155 lines
16 KiB
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155 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.264.4304" ID-GBIF-Dataset="869fb7d2-f383-44d3-897c-c6c0945e11a0" ID-PMC="PMC3668377" ID-Pensoft-Pub="1313-2970-264-85" ID-PubMed="23730179" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-264-85" ModsDocOrigin="ZooKeys 264" ModsDocTitle="Five new species and three new subspecies of Erebidae and Noctuidae (Insecta, Lepidoptera) from Northwestern North America, with notes on Chytolita Grote (Erebidae) and Hydraecia Guenée (Noctuidae)" checkinTime="1451247710553" checkinUser="pensoft" docAuthor="Crabo, Lars G., Davis, Melanie, Hammond, Paul, Tomas Mustelin, & Jon Shepard," docDate="2013" docId="9F7C3C82D189B2C70CDCD7CE2902F53C" docLanguage="en" docName="ZooKeys 264: 85-123" docOrigin="ZooKeys 264" docSource="http://dx.doi.org/10.3897/zookeys.264.4304" docTitle="Resapamea mammuthus Crabo, sp. n." docType="treatment" docVersion="5" lastPageNumber="105" masterDocId="FFA4FFA8FFD7DC68A95BFF963540FFDB" masterDocTitle="Five new species and three new subspecies of Erebidae and Noctuidae (Insecta, Lepidoptera) from Northwestern North America, with notes on Chytolita Grote (Erebidae) and Hydraecia Guenee (Noctuidae)" masterLastPageNumber="123" masterPageNumber="85" pageNumber="103" updateTime="1668155270344" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five new species and three new subspecies of Erebidae and Noctuidae (Insecta, Lepidoptera) from Northwestern North America, with notes on Chytolita Grote (Erebidae) and Hydraecia Guenee (Noctuidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Crabo, Lars G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Davis, Melanie</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hammond, Paul</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tomas Mustelin,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jon Shepard,</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>264</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>85</mods:start>
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<mods:end>123</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.264.4304</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.264.4304</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-264-85</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040360" LSID="urn:lsid:zoobank.org:act:5D5927C7-717D-4DC4-B855-37389C705002" httpUri="http://treatment.plazi.org/id/9F7C3C82D189B2C70CDCD7CE2902F53C" lastPageId="20" lastPageNumber="105" pageId="18" pageNumber="103">
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<subSubSection pageId="18" pageNumber="103" type="nomenclature">
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<paragraph pageId="18" pageNumber="103">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:5D5927C7-717D-4DC4-B855-37389C705002" authority="Crabo" class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus Crabo</taxonomicName>
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<taxonomicNameLabel pageId="18" pageNumber="103">sp. n.</taxonomicNameLabel>
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Figs 17, 41
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="103" type="type material">
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<paragraph pageId="18" pageNumber="103">Type material.</paragraph>
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<paragraph pageId="18" pageNumber="103">
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Holotype Male. Canada, Yukon Territory, Old Crow, 5.VII.1983, R. J. Cannings leg./ Malaise trap. Forest edge on S-facing bluff/ Database CNC LEP 0094163/ SLIDE
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<taxonomicName class="Insecta" family="Noctuidae" genus="Luperina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Luperina" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="genus">Luperina</taxonomicName>
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male ER8824. CNC. Paratypes None.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="103" type="etymology">
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<paragraph pageId="18" pageNumber="103">Etymology.</paragraph>
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<paragraph pageId="18" pageNumber="103">
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The name is derived from the genus of the wooly
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<normalizedToken originalValue="mammoth–">mammoth-</normalizedToken>
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<taxonomicName class="Mammalia" family="Elephantidae" genus="Mammuthus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mammuthus" order="Proboscidea" pageId="18" pageNumber="103" phylum="Chordata" rank="genus">Mammuthus</taxonomicName>
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. It is befitting of the moth because its Beringian distribution and relatively large size for the genus. It is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="103" type="diagnosis">
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<paragraph pageId="18" pageNumber="103">Diagnosis.</paragraph>
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<paragraph pageId="18" pageNumber="103">
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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is unlikely to be confused with most other species of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="genus">Resapamea</taxonomicName>
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in North America due to its northerly distribution and orange-tan color. It is superficially similar to
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea hedeni" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="hedeni">Resapamea hedeni</taxonomicName>
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(Graeser) (Fig. 18), which occurs in Asia and might also occur in Alaska (see Remarks, below). The male genitalia of these species differ in the shape of the distal uncus. It is truncated in
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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with a small T-shaped expansion at the tip (Fig. 41 inset) and tapered to a point in
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea hedeni" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="hedeni">Resapamea hedeni</taxonomicName>
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(Fig. 42) as well as in all other North American
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="genus">Resapamea</taxonomicName>
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species. The vesica of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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differs from those of all other
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="genus">Resapamea</taxonomicName>
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discussed in this paper in lacking the subbasal diverticulum and medial cornuti. It differs from that of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea hedeni" order="Lepidoptera" pageId="18" pageNumber="103" phylum="Arthropoda" rank="species" species="hedeni">Resapamea hedeni</taxonomicName>
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in lacking a subbasal serrate (
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<normalizedToken originalValue="cock’s">cock's</normalizedToken>
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comb) cornutus.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="19" lastPageNumber="104" pageId="18" pageNumber="103" type="description">
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<paragraph pageId="18" pageNumber="103">Description.</paragraph>
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<paragraph lastPageId="19" lastPageNumber="104" pageId="18" pageNumber="103">
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Head - Antenna of male nearly filiform, with slight constriction at base of each segment, covered ventrally by short fine cilia. Antenna of female unknown. Scape orange tan, with dorsal tuft. Eye rounded, smooth. Labial palp covered laterally by short flat tan scales, lengthening to a brush-like fringe on ventral surface of first two segments. Frons smooth, covered in narrow orange-tan scales. Top of head covered in long narrow orange-tan scales. Thorax - Vestiture of collar, thorax, and tegula long,
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<pageBreakToken pageId="19" pageNumber="104" start="start">narrow</pageBreakToken>
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, apically notched orange-tan scales, appearing medium-dark orange-tan [central thorax of holotype partially mildewed]. Legs light tan; with three ventral rows of spiniform setae on basitarsus and four irregular rows on other tarsal segments. Wings - Forewing length: male 21.5 mm. Forewing with a mixture of tan, orange-tan, gray-tan, light-gray, brown-gray, and gray scales, appearing medium-dark orange tan, grayer near anterior and posterior margins and darker gray-brown in terminal area; veins near costa, distal to postmedial line, and near posterior margin gray but not strongly contrasting; an ill-defined dark mark in medial area distal to lower reniform spot. Basal, antemedial, and postmedial lines faint, ill-defined dark gray with adjacent light orange tan. Basal line only evident near costa. Antemedial line evident on costa and posterior to claviform spot, forming an oblique dark mark on costa and a zigzag line from claviform spot to posterior margin. Medial line absent. Postmedial line very faint, ill-defined, smooth, strongly oblique toward base anterior to reniform spot, straight and parallel to outer margin lateral to spot, and slightly angled and concave toward base below spot to meet posterior margin at a right angle. Subterminal line light orange tan, faint, undulating; preceded by a faint indistinct shade of dark gray that is strongest opposite cell and in fold. Terminal line thin, dark gray. Orbicular spot round, outlined by ill-defined faint gray and filled with light orange tan. Reniform spot moderately large, kidney shaped with strong lateral indent, dark gray along medial and lateral sides and open anteriorly and posteriorly, filled with cream, slightly grayer at posterior end. Claviform spot black, ill defined, strong anteriorly and weak posteriorly, narrow, filled with ground color. Fringe gray tan, with a lighter tan base and gray medial line. Hindwing light gray tan with gray suffusion, very faint postmedial line, marginal band, terminal line, veins, and chevron-shaped discal spot. Hindwing fringe slightly lighter than hindwing ground color. Abdomen - tan [abdomen of holotype mildewed]. Male genitalia - Uncus cylindrical at base, evenly downcurved, distal portion slightly dorsoventrally flattened and truncated at apex with small lateral projections to appear T-shaped (Fig. 41 inset). Tegumen with large penicillus lobes. Juxta shield shaped, 0.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as high as wide, with V-shaped ventral margin. Valve S-shaped, 5.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide (measured at mid-valve), widest at base and cucullus, mid-section 2/3 as wide as base and tapering slightly to narrow neck at base of cucullus; stout sclerotized knob-like basal saccular process extending dorsolaterally from base to just dorsal to costal attachment of valve, medial margin of this process irregular and apex rounded. Sacculus reaching 2/3 of distance to costal margin and extending distally to mid-valve. Clasper a smooth ridge. Ampulla short, round. Digitus a weak ridge, partially covered by medial cucullus. Cucullus well developed with rounded apical and anal ends, 2
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<normalizedToken originalValue="×">x</normalizedToken>
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as wide as mid-valve; mesial surface covered by fine setae; corona of stout curved setae, dorsal half partially double. Aedeagus tubular, 3.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with short linear extension onto ventral vesica bearing a loose row of very small spines. Vesica 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as aedeagus, bent 135° toward right at base to project anteriorly and toward right, basal two-thirds bulbous and distal half tapering, with a single conical membranous diverticulum on anterior side of distal vesica projecting anteriorly and a subapical posterior patch of variable-sized spine-like cornuti directed basad. Female genitalia - Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="105" type="distribution">
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<paragraph pageId="20" pageNumber="105">
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<pageBreakToken pageId="20" pageNumber="105" start="start">Distribution</pageBreakToken>
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and biology.
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</paragraph>
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<paragraph pageId="20" pageNumber="105">This species is known only from the type locality at Old Crow, Yukon Territory. The habitat is described as forest edge on a south-facing hillside on the specimen label. The holotype was collected during early July. The early stages are unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="105" type="remarks">
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<paragraph pageId="20" pageNumber="105">Remarks.</paragraph>
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<paragraph pageId="20" pageNumber="105">
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Two unidentified
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="genus">Resapamea</taxonomicName>
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females in the CNC, one from Unalakleet, Alaska (Fig. 19) and the other from Reindeer Station, Aklavik, Northwest Territories (Fig. 20), resemble
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea hedeni" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="hedeni">Resapamea hedeni</taxonomicName>
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. We exclude them from the type series of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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because their identity is uncertain until either population can be associated with males or until females of the Old Crow population of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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are found. Their superficial appearances suggest that the Aklavik specimen is the female of
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea mammuthus" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="mammuthus">Resapamea mammuthus</taxonomicName>
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and that the Unalakleet specimen is
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<taxonomicName class="Insecta" family="Noctuidae" genus="Resapamea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Resapamea hedeni" order="Lepidoptera" pageId="20" pageNumber="105" phylum="Arthropoda" rank="species" species="hedeni">Resapamea hedeni</taxonomicName>
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or a closely related species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |