121 lines
10 KiB
XML
121 lines
10 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1311" ID-GBIF-Dataset="dd79f0ad-b6c3-4508-b729-703f7f09dac6" ID-Pensoft-Pub="1313-2970-130-41" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-41" ModsDocOrigin="ZooKeys 130" ModsDocTitle="New Middle Permian palaeopteran insects from Lodève Basin in southern France (Ephemeroptera, Diaphanopterodea, Megasecoptera)" checkinTime="1451249897177" checkinUser="pensoft" docAuthor="Prokop, Jakub & Nel, Andre" docDate="2011" docId="71D043E76D81F13D05C9AECAC55422FB" docLanguage="en" docName="ZooKeys 130: 41-55" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1311" docTitle="Genus Prokop & Nel, 2011, indet." docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="C647FFB9482FFFB2FFC6F623763DF04A" masterDocTitle="New Middle Permian palaeopteran insects from Lodeve Basin in southern France (Ephemeroptera, Diaphanopterodea, Megasecoptera)" masterLastPageNumber="55" masterPageNumber="41" pageNumber="42" updateTime="1643550256617" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Middle Permian palaeopteran insects from Lodeve Basin in southern France (Ephemeroptera, Diaphanopterodea, Megasecoptera)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prokop, Jakub</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nel, Andre</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>130</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>41</mods:start>
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<mods:end>55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.130.1311</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1311</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-130-41</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152030973" LSID="urn:lsid:plazi:treatment:71D043E76D81F13D05C9AECAC55422FB" httpUri="http://treatment.plazi.org/id/71D043E76D81F13D05C9AECAC55422FB" lastPageNumber="42" pageId="1" pageNumber="42">
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<subSubSection pageId="1" pageNumber="42" type="nomenclature">
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<paragraph pageId="1" pageNumber="42">
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<taxonomicName lsidName="Genus" pageId="1" pageNumber="42" rank="genus">Genus</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="42">indet.</taxonomicNameLabel>
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Figs 1
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="material">
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<paragraph pageId="1" pageNumber="42">Material.</paragraph>
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<paragraph pageId="1" pageNumber="42">
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Specimen Ld LAP 483 (Lapeyrie collection, imprint of proximal part of forewing), stored at the
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<normalizedToken originalValue="Musée">Musee</normalizedToken>
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of
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<normalizedToken originalValue="Lodève">Lodeve</normalizedToken>
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, France.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="age and locality">
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<paragraph pageId="1" pageNumber="42">Age and locality.</paragraph>
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<paragraph pageId="1" pageNumber="42">
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Middle Permian, Guadalupian,
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<normalizedToken originalValue="Lodève">Lodeve</normalizedToken>
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Basin, Salagou Formation,
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<normalizedToken originalValue="Lodève">Lodeve</normalizedToken>
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,
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<normalizedToken originalValue="Hérault">Herault</normalizedToken>
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, France (
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<bibRefCitation author="Garric, J" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle d'Autun" pageId="8" pageNumber="49" pagination="7 - 48" title="Les rigoles fossiliferes du Saxonien du bassin de Lodeve (Languedoc, France)." volume="174" year="2000">Garric 2000</bibRefCitation>
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;
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<bibRefCitation author="Bethoux, O" journalOrPublisher="Geobios" pageId="8" pageNumber="49" pagination="293 - 302" title="Discovery of the genus Iasvia Zalessky, 1934 in the Upper Permian of France (Lodeve basin) (Orthoptera: Ensifera: Oedischiidae)." volume="35" year="2002">
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<normalizedToken originalValue="Béthoux">Bethoux</normalizedToken>
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et al. 2002
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="description">
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<paragraph pageId="1" pageNumber="42">Description.</paragraph>
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<paragraph pageId="1" pageNumber="42">Counter-imprint of a fore wing without trace of preserved coloration, strongly developed corrugation of longitudinal veins; basal part of fore wing 22.7 mm long and 11.6 mm wide, estimated total length about 45 mm; area between ScP and C rather broad with numerous simple cross-veins; concave ScP straight and basally running close to radial and medial veins; RA nearly straight; RA and RP extremely approximate between wing base and a point situated 17.5 mm distally at which RP strongly diverges from RA; convex MA diverging from MP nearly at right angle and directed towards radial veins, 11.3 mm from wing base; MA distally closely parallel to RA and RP for 5.4 mm, then nearly touching RP at one point; RP emerging 6.4 mm distal of base of MA; concave MP nearly straight; convex CuA diverging from CuP close to wing base and running parallel to medial veins, CuA with two visible terminal branches; simple CuP strongly concave and only weakly curved; anal area partly preserved, first anal vein of neutral polarity ending with two main branches, second anal vein convex and distally pectinate with several branches connected by rather dense network of cells; along posterior wing margin a broad area between CuP and first anal vein; a small pentagonal elongate cell below second anal vein, near wing base.</paragraph>
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<caption pageId="1" pageNumber="42">
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<paragraph pageId="1" pageNumber="42">
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Figure 1. Genus and species indet.(
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<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Ephemeroptera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ephemeroptera" order="Malvales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">Ephemeroptera</taxonomicName>
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:
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<taxonomicName family="Syntonopteridae" lsidName="" pageId="1" pageNumber="42" rank="family">Syntonopteridae</taxonomicName>
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), specimen Ld LAP 483: A photograph of forewing B drawing of forewing (Scale bar represents 2 mm).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="42" type="discussion">
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<paragraph pageId="1" pageNumber="42">Discussion.</paragraph>
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<paragraph pageId="1" pageNumber="42">
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This fossilbears a combination of the main characters of ephemeropterid family
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<taxonomicName family="Syntonopteridae" lsidName="" pageId="1" pageNumber="42" rank="family">Syntonopteridae</taxonomicName>
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, i.e., a strong corrugation of the main longitudinal veins connected by mainly simple transverse crossveins also present in
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<taxonomicName genus="Odonatoptera" lsidName="Odonatoptera" pageId="1" pageNumber="42" rank="genus">Odonatoptera</taxonomicName>
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and some Palaeodictyopterida (e.g.,
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<taxonomicName family="Breyeriidae" lsidName="" pageId="1" pageNumber="42" rank="family">Breyeriidae</taxonomicName>
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), MA with a strong anterior curve at its base, shortly connected with RP distally; CuA with a short terminal twigging and anal area with well-defined cell(s). We can argue that this fossil is a forewing fragment for the absence of a characteristic constriction of concave vein IN- between AA1+2 and AA3+4 (
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<bibRefCitation author="Prokop, J" journalOrPublisher="Organisms Diversity & Evolution" pageId="9" pageNumber="50" pagination="331 - 340" title="On the phylogenetic position of the palaeopteran Syntonopteroidea (Insecta: Ephemeroptera), with a new species from the Upper Carboniferous of England." url="doi: 10.1007/s13127-010-0022-2" volume="10" year="2010">Prokop et al. 2010</bibRefCitation>
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). The specimen can be possibly attributed to a new genus and species for the long part of MA closely parallel to extremely approximate bases of RA and RP; a broad area between CuP and first anal vein near posterior wing margin, and CuP simple (differences with all other genera
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<taxonomicName class="Insecta" family="Lithoneuridae" genus="Lithoneura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lithoneura" order="Ephemeroptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Lithoneura</taxonomicName>
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Carpenter, 1938,
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<taxonomicName class="Insecta" family="Syntonopteridae" genus="Syntonoptera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Syntonoptera" order="Ephemeroptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Syntonoptera</taxonomicName>
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Handlirsch, 1911,
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<taxonomicName class="Insecta" family="Lithoneuridae" genus="Anglolithoneura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Anglolithoneura" order="Ephemeroptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Anglolithoneura</taxonomicName>
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Prokop & al., 2010, except maybe
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<taxonomicName class="Insecta" family="Syntonopteridae" genus="Gallolithoneura" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gallolithoneura" order="Ephemeroptera" pageId="1" pageNumber="42" phylum="Arthropoda" rank="genus">Gallolithoneura</taxonomicName>
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Garrouste et al., 2009 based on a rather incomplete wing) (
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<bibRefCitation author="Garrouste, R" journalOrPublisher="CR Palevol" pageId="9" pageNumber="50" pagination="49 - 57" title="New fossil arthropods (Notostraca and Insecta: Syntonopterida) in the Continental Middle Permian of Provence (Bas-Argens basin, France)." url="doi: 10.1016/j.crpv.2008.11.008" volume="8" year="2009">Garrouste et al. 2009</bibRefCitation>
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). However, we prefer to maintain this specimen as
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<taxonomicName family="Syntonopteridae" lsidName="" pageId="1" pageNumber="42" rank="family">Syntonopteridae</taxonomicName>
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gen. et sp. indet. for its incompleteness. Nevertheless this fossil represents the first undisputed record of mayflies from Salagou Formation in the
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<normalizedToken originalValue="Lodève">Lodeve</normalizedToken>
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Basin.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |