" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beaumontii">Pratinus beaumontii</taxonomicName>
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(Att.) -
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<bibRefCitation id="5A692A50085B2517052D8F7BFC93F857" pageId="5" pageNumber="6">Attems 1937</bibRefCitation>
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: 122 (M) [nec].
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</paragraph>
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</subSubSection>
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<subSubSection id="E156C9C3CABDB18DD199CF3E61B1E88F" pageId="5" pageNumber="6" type="material examined">
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<paragraph id="2A5053A014B72DB85484D31623C0F1DF" pageId="5" pageNumber="6">Material examined.</paragraph>
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<paragraph id="1DA56433779BB5F8CA5BA6616532756A" pageId="5" pageNumber="6">
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Syntypes of
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<taxonomicName id="A621979DE217C77E948FF962D3F7A588" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica subsp. spinala" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="hydrobiologica" subSpecies="spinala">Orthomorpha hydrobiologica spinala</taxonomicName>
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: 4 ♂, 1 ♀ (
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<normalizedToken id="787E6577FEDE2B93A56C6A5FE669D1A4" originalValue="NHMW–">NHMW-</normalizedToken>
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3510), Indonesia, "Karimon Djawa Inseln" (= Pulau Karimunjawa Island, north of Java), V.1926, leg. Dammerman; 1 ♀ of
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<taxonomicName id="1C7D32180BA71C8E806845680BEF72A0" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica subsp. spinala" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subSpecies" species="hydrobiologica" subSpecies="spinala">Orthomorpha hydrobiologica spinala</taxonomicName>
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(
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<normalizedToken id="B04FBE71C16C0F829A8A94890F14323D" originalValue="NHMW–">NHMW-</normalizedToken>
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8001), Indonesia, Java, Tjibodas, no date, leg. W. S. S. van Benthem-Jutting, det. C. Attems.
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</paragraph>
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</subSubSection>
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<subSubSection id="BACB9345F3467125A0478A8DBF1696CB" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6" type="redescription">
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<paragraph id="FCD56193A651ECE53C5D33FF785B1D55" pageId="5" pageNumber="6">Redescription.</paragraph>
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<paragraph id="12699ABAC49C2F629FD4EFEE0E37030C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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Length 33-38 mm (♂), 36-38 mm (♀), width of midbody pro- and metazona 2.7-2.8 and 4.0-4.2 mm (♂), 3.1-3.3 and 4.3-4.4 mm (♀), respe
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<pageBreakToken id="C8A4F64776BF3BBAC77587F919EACC64" pageId="6" pageNumber="7" start="start">ctively</pageBreakToken>
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. Coloration of alcohol material upon long-term preservation rather uniformly brown with contrasting pale yellowish paraterga, venter and legs light yellow-brown (Fig. 2).
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</paragraph>
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<paragraph id="8F28473CCB8FC08336B3074108792E73" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
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Head usual, clypeolabral region sparsely setose, surface of vertex smooth; epicranial suture distinct. Antennae moderately long (Fig. 2A & B), reaching behind midway of body segment 3 (♂) or beyond segment 2 (♀). Head in width <collum <segments 3 and 4 <segment 2 <segments 5-16(17), gently and gradually tapering thereafter. Collum with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3
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<pageBreakToken id="7E49C9CCB1D779107838BB069A5927B9" pageId="7" pageNumber="8" start="start">posterior</pageBreakToken>
|
||
setae; caudal corner of paraterga dentiform, pointed, directed caually (Fig. 2A, B & J). Tegument smooth and shining, prozona very finely shagreened, metaterga slightly rugulose; surface below paraterga smooth. Postcollum metaterga with two transverse rows of setae, these being always abraded and traceable as insertion points: 2+2 in anterior (pre-sulcus) row, 3+3 in posterior (postsulcus) one. Axial line barely visible both on pro- and metazona. Paraterga very strongly developed (Fig. 2A-G, J-L), especially so in ♂, subhorizontal, always lying below dorsum, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, always clearly projecting well behind tergal margin. Calluses delimited both dorsally and ventrally, only on segment 2 without ventral sulcus, thin, especially so on poreless segments. Paraterga 2 broad, anterior edge rounded, lateral edge with two small, but evident incisions in anterior 1/3; posterior edge evidently concave (Fig. 2A & J). Paraterga 3 and 4 subequal,
|
||
<pageBreakToken id="83328A200339A5050EE0DADDE6CC25C7" pageId="8" pageNumber="9" start="start">like</pageBreakToken>
|
||
subsequent paraterga, anterior edge slightly rounded, bordered and fused to callus, lateral edge with a small incision in anterior third. Paraterga 15-19 with tip of caudal corner evidently curved mesad. Ozopores evident, lateral, lying in an ovoid groove at about 1/3 of metazonital length. Transverse sulcus present on metaterga 5-18, shallow, not reaching bases of paraterga, finely beaded at bottom (Fig. 2A, C, F, J-L). Stricture between pro- and metazona narrow, shallow, beaded at bottom down to base of paraterga (Fig. 2D & E). Pleurosternal carinae complete crests only on segment 2 or segments 2 and 3, with a small, sharp, caudal tooth on segments 3-7(8) (♂) or 4-6 (♀), thereafter with a very small caudal denticle until segment 15 (♂, ♀). Epiproct (Fig. 2E-G & L) conical, flattened dorsoventrally, apical papillae well-developed, acute and directed ventrad; tip subtruncate; pre-apical papillae small, but visible. Hypoproct (Fig. 2G) subtriangular, setiferous knobs at caudal edge well-separated.
|
||
</paragraph>
|
||
<paragraph id="16F65134554BD344DC24E57537FA14AC" pageId="8" pageNumber="9">Sterna sparsely setose, without modifications, but with a pair of small, rounded, completely separated, setose cones between ♂ coxae 4 (Fig. 2H & I). No conspicuous ridge in front of gonopod aperture. Legs long and slender, slightly incrassate in ♂, midbody ones ca 1.2-1.3 (♂) or 0.8-0.9 times (♀) as long as body height, prefemora without modifications, tarsal brushes present until legs of segment 9.</paragraph>
|
||
<paragraph id="259BC4CC85E37D2D4102FEF454B77547" pageId="8" pageNumber="9">
|
||
Gonopods (Fig. 3) simple. Coxa long and slender, with several setae distodorsally. Prefemoral (= densely setose) portion more than 3 times shorter than femorite (measured until beginning of solenomere, including
|
||
<normalizedToken id="29DF44141E624EA84F0B48B1EB27DE3B" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
||
part lying beyond lateral sulcus). Femorite slender, slightly curved and not enlarged distad,
|
||
<normalizedToken id="BA83B2C7D34A2823CA7A5ADEC6E795D9" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
||
part demarcated by an oblique lateral sulcus; tip of solenophore evidently trifid, middle denticle much smaller than both a terminal tooth and a subterminal lobule; solenomere about as long as solenophore, flagelliform.
|
||
</paragraph>
|
||
<caption id="0A18A779379D2F2D7E54DDE70CFED75B" pageId="8" pageNumber="9">
|
||
<paragraph id="C44665B2B38F3BA8199613C61E265486" pageId="8" pageNumber="9">
|
||
Figure 1.
|
||
<taxonomicName id="CDC55D01E53135324B2DBE0AC3BFE902" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
(Le Guillou, 1841), ♀ holotype. A, B segments 2-5, dorsal and lateral views, respectively C, D segments 10 and 11, dorsal and lateral views, respectively E segments 16-20, dorsal view F posterior part of body, ventral view (after
|
||
<bibRefCitation id="32D58A476581AD1EBB15ECDE8A32C742" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D4248C11CA50F36311094C5738080358" pageId="8" pageNumber="9">
|
||
<paragraph id="F68D2601AECBD4D01D20DD4FE2EC333F" pageId="8" pageNumber="9">
|
||
Figure 2.
|
||
<taxonomicName id="85D3B24C2A3594989DDCCAC88561E0E8" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
(Le Guillou, 1841), ♂ (
|
||
<normalizedToken id="9C15534DBB6776CB7A5B4B9519F034FD" originalValue="A–I">A-I</normalizedToken>
|
||
) and ♀ (
|
||
<normalizedToken id="D13CD22FEC6250B3570B0A5713F6F5BF" originalValue="J–L">J-L</normalizedToken>
|
||
) syntypes of
|
||
<taxonomicName id="AF442E4DA9D69E32E8ACA8EAAA27EB25" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica subsp. spinala" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="hydrobiologica" subSpecies="spinala">Orthomorpha hydrobiologica spinala</taxonomicName>
|
||
Attems, 1932. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
|
||
<normalizedToken id="01A22E99D68EA759C1174F7E4F079032" originalValue="E–G">E-G</normalizedToken>
|
||
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B959CF29631BAC88F9E4AA9302593269" pageId="8" pageNumber="9">
|
||
<paragraph id="3D484B8CD2EC6382B1299A7AFB3E2E53" pageId="8" pageNumber="9">
|
||
Figure 3.
|
||
<taxonomicName id="27894771FF3D98E03F4FA4E5A8CE491A" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
(Le Guillou, 1841), ♂ syntype of
|
||
<taxonomicName id="244E8F0B6FC2C9660AAE3EFB99E61DBF" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica subsp. spinala" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="hydrobiologica" subSpecies="spinala">Orthomorpha hydrobiologica spinala</taxonomicName>
|
||
Attems, 1932. A,B right gonopod, lateral and mesal views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5CF00DB512EA77638EE7B89B6B3AF61D" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="remarks">
|
||
<paragraph id="AB55ADFF84016F84FD2145D82DF025CB" pageId="8" pageNumber="9">Remarks.</paragraph>
|
||
<paragraph id="A83D8FCA678A3100A1B88CFDF6FB1ECE" pageId="8" pageNumber="9">
|
||
A complete historical review of the typification of
|
||
<taxonomicName id="3996417DD2D1F61A546306252C1A5D24" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
has long been provided by
|
||
<bibRefCitation id="85ADE944F3A919EE77C3B975639D861F" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel (1963)</bibRefCitation>
|
||
. Despite some confusion,
|
||
<taxonomicName id="706B8B736D64D89D4D7928D5EBD15898" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
was properly typified by
|
||
<bibRefCitation id="0771DAED2000A6C4A39D724F4F254AF2" pageId="8" pageNumber="9">Pocock (1909)</bibRefCitation>
|
||
, with
|
||
<taxonomicName id="823940A84E7F9D4E20B6F7F76A03F3F3" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
(Le Guillou, 1841) serving as the type-species.
|
||
</paragraph>
|
||
<paragraph id="BB1E07435981B694B3201A6CE75A9B71" pageId="8" pageNumber="9">
|
||
Originally described as a subspecies of
|
||
<taxonomicName id="465ED7AF8303D9D7E8E7D2EE38E38FD6" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha hydrobiologica" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hydrobiologica">Orthomorpha hydrobiologica</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="8C11F3EE705FF20635143A0260111D6D" author="Attems, C" journalOrPublisher="Treubia" pageId="114" pageNumber="115" pagination="29 - 41" title="Neue Polydesmiden des Museums Buitenzorg." volume="14" year="1932">Attems 1932</bibRefCitation>
|
||
), not as a variety as mistakenly quoted by
|
||
<bibRefCitation id="8A13DCE45F788CE3677847AF1CB94494" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel (1968)</bibRefCitation>
|
||
, spinala has since been treated as a full species (
|
||
<bibRefCitation id="22DD503C441BBF70DBD85A5156728BB1" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
|
||
). The above samples, especially the only available ♀ syntype, agree in almost every detail with the very accurate redescription of the
|
||
<taxonomicName id="1E5DCB6642984CBB25A350F7D5A48A09" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
holotype provided by
|
||
<bibRefCitation id="41B4009D3C8A4BF33D0B4661A1490861" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel (1963)</bibRefCitation>
|
||
, making a restudy of the holotype superfluous. The few differences, such as size (4.4 vs 5.0 mm), coloration (brown vs blackish), the shape of the caudal tooth on pleurosternal carinae (sharp teeth vs triangular lappets), the presence of an anterolateral denticle on paraterga (very small vs virtually missing) etc., are deemed too minor, rather reflecting individual or population-level variation, to consider
|
||
<taxonomicName id="22F6EB1164D7CADC729A3986966BD478" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha spinala" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinala">Orthomorpha spinala</taxonomicName>
|
||
as being distinct from
|
||
<taxonomicName id="9DD06F180FE22990AE3830B90E4BA454" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
at the species level. Hence the new synonymy advanced. In addition, the type series of
|
||
<taxonomicName id="0E32B37834EE2C95BEABA459A845A661" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha spinala" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="spinala">Orthomorpha spinala</taxonomicName>
|
||
derives from an islet lying nearly halfway between Java and the beaumontii type locality, Borneo.
|
||
</paragraph>
|
||
<paragraph id="B5DEA0C855C10248BE803E1C8C619B69" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<bibRefCitation id="F1F233DFC166B89DD37239E6BBA0975B" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel (1963)</bibRefCitation>
|
||
, when trying to find the closest match among the known
|
||
<taxonomicName id="25679059673BE031CD41F53F42F4C5E8" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
species to the holotype of
|
||
<taxonomicName id="7D830035AF1F59B168FD7A5365F1CFF3" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
, emphasized its especially strong similarities to
|
||
<taxonomicName id="F53C4D7C0BE28A943DC413881633BED7" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha weberi" order="Polydesmida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="weberi">Orthomorpha weberi</taxonomicName>
|
||
. Slight differences were only noted in the shape of the paraterga. It
|
||
<pageBreakToken id="3C93230C3CDE762B97F934B14C767A80" pageId="9" pageNumber="10" start="start">was</pageBreakToken>
|
||
this that allowed Jeekel to unequivocally conserve the concept of
|
||
<taxonomicName id="68DE616473CE7803C7E546C418623F56" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
. Zoogeographically, the strong morphological similarities between
|
||
<taxonomicName id="93119496D3D39C9DA8DB23377C09A901" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
and
|
||
<taxonomicName id="C8E9F841FEACD57765BE4BE2681F2EF9" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha weberi" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="weberi">Orthomorpha weberi</taxonomicName>
|
||
make sense, because the latter species is endemic to Java, Indonesia.
|
||
</paragraph>
|
||
<paragraph id="F61EB6A0A527856CC00B2C7C6DE22FF4" pageId="9" pageNumber="10">
|
||
With the above synonymization, the nomenclature of
|
||
<taxonomicName id="AA7C4FB74FC322675E4DF2ADFBB1E920" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
becomes stabilized, confirming this
|
||
<normalizedToken id="A2C3BA9FEEA3B9054DF6329A6D981BC1" originalValue="genus’">genus'</normalizedToken>
|
||
present scope. The identity of its type-species,
|
||
<taxonomicName id="E1F65586D483E969E70BC9AF3965F633" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
, has been refined, based on male characters as well.
|
||
</paragraph>
|
||
<paragraph id="33A8EA5C2EFFE1E3ED258688D94E3754" pageId="9" pageNumber="10">
|
||
<bibRefCitation id="0CD6AE33AF9AAD69F2A711D372EF39CD" author="Bollman, CH" journalOrPublisher="Bulletin of the United States National Museum" pageId="114" pageNumber="115" pagination="150 - 152" title=": Notes upon the North American myriapods described by C. L. Koch." volume="46" year="1893">Bollman (1893)</bibRefCitation>
|
||
, when proposing
|
||
<taxonomicName id="DCD3082EB6FDE0AF07121C243C67809F" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
||
as a replacement name for the preoccupied
|
||
<taxonomicName id="56EAF91A72B6E2E4C617E9F9379749DE" class="Diplopoda" family="Paradoxosomatidae" genus="Paradesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paradesmus" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Paradesmus</taxonomicName>
|
||
De Saussure, 1859, synonymized
|
||
<taxonomicName id="2F2AE4105016905BBBC6444027F0ADE8" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha beaumontii" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="beaumontii">Orthomorpha beaumontii</taxonomicName>
|
||
with
|
||
<taxonomicName id="F736FA2B1A3E1D8A7330DA77E601356B" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha spectabilis" order="Polydesmida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="spectabilis">Orthomorpha spectabilis</taxonomicName>
|
||
(Karsch, 1881), the latter species from Java. Apparently because he provided no evidence whatsoever to substantiate his synonymization, it has since been neglected, spectabilis still remaining a dubious name (e.g.
|
||
<bibRefCitation id="00013644F863F75EB482A35AE5AD4348" pageId="9" pageNumber="10">Attems 1937</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="657F15D4C1439DE0B78B09270423EF3D" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A9497BE058CEB6314FBFEE74F27B2E6D" pageId="9" pageNumber="10">
|
||
(
|
||
<bibRefCitation id="81376AF70D08D0D956ADD89DD21366B8" author="Attems, C" journalOrPublisher="Denkschriften der Kaiserlichen Akademie der Wissenschaften zu Wien, mathematisch-naturwissenschaftliche Classe" pageId="114" pageNumber="115" pagination="221 - 482" title="System der Polydesmiden. I. Theil." volume="67" year="1898">Attems (1898</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="BEEF834F2718A168D9DDCCC4D798C9E9" pageId="9" pageNumber="10">1914</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="756471E6258FA470D135AC6309CA15B6" pageId="9" pageNumber="10">1937</bibRefCitation>
|
||
) referred to beaumontii a sample from Java, Indonesia which he had received from the Berlin Museum, thus providing a second record of this species. However, because the gonopod tip of that sample shows a remarkably small subterminal lappet, while the paraterga and pleurosternal carinae slightly differ in shape from those of beaumontii, there can be no doubt of
|
||
<normalizedToken id="CF0BF5A380077299D0C7CDA643E573AB" originalValue="Attems’">Attems'</normalizedToken>
|
||
misidentification. Hence our references to it as such in the catalogue section above.
|
||
</paragraph>
|
||
<subSection id="C14DBDC754E370CC62C41E7570EE9595" pageId="9" pageNumber="10" type="species with only a single terminal lobule on the gonopod tip">
|
||
<paragraph id="D98CA058BB1F3142F90552314BF68364" pageId="9" pageNumber="10">Species with only a single terminal lobule on the gonopod tip</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |