172 lines
19 KiB
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172 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="65B46BACD1ABB1B84A5DF1A342466211" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Pterodactylus sagittirostris Owen 1874, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="49" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="49" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044980" LSID="urn:lsid:plazi:treatment:65B46BACD1ABB1B84A5DF1A342466211" httpUri="http://treatment.plazi.org/id/65B46BACD1ABB1B84A5DF1A342466211" lastPageNumber="49" pageId="48" pageNumber="49">
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<subSubSection pageId="48" pageNumber="49" type="nomenclature">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName LSID="65B46BAC-D1AB-B1B8-4A5D-F1A342466211" authority="Owen, 1874" authorityName="Owen" authorityYear="1874" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pterodactylus sagittirostris" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Pterodactylus sagittirostris Owen, 1874</taxonomicName>
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<taxonomicNameLabel pageId="48" pageNumber="49">(nomen dubium)</taxonomicNameLabel>
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pterodactylus sagittirostris, holotype NHMUK PV R 1823 (upper Berriasian / Valangianian, Hastings Group), part of the mandibular rami. A right lateral view B respective line drawing C left lateral view D respective line drawing E ventral view. Abbreviation: d - dentary. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22088" pageId="48" pageNumber="49">Fig. 20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="reference_group">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Pterodactylus sagittirostris</taxonomicName>
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Owen:
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<bibRefCitation author="Owen, R" journalOrPublisher="Palaeontographical Society, London" pageId="68" pageNumber="69" refId="B67" refString="Owen, R, 1874. Monograph on the fossil Reptilia of the Mesozoic Formations. Palaeontographical Society, London, 14 pp." title="Monograph on the fossil Reptilia of the Mesozoic Formations." year="1874">Owen 1874</bibRefCitation>
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: p. 3, pl. II, fig 1-8.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Lonchodectes sagittirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 538.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Ornithocheirus sagittirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Arthaber, G" journalOrPublisher="Palaeontologische Zeitschrift" pageId="64" pageNumber="65" pagination="1 - 47" refId="B2" refString="Arthaber, G, 1922. Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier. Palaeontologische Zeitschrift 4: 1 - 47" title="Ueber Entwicklung, Ausbildung und Austerben der Flugsaurier." volume="4" year="1922">Arthaber 1922</bibRefCitation>
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: p. 16.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Ornithocheirus sagittirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Lonchodectes sagittirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: 209.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris" status="(nomen dubium)">Lonchodectes sagittirostris</taxonomicName>
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(Owen):
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<bibRefCitation author="Martill, DM" editor="Batten, DJ" journalOrPublisher="The Palaeontological Association, London" pageId="67" pageNumber="68" pagination="370 - 390" refId="B54" refString="Martill, DM, Sweetman, SC, Witton, MP, 2011. Pterosaurs. In: Batten, DJ, Ed., English Wealden fossils. The Palaeontological Association, London: 370 - 390" title="Pterosaurs." volumeTitle="English Wealden fossils." year="2011">Martill, Sweetman and Witton 2011</bibRefCitation>
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: p. 385, fig. 25.12
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="holotype">
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<paragraph pageId="48" pageNumber="49">Holotype.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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NHMUK PV R 1823, partial mandibular rami (
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<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Pterodactylus sagittirostris, holotype NHMUK PV R 1823 (upper Berriasian / Valangianian, Hastings Group), part of the mandibular rami. A right lateral view B respective line drawing C left lateral view D respective line drawing E ventral view. Abbreviation: d - dentary. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum." httpUri="https://binary.pensoft.net/fig/22088" pageId="48" pageNumber="49">Fig. 20A-E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type locality">
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<paragraph pageId="48" pageNumber="49">Type locality.</paragraph>
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<paragraph pageId="48" pageNumber="49">St.-Leonards-on-Sea, Sussex. England.</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="type horizon">
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<paragraph pageId="48" pageNumber="49">Type horizon.</paragraph>
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<paragraph pageId="48" pageNumber="49">Hastings Group, Wealden (late Berriasian / Valanginian).</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="description">
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<paragraph pageId="48" pageNumber="49">Description.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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The holotype of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris">
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<emphasis italics="true" pageId="48" pageNumber="49">Pterodactylus sagittirostris</emphasis>
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</taxonomicName>
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consists of partial associated mandibular rami. The specimen is not comparable to any of the species referred in
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<taxonomicName authorityName="Hooley" authorityYear="1914" class="Reptilia" family="Lonchodectidae" genus="Lonchodectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="48" pageNumber="49">Lonchodectes</emphasis>
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</taxonomicName>
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by
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley (1914)</bibRefCitation>
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and
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
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because they are mostly based on jaw tips. Its teeth are elongated and differ from the shorter ones present in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Lonchodraconidae" genus="Lonchodraco" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="giganteus">
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<emphasis italics="true" pageId="48" pageNumber="49">Lonchodraco giganteus</emphasis>
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</taxonomicName>
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, and hence
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris">
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<emphasis italics="true" pageId="48" pageNumber="49">Pterodactylus sagittirostris</emphasis>
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</taxonomicName>
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cannot be referred to
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Lonchodraconidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="family">Lonchodraconidae</taxonomicName>
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(which corresponds more or less to the
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<taxonomicName baseAuthorityName="Hooley" baseAuthorityYear="1914" class="Reptilia" family="Lonchodectidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="family">Lonchodectidae</taxonomicName>
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<emphasis italics="true" pageId="48" pageNumber="49">sensu</emphasis>
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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). An apparent elevation of the alveolar margin, forming a collar around the teeth, rather seems to be an artifact of the partial preparation of the specimen; in the right ramus, which has been more extensively prepared, this collar is, in fact, smaller and resembles the one observed in other pterosaurs, such as anhanguerids. This specimen presents no diagnostic characters or character combination, and therefore
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris">
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<emphasis italics="true" pageId="48" pageNumber="49">Pterodactylus sagittirostris</emphasis>
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</taxonomicName>
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is considered a
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<emphasis italics="true" pageId="48" pageNumber="49">nomen dubium</emphasis>
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.
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</paragraph>
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<caption httpUri="https://binary.pensoft.net/fig/22088" pageId="48" pageNumber="49" start="Figure 20" startId="F20">
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<paragraph pageId="48" pageNumber="49">
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<emphasis bold="true" pageId="48" pageNumber="49">Figure 20.</emphasis>
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Pterodactylidae" genus="Pterodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="sagittirostris">
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<emphasis italics="true" pageId="48" pageNumber="49">Pterodactylus sagittirostris</emphasis>
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</taxonomicName>
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, holotype NHMUK PV R 1823 (upper Berriasian / Valangianian, Hastings Group), part of the mandibular rami.
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<emphasis bold="true" pageId="48" pageNumber="49">A</emphasis>
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right lateral view
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<emphasis bold="true" pageId="48" pageNumber="49">B</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="48" pageNumber="49">C</emphasis>
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left lateral view
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<emphasis bold="true" pageId="48" pageNumber="49">D</emphasis>
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respective line drawing
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<emphasis bold="true" pageId="48" pageNumber="49">E</emphasis>
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ventral view. Abbreviation:
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<emphasis bold="true" pageId="48" pageNumber="49">d</emphasis>
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- dentary. Arrows indicate alveoli or teeth. Scale bar = 10 mm. Photos courtesy of The Natural History Museum.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |