937 lines
112 KiB
XML
937 lines
112 KiB
XML
<document id="05912E02B7514F3140443AF92B05450A" ID-DOI="10.5281/zenodo.177415" ID-GBIF-Dataset="1407b573-76bc-4071-b12d-eb974eadfa2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="177415" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459860034900" checkinUser="plazi" docAuthor="Loiselle, Paul V. & Rodriguez, Damaris" docDate="2007" docId="2F6C5E66FF9DCC1F70E1FE173381FCD5" docLanguage="en" docName="zt01520p018.pdf" docOrigin="Zootaxa 1520" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Bedotia geayi Pellegrin 1907" docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="D355261EFF9ACC107076FFB83037FFE7" masterDocTitle="A new species of Bedotia (Teleostei: Atherinomorpha: Bedotiidae) from the Rianila drainage of Eastern Madagascar, with redescriptions of Bedotia madagascariensis and Bedotia geayi" masterLastPageNumber="18" masterPageNumber="1" pageNumber="8" updateTime="1698225974792" updateUser="plazi">
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<mods:title id="1C10BB7B51BD3DAC304E8A7F7AF26B11">A new species of Bedotia (Teleostei: Atherinomorpha: Bedotiidae) from the Rianila drainage of Eastern Madagascar, with redescriptions of Bedotia madagascariensis and Bedotia geayi</mods:title>
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<mods:namePart id="3D095E12E47628C41B35810B4AE37DF3">Loiselle, Paul V.</mods:namePart>
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<mods:namePart id="71DAB804CBF85C61C164CC84F5C8A525">Rodriguez, Damaris</mods:namePart>
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<mods:typeOfResource id="BE94DACDD03825B4644D6AA6171662E5">text</mods:typeOfResource>
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<mods:title id="A5D9F8726B012859E3EF98F65740BF57">Zootaxa</mods:title>
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<mods:part id="DAA5603CC7CA303A48A4B872C5E2874B">
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<mods:date id="8AB05B44E209FCA9077ECED2424F4E1D">2007</mods:date>
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<mods:number id="E8CC9A8BE7710FFF282CA3C511054580">1520</mods:number>
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<mods:identifier id="27287F695A71C1AD92FFCB2D95D92E31" type="DOI">10.5281/zenodo.177415</mods:identifier>
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<mods:identifier id="362FE7441695E160EA859DDDEEBFD0F9" type="ISSN">1175-5326</mods:identifier>
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<treatment id="2F6C5E66FF9DCC1F70E1FE173381FCD5" ID-DOI="http://doi.org/10.5281/zenodo.6241663" ID-GBIF-Taxon="119368057" ID-Zenodo-Dep="6241663" LSID="urn:lsid:plazi:treatment:2F6C5E66FF9DCC1F70E1FE173381FCD5" httpUri="http://treatment.plazi.org/id/2F6C5E66FF9DCC1F70E1FE173381FCD5" lastPageId="15" lastPageNumber="16" pageId="7" pageNumber="8">
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<subSubSection id="EFDFBCFBFF9DCC1770E1FE1731C3FE2E" box="[151,500,431,457]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="A77AEF70FF9DCC1770E1FE1731C3FE2E" blockId="7.[151,500,431,457]" box="[151,500,431,457]" pageId="7" pageNumber="8">
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<heading id="FC32581CFF9DCC1770E1FE1731C3FE2E" bold="true" box="[151,500,431,457]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="60C594F3FF9DCC1770E1FE1731C3FE2E" ID-CoL="68FQ3" authority="Pellegrin 1907" authorityName="Pellegrin" authorityYear="1907" box="[151,500,431,457]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="geayi">
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<emphasis id="95B13362FF9DCC1770E1FE1731C3FE2E" bold="true" box="[151,500,431,457]" pageId="7" pageNumber="8">
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<emphasis id="95B13362FF9DCC1770E1FE17310CFE2E" bold="true" box="[151,315,431,457]" italics="true" pageId="7" pageNumber="8">Bedotia geayi</emphasis>
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<bibRefCitation id="C3549281FF9DCC177134FE1731C3FE2E" author="Pellegrin" box="[322,500,431,457]" pageId="7" pageNumber="18" refString="Pellegrin, J. (1907) Liste des poissons recueillis a Madagascar par M. F. Geay. Description d'une espece nouvelle. Bulletin du Musee Nationale d' Histoire Naturelle, 30, 201 - 206." type="journal article" year="1907">Pellegrin 1907</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFDFBCFBFF9DCC1770E1FE423109FD96" pageId="7" pageNumber="8" type="reference_group">
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<paragraph id="A77AEF70FF9DCC1770E1FE423109FD96" blockId="7.[151,1436,506,625]" pageId="7" pageNumber="8">
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<treatmentCitationGroup id="87D5C85EFF9DCC1770E1FE423109FD96" infered="true" pageId="7" pageNumber="8">
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<taxonomicName id="60C594F3FF9DCC1770E1FE42323CFDF6" ID-CoL="68FQ3" authority="Pellegrin 1907: 205" authorityName="Pellegrin" authorityPageNumber="205" authorityYear="1907" box="[151,523,506,529]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="geayi">
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<emphasis id="95B13362FF9DCC1770E1FE42311BFDF6" box="[151,300,506,529]" italics="true" pageId="7" pageNumber="8">Bedotia geayi</emphasis>
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<treatmentCitation id="2664C961FF9DCC177145FE42323CFDF6" author="Pellegrin" box="[307,523,506,529]" page="205" pageId="7" pageNumber="8" year="1907">
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<bibRefCitation id="C3549281FF9DCC177145FE4231E2FDF6" author="Pellegrin" box="[307,469,506,529]" pageId="7" pageNumber="18" refString="Pellegrin, J. (1907) Liste des poissons recueillis a Madagascar par M. F. Geay. Description d'une espece nouvelle. Bulletin du Musee Nationale d' Histoire Naturelle, 30, 201 - 206." type="journal article" year="1907">Pellegrin 1907</bibRefCitation>
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: 205
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</treatmentCitation>
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</taxonomicName>
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. Morafeno, dans les placers, à une altitude de
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<quantity id="603D4295FF9DCC177474FE423471FDF6" box="[1026,1094,506,529]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="m" value="300.0">300 m</quantity>
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environs, aux sources des ruisseaux de la Haute-Maha, affluent du Bas-Mananjary. (Morafeno, at approximately
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<quantity id="603D4295FF9DCC177442FDA23440FDD6" box="[1076,1143,538,561]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="m" value="300.0">300 m</quantity>
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altitude, over sand bottoms at the sources of the headwaters of the Upper Maha, a tributary of the Lower Mananjary).
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<typeStatus id="787E51D2FF9DCC1774EAFD823535FDB6" box="[1180,1282,570,593]" pageId="7" pageNumber="8" type="syntype">Syntypes</typeStatus>
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: MNHN 1907 35-37 (11).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="EFDFBCFBFF9DCC1970E1FD1A3179F90D" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="A77AEF70FF9DCC1770E1FD1A313BFD5B" blockId="7.[151,1437,674,940]" box="[151,268,674,700]" pageId="7" pageNumber="8">Diagnosis</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0FD72357CFC4B" blockId="7.[151,1437,674,940]" pageId="7" pageNumber="8">
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<taxonomicName id="60C594F3FF9DCC1770B0FD723150FD04" box="[198,359,714,739]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="geayi">
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<emphasis id="95B13362FF9DCC1770B0FD723150FD04" box="[198,359,714,739]" italics="true" pageId="7" pageNumber="8">Bedotia geayi</emphasis>
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</taxonomicName>
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of both sexes can be distinguished from laterally striped congeners by the presence of a discrete black basal spot on the caudal fin base in both living and preserved specimens. Living males have red dorsal and anal fin margins and a red caudal fin, as well as a large red spot on the chin, while a narrow, more or less well defined black margin is present in the caudal fin of preserved individuals. Elevated second dorsal [11–14 (mode: 12)] and anal [17–19 (mode: 18)] ray counts are likewise diagnostic for preserved material. Morphometric characters that set this species apart from
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<taxonomicName id="60C594F3FF9DCC177346FC2A3413FC4C" box="[816,1060,914,939]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="madagascariensis">
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<emphasis id="95B13362FF9DCC177346FC2A3413FC4C" box="[816,1060,914,939]" italics="true" pageId="7" pageNumber="8">B. madagascariensis</emphasis>
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</taxonomicName>
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are presented in
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<tableCitation id="EA47DACBFF9DCC17749AFC2A3572FC4B" box="[1260,1349,914,940]" captionStart="TABLE 3" captionStartId="10.[151,239,431,455]" captionTargetBox="[167,1331,541,991]" captionTargetPageId="10" captionText="TABLE 3. Summary of Distinguishing Characters for Bedotia madagascariensis and B. geayi." httpUri="http://table.plazi.org/id/F3BABFF8FF90CC1A70E1FE173452FE20" pageId="7" pageNumber="8" tableUuid="F3BABFF8FF90CC1A70E1FE173452FE20">Table 3</tableCitation>
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.
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</paragraph>
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<paragraph id="A77AEF70FF9DCC1770E1FBB23117FBC3" blockId="7.[151,1437,1034,1980]" box="[151,288,1034,1060]" pageId="7" pageNumber="8">Description</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0FB8A30F3FAF3" blockId="7.[151,1437,1034,1980]" pageId="7" pageNumber="8">
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Morphological measurements and meristic counts are given in
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<tableCitation id="EA47DACBFF9DCC1773DFFB8A3434FBAB" box="[937,1027,1074,1100]" captionStart="TABLE 2" captionStartId="8.[151,239,151,175]" captionTargetBox="[153,1403,213,1007]" captionTargetPageId="8" captionText="TABLE 2. Morphometric data for Bedotia geayi as % SL." httpUri="http://table.plazi.org/id/F3BABFF8FF92CC1870E1FF2F333CFF48" pageId="7" pageNumber="8" tableUuid="F3BABFF8FF92CC1870E1FF2F333CFF48">Table 2</tableCitation>
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||
. The largest specimen examined is a 74.0 mm SL male.
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<taxonomicName id="60C594F3FF9DCC1771E7FBE33204FB93" box="[401,563,1115,1140]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="geayi">
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<emphasis id="95B13362FF9DCC1771E7FBE33204FB93" box="[401,563,1115,1140]" italics="true" pageId="7" pageNumber="8">Bedotia geayi</emphasis>
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</taxonomicName>
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||
are gracile, relatively long-bodied fishes somewhat deeper-bodied anteriorly and showing a rather straight ventral outline. Dorsal outline of head and nape moderately curved to first dorsal fin. Head length divisible 3.3-4.5 times in the standard length. First dorsal fin origin is posterior to the vertical through pelvic-fin insertion, while that of second is posterior to the vertical through the anal fin origin.
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</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0FA9A34A7FA53" blockId="7.[151,1437,1034,1980]" pageId="7" pageNumber="8">
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Snout slightly indented behind premaxillary pedicels. Snout length divisible 2.9–4.1 times in the head length. Lower jaw is slightly prognathous and angled at about 40°–45° to horizontal when mouth is closed. Premaxilla and maxilla reach the anterior margin of the orbit. Premaxilla with a distinct lateral "
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<taxonomicName id="60C594F3FF9DCC177537FACB35ABFA6B" box="[1345,1436,1395,1420]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
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<emphasis id="95B13362FF9DCC177537FACB35ABFA6B" box="[1345,1436,1395,1420]" italics="true" pageId="7" pageNumber="8">Bedotia</emphasis>
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</taxonomicName>
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notch". Orbital diameter divisible
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<quantity id="603D4295FF9DCC177251FA22328DFA53" box="[551,698,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="8.4455" metricValueMax="9.5758" metricValueMin="7.3152" pageId="7" pageNumber="8" unit="in" value="3.325" valueMax="3.77" valueMin="2.88">2.88–3.77 in</quantity>
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the head,
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<quantity id="603D4295FF9DCC177344FA2233F1FA53" box="[818,966,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="2.4638" metricValueMax="2.5654" metricValueMin="2.3622" pageId="7" pageNumber="8" unit="in" value="0.97" valueMax="1.01" valueMin="0.93">0.93–1.01 in</quantity>
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the snout length.
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</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0FA7A3440F99B" blockId="7.[151,1437,1034,1980]" pageId="7" pageNumber="8">
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Teeth. Anteriorly both upper and lower jaws bear 4 to 6 rows of numerous small, strongly recurved unicuspid teeth. The outermost row of teeth is poorly differentiated from those of the inner band. The lower jaw and the premaxilla posterior to the
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<taxonomicName id="60C594F3FF9DCC177237F9AB32ABF9CB" box="[577,668,1555,1580]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
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<emphasis id="95B13362FF9DCC177237F9AB32ABF9CB" box="[577,668,1555,1580]" italics="true" pageId="7" pageNumber="8">Bedotia</emphasis>
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</taxonomicName>
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notch each have a single row of teeth. A single row of teeth is present along the anteroventral face of vomer. Small patches of endopterygoid teeth are also present. No ectopterygoid teeth present, at least in individuals of sizes available for examination.
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</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0F93234DCF92B" blockId="7.[151,1437,1034,1980]" pageId="7" pageNumber="8">Gill Rakers. Two or three stout hypobranchial rakers and 10-13 (mode: 11) elongate ceratobranchial rakers are present on the lower limb of the first branchial arch. All rakers are strongly denticulate.</paragraph>
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<paragraph id="A77AEF70FF9DCC1770B0F9623226F85B" blockId="7.[151,1437,1034,1980]" pageId="7" pageNumber="8">Scales. Body is fully covered with large, regularly imbricate, cycloid scales. Predorsal scales along the dorsal midline: 13–14 (mode: 14). Scales along the midlateral axis from just behind the operculum, above the pectoral fin, to the end of the hypural plate: 31–35 (modes: 32, 33). Scales in transverse series between the origins of the anal and the second dorsal fin (including a very small scale adjacent to each fin): 9. Scales separating the first and second dorsal fins: 3. Circumpeduncular scales: 12. Dorsal, anal, and caudal scale sheaths and axillary pelvic scales are absent.</paragraph>
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<caption id="F3BABFF8FF92CC1870E1FF2F333CFF48" ID-Table-UUID="F3BABFF8FF92CC1870E1FF2F333CFF48" box="[151,779,151,175]" httpUri="http://table.plazi.org/id/F3BABFF8FF92CC1870E1FF2F333CFF48" pageId="8" pageNumber="9" targetBox="[153,1403,213,1007]" targetIsTable="true" targetPageId="8">
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<paragraph id="A77AEF70FF92CC1870E1FF2F333CFF48" blockId="8.[151,779,151,175]" box="[151,779,151,175]" pageId="8" pageNumber="9">
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||
<emphasis id="95B13362FF92CC1870E1FF2F313FFF48" bold="true" box="[151,264,151,175]" pageId="8" pageNumber="9">TABLE 2.</emphasis>
|
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Morphometric data for
|
||
<taxonomicName id="60C594F3FF92CC187267FF203293FF48" box="[529,676,152,175]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="geayi">
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||
<emphasis id="95B13362FF92CC187267FF203293FF48" box="[529,676,152,175]" italics="true" pageId="8" pageNumber="9">Bedotia geayi</emphasis>
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||
</taxonomicName>
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||
as % SL.
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||
</paragraph>
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||
</caption>
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<paragraph id="A77AEF70FF92CC18710CFF6D354CFC08" pageId="8" pageNumber="9">
|
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<table id="D5C51DD0FF9233EF70EFFF6D354CFC08" box="[153,1403,213,1007]" gridcols="5" gridrows="17" pageId="8" pageNumber="9">
|
||
<tr id="19F5ED32FF9233EF70EFFF6D354CFF0A" box="[153,1403,213,237]" gridrow="0" pageId="8" pageNumber="9">
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<th id="5A24844EFF9233EF70EFFF6D31D8FF0A" box="[153,495,213,237]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">
|
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<emphasis id="95B13362FF92CC18710CFF6D31D8FF0A" bold="true" box="[378,495,213,237]" pageId="8" pageNumber="9">Character</emphasis>
|
||
</th>
|
||
<th id="5A24844EFF9233EF7288FF6D332EFF0A" box="[766,793,213,237]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis id="95B13362FF92CC187377FF6D3323FF0A" bold="true" box="[769,788,213,237]" pageId="8" pageNumber="9">N</emphasis>
|
||
</th>
|
||
<th id="5A24844EFF9233EF73D2FF6D3438FF0A" box="[932,1039,213,237]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
|
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<emphasis id="95B13362FF92CC1873C3FF6D33CAFF0A" bold="true" box="[949,1021,213,237]" pageId="8" pageNumber="9">Range</emphasis>
|
||
</th>
|
||
<th id="5A24844EFF9233EF74D8FF6D34D8FF0A" box="[1198,1263,213,237]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis id="95B13362FF92CC1874D8FF6D34D8FF0A" bold="true" box="[1198,1263,213,237]" pageId="8" pageNumber="9">Mean</emphasis>
|
||
</th>
|
||
<th id="5A24844EFF9233EF752FFF6D354CFF0A" box="[1369,1403,213,237]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">
|
||
<emphasis id="95B13362FF92CC18752FFF6D354CFF0A" bold="true" box="[1369,1403,213,237]" pageId="8" pageNumber="9">SD</emphasis>
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||
</th>
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</tr>
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<tr id="19F5ED32FF9233EF70EFFEBB354CFEFD" box="[153,1403,259,282]" gridrow="1" pageId="8" pageNumber="9" rowspan-3="1" rowspan-4="1">
|
||
<th id="5A24844EFF9233EF70EFFEBB31D8FEFD" box="[153,495,259,282]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">Standard length [mm]</th>
|
||
<td id="5A24844EFF9233EF7288FEBB332EFEFD" box="[766,793,259,282]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FEBB3438FEFD" box="[932,1039,259,282]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">43.9–74.0</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFE8B354CFEAD" box="[153,1403,307,330]" gridrow="2" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFE8B31D8FEAD" box="[153,495,307,330]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">Head length</th>
|
||
<td id="5A24844EFF9233EF7288FE8B332EFEAD" box="[766,793,307,330]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FE8B3438FEAD" box="[932,1039,307,330]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">22.0–29.8</td>
|
||
<td id="5A24844EFF9233EF74D8FE8B34D8FEAD" box="[1198,1263,307,330]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">27.8</td>
|
||
<td id="5A24844EFF9233EF752FFE8B354CFEAD" box="[1369,1403,307,330]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">2.0</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFED9354CFE9F" box="[153,1403,353,376]" gridrow="3" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFED931D8FE9F" box="[153,495,353,376]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">Snout length</th>
|
||
<td id="5A24844EFF9233EF7288FED9332EFE9F" box="[766,793,353,376]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FED93438FE9F" box="[932,1039,353,376]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">7.9–10.2</td>
|
||
<td id="5A24844EFF9233EF74D8FED934D8FE9F" box="[1198,1263,353,376]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">8.5</td>
|
||
<td id="5A24844EFF9233EF752FFED9354CFE9F" box="[1369,1403,353,376]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">0.2</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFE36354CFE42" box="[153,1403,398,421]" gridrow="4" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFE3631D8FE42" box="[153,495,398,421]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Lower jaw length</th>
|
||
<td id="5A24844EFF9233EF7288FE36332EFE42" box="[766,793,398,421]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FE363438FE42" box="[932,1039,398,421]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">13.4–16.2</td>
|
||
<td id="5A24844EFF9233EF74D8FE3634D8FE42" box="[1198,1263,398,421]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">14.6</td>
|
||
<td id="5A24844EFF9233EF752FFE36354CFE42" box="[1369,1403,398,421]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">0.9</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFE04354CFE34" box="[153,1403,444,467]" gridrow="5" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFE0431D8FE34" box="[153,495,444,467]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">Orbital diameter</th>
|
||
<td id="5A24844EFF9233EF7288FE04332EFE34" box="[766,793,444,467]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FE043438FE34" box="[932,1039,444,467]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">7.6–9.6</td>
|
||
<td id="5A24844EFF9233EF74D8FE0434D8FE34" box="[1198,1263,444,467]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">8.6</td>
|
||
<td id="5A24844EFF9233EF752FFE04354CFE34" box="[1369,1403,444,467]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">0.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFE51354CFDE7" box="[153,1403,489,512]" gridrow="6" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFE5131D8FDE7" box="[153,495,489,512]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">Interorbital distance</th>
|
||
<td id="5A24844EFF9233EF7288FE51332EFDE7" box="[766,793,489,512]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FE513438FDE7" box="[932,1039,489,512]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">9.7–11.6</td>
|
||
<td id="5A24844EFF9233EF74D8FE5134D8FDE7" box="[1198,1263,489,512]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">10.8</td>
|
||
<td id="5A24844EFF9233EF752FFE51354CFDE7" box="[1369,1403,489,512]" gridcol="4" gridrow="6" pageId="8" pageNumber="9">0.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFDAE354CFDCA" box="[153,1403,534,557]" gridrow="7" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFDAE31D8FDCA" box="[153,495,534,557]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">Pectoral fin length</th>
|
||
<td id="5A24844EFF9233EF7288FDAE332EFDCA" box="[766,793,534,557]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FDAE3438FDCA" box="[932,1039,534,557]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">10.4–20.3</td>
|
||
<td id="5A24844EFF9233EF74D8FDAE34D8FDCA" box="[1198,1263,534,557]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">15.9</td>
|
||
<td id="5A24844EFF9233EF752FFDAE354CFDCA" box="[1369,1403,534,557]" gridcol="4" gridrow="7" pageId="8" pageNumber="9">3.4</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFDFC354CFD84" box="[153,1403,580,611]" gridrow="8" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFDFC31D8FD84" box="[153,495,580,611]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">
|
||
Dorsal1
|
||
<emphasis id="95B13362FF92CC187087FDFC30C9FDBB" bold="true" box="[241,254,580,604]" pageId="8" pageNumber="9">–</emphasis>
|
||
anal fin depth
|
||
</th>
|
||
<td id="5A24844EFF9233EF7288FDFC332EFD84" box="[766,793,580,611]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">14</td>
|
||
<td id="5A24844EFF9233EF73D2FDFC3438FD84" box="[932,1039,580,611]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">18.4–25.4</td>
|
||
<td id="5A24844EFF9233EF74D8FDFC34D8FD84" box="[1198,1263,580,611]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">23.0</td>
|
||
<td id="5A24844EFF9233EF752FFDFC354CFD84" box="[1369,1403,580,611]" gridcol="4" gridrow="8" pageId="8" pageNumber="9">2.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFDC1354CFD7E" box="[153,1403,633,665]" gridrow="9" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFDC131D8FD7E" box="[153,495,633,665]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">
|
||
Dorsal2
|
||
<emphasis id="95B13362FF92CC187087FDC130C9FD76" bold="true" box="[241,254,633,657]" pageId="8" pageNumber="9">–</emphasis>
|
||
anal fin depth
|
||
</th>
|
||
<td id="5A24844EFF9233EF7288FDC1332EFD7E" box="[766,793,633,665]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">14</td>
|
||
<td id="5A24844EFF9233EF73D2FDC13438FD7E" box="[932,1039,633,665]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">19.9–32.2</td>
|
||
<td id="5A24844EFF9233EF74D8FDC134D8FD7E" box="[1198,1263,633,665]" gridcol="3" gridrow="9" pageId="8" pageNumber="9">24.5</td>
|
||
<td id="5A24844EFF9233EF752FFDC1354CFD7E" box="[1369,1403,633,665]" gridcol="4" gridrow="9" pageId="8" pageNumber="9">3.2</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFD17354CFD29" box="[153,1403,687,718]" gridrow="10" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFD1731D8FD29" box="[153,495,687,718]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">Pre-dorsal1 distance</th>
|
||
<td id="5A24844EFF9233EF7288FD17332EFD29" box="[766,793,687,718]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FD173438FD29" box="[932,1039,687,718]" gridcol="2" gridrow="10" pageId="8" pageNumber="9">52.4–58.7</td>
|
||
<td id="5A24844EFF9233EF74D8FD1734D8FD29" box="[1198,1263,687,718]" gridcol="3" gridrow="10" pageId="8" pageNumber="9">54.5</td>
|
||
<td id="5A24844EFF9233EF752FFD17354CFD29" box="[1369,1403,687,718]" gridcol="4" gridrow="10" pageId="8" pageNumber="9">1.7</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFD5C354CFCE3" box="[153,1403,740,772]" gridrow="11" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFD5C31D8FCE3" box="[153,495,740,772]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">Pre-dorsal 2 distance</th>
|
||
<td id="5A24844EFF9233EF7288FD5C332EFCE3" box="[766,793,740,772]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FD5C3438FCE3" box="[932,1039,740,772]" gridcol="2" gridrow="11" pageId="8" pageNumber="9">64.7–69.2</td>
|
||
<td id="5A24844EFF9233EF74D8FD5C34D8FCE3" box="[1198,1263,740,772]" gridcol="3" gridrow="11" pageId="8" pageNumber="9">66.9</td>
|
||
<td id="5A24844EFF9233EF752FFD5C354CFCE3" box="[1369,1403,740,772]" gridcol="4" gridrow="11" pageId="8" pageNumber="9">1.7</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFCA2354CFCD6" box="[153,1403,794,817]" gridrow="12" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFCA231D8FCD6" box="[153,495,794,817]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">Pre-pelvic distance</th>
|
||
<td id="5A24844EFF9233EF7288FCA2332EFCD6" box="[766,793,794,817]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FCA23438FCD6" box="[932,1039,794,817]" gridcol="2" gridrow="12" pageId="8" pageNumber="9">40.9–46.5</td>
|
||
<td id="5A24844EFF9233EF74D8FCA234D8FCD6" box="[1198,1263,794,817]" gridcol="3" gridrow="12" pageId="8" pageNumber="9">43.7</td>
|
||
<td id="5A24844EFF9233EF752FFCA2354CFCD6" box="[1369,1403,794,817]" gridcol="4" gridrow="12" pageId="8" pageNumber="9">1.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFCF0354CFC80" box="[153,1403,840,871]" gridrow="13" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFCF031D8FC80" box="[153,495,840,871]" gridcol="0" gridrow="13" pageId="8" pageNumber="9">Base dorsal2</th>
|
||
<td id="5A24844EFF9233EF7288FCF0332EFC80" box="[766,793,840,871]" gridcol="1" gridrow="13" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FCF03438FC80" box="[932,1039,840,871]" gridcol="2" gridrow="13" pageId="8" pageNumber="9">16.9–21.4</td>
|
||
<td id="5A24844EFF9233EF74D8FCF034D8FC80" box="[1198,1263,840,871]" gridcol="3" gridrow="13" pageId="8" pageNumber="9">19.4</td>
|
||
<td id="5A24844EFF9233EF752FFCF0354CFC80" box="[1369,1403,840,871]" gridcol="4" gridrow="13" pageId="8" pageNumber="9">1.3</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFCC5354CFC73" box="[153,1403,893,916]" gridrow="14" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFCC531D8FC73" box="[153,495,893,916]" gridcol="0" gridrow="14" pageId="8" pageNumber="9">Base anal</th>
|
||
<td id="5A24844EFF9233EF7288FCC5332EFC73" box="[766,793,893,916]" gridcol="1" gridrow="14" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FCC53438FC73" box="[932,1039,893,916]" gridcol="2" gridrow="14" pageId="8" pageNumber="9">25.1–30.0</td>
|
||
<td id="5A24844EFF9233EF74D8FCC534D8FC73" box="[1198,1263,893,916]" gridcol="3" gridrow="14" pageId="8" pageNumber="9">27.2</td>
|
||
<td id="5A24844EFF9233EF752FFCC5354CFC73" box="[1369,1403,893,916]" gridcol="4" gridrow="14" pageId="8" pageNumber="9">1.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFC13354CFC25" box="[153,1403,939,962]" gridrow="15" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFC1331D8FC25" box="[153,495,939,962]" gridcol="0" gridrow="15" pageId="8" pageNumber="9">Length caudal peduncle</th>
|
||
<td id="5A24844EFF9233EF7288FC13332EFC25" box="[766,793,939,962]" gridcol="1" gridrow="15" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FC133438FC25" box="[932,1039,939,962]" gridcol="2" gridrow="15" pageId="8" pageNumber="9">13.9–17.7</td>
|
||
<td id="5A24844EFF9233EF74D8FC1334D8FC25" box="[1198,1263,939,962]" gridcol="3" gridrow="15" pageId="8" pageNumber="9">16.2</td>
|
||
<td id="5A24844EFF9233EF752FFC13354CFC25" box="[1369,1403,939,962]" gridcol="4" gridrow="15" pageId="8" pageNumber="9">1.2</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9233EF70EFFC60354CFC08" box="[153,1403,984,1007]" gridrow="16" pageId="8" pageNumber="9">
|
||
<th id="5A24844EFF9233EF70EFFC6031D8FC08" box="[153,495,984,1007]" gridcol="0" gridrow="16" pageId="8" pageNumber="9">Depth caudal peduncle</th>
|
||
<td id="5A24844EFF9233EF7288FC60332EFC08" box="[766,793,984,1007]" gridcol="1" gridrow="16" pageId="8" pageNumber="9">17</td>
|
||
<td id="5A24844EFF9233EF73D2FC603438FC08" box="[932,1039,984,1007]" gridcol="2" gridrow="16" pageId="8" pageNumber="9">9.0–15.1</td>
|
||
<td id="5A24844EFF9233EF74D8FC6034D8FC08" box="[1198,1263,984,1007]" gridcol="3" gridrow="16" pageId="8" pageNumber="9">10.8</td>
|
||
<td id="5A24844EFF9233EF752FFC60354CFC08" box="[1369,1403,984,1007]" gridcol="4" gridrow="16" pageId="8" pageNumber="9">1.4</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870B0FBE8340DFB05" blockId="8.[151,1437,1104,1450]" pageId="8" pageNumber="9">Fins. First dorsal fin with 5 weak spines. Second dorsal fin rays: 11–14 (mode: 12), the first 4 or 5 unbranched. Anal fin rays 17–19 (mode: 18), usually the first 3 or 4 unbranched. Pectoral fins short, high-set with 12 rays, the longest barely reaching the vertical from the pelvic fin insertion. Pelvic fins with one weak spine and five strongly bifurcate, branched rays. Caudal fin weakly emarginate.</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870B0FB4834CEFAD5" blockId="8.[151,1437,1104,1450]" pageId="8" pageNumber="9">Vertebrae. Total vertebral count taken from radiographs: 34–37 (mode: 36), and a terminal, hypural-bearing half centrum. Pre-caudal vertebrae: 18–20 (mode: 19). Caudal vertebrae: 15–17 (mode: 17).</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870B0FAF83481FA4D" blockId="8.[151,1437,1104,1450]" pageId="8" pageNumber="9">Viscera and Diet. Gut extremely short, intestinal length only about one-third body length. Examination of feces produced by newly caught specimens within two to four hours of capture revealed the remains of terrestrial insects, suggesting that this species relies primarily upon allochthonous food sources.</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870E1FA583122FA1D" blockId="8.[151,1436,1504,1930]" box="[151,277,1504,1530]" pageId="8" pageNumber="9">Coloration</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870B7F9B032EBF925" blockId="8.[151,1436,1504,1930]" pageId="8" pageNumber="9">
|
||
Living specimens:
|
||
<figureCitation id="3FFEF3F5FF92CC1871E9F9B03235F9C5" box="[415,514,1544,1570]" captionStart="FIGURE 4" captionStartId="9.[151,255,589,613]" captionTargetBox="[151,1434,181,570]" captionTargetId="figure@9.[151,1436,179,571]" captionTargetPageId="9" captionText="FIGURE 4. Sexually quiescent wild-caught male Bedotia geayi, Andranomaintso Creek. Not preserved." httpUri="https://zenodo.org/record/177418/files/figure.png" pageId="8" pageNumber="9">Figure 4</figureCitation>
|
||
depicts a sexually quiescent male
|
||
<emphasis id="95B13362FF92CC1873E3F9B03409F9C6" box="[917,1086,1544,1569]" italics="true" pageId="8" pageNumber="9">
|
||
<taxonomicName id="60C594F3FF92CC1873E3F9B0340DF9C6" box="[917,1082,1544,1569]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="geayi">Bedotia geayi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It does not show the diagnostic large red spot on the chin. The pectorals are hyaline in both sexes, but the color pattern of the unpaired fins and ventrals is sexually dimorphic.
|
||
<figureCitation id="3FFEF3F5FF92CC187237F9E03291F995" box="[577,678,1624,1650]" captionStart="FIGURE 5" captionStartId="9.[151,255,1189,1213]" captionTargetBox="[151,1434,701,1130]" captionTargetId="figure@9.[151,1436,701,1130]" captionTargetPageId="9" captionText="FIGURE 5. Adult wild-caught female Bedotia geayi, Andranomaintso Creek. Not preserved." httpUri="https://zenodo.org/record/177419/files/figure.png" pageId="8" pageNumber="9">Figure 5</figureCitation>
|
||
depicts an adult female. The clear yellow halo surrounding the black spot at the base of the caudal is diagnostic. None of the populations to date sampled is characterized by polymorphism with regard to male fin coloration.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF92CC1870CBF9683299F86D" blockId="8.[151,1436,1504,1930]" pageId="8" pageNumber="9">
|
||
Preserved specimens: Color pattern of the body as in
|
||
<emphasis id="95B13362FF92CC187347F968341AF90E" box="[817,1069,1744,1769]" italics="true" pageId="8" pageNumber="9">
|
||
<taxonomicName id="60C594F3FF92CC187347F968341EF90E" box="[817,1065,1744,1769]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="madagascariensis">B. madagascariensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but the midlateral stripe terminates in a distinct black spot on the base of the caudal fin. Both dorsal fins and ventrals clear grey in males, hyaline in females. Anal in males clear grey, often with a narrow black edging along its posterior half, entirely hyaline in females. Caudal uniform clear grey with a variably present narrow black distal margin in males, hyaline basally, clear grey distally in females.
|
||
</paragraph>
|
||
<caption id="F3BABFF8FF93CC1970E1FDF534C4FD83" box="[151,1267,589,613]" httpUri="https://zenodo.org/record/177418/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1434,181,570]" targetPageId="9">
|
||
<paragraph id="A77AEF70FF93CC1970E1FDF534C4FD83" blockId="9.[151,1267,589,613]" box="[151,1267,589,613]" pageId="9" pageNumber="10">
|
||
<emphasis id="95B13362FF93CC1970E1FDF5312EFD82" bold="true" box="[151,281,589,613]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
|
||
Sexually quiescent wild-caught male
|
||
<taxonomicName id="60C594F3FF93CC1972D9FDF5344AFD83" authority="Andranomaintso Creek. Not" authorityName="Andranomaintso Creek. Not" box="[687,1149,589,612]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC1972D9FDF53374FD83" box="[687,835,589,612]" italics="true" pageId="9" pageNumber="10">Bedotia geayi</emphasis>
|
||
, Andranomaintso Creek. Not
|
||
</taxonomicName>
|
||
preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F3BABFF8FF93CC1970E1FB1D34B4FB5B" box="[151,1155,1189,1213]" httpUri="https://zenodo.org/record/177419/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1434,701,1130]" targetPageId="9">
|
||
<paragraph id="A77AEF70FF93CC1970E1FB1D34B4FB5B" blockId="9.[151,1155,1189,1213]" box="[151,1155,1189,1213]" pageId="9" pageNumber="10">
|
||
<emphasis id="95B13362FF93CC1970E1FB1D312EFB5A" bold="true" box="[151,281,1189,1213]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
|
||
Adult wild-caught female
|
||
<taxonomicName id="60C594F3FF93CC19724EFB1E343AFB5B" authority="Andranomaintso Creek. Not" authorityName="Andranomaintso Creek. Not" box="[568,1037,1189,1213]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC19724EFB1E32FCFB5A" box="[568,715,1190,1213]" italics="true" pageId="9" pageNumber="10">Bedotia geayi</emphasis>
|
||
, Andranomaintso Creek. Not
|
||
</taxonomicName>
|
||
preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A77AEF70FF93CC1970E1FB4830D5FAED" blockId="9.[151,1437,1264,1690]" box="[151,226,1264,1290]" pageId="9" pageNumber="10">Range</paragraph>
|
||
<paragraph id="A77AEF70FF93CC1970B0FAA034F9FA65" blockId="9.[151,1437,1264,1690]" pageId="9" pageNumber="10">
|
||
The eleven
|
||
<typeStatus id="787E51D2FF93CC197122FAA0318CFAD5" box="[340,443,1304,1330]" pageId="9" pageNumber="10" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="60C594F3FF93CC197191FAA1327DFAD5" box="[487,586,1305,1330]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC197191FAA1327DFAD5" box="[487,586,1305,1330]" italics="true" pageId="9" pageNumber="10">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
, measuring 48.0–74.0 mm SL, were collected from the Maha River, a north bank tributary of the Mananjary River near the town of Morafeno, at an altitude of c.
|
||
<specimenCount id="B1C324F9FF93CC1974A2FAF83510FABD" box="[1236,1319,1344,1370]" pageId="9" pageNumber="10" type="generic">
|
||
<quantity id="603D4295FF93CC1974A2FAF83515FABD" box="[1236,1314,1344,1370]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="300.0">300 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
This species has also been collected from several south bank tributaries of the Mananjary (Figure 3).
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF93CC1970B0FA28343CF97D" blockId="9.[151,1437,1264,1690]" pageId="9" pageNumber="10">
|
||
Regrettably, neither preserved material of the
|
||
<taxonomicName id="60C594F3FF93CC197374FA29336AFA4D" box="[770,861,1425,1450]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF93CC197374FA29336AFA4D" box="[770,861,1425,1450]" italics="true" pageId="9" pageNumber="10">Bedotia</emphasis>
|
||
</taxonomicName>
|
||
populations found between the Fanantara and Mangoro Rivers nor data on their life colors are available for analysis. This is unfortunate, as living individuals of
|
||
<taxonomicName id="60C594F3FF93CC197095FA583171FA1E" box="[227,326,1504,1529]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC197095FA583171FA1E" box="[227,326,1504,1529]" italics="true" pageId="9" pageNumber="10">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished from
|
||
<taxonomicName id="60C594F3FF93CC1972A8FA5833E3FA1E" box="[734,980,1504,1529]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="95B13362FF93CC1972A8FA5833E3FA1E" box="[734,980,1504,1529]" italics="true" pageId="9" pageNumber="10">B. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
by differences in the pigmentation of their unpaired fins noted in each species’ diagnosis while meristic and morphometric characters presented in
|
||
<tableCitation id="EA47DACBFF93CC1970E1F98830D9F9AD" box="[151,238,1584,1610]" captionStart="TABLE 3" captionStartId="10.[151,239,431,455]" captionTargetBox="[167,1331,541,991]" captionTargetPageId="10" captionText="TABLE 3. Summary of Distinguishing Characters for Bedotia madagascariensis and B. geayi." httpUri="http://table.plazi.org/id/F3BABFF8FF90CC1A70E1FE173452FE20" pageId="9" pageNumber="10" tableUuid="F3BABFF8FF90CC1A70E1FE173452FE20">Table 3</tableCitation>
|
||
permit differentiation of preserved material. Until further sampling corrects this deficiency, the northern range limit of
|
||
<taxonomicName id="60C594F3FF93CC197102F9E031E2F996" box="[372,469,1624,1649]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC197102F9E031E2F996" box="[372,469,1624,1649]" italics="true" pageId="9" pageNumber="10">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
cannot be precisely determined. In the Namorona River, the basin immediately to the south of the Manajary,
|
||
<taxonomicName id="60C594F3FF93CC1971A7F9383207F97E" box="[465,560,1664,1689]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC1971A7F9383207F97E" box="[465,560,1664,1689]" italics="true" pageId="9" pageNumber="10">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
is replaced by an undescribed congener.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF93CC1970E1F9683179F90D" blockId="9.[151,1437,1744,2010]" box="[151,334,1744,1770]" pageId="9" pageNumber="10">Natural History</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFDFBCFBFF93CC1A70B0F94032D5FE9E" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="reference_group">
|
||
<paragraph id="A77AEF70FF93CC1970B0F94033A1F83D" blockId="9.[151,1437,1744,2010]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="60C594F3FF93CC1970B0F940315EF8F6" ID-CoL="68FQ3" box="[198,361,1784,1809]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF93CC1970B0F940315EF8F6" box="[198,361,1784,1809]" italics="true" pageId="9" pageNumber="10">Bedotia geayi</emphasis>
|
||
</taxonomicName>
|
||
inhabits small streams flowing under partial or complete forest cover at altitudes of 300 to 650 meters above sea level (
|
||
<bibRefCitation id="C3549281FF93CC1971A9F89832A0F8DD" author="Pellegrin" box="[479,663,1824,1850]" pageId="9" pageNumber="18" refString="Pellegrin, J. (1907) Liste des poissons recueillis a Madagascar par M. F. Geay. Description d'une espece nouvelle. Bulletin du Musee Nationale d' Histoire Naturelle, 30, 201 - 206." type="journal article" year="1907">Pellegrin, 1907</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C3549281FF93CC1972D4F89833DDF8DD" author="Reinthal" box="[674,1002,1824,1850]" pageId="9" pageNumber="18" refString="Reinthal, P. N. & Stiassny, M. L. J. (1991) The freshwater fishes of Madagascar: a study of an endangered fauna with recommendations for a conservation strategy. Conservation Biology, 5, 231 - 243." type="journal article" year="1991">Reinthal and Stiassny, 1991</bibRefCitation>
|
||
). This species has not been collected from low altitude habitats in the Mananjary basin. This is may be due to the fact that the immediate hinterland of the town of Mananjary has undergone extensive anthropogenic modification. This process favors exotic species, which are better adapted to deal with increased silt loads and higher water temperatures that follow deforestation than are the majority of Madagascar’s native fishes.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A70B0FF2032D5FE9E" blockId="10.[151,1437,152,378]" pageId="10" pageNumber="11">
|
||
Although the streams where it occurs frequently have a strong current,
|
||
<taxonomicName id="60C594F3FF90CC1A747DFF20345BFF56" ID-CoL="68FQ3" box="[1035,1132,152,177]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A747DFF20345BFF56" box="[1035,1132,152,177]" italics="true" pageId="10" pageNumber="11">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
prefers their well-shaded, quieter sections. Like the preceding species, it is indifferent to the composition of the riparian vegetation. This species has been observed swimming in loose, size-graded associations of up to a dozen individuals. Juveniles are usually found in the shallows, while adults frequent deeper water away from the banks. The scant information available on the natural history of
|
||
<taxonomicName id="60C594F3FF90CC1A722AFE80328CFEB6" ID-CoL="68FQ3" box="[604,699,312,337]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A722AFE80328CFEB6" box="[604,699,312,337]" italics="true" pageId="10" pageNumber="11">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
suggests that its dietary pattern, enemies and reproductive pattern are identical to those of
|
||
<emphasis id="95B13362FF90CC1A7190FED832D5FE9E" box="[486,738,352,377]" italics="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A7190FED832E9FE9E" ID-CoL="5WG22" box="[486,734,352,377]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="madagascariensis">B. madagascariensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F3BABFF8FF90CC1A70E1FE173452FE20" ID-Table-UUID="F3BABFF8FF90CC1A70E1FE173452FE20" box="[151,1125,431,455]" httpUri="http://table.plazi.org/id/F3BABFF8FF90CC1A70E1FE173452FE20" pageId="10" pageNumber="11" targetBox="[167,1331,541,991]" targetIsTable="true" targetPageId="10">
|
||
<paragraph id="A77AEF70FF90CC1A70E1FE173452FE20" blockId="10.[151,1125,431,455]" box="[151,1125,431,455]" pageId="10" pageNumber="11">
|
||
<emphasis id="95B13362FF90CC1A70E1FE17313FFE20" bold="true" box="[151,264,431,455]" pageId="10" pageNumber="11">TABLE 3.</emphasis>
|
||
Summary of Distinguishing Characters for
|
||
<taxonomicName id="60C594F3FF90CC1A72A8FE0A33ECFE20" box="[734,987,434,455]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="95B13362FF90CC1A72A8FE0A33ECFE20" box="[734,987,434,455]" italics="true" pageId="10" pageNumber="11">Bedotia madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="95B13362FF90CC1A7397FE09343FFE21" bold="true" box="[993,1032,433,454]" pageId="10" pageNumber="11">and</emphasis>
|
||
<taxonomicName id="60C594F3FF90CC1A7478FE0A3457FE20" box="[1038,1120,432,455]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A7478FE0A3414FE20" box="[1038,1059,434,455]" italics="true" pageId="10" pageNumber="11">B.</emphasis>
|
||
geayi
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="EFDFBCFBFF90CC1A73CAFE55324CF9F4" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="A77AEF70FF90CC1A73CAFE5531B6FBBC" blockId="10.[956,1039,493,517]" lastBlockId="10.[151,1436,1089,1435]" pageId="10" pageNumber="11">
|
||
<emphasis id="95B13362FF90CC1A73CAFE553438FDE2" bold="true" box="[956,1039,493,517]" pageId="10" pageNumber="11">Species</emphasis>
|
||
Conservation Status
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A7094FDA53504FC38" pageId="10" pageNumber="11">
|
||
<table id="D5C51DD0FF9033EF70D1FDA53504FC38" box="[167,1331,541,991]" gridcols="3" gridrows="7" pageId="10" pageNumber="11">
|
||
<tr id="19F5ED32FF9033EF70D1FDA53504FDD2" box="[167,1331,541,565]" gridrow="0" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FDA531CAFDD2" box="[167,509,541,565]" gridcol="0" gridrow="0" pageId="10" pageNumber="11">
|
||
<emphasis id="95B13362FF90CC1A7094FDA53160FDD2" bold="true" box="[226,343,541,565]" pageId="10" pageNumber="11">Character</emphasis>
|
||
</th>
|
||
<emphasis id="95B13362FF90CC1A72E0FDA63537FDD2" box="[662,1280,542,565]" italics="true" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF72E0FDA53344FDD2" box="[662,883,541,565]" gridcol="1" gridrow="0" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A72E0FDA63344FDD2" box="[662,883,542,565]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="95B13362FF90CC1A72E0FDA63344FDD2" box="[662,883,542,565]" italics="true" pageId="10" pageNumber="11">B. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="5A24844EFF9033EF7403FDA53504FDD2" box="[1141,1331,541,565]" gridcol="2" gridrow="0" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A74DEFDA63537FDD2" box="[1192,1280,542,565]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A74DEFDA63537FDD2" box="[1192,1280,542,565]" italics="true" pageId="10" pageNumber="11">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</emphasis>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FDE93504FD62" box="[167,1331,593,645]" gridrow="1" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FDE931CAFD62" box="[167,509,593,645]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">Head length [% SL]</th>
|
||
<td id="5A24844EFF9033EF72E0FDE93344FD62" box="[662,883,593,645]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">27.2–31.8 (mean: 30.0 ± 1.1)</td>
|
||
<td id="5A24844EFF9033EF7403FDE93504FD62" box="[1141,1331,593,645]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">22.0–29.8 (mean 27.8 ± 2.0)</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FD243504FD37" box="[167,1331,668,720]" gridrow="2" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FD2431CAFD37" box="[167,509,668,720]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">Snout length [% SL]</th>
|
||
<td id="5A24844EFF9033EF72E0FD243344FD37" box="[662,883,668,720]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">8.0–10.2 (mean: 9.4 ± 0.5)</td>
|
||
<td id="5A24844EFF9033EF7403FD243504FD37" box="[1141,1331,668,720]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">7.1–10.1 (mean: 8.5 ± 0.2)</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FD5F3504FCFC" box="[167,1331,743,795]" gridrow="3" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FD5F31CAFCFC" box="[167,509,743,795]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">Dorsal2 base [% SL]</th>
|
||
<td id="5A24844EFF9033EF72E0FD5F3344FCFC" box="[662,883,743,795]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">15.9 - 18.7 (mean: 17.6 ± 0.9)</td>
|
||
<td id="5A24844EFF9033EF7403FD5F3504FCFC" box="[1141,1331,743,795]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">16.9–21.4 (mean 19.4 ± 1.3)</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FC8A3504FC81" box="[167,1331,818,870]" gridrow="4" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FC8A31CAFC81" box="[167,509,818,870]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">Anal fin base [% SL]</th>
|
||
<td id="5A24844EFF9033EF72E0FC8A3344FC81" box="[662,883,818,870]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">22.3–27.2 (mean: 24.7 ± 1.4)</td>
|
||
<td id="5A24844EFF9033EF7403FC8A3504FC81" box="[1141,1331,818,870]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">25.1–30.0 (mean 27.2 ± 1.6)</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FCC53504FC55" box="[167,1331,893,946]" gridrow="5" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FCC531CAFC55" box="[167,509,893,946]" gridcol="0" gridrow="5" pageId="10" pageNumber="11">Caudal peduncle length [% SL]</th>
|
||
<td id="5A24844EFF9033EF72E0FCC53344FC55" box="[662,883,893,946]" gridcol="1" gridrow="5" pageId="10" pageNumber="11">16.1–20.4 (mean: 17.4 ± 1.1)</td>
|
||
<td id="5A24844EFF9033EF7403FCC53504FC55" box="[1141,1331,893,946]" gridcol="2" gridrow="5" pageId="10" pageNumber="11">13.9 - 17.7 (mean 16.2 ± 1.2)</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9033EF70D1FC703504FC38" box="[167,1331,968,991]" gridrow="6" pageId="10" pageNumber="11">
|
||
<th id="5A24844EFF9033EF70D1FC7031CAFC38" box="[167,509,968,991]" gridcol="0" gridrow="6" pageId="10" pageNumber="11">Number of anal fin rays</th>
|
||
<td id="5A24844EFF9033EF72E0FC703344FC38" box="[662,883,968,991]" gridcol="1" gridrow="6" pageId="10" pageNumber="11">14–17 (mode: 16)]</td>
|
||
<td id="5A24844EFF9033EF7403FC703504FC38" box="[1141,1331,968,991]" gridcol="2" gridrow="6" pageId="10" pageNumber="11">17–19 (mode: 18)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A70B0FBD130ECFA7C" blockId="10.[151,1436,1089,1435]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A70B0FBD13150FB65" box="[198,359,1129,1154]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A70B0FBD13150FB65" box="[198,359,1129,1154]" italics="true" pageId="10" pageNumber="11">Bedotia geayi</emphasis>
|
||
</taxonomicName>
|
||
remains abundant in those localities where it does occur. While
|
||
<taxonomicName id="60C594F3FF90CC1A7423FBD13511FB65" box="[1109,1318,1129,1154]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="95B13362FF90CC1A7423FBD13511FB65" box="[1109,1318,1129,1154]" italics="true" pageId="10" pageNumber="11">Channa maculata</emphasis>
|
||
</taxonomicName>
|
||
is present in the immediately adjacent Namorona basin, it has not to date been reported from that of the Mananjary. The presence of exotic poeciliids, notably the highly predatory
|
||
<emphasis id="95B13362FF90CC1A7332FB01340CFB35" box="[836,1083,1209,1234]" italics="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A7332FB013400FB35" box="[836,1079,1209,1234]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="holbrooki">Gambusia holbrooki</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
in the Manajary basin is presently a greater cause for concern. However, if for the moment
|
||
<taxonomicName id="60C594F3FF90CC1A730EFB5933EEFB1D" box="[888,985,1249,1274]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="geayi">
|
||
<emphasis id="95B13362FF90CC1A730EFB5933EEFB1D" box="[888,985,1249,1274]" italics="true" pageId="10" pageNumber="11">B. geayi</emphasis>
|
||
</taxonomicName>
|
||
appears to be in no immediate danger of extirpation, its apparently circumscribed distribution suggests particular vulnerability to both further degradation of its habitat and the probable future translocation of
|
||
<taxonomicName id="60C594F3FF90CC1A7310FA8933C0FAAD" box="[870,1015,1329,1354]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="95B13362FF90CC1A7310FA8933C0FAAD" box="[870,1015,1329,1354]" italics="true" pageId="10" pageNumber="11">C. maculata</emphasis>
|
||
</taxonomicName>
|
||
. Following the criteria established by the I.U.C.N., it should be considered a species of special concern whose status needs to be regularly monitored.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A70E1FA41324CF9F4" blockId="10.[151,635,1528,1555]" box="[151,635,1528,1555]" pageId="10" pageNumber="11">
|
||
<emphasis id="95B13362FF90CC1A70E1FA4131D8F9F5" bold="true" box="[151,495,1528,1555]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="60C594F3FF90CC1A70E1FA4131BDF9F4" box="[151,394,1529,1555]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF90CC1A70E1FA4131BDF9F4" bold="true" box="[151,394,1529,1555]" italics="true" pageId="10" pageNumber="11">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF90CC1A71E4FA4031D8F9F5" box="[402,495,1528,1554]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="3FFEF3F5FF90CC1A7270FA41325BF9F4" box="[518,620,1529,1555]" captionStart="FIGURE 6" captionStartId="11.[151,255,660,684]" captionTargetBox="[180,1406,263,649]" captionTargetId="figure@11.[179,1409,262,651]" captionTargetPageId="11" captionText="FIGURE 6. Bedotia leucopteron sp. nov., holotype, AMNH 231236." httpUri="https://zenodo.org/record/177420/files/figure.png" pageId="10" pageNumber="11">Figure 6</figureCitation>
|
||
.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EFDFBCFBFF90CC1F70E1F9F33381FCD5" lastPageId="15" lastPageNumber="16" pageId="10" pageNumber="11" type="materials_examined">
|
||
<paragraph id="A77AEF70FF90CC1A70E1F9F3312AF942" blockId="10.[151,1436,1610,1797]" pageId="10" pageNumber="11">
|
||
<emphasis id="95B13362FF90CC1A70E1F9F3313FF984" bold="true" box="[151,264,1611,1635]" pageId="10" pageNumber="11">
|
||
<typeStatus id="787E51D2FF90CC1A70E1F9F33134F984" box="[151,259,1611,1635]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<collectionCode id="C1D477B5FF90CC1A717BF9F43168F984" LSID="urn:lsid:biocol.org:col:34925" box="[269,351,1612,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="10" pageNumber="11">AMNH</collectionCode>
|
||
<date id="D37BC9B0FF90CC1A7113F9F4318FF984" box="[357,440,1612,1635]" pageId="10" pageNumber="11" value="63-12-23">231263</date>
|
||
. Sandrakatrana Stream at Ampasimbe Village, Toamasina Province,
|
||
<collectingCountry id="DFD2AFE0FF90CC1A74E7F9F43525F984" box="[1169,1298,1612,1635]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
|
||
(18o 56' 26S, 48o 41'
|
||
<geoCoordinate id="C2F189B7FF90CC1A7163F9D6317EF962" box="[277,329,1646,1669]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="0.016666668">0 1 E</geoCoordinate>
|
||
). Altitude
|
||
<specimenCount id="B1C324F9FF90CC1A71CFF9D63234F962" box="[441,515,1646,1669]" pageId="10" pageNumber="11" type="generic">
|
||
<quantity id="603D4295FF90CC1A71CFF9D631C8F962" box="[441,511,1646,1669]" metricMagnitude="2" metricUnit="m" metricValue="1.26" pageId="10" pageNumber="11" unit="m" value="126.0">126 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Rianila drainage. Collected
|
||
<date id="D37BC9B0FF90CC1A734EF9D633DDF962" box="[824,1002,1646,1669]" pageId="10" pageNumber="11" value="2000-10-21">21 October 2000</date>
|
||
by villagers; single male specimen,
|
||
<quantity id="603D4295FF90CC1A751BF9D630C6F942" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" pageId="10" pageNumber="11" unit="mm" value="64.1">64.1 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A70E1F91631E7F8E2" blockId="10.[151,1436,1610,1797]" pageId="10" pageNumber="11">
|
||
<typeStatus id="787E51D2FF90CC1A70E1F916313EF921" box="[151,265,1710,1734]" pageId="10" pageNumber="11" type="paratype">
|
||
<emphasis id="95B13362FF90CC1A70E1F916313EF921" bold="true" box="[151,265,1710,1734]" pageId="10" pageNumber="11">Paratypes</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<collectionCode id="C1D477B5FF90CC1A7151F916314FF922" LSID="urn:lsid:biocol.org:col:34925" box="[295,376,1710,1733]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="10" pageNumber="11">AMNH</collectionCode>
|
||
<date id="D37BC9B0FF90CC1A71F2F91631E0F922" box="[388,471,1710,1733]" pageId="10" pageNumber="11" value="65-12-23">231265</date>
|
||
. Same locality and collection data as
|
||
<typeStatus id="787E51D2FF90CC1A73FFF91633DDF922" box="[905,1002,1710,1733]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
. Eleven specimens,
|
||
<quantity id="603D4295FF90CC1A74A6F916355CF922" box="[1232,1387,1710,1733]" metricMagnitude="-2" metricUnit="m" metricValue="4.455" metricValueMax="5.86" metricValueMin="3.05" pageId="10" pageNumber="11" unit="mm" value="44.55" valueMax="58.6" valueMin="30.5">30.5-58.6 mm</quantity>
|
||
SL.
|
||
<collectionCode id="C1D477B5FF90CC1A70B1F976312EF902" LSID="urn:lsid:biocol.org:col:34988" box="[199,281,1742,1765]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
|
||
1942 0 0 81. Beforona, Toamasina Province,
|
||
<collectingCountry id="DFD2AFE0FF90CC1A7283F9763343F902" box="[757,884,1742,1765]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
|
||
. Rianila drainage. Collected by R. Decary;
|
||
<specimenCount id="B1C324F9FF90CC1A753EF9763135F8E2" pageId="10" pageNumber="11" type="generic">8 specimens</specimenCount>
|
||
,
|
||
<quantity id="603D4295FF90CC1A717AF9563195F8E2" box="[268,418,1774,1797]" metricMagnitude="-2" metricUnit="m" metricValue="6.42" metricValueMax="7.19" metricValueMin="5.65" pageId="10" pageNumber="11" unit="mm" value="64.2" valueMax="71.9" valueMin="56.5">56.5-71.9 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1A70E1F88E313BF8B7" blockId="10.[151,1436,1846,2032]" box="[151,268,1846,1872]" pageId="10" pageNumber="11">Diagnosis</paragraph>
|
||
<paragraph id="A77AEF70FF90CC1B70B0F8E6341FFF3B" blockId="10.[151,1436,1846,2032]" lastBlockId="11.[151,1436,152,220]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Living
|
||
<taxonomicName id="60C594F3FF90CC1A716AF8E7323DF89F" box="[284,522,1887,1912]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF90CC1A716AF8E7323DF89F" box="[284,522,1887,1912]" italics="true" pageId="10" pageNumber="11">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
are readily distinguished from congeners by their metallic blue and gold base coloration, the presence of numerous small irregularly distributed black spots on the flanks rather than a pair of discrete black lateral stripes and the broad, iridescent white margins of the vertical fins in specimens> 25.0 mm SL. The distinctive melanophore pattern of the flanks, opaque white margins of the unpaired fins, deeper body and markedly posterior insertion of the second dorsal fin, as reflected by a snout to D2 distance of 66.2– 71.9 (mean: 68.5 ± 1.3) % SL are likewise diagnostic in preserved specimens.
|
||
</paragraph>
|
||
<caption id="F3BABFF8FF91CC1B70E1FD2C334FFD4C" box="[151,888,660,684]" httpUri="https://zenodo.org/record/177420/files/figure.png" pageId="11" pageNumber="12" targetBox="[180,1406,263,649]" targetPageId="11">
|
||
<paragraph id="A77AEF70FF91CC1B70E1FD2C334FFD4C" blockId="11.[151,888,660,684]" box="[151,888,660,684]" pageId="11" pageNumber="12">
|
||
<emphasis id="95B13362FF91CC1B70E1FD2C312DFD4B" bold="true" box="[151,282,660,684]" pageId="11" pageNumber="12">FIGURE 6.</emphasis>
|
||
<taxonomicName id="60C594F3FF91CC1B7156FD2D31C0FD4B" box="[288,503,661,684]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF91CC1B7156FD2D31C0FD4B" box="[288,503,661,684]" italics="true" pageId="11" pageNumber="12">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF91CC1B7188FD2C3264FD4C" box="[510,595,660,683]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, AMNH 231236.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A77AEF70FF91CC1B70E1FD673117FD1E" blockId="11.[151,1436,735,1001]" box="[151,288,735,761]" pageId="11" pageNumber="12">Description</paragraph>
|
||
<paragraph id="A77AEF70FF91CC1B70B0FCBF3527FC0E" blockId="11.[151,1436,735,1001]" pageId="11" pageNumber="12">
|
||
Morphological measurements and meristic counts are given in
|
||
<tableCitation id="EA47DACBFF91CC1B73C7FCBF343BFCC6" box="[945,1036,775,801]" captionStart="TABLE 4" captionStartId="11.[151,239,1053,1077]" captionTargetBox="[167,1393,1117,1907]" captionTargetPageId="11" captionText="TABLE 4. Morphometric data for Bedotia leucopteron sp. nov. as % SL." httpUri="http://table.plazi.org/id/F3BABFF8FF91CC1B70E1FBA53396FBD3" pageId="11" pageNumber="12" tableUuid="F3BABFF8FF91CC1B70E1FBA53396FBD3">Table 4</tableCitation>
|
||
.
|
||
<taxonomicName id="60C594F3FF91CC1B746EFCB03530FCC6" box="[1048,1287,776,801]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF91CC1B746EFCB03530FCC6" box="[1048,1287,776,801]" italics="true" pageId="11" pageNumber="12">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
can grow to 110.0 mm SL in captivity. The largest specimen examined in this study is a
|
||
<quantity id="603D4295FF91CC1B7450FC9734B8FCAE" box="[1062,1167,815,841]" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" pageId="11" pageNumber="12" unit="mm" value="64.1">64.1 mm</quantity>
|
||
SL male.
|
||
<taxonomicName id="60C594F3FF91CC1B757DFC8830C9FC96" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF91CC1B757DFC8830C9FC96" italics="true" pageId="11" pageNumber="12">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
is a robust fish somewhat deeper bodied than either of the two preceding species and showing a moderately curved ventral outline. Dorsal outline of head and nape moderately curved to first dorsal fin. Head length divisible 3.1–3.5 times in the standard length. First dorsal fin origin is posterior to the vertical through the pelvic fin insertion, while that of second is posterior to the vertical through the anal fin origin.
|
||
</paragraph>
|
||
<caption id="F3BABFF8FF91CC1B70E1FBA53396FBD3" ID-Table-UUID="F3BABFF8FF91CC1B70E1FBA53396FBD3" box="[151,929,1053,1077]" httpUri="http://table.plazi.org/id/F3BABFF8FF91CC1B70E1FBA53396FBD3" pageId="11" pageNumber="12" targetBox="[167,1393,1117,1907]" targetIsTable="true" targetPageId="11">
|
||
<paragraph id="A77AEF70FF91CC1B70E1FBA53396FBD3" blockId="11.[151,929,1053,1077]" box="[151,929,1053,1077]" pageId="11" pageNumber="12">
|
||
<emphasis id="95B13362FF91CC1B70E1FBA5313FFBD2" bold="true" box="[151,264,1053,1077]" pageId="11" pageNumber="12">TABLE 4.</emphasis>
|
||
Morphometric data for
|
||
<taxonomicName id="60C594F3FF91CC1B727CFBA632D5FBD2" box="[522,738,1054,1077]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF91CC1B727CFBA632D5FBD2" box="[522,738,1054,1077]" italics="true" pageId="11" pageNumber="12">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF91CC1B729EFBA5330CFBD3" box="[744,827,1053,1076]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
as % SL.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A77AEF70FF91CC1B71F7FBE53546F894" pageId="11" pageNumber="12">
|
||
<table id="D5C51DD0FF9133EF70D1FBE53546F894" box="[167,1393,1117,1907]" gridcols="5" gridrows="16" pageId="11" pageNumber="12">
|
||
<tr id="19F5ED32FF9133EF70D1FBE53546FB93" box="[167,1393,1117,1140]" gridrow="0" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FBE531DEFB93" box="[167,489,1117,1140]" gridcol="0" gridrow="0" pageId="11" pageNumber="12">Character</th>
|
||
<th id="5A24844EFF9133EF728BFBE5332FFB93" box="[765,792,1117,1140]" gridcol="1" gridrow="0" pageId="11" pageNumber="12">N</th>
|
||
<th id="5A24844EFF9133EF73D7FBE5343BFB93" box="[929,1036,1117,1140]" gridcol="2" gridrow="0" pageId="11" pageNumber="12">Range</th>
|
||
<th id="5A24844EFF9133EF74D1FBE534D3FB93" box="[1191,1252,1117,1140]" gridcol="3" gridrow="0" pageId="11" pageNumber="12">Mean</th>
|
||
<th id="5A24844EFF9133EF7538FBE53546FB93" box="[1358,1393,1117,1140]" gridcol="4" gridrow="0" pageId="11" pageNumber="12">SD</th>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FB333546FB45" box="[167,1393,1163,1186]" gridrow="1" pageId="11" pageNumber="12" rowspan-3="1" rowspan-4="1">
|
||
<th id="5A24844EFF9133EF70D1FB3331DEFB45" box="[167,489,1163,1186]" gridcol="0" gridrow="1" pageId="11" pageNumber="12">Standard length [mm]</th>
|
||
<td id="5A24844EFF9133EF728BFB33332FFB45" box="[765,792,1163,1186]" gridcol="1" gridrow="1" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FB33343BFB45" box="[929,1036,1163,1186]" gridcol="2" gridrow="1" pageId="11" pageNumber="12">25.0–64.1</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FB033546FB35" box="[167,1393,1211,1234]" gridrow="2" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FB0331DEFB35" box="[167,489,1211,1234]" gridcol="0" gridrow="2" pageId="11" pageNumber="12">Head length</th>
|
||
<td id="5A24844EFF9133EF728BFB03332FFB35" box="[765,792,1211,1234]" gridcol="1" gridrow="2" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FB03343BFB35" box="[929,1036,1211,1234]" gridcol="2" gridrow="2" pageId="11" pageNumber="12">21.6–33.1</td>
|
||
<td id="5A24844EFF9133EF74D1FB0334D3FB35" box="[1191,1252,1211,1234]" gridcol="3" gridrow="2" pageId="11" pageNumber="12">29.8</td>
|
||
<td id="5A24844EFF9133EF7538FB033546FB35" box="[1358,1393,1211,1234]" gridcol="4" gridrow="2" pageId="11" pageNumber="12">2.7</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FB503546FB18" box="[167,1393,1256,1279]" gridrow="3" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FB5031DEFB18" box="[167,489,1256,1279]" gridcol="0" gridrow="3" pageId="11" pageNumber="12">Snout length</th>
|
||
<td id="5A24844EFF9133EF728BFB50332FFB18" box="[765,792,1256,1279]" gridcol="1" gridrow="3" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FB50343BFB18" box="[929,1036,1256,1279]" gridcol="2" gridrow="3" pageId="11" pageNumber="12">7.2–10.6</td>
|
||
<td id="5A24844EFF9133EF74D1FB5034D3FB18" box="[1191,1252,1256,1279]" gridcol="3" gridrow="3" pageId="11" pageNumber="12">8.5</td>
|
||
<td id="5A24844EFF9133EF7538FB503546FB18" box="[1358,1393,1256,1279]" gridcol="4" gridrow="3" pageId="11" pageNumber="12">0.9</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FAAE3546FACA" box="[167,1393,1302,1325]" gridrow="4" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FAAE31DEFACA" box="[167,489,1302,1325]" gridcol="0" gridrow="4" pageId="11" pageNumber="12">Lower jaw length</th>
|
||
<td id="5A24844EFF9133EF728BFAAE332FFACA" box="[765,792,1302,1325]" gridcol="1" gridrow="4" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FAAE343BFACA" box="[929,1036,1302,1325]" gridcol="2" gridrow="4" pageId="11" pageNumber="12">13.9–16.7</td>
|
||
<td id="5A24844EFF9133EF74D1FAAE34D3FACA" box="[1191,1252,1302,1325]" gridcol="3" gridrow="4" pageId="11" pageNumber="12">15.2</td>
|
||
<td id="5A24844EFF9133EF7538FAAE3546FACA" box="[1358,1393,1302,1325]" gridcol="4" gridrow="4" pageId="11" pageNumber="12">0.9</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FAFB3546FABD" box="[167,1393,1347,1370]" gridrow="5" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FAFB31DEFABD" box="[167,489,1347,1370]" gridcol="0" gridrow="5" pageId="11" pageNumber="12">Orbital diameter</th>
|
||
<td id="5A24844EFF9133EF728BFAFB332FFABD" box="[765,792,1347,1370]" gridcol="1" gridrow="5" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FAFB343BFABD" box="[929,1036,1347,1370]" gridcol="2" gridrow="5" pageId="11" pageNumber="12">8.6–11.6</td>
|
||
<td id="5A24844EFF9133EF74D1FAFB34D3FABD" box="[1191,1252,1347,1370]" gridcol="3" gridrow="5" pageId="11" pageNumber="12">9.9</td>
|
||
<td id="5A24844EFF9133EF7538FAFB3546FABD" box="[1358,1393,1347,1370]" gridcol="4" gridrow="5" pageId="11" pageNumber="12">0.8</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FAC93546FA6F" box="[167,1393,1393,1416]" gridrow="6" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FAC931DEFA6F" box="[167,489,1393,1416]" gridcol="0" gridrow="6" pageId="11" pageNumber="12">Interorbital distance</th>
|
||
<td id="5A24844EFF9133EF728BFAC9332FFA6F" box="[765,792,1393,1416]" gridcol="1" gridrow="6" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FAC9343BFA6F" box="[929,1036,1393,1416]" gridcol="2" gridrow="6" pageId="11" pageNumber="12">9.7–12.8</td>
|
||
<td id="5A24844EFF9133EF74D1FAC934D3FA6F" box="[1191,1252,1393,1416]" gridcol="3" gridrow="6" pageId="11" pageNumber="12">11.4</td>
|
||
<td id="5A24844EFF9133EF7538FAC93546FA6F" box="[1358,1393,1393,1416]" gridcol="4" gridrow="6" pageId="11" pageNumber="12">0.8</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FA263546FA52" box="[167,1393,1438,1461]" gridrow="7" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FA2631DEFA52" box="[167,489,1438,1461]" gridcol="0" gridrow="7" pageId="11" pageNumber="12">Pectoral fin length</th>
|
||
<td id="5A24844EFF9133EF728BFA26332FFA52" box="[765,792,1438,1461]" gridcol="1" gridrow="7" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7FA26343BFA52" box="[929,1036,1438,1461]" gridcol="2" gridrow="7" pageId="11" pageNumber="12">13.9–20.7</td>
|
||
<td id="5A24844EFF9133EF74D1FA2634D3FA52" box="[1191,1252,1438,1461]" gridcol="3" gridrow="7" pageId="11" pageNumber="12">17.5</td>
|
||
<td id="5A24844EFF9133EF7538FA263546FA52" box="[1358,1393,1438,1461]" gridcol="4" gridrow="7" pageId="11" pageNumber="12">2.2</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1FA733546F9FC" box="[167,1393,1483,1563]" gridrow="8" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1FA7331DEF9FC" box="[167,489,1483,1563]" gridcol="0" gridrow="8" pageId="11" pageNumber="12">
|
||
Dorsal1
|
||
<emphasis id="95B13362FF91CC1B7089FA73313BFA04" bold="true" box="[255,268,1483,1507]" pageId="11" pageNumber="12">–</emphasis>
|
||
anal fin depth Dorsal 2
|
||
<emphasis id="95B13362FF91CC1B7089FA43313BF9F4" bold="true" box="[255,268,1531,1555]" pageId="11" pageNumber="12">–</emphasis>
|
||
anal fin depth
|
||
</th>
|
||
<td id="5A24844EFF9133EF728BFA73332FF9FC" box="[765,792,1483,1563]" gridcol="1" gridrow="8" pageId="11" pageNumber="12">15 15</td>
|
||
<td id="5A24844EFF9133EF73D7FA73343BF9FC" box="[929,1036,1483,1563]" gridcol="2" gridrow="8" pageId="11" pageNumber="12">22.9–26.7 23.3–27.5</td>
|
||
<td id="5A24844EFF9133EF74D1FA7334D3F9FC" box="[1191,1252,1483,1563]" gridcol="3" gridrow="8" pageId="11" pageNumber="12">24.4 25.8</td>
|
||
<td id="5A24844EFF9133EF7538FA733546F9FC" box="[1358,1393,1483,1563]" gridcol="4" gridrow="8" pageId="11" pageNumber="12">1.1 1.2</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F98B3546F9B4" box="[167,1393,1587,1619]" gridrow="9" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F98B31DEF9B4" box="[167,489,1587,1619]" gridcol="0" gridrow="9" pageId="11" pageNumber="12">Pre-dorsal 1 distance</th>
|
||
<td id="5A24844EFF9133EF728BF98B332FF9B4" box="[765,792,1587,1619]" gridcol="1" gridrow="9" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F98B343BF9B4" box="[929,1036,1587,1619]" gridcol="2" gridrow="9" pageId="11" pageNumber="12">53.8–59.3</td>
|
||
<td id="5A24844EFF9133EF74D1F98B34D3F9B4" box="[1191,1252,1587,1619]" gridcol="3" gridrow="9" pageId="11" pageNumber="12">57.1</td>
|
||
<td id="5A24844EFF9133EF7538F98B3546F9B4" box="[1358,1393,1587,1619]" gridcol="4" gridrow="9" pageId="11" pageNumber="12">1.6</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F9D13546F96F" box="[167,1393,1641,1672]" gridrow="10" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F9D131DEF96F" box="[167,489,1641,1672]" gridcol="0" gridrow="10" pageId="11" pageNumber="12">Pre-dorsal2 distance</th>
|
||
<td id="5A24844EFF9133EF728BF9D1332FF96F" box="[765,792,1641,1672]" gridcol="1" gridrow="10" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F9D1343BF96F" box="[929,1036,1641,1672]" gridcol="2" gridrow="10" pageId="11" pageNumber="12">66.2–71.9</td>
|
||
<td id="5A24844EFF9133EF74D1F9D134D3F96F" box="[1191,1252,1641,1672]" gridcol="3" gridrow="10" pageId="11" pageNumber="12">68.5</td>
|
||
<td id="5A24844EFF9133EF7538F9D13546F96F" box="[1358,1393,1641,1672]" gridcol="4" gridrow="10" pageId="11" pageNumber="12">1.8</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F9263546F952" box="[167,1393,1694,1717]" gridrow="11" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F92631DEF952" box="[167,489,1694,1717]" gridcol="0" gridrow="11" pageId="11" pageNumber="12">Pre-pelvic distance</th>
|
||
<td id="5A24844EFF9133EF728BF926332FF952" box="[765,792,1694,1717]" gridcol="1" gridrow="11" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F926343BF952" box="[929,1036,1694,1717]" gridcol="2" gridrow="11" pageId="11" pageNumber="12">42.4–47.7</td>
|
||
<td id="5A24844EFF9133EF74D1F92634D3F952" box="[1191,1252,1694,1717]" gridcol="3" gridrow="11" pageId="11" pageNumber="12">44.4</td>
|
||
<td id="5A24844EFF9133EF7538F9263546F952" box="[1358,1393,1694,1717]" gridcol="4" gridrow="11" pageId="11" pageNumber="12">1.7</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F9743546F90C" box="[167,1393,1740,1771]" gridrow="12" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F97431DEF90C" box="[167,489,1740,1771]" gridcol="0" gridrow="12" pageId="11" pageNumber="12">Base dorsal2</th>
|
||
<td id="5A24844EFF9133EF728BF974332FF90C" box="[765,792,1740,1771]" gridcol="1" gridrow="12" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F974343BF90C" box="[929,1036,1740,1771]" gridcol="2" gridrow="12" pageId="11" pageNumber="12">14.7–20.2</td>
|
||
<td id="5A24844EFF9133EF74D1F97434D3F90C" box="[1191,1252,1740,1771]" gridcol="3" gridrow="12" pageId="11" pageNumber="12">18.4</td>
|
||
<td id="5A24844EFF9133EF7538F9743546F90C" box="[1358,1393,1740,1771]" gridcol="4" gridrow="12" pageId="11" pageNumber="12">1.5</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F8B93546F8FF" box="[167,1393,1793,1816]" gridrow="13" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F8B931DEF8FF" box="[167,489,1793,1816]" gridcol="0" gridrow="13" pageId="11" pageNumber="12">Base anal</th>
|
||
<td id="5A24844EFF9133EF728BF8B9332FF8FF" box="[765,792,1793,1816]" gridcol="1" gridrow="13" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F8B9343BF8FF" box="[929,1036,1793,1816]" gridcol="2" gridrow="13" pageId="11" pageNumber="12">24.1–28.6</td>
|
||
<td id="5A24844EFF9133EF74D1F8B934D3F8FF" box="[1191,1252,1793,1816]" gridcol="3" gridrow="13" pageId="11" pageNumber="12">26.8</td>
|
||
<td id="5A24844EFF9133EF7538F8B93546F8FF" box="[1358,1393,1793,1816]" gridcol="4" gridrow="13" pageId="11" pageNumber="12">1.3</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F8973546F8A1" box="[167,1393,1839,1862]" gridrow="14" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F89731DEF8A1" box="[167,489,1839,1862]" gridcol="0" gridrow="14" pageId="11" pageNumber="12">Length caudal peduncle</th>
|
||
<td id="5A24844EFF9133EF728BF897332FF8A1" box="[765,792,1839,1862]" gridcol="1" gridrow="14" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F897343BF8A1" box="[929,1036,1839,1862]" gridcol="2" gridrow="14" pageId="11" pageNumber="12">15.1–19.4</td>
|
||
<td id="5A24844EFF9133EF74D1F89734D3F8A1" box="[1191,1252,1839,1862]" gridcol="3" gridrow="14" pageId="11" pageNumber="12">17.3</td>
|
||
<td id="5A24844EFF9133EF7538F8973546F8A1" box="[1358,1393,1839,1862]" gridcol="4" gridrow="14" pageId="11" pageNumber="12">1.4</td>
|
||
</tr>
|
||
<tr id="19F5ED32FF9133EF70D1F8E43546F894" box="[167,1393,1884,1907]" gridrow="15" pageId="11" pageNumber="12">
|
||
<th id="5A24844EFF9133EF70D1F8E431DEF894" box="[167,489,1884,1907]" gridcol="0" gridrow="15" pageId="11" pageNumber="12">Depth caudal peduncle</th>
|
||
<td id="5A24844EFF9133EF728BF8E4332FF894" box="[765,792,1884,1907]" gridcol="1" gridrow="15" pageId="11" pageNumber="12">15</td>
|
||
<td id="5A24844EFF9133EF73D7F8E4343BF894" box="[929,1036,1884,1907]" gridcol="2" gridrow="15" pageId="11" pageNumber="12">10.3–12.3</td>
|
||
<td id="5A24844EFF9133EF74D1F8E434D3F894" box="[1191,1252,1884,1907]" gridcol="3" gridrow="15" pageId="11" pageNumber="12">11.3</td>
|
||
<td id="5A24844EFF9133EF7538F8E43546F894" box="[1358,1393,1884,1907]" gridcol="4" gridrow="15" pageId="11" pageNumber="12">0.6</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF91CC1B70B0F81435AAF809" blockId="11.[151,1437,1964,2030]" pageId="11" pageNumber="12">Snout slightly indented behind the premaxillary pedicels. Snout length divisible 3.1–4.0 in the head length. Lower jaw is slightly prognathous and angled at about 40°–45° to horizontal when mouth is closed.</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70E1FF203565FF3D" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">
|
||
Premaxilla and maxilla extend posterior to the anterior margin of the orbit. Premaxilla with a distinct lateral "
|
||
<taxonomicName id="60C594F3FF96CC1C70D5FF7830C9FF3E" box="[163,254,192,217]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF96CC1C70D5FF7830C9FF3E" box="[163,254,192,217]" italics="true" pageId="12" pageNumber="13">Bedotia</emphasis>
|
||
</taxonomicName>
|
||
notch". Orbital diameter divisible 2.8–3.4 times in the head, 0.7–1.1 times in the snout length.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FF503495FE45" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">
|
||
Teeth. Anteriorly both upper and lower jaws bear 4 to 6 rows of numerous small, strongly recurved unicuspid teeth. The outermost row of teeth is poorly differentiated from those of the inner band. The lower jaw and the premaxilla posterior to the
|
||
<taxonomicName id="60C594F3FF96CC1C7237FE8032ABFEB6" box="[577,668,312,337]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF96CC1C7237FE8032ABFEB6" box="[577,668,312,337]" italics="true" pageId="12" pageNumber="13">Bedotia</emphasis>
|
||
</taxonomicName>
|
||
notch each have a single row of teeth. A single row of teeth is present along the anteroventral face of vomer. A small patch of endopterygoid teeth is present. No palatine or ectopterygoid teeth are present, at least in individuals of sizes available for examination.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FE0834DCFE15" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">Gill Rakers. Two or three stout hypobranchial rakers and 11–12 (mode: 11) elongate ceratobranchial rakers are present on the lower limb of the first branchial arch. All rakers are strongly denticulate.</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FDB83296FD05" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">Scales. Body is fully covered with large, regularly imbricate, cycloid scales. Predorsal scales along the dorsal midline: 14 or 15 (modal value: 15). Scales along the midlateral axis from just behind the operculum, above the pectoral fin, to the end of the hypural plate: 32–35 (mode: 34). Scales in transverse series between the anal fin and the second dorsal fin (including a very small scale adjacent to each fin): 9. Scales between the first and second dorsal fins: 2. Circumpeduncular scales: 10–12 (mode: 10). Dorsal, anal, and caudal scale sheaths and axillary pelvic scales are absent.</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FD483448FC65" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">Fins. First dorsal fin with 4 weak spines. Second dorsal fin rays: 10–12 (mode: 11), the first 4 or 5 unbranched. Anal fin rays: 15–17 (mode: 17), the first 4 or 5 unbranched. High-set pectoral fins with 12-13 (mode: 12) rays, the longest extending well beyond the vertical to the pelvic fin insertion. Pelvic fins with one weak spine and five strongly bifurcate, branched rays. Caudal fin weakly emarginate.</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FC2834CEFC35" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">Vertebrae. Total vertebral count taken from radiographs: 34–36 (mode: 35) and a terminal, hypural-bearing half centrum. Pre-caudal vertebrae: 18–19 (mode: 18). Caudal vertebrae: 16–17 (mode: 17).</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70B0FC5832F4FB95" blockId="12.[151,1436,152,1138]" pageId="12" pageNumber="13">Viscera and Diet. Gut extremely short, intestinal length only about one-third body length. Examination of feces produced by newly caught specimens within two to four hours of capture revealed the remains of both aquatic insect larvae and terrestrial insect imagos, suggesting that this species opportunistically exploits both autochthonous and allochthonous food sources.</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70E1FB103122FB25" blockId="12.[151,1436,1192,1338]" box="[151,277,1192,1218]" pageId="12" pageNumber="13">Coloration</paragraph>
|
||
<paragraph id="A77AEF70FF96CC1C70CAFB6833EBFADD" blockId="12.[151,1436,1192,1338]" pageId="12" pageNumber="13">
|
||
Living specimens:
|
||
<figureCitation id="3FFEF3F5FF96CC1C71EAFB6831C9FB0D" box="[412,510,1232,1258]" captionStart="FIGURE 7" captionStartId="12.[151,255,1955,1979]" captionTargetBox="[151,1435,1381,1945]" captionTargetId="figure@12.[151,1436,1379,1947]" captionTargetPageId="12" captionText="FIGURE 7. Sexually quiescent wild-caught male Bedotia leucopteron sp. nov. Lazana River. Not preserved." httpUri="https://zenodo.org/record/177421/files/figure.png" pageId="12" pageNumber="13">Figure 7</figureCitation>
|
||
depicts a male,
|
||
<figureCitation id="3FFEF3F5FF96CC1C72CAFB683328FB0D" box="[700,799,1232,1258]" captionStart="FIGURE 8" captionStartId="13.[151,255,1144,1168]" captionTargetBox="[151,1434,661,1099]" captionTargetId="figure@13.[151,1436,658,1101]" captionTargetPageId="13" captionText="FIGURE 8. Adult wild-caught female Bedotia leucopteron sp. nov., Lazana River. Not preserved." httpUri="https://zenodo.org/record/177422/files/figure.png" pageId="12" pageNumber="13">Figure 8</figureCitation>
|
||
a female and Figure
|
||
<specimenCount id="B1C324F9FF96CC1C746EFB6834BBFB0D" box="[1048,1164,1232,1258]" pageId="12" pageNumber="13" type="juvenile">9 juvenile</specimenCount>
|
||
<taxonomicName id="60C594F3FF96CC1C74E2FB683577FB0E" box="[1172,1344,1232,1257]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF96CC1C74E2FB683577FB0E" box="[1172,1344,1232,1257]" italics="true" pageId="12" pageNumber="13">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
. These photographs do not show the narrow salmon pink mid-dorsal line in specimens>
|
||
<quantity id="603D4295FF96CC1C7416FB403483FAF5" box="[1120,1204,1272,1298]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="mm" value="25.0">25 mm</quantity>
|
||
TL. This species is not characterized by color polymorphism with respect to fin coloration.
|
||
</paragraph>
|
||
<caption id="F3BABFF8FF96CC1C70E1F81B3528F85C" box="[151,1311,1955,1979]" httpUri="https://zenodo.org/record/177421/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1435,1381,1945]" targetPageId="12">
|
||
<paragraph id="A77AEF70FF96CC1C70E1F81B3528F85C" blockId="12.[151,1311,1955,1979]" box="[151,1311,1955,1979]" pageId="12" pageNumber="13">
|
||
<emphasis id="95B13362FF96CC1C70E1F81B312EF85C" bold="true" box="[151,281,1955,1979]" pageId="12" pageNumber="13">FIGURE 7.</emphasis>
|
||
Sexually quiescent wild-caught male
|
||
<taxonomicName id="60C594F3FF96CC1C72D9F81C33B0F85C" box="[687,903,1956,1979]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF96CC1C72D9F81C33B0F85C" box="[687,903,1956,1979]" italics="true" pageId="12" pageNumber="13">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF96CC1C73F8F81C33D6F85C" box="[910,993,1956,1979]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Lazana River. Not preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A77AEF70FF97CC1D70B4FF2030FDFE2D" blockId="13.[151,1436,152,618]" pageId="13" pageNumber="14">Preserved specimens: Top of the head, dorsum and upper third of the flanks light brown, each scale with a darker brown margin. Flanks beige, shading to off-white on the venter. A grey band two scale rows wide extends along the midlateral line from the base of the caudal fin to a point just above the origin of the ventral fins. The posterior half of the flanks irregularly speckled with small black spots. The basal half of both dorsal fins and the anal fin clear grey, marked with black inter-radial streaks. Their distal half is opaque white. The median region of the caudal is clear grey marked with black inter-radial streaks, producing a triangular dark basal zone. The remainder of the caudal is opaque white. The ventral fins are opaque white, the pectorals hyaline.</paragraph>
|
||
<paragraph id="A77AEF70FF97CC1D70B0FE6033EFFD8D" blockId="13.[151,1436,152,618]" pageId="13" pageNumber="14">Females differ from males in having clearer traces of the median and subpectoral lateral bands and fewer black dots on the posterior half of the flanks. The vertical fins are hyaline basally with dusky grey inter-radial streaks and narrower opaque white distal margins. Those of specimens> 50.0 mm SL may be irregularly sprinkled with small black dots. The ventrals and pectorals are hyaline.</paragraph>
|
||
<caption id="F3BABFF8FF97CC1D70E1FBC03499FB77" box="[151,1198,1144,1168]" httpUri="https://zenodo.org/record/177422/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1434,661,1099]" targetPageId="13">
|
||
<paragraph id="A77AEF70FF97CC1D70E1FBC03499FB77" blockId="13.[151,1198,1144,1168]" box="[151,1198,1144,1168]" pageId="13" pageNumber="14">
|
||
<emphasis id="95B13362FF97CC1D70E1FBC0312FFB77" bold="true" box="[151,280,1144,1168]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
|
||
Adult wild-caught female
|
||
<taxonomicName id="60C594F3FF97CC1D724EFBC13327FB77" box="[568,784,1145,1168]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF97CC1D724EFBC13327FB77" box="[568,784,1145,1168]" italics="true" pageId="13" pageNumber="14">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF97CC1D7361FBC1335CFB77" box="[791,875,1145,1168]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, Lazana River. Not preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A77AEF70FF97CC1D70E1F87E3422F83A" blockId="13.[151,1045,1990,2018]" box="[151,1045,1990,2018]" pageId="13" pageNumber="14">
|
||
<emphasis id="95B13362FF97CC1D70E1F87E312DF839" bold="true" box="[151,282,1990,2014]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
|
||
Juvenile captive-bred (F1)
|
||
<taxonomicName id="60C594F3FF97CC1D7236F87F332FF839" box="[576,792,1991,2014]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="leucopteron" status="sp. nov.">
|
||
<emphasis id="95B13362FF97CC1D7236F87F332FF839" box="[576,792,1991,2014]" italics="true" pageId="13" pageNumber="14">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8E828E19FF97CC1D7369F87E3346F83A" box="[799,881,1990,2013]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Not preserved.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70E1FF20312EFF55" blockId="14.[151,1436,152,258]" box="[151,281,152,178]" pageId="14" pageNumber="15">Etymology</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0FF7834A1FEE5" blockId="14.[151,1436,152,258]" pageId="14" pageNumber="15">
|
||
The species name, derived from the Greek
|
||
<emphasis id="95B13362FF94CC1E72B5FF783339FF3E" box="[707,782,192,217]" italics="true" pageId="14" pageNumber="15">leukos</emphasis>
|
||
, white and
|
||
<emphasis id="95B13362FF94CC1E73EFFF7833D2FF3E" box="[921,997,192,217]" italics="true" pageId="14" pageNumber="15">pteron</emphasis>
|
||
, fin, refers to the iridescent white fin coloration particularly evident in adult males. It is to be treated as a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70E1FE8030D5FEB5" blockId="14.[151,1436,312,498]" box="[151,226,312,338]" pageId="14" pageNumber="15">Range</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0FED831C6FE15" blockId="14.[151,1436,312,498]" pageId="14" pageNumber="15">
|
||
The
|
||
<typeStatus id="787E51D2FF94CC1E708DFED83119FE9D" box="[251,302,352,378]" pageId="14" pageNumber="15">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="60C594F3FF94CC1E71EAFED832A7FE9E" box="[412,656,352,377]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E71EAFED832A7FE9E" box="[412,656,352,377]" italics="true" pageId="14" pageNumber="15">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
was collected from the middle reaches of the Iaroka-Rianila basin. Further material has been collected at localities within this drainage situated along Route Nationale 2 between
|
||
<quantity id="603D4295FF94CC1E70E1FE0830D2FE2D" box="[151,229,432,458]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="100.0">100 m</quantity>
|
||
and
|
||
<quantity id="603D4295FF94CC1E7153FE083143FE2D" box="[293,372,432,458]" metricMagnitude="2" metricUnit="m" metricValue="8.43" pageId="14" pageNumber="15" unit="m" value="843.0">843 m</quantity>
|
||
above sea level (Figure 3). Additional collecting is required to ascertain whether it is present in adjacent drainages.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70E1FD903179FDA5" blockId="14.[151,1437,552,1507]" box="[151,334,552,578]" pageId="14" pageNumber="15">Natural History</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0FDE833FDFCD5" blockId="14.[151,1437,552,1507]" pageId="14" pageNumber="15">
|
||
All the streams from which
|
||
<taxonomicName id="60C594F3FF94CC1E7279FDE8328CFD8E" box="[527,699,592,617]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E7279FDE8328CFD8E" box="[527,699,592,617]" italics="true" pageId="14" pageNumber="15">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
has been collected flow under degraded forest cover.
|
||
<collectingCountry id="DFD2AFE0FF94CC1E7543FDE83561FD8D" box="[1333,1366,592,618]" name="American Samoa" pageId="14" pageNumber="15">As</collectingCountry>
|
||
is the case with the preceding species, its occurrence is not influenced by the composition of the riparian vegetation. The Lazana River at Beforona flows strongly even during the driest month of the year and according to local residents, becomes torrential during the rainy season. Recurrent flooding has undercut the banks and caused many trees to fall into the river. Size-graded schools of fifty to one hundred
|
||
<taxonomicName id="60C594F3FF94CC1E7450FD4834E3FCEE" box="[1062,1236,752,777]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E7450FD4834E3FCEE" box="[1062,1236,752,777]" italics="true" pageId="14" pageNumber="15">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
can be observed from its banks swimming through these tangles of waterlogged wood.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0FCF833EDFB9C" blockId="14.[151,1437,552,1507]" pageId="14" pageNumber="15">
|
||
While their waters are very low in dissolved solids (GH: <17.1–35.0 ppm; electrical conductivity: 10.0– 21.0 Μmho/cm2),
|
||
<collectingCountry id="DFD2AFE0FF94CC1E711AFCD231A7FC63" box="[364,400,874,900]" name="Philippines" pageId="14" pageNumber="15">pH</collectingCountry>
|
||
values in these streams range from 6.0–7.0 and do not reach the extremes characteristic of coastal black water habitats. Water temperature in such habitats is strongly influenced by altitude. That of the Sahamamy River, the lowest altitude at which this species has been collected, measured 26o C, while values of 20o C. and 17o C respectively were measured in the Lazana River and in Amalabe Creek during the month of October. Residents of both Amalabe and Beforona stated that water temperatures of their respective streams did not vary noticeably on a seasonal basis. Nevertheless, October is early in the austral spring, so it is possible that summer temperatures in both streams are slightly warmer.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0FB3131ACFA04" blockId="14.[151,1437,552,1507]" pageId="14" pageNumber="15">
|
||
This species coexists with
|
||
<taxonomicName id="60C594F3FF94CC1E7189FB3132D8FB45" box="[511,751,1161,1186]" class="Actinopterygii" family="Poeciliidae" genus="Gambusia" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="holbrooki">
|
||
<emphasis id="95B13362FF94CC1E7189FB3132D8FB45" box="[511,751,1161,1186]" italics="true" pageId="14" pageNumber="15">Gambusia holbrooki</emphasis>
|
||
</taxonomicName>
|
||
in Sandrakatrana Creek and
|
||
<taxonomicName id="60C594F3FF94CC1E743CFB313564FB45" box="[1098,1363,1161,1186]" class="Actinopterygii" family="Bedotiidae" genus="Xiphoporus" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="95B13362FF94CC1E743CFB313564FB45" box="[1098,1363,1161,1186]" italics="true" pageId="14" pageNumber="15">Xiphoporus maculatus</emphasis>
|
||
</taxonomicName>
|
||
in the Sahamany River. According to local residents,
|
||
<taxonomicName id="60C594F3FF94CC1E72C8FB0933B8FB2D" box="[702,911,1201,1226]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="95B13362FF94CC1E72C8FB0933B8FB2D" box="[702,911,1201,1226]" italics="true" pageId="14" pageNumber="15">Channa maculata</emphasis>
|
||
</taxonomicName>
|
||
also inhabits the quieter reaches of the latter stream. In the Lazana River,
|
||
<taxonomicName id="60C594F3FF94CC1E7187FB6132A9FB15" box="[497,670,1241,1266]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E7187FB6132A9FB15" box="[497,670,1241,1266]" italics="true" pageId="14" pageNumber="15">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
shares its habitat with
|
||
<taxonomicName id="60C594F3FF94CC1E73CAFB613462FB15" box="[956,1109,1241,1266]" class="Actinopterygii" family="Bedotiidae" genus="Xiphoporus" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="95B13362FF94CC1E73CAFB613462FB15" box="[956,1109,1241,1266]" italics="true" pageId="14" pageNumber="15">X. maculatus</emphasis>
|
||
</taxonomicName>
|
||
, an undescribed eleotrid of the genus
|
||
<taxonomicName id="60C594F3FF94CC1E7163FAB931B8FAFD" box="[277,399,1281,1306]" class="Actinopterygii" family="Eleotridae" genus="Ratsirakia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF94CC1E7163FAB931B8FAFD" box="[277,399,1281,1306]" italics="true" pageId="14" pageNumber="15">Ratsirakia</emphasis>
|
||
</taxonomicName>
|
||
, the tadpoles of several frog species, diving beetles, a
|
||
<taxonomicName id="60C594F3FF94CC1E7444FAB934D8FAFD" box="[1074,1263,1281,1306]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="95B13362FF94CC1E7444FAB934D8FAFD" box="[1074,1263,1281,1306]" italics="true" pageId="14" pageNumber="15">Macrobrachium</emphasis>
|
||
</taxonomicName>
|
||
species and a freshwater crab. The Amalabe Creek population occurs syntopically with two other bedotiids,
|
||
<taxonomicName id="60C594F3FF94CC1E7493FA9130ECFA8D" class="Actinopterygii" family="Bedotiidae" genus="Rheocles" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="alaotrensis">
|
||
<emphasis id="95B13362FF94CC1E7493FA9130ECFA8D" italics="true" pageId="14" pageNumber="15">Rheocles alaotrensis</emphasis>
|
||
</taxonomicName>
|
||
and an undescribed
|
||
<taxonomicName id="60C594F3FF94CC1E71B9FAE9320EFA8D" box="[463,569,1361,1386]" class="Actinopterygii" family="Bedotiidae" genus="Rheocles" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF94CC1E71B9FAE9320EFA8D" box="[463,569,1361,1386]" italics="true" pageId="14" pageNumber="15">Rheocles</emphasis>
|
||
</taxonomicName>
|
||
species, the same
|
||
<taxonomicName id="60C594F3FF94CC1E7362FAE933B9FA8D" box="[788,910,1361,1386]" class="Actinopterygii" family="Eleotridae" genus="Ratsirakia" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="95B13362FF94CC1E7362FAE933B9FA8D" box="[788,910,1361,1386]" italics="true" pageId="14" pageNumber="15">Ratsirakia</emphasis>
|
||
</taxonomicName>
|
||
present in the Lazana River, and
|
||
<taxonomicName id="60C594F3FF94CC1E756DFAE9308EFA75" class="Actinopterygii" family="Bedotiidae" genus="Xiphoporus" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="95B13362FF94CC1E756DFAE9308EFA75" italics="true" pageId="14" pageNumber="15">X. maculatus</emphasis>
|
||
</taxonomicName>
|
||
. According to local residents, eels are the only predatory fish present at these altitudes.
|
||
<taxonomicName id="60C594F3FF94CC1E7537FAC13115FA5D" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E7537FAC13115FA5D" italics="true" pageId="14" pageNumber="15">Bedotia leucopteron</emphasis>
|
||
</taxonomicName>
|
||
is thus chiefly at risk from kingfishers and the skilfully wielded
|
||
<emphasis id="95B13362FF94CC1E7458FA1934AEFA5D" box="[1070,1177,1441,1466]" italics="true" pageId="14" pageNumber="15">tandroho</emphasis>
|
||
(woven reed baskets) of artisanal fisherfolk.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70E1F9A131B6F9D4" blockId="14.[151,1437,1561,1867]" box="[151,385,1561,1587]" pageId="14" pageNumber="15">Conservation Status</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70B0F9F93227F8AC" blockId="14.[151,1437,1561,1867]" pageId="14" pageNumber="15">
|
||
Although Mantadia-Andasibe National Park affords the watershed of the of the Iaroka-Rianila basin a degree of protection, the middle reaches of this basin are characterized by much reduced and at best degraded forest cover. Nevertheless,
|
||
<taxonomicName id="60C594F3FF94CC1E71AFF92932B1F94D" box="[473,646,1681,1706]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E71AFF92932B1F94D" box="[473,646,1681,1706]" italics="true" pageId="14" pageNumber="15">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
is quite abundant in those localities where it occurs. Like the preceding species, it does not appear at present to be seriously endangered, notwithstanding the presence of potentially competitive and/or predatory naturalized poeciliids throughout its altitudinal distribution and of the highly predatory
|
||
<taxonomicName id="60C594F3FF94CC1E71FDF8B1326BF8C5" box="[395,604,1801,1826]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculata">
|
||
<emphasis id="95B13362FF94CC1E71FDF8B1326BF8C5" box="[395,604,1801,1826]" italics="true" pageId="14" pageNumber="15">Channa maculata</emphasis>
|
||
</taxonomicName>
|
||
at its lower end. It should be classified as vulnerable following the criteria established by the I.U.C.N.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1E70E1F839312FF87C" blockId="14.[151,1436,1921,2027]" box="[151,280,1921,1947]" pageId="14" pageNumber="15">Discussion</paragraph>
|
||
<paragraph id="A77AEF70FF94CC1F70B0F8113414FD8D" blockId="14.[151,1436,1921,2027]" lastBlockId="15.[151,1436,152,818]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
The lateral melanophore pattern of adult
|
||
<taxonomicName id="60C594F3FF94CC1E72DBF811336DF825" box="[685,858,1961,1986]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF94CC1E72DBF811336DF825" box="[685,858,1961,1986]" italics="true" pageId="14" pageNumber="15">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles of that of
|
||
<taxonomicName id="60C594F3FF94CC1E74A3F811311DF80C" authority="Sparks 2001" authorityName="Sparks" authorityYear="2001" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="masoala">
|
||
<emphasis id="95B13362FF94CC1E74A3F81135ACF825" box="[1237,1435,1961,1986]" italics="true" pageId="14" pageNumber="15">Bedotia masoala</emphasis>
|
||
<bibRefCitation id="C3549281FF94CC1E70E1F869311DF80C" author="Sparks" box="[151,298,2001,2027]" pageId="14" pageNumber="18" refString="Sparks, J. S. (2001) Bedotia masoala, a new species of atherinoid rainbowfish (Teleostei: Atherinoidei) From the Masoala Peninsula, northeastern Madagascar. Copeia, (2), 482 - 489." type="journal article" year="2001">Sparks 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="60C594F3FF94CC1E7140F869337FF80C" authority="Stiassny and Harrison 2000" authorityName="Stiassny and Harrison" authorityYear="2000" box="[310,840,2001,2027]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="marojejy">
|
||
<emphasis id="95B13362FF94CC1E7140F86931C8F80D" box="[310,511,2001,2026]" italics="true" pageId="14" pageNumber="15">Bedotia marojejy</emphasis>
|
||
<bibRefCitation id="C3549281FF94CC1E7270F869337FF80C" author="Stiassny" box="[518,840,2001,2027]" pageId="14" pageNumber="18" refString="Stiassny, M. L. J. and I. J. Harrison. (2000) Notes on a small collection of fishes from the Parc National de Marojejy, northeastern Madagascar, with a description of a new species of the endemic genus Bedotia (Atherinomorpha: Bedotiidae). In: S. M. Goodman (Ed.) A Floral and Faunal Inventory of the Parc National de Marojejy, Madagascar with Reference to Elevational Variation. Fieldiana (Zoology), (97), 143 - 156." type="journal article" year="2000">Stiassny and Harrison 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
and three undescribed congeners restricted respectively to the basins of the Sambava, Bemarivo and Mahanara du Nord Rivers in northeastern
|
||
<collectingCountry id="DFD2AFE0FF95CC1F7495FF203544FF55" box="[1251,1395,152,178]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
|
||
. Its ontogeny in
|
||
<taxonomicName id="60C594F3FF95CC1F7158FF7831EAFF3E" box="[302,477,192,217]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF95CC1F7158FF7831EAFF3E" box="[302,477,192,217]" italics="true" pageId="15" pageNumber="16">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
, however, is quite different from that of the adult color pattern of this northern quintet of species, which for convenience sake we will refer to subsequently as the
|
||
<emphasis id="95B13362FF95CC1F7401FF5034EDFEE6" box="[1143,1242,232,257]" italics="true" pageId="15" pageNumber="16">karikary</emphasis>
|
||
group, from the
|
||
<collectingCountry id="DFD2AFE0FF95CC1F70E1FEA8313EFECD" box="[151,265,272,298]" name="Madagascar" pageId="15" pageNumber="16">Malagasy</collectingCountry>
|
||
word for speckled. Juvenile
|
||
<taxonomicName id="60C594F3FF95CC1F721FFEA8332FFECE" box="[617,792,272,297]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF95CC1F721FFEA8332FFECE" box="[617,792,272,297]" italics="true" pageId="15" pageNumber="16">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
develop well-defined black lateral stripes within two weeks of hatching which become more intense until the young reach c. 30.0 mm SL. By this time, the stripes are more or less broadly edged in metallic gold.
|
||
<collectingCountry id="DFD2AFE0FF95CC1F72BFFED832DEFE9D" box="[713,745,352,378]" name="American Samoa" pageId="15" pageNumber="16">As</collectingCountry>
|
||
the fish grow larger, the black lateral stripes begin to break up and the zone of metallic gold progressively expands while undergoing a comparable degree of fragmentation. The end result of this process is the distinctive adult color pattern seen in both sexes of
|
||
<taxonomicName id="60C594F3FF95CC1F749EFE0835A2FE2E" box="[1256,1429,432,457]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF95CC1F749EFE0835A2FE2E" box="[1256,1429,432,457]" italics="true" pageId="15" pageNumber="16">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
. In the three representatives of the
|
||
<emphasis id="95B13362FF95CC1F7234FE603292FE16" box="[578,677,472,497]" italics="true" pageId="15" pageNumber="16">karikary</emphasis>
|
||
group bred to date, juveniles develop indistinct dusky lateral stripes shortly after hatching. In an attenuated form, these stripes persist into adulthood in females, but by the time males attain 3.0 cm SL, they have been replaced by a more or less well defined series of diffuse black spots, whose intensity and size decrease progressively as the fish grow larger.
|
||
</paragraph>
|
||
<paragraph id="A77AEF70FF95CC1F70B0FDC03381FCD5" blockId="15.[151,1436,152,818]" pageId="15" pageNumber="16">
|
||
The different fashion in which their superficially similar adult coloration develops in
|
||
<taxonomicName id="60C594F3FF95CC1F74CAFDC0355EFD76" box="[1212,1385,632,657]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF95CC1F74CAFDC0355EFD76" box="[1212,1385,632,657]" italics="true" pageId="15" pageNumber="16">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
and the species of the
|
||
<emphasis id="95B13362FF95CC1F711EFD1831FCFD5E" box="[360,459,672,697]" italics="true" pageId="15" pageNumber="16">karikary</emphasis>
|
||
group suggests that a spotted adult color pattern has evolved independently at least twice in the genus. This hypothesis is supported by the results of a recent genetic study of the
|
||
<taxonomicName id="60C594F3FF95CC1F756BFD7031F0FCED" authority="Sparks and Smith, 2004" authorityName="Sparks and Smith" authorityYear="2004" class="Actinopterygii" family="Bedotiidae" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="family">
|
||
Bedotiidae (
|
||
<bibRefCitation id="C3549281FF95CC1F70E9FD483188FCED" author="Sparks" box="[159,447,752,778]" pageId="15" pageNumber="18" refString="Sparks, J. S. & Smith, W. L. (2004) Phylogeny and biogeography of the Malagasy and Australasian rainbowfishes (Teleostei: Melanotaenioidei): Gondwanan vicariance and evolution in freshwater. Molecular Phylogenetics and Evolution, 33, 719 - 734." type="journal article" year="2004">Sparks and Smith, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which recovers the
|
||
<emphasis id="95B13362FF95CC1F72CDFD483328FCEE" box="[699,799,752,777]" italics="true" pageId="15" pageNumber="16">karikary</emphasis>
|
||
group as an assemblage of closely related species but places
|
||
<taxonomicName id="60C594F3FF95CC1F7091FCA031ECFCD6" box="[231,475,792,817]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="madagascariensis">
|
||
<emphasis id="95B13362FF95CC1F7091FCA031ECFCD6" box="[231,475,792,817]" italics="true" pageId="15" pageNumber="16">B. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
as the closest relative of
|
||
<taxonomicName id="60C594F3FF95CC1F7372FCA03398FCD6" box="[772,943,792,817]" class="Actinopterygii" family="Bedotiidae" genus="Bedotia" kingdom="Animalia" order="Atheriniformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="leucopteron">
|
||
<emphasis id="95B13362FF95CC1F7372FCA03398FCD6" box="[772,943,792,817]" italics="true" pageId="15" pageNumber="16">B. leucopteron</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |