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<document id="9F452B012C2CE7D9A629A3A166A728CA" ID-CLB-Dataset="7356" ID-DOI="10.1051/acarologia/20122042" ID-GBIF-Dataset="1bbcbc7b-56d5-4f1e-b8e9-cb5760c07aae" ID-ISSN="2107-7207" ID-Zenodo-Dep="4639918" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616776124528" checkinUser="carolina" docAuthor="Bertrand, M., Bagheri, M., Akbari, A., Yazdanian, M., Irani-Nejad, K. H., Mohajer, S. S. &amp; Saboori, A." docDate="2012" docId="03BE87E14367FFD57AECFA5BFDF68889" docLanguage="en" docName="Acarologia.52.3.233-245.pdf" docOrigin="Acarologia 52 (3)" docSource="http://dx.doi.org/10.1051/acarologia/20122042" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Labidostoma (Labidostoma) intermedia Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="240" masterDocId="FF87FF994362FFDD7A57FF86FFD48B79" masterDocTitle="A New Iranian Species Of The Subgenus Labidostoma (Prostigmata: Labidostomatidae), With New Biogeographic Data On The Integrum Species Group" masterLastPageNumber="245" masterPageNumber="233" pageNumber="237" updateTime="1698935121063" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
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<mods:title id="20153287AE0420F458A6DF555421F5DE">A New Iranian Species Of The Subgenus Labidostoma (Prostigmata: Labidostomatidae), With New Biogeographic Data On The Integrum Species Group</mods:title>
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<mods:namePart id="29DDE810DAD7D4F46E36A0A3D7B7E843">Bertrand, M.</mods:namePart>
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<mods:namePart id="090D51B778F8981DB0C588BC191AE085">Bagheri, M.</mods:namePart>
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<mods:namePart id="E75E4F624C883B830373F770E8D8AD7A">Akbari, A.</mods:namePart>
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<mods:namePart id="B69D81F1909C23470772850C42FB78A3">Yazdanian, M.</mods:namePart>
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<mods:namePart id="EB3AB406DDD6D224B52EF534C54638E5">Irani-Nejad, K. H.</mods:namePart>
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<mods:namePart id="43CECBC765FCA605D27D42CBCA706635">Mohajer, S. S.</mods:namePart>
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<mods:namePart id="1DAAC59A04954595C5C144D0AA5E6E50">Saboori, A.</mods:namePart>
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<mods:title id="D65A37EDA4823ED3461B6C429F5D6A67">Acarologia</mods:title>
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<mods:date id="F64FFE8AB44E839623C14D9E94145F63">2012</mods:date>
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<treatment id="03BE87E14367FFD57AECFA5BFDF68889" ID-DOI="http://doi.org/10.5281/zenodo.4694012" ID-GBIF-Taxon="181009821" ID-Zenodo-Dep="4694012" LSID="urn:lsid:plazi:treatment:03BE87E14367FFD57AECFA5BFDF68889" httpUri="http://treatment.plazi.org/id/03BE87E14367FFD57AECFA5BFDF68889" lastPageId="8" lastPageNumber="240" pageId="5" pageNumber="237">
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<paragraph id="8BA836F74367FFD87AECFA5BFD0A8E8C" blockId="5.[187,734,1501,1525]" box="[187,734,1501,1525]" pageId="5" pageNumber="237">
<heading id="D0E0819B4367FFD87AECFA5BFD0A8E8C" bold="true" box="[187,734,1501,1525]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="237" reason="2">
<emphasis id="B963EAE54367FFD87AECFA5BFD0A8E8C" bold="true" box="[187,734,1501,1525]" pageId="5" pageNumber="237">
<taxonomicName id="4C174D744367FFD87AECFA5BFD418E8C" authority="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori, 2012" authorityName="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori" authorityYear="2012" box="[187,661,1501,1525]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="5" pageNumber="237" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Labidostoma">
<emphasis id="B963EAE54367FFD87AECFA5BFE8C8E8C" bold="true" box="[187,344,1501,1525]" italics="true" pageId="5" pageNumber="237">Labidostoma</emphasis>
(
<emphasis id="B963EAE54367FFD87B3FFA5BFDD18E8C" bold="true" box="[360,517,1501,1525]" italics="true" pageId="5" pageNumber="237">Labidostoma</emphasis>
)
<emphasis id="B963EAE54367FFD87842FA5BFD418E8C" bold="true" box="[533,661,1501,1525]" italics="true" pageId="5" pageNumber="237">intermedia</emphasis>
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<taxonomicNameLabel id="A250579E4367FFD878CBFA5BFD0A8E8C" box="[668,734,1501,1525]" pageId="5" pageNumber="237" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BA836F74367FFD87AC8F990FE808C79" blockId="5.[159,762,1557,1948]" pageId="5" pageNumber="237">
<materialsCitation id="3B7F3CAA4367FFD87AC8F990FE498D0A" ID-GBIF-Occurrence="3072648302" collectingDate="2011-03-01" country="Iran" county="Gorgan" latitude="36.833332" location="Zoology Museum" longLatPrecision="1185" longitude="54.5" municipality="Berlese" pageId="5" pageNumber="237" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Golestan" typeStatus="holotype">
Types:
<typeStatus id="54AC88554367FFD87AA2F993FEB08D54" box="[245,356,1557,1581]" pageId="5" pageNumber="237" type="holotype">Holotype</typeStatus>
<specimenCount id="9D11FD7E4367FFD87B3BF990FEAE8D54" box="[364,378,1558,1581]" pageId="5" pageNumber="237" type="female"></specimenCount>
, MNHN-Ac1152,
<collectingCountry id="F30076674367FFD87805F990FD508D54" box="[594,644,1558,1581]" name="Iran" pageId="5" pageNumber="237">Iran</collectingCountry>
,
<collectingRegion id="49D3F8154367FFD878C5F993FEDD8D29" country="Iran" name="Golestan" pageId="5" pageNumber="237">Golestan Province</collectingRegion>
, Gorgan (
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,
<geoCoordinate id="EE2350304367FFD87BB3F9BEFDEA8D29" box="[484,574,1592,1616]" degrees="54" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="237" precision="925" value="54.5">54°30E</geoCoordinate>
), Berlese extraction of soil,
<date id="FFA910374367FFD87B7CF9DDFE4D8D0A" box="[299,409,1627,1651]" pageId="5" pageNumber="237" value="2011-03-01">
<collectingDate id="EFEDE9DF4367FFD87B7CF9DDFE4D8D0A" box="[299,409,1627,1651]" pageId="5" pageNumber="237" value="2011-03-01">1.03.2011</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="3B7F3CAA4367FFD87BF0F9DDFE848C79" ID-GBIF-Occurrence="3072648303" collectingDate="2011-03-01" country="Iran" county="Gorgan" latitude="36.833332" location="Zoology Museum" longLatPrecision="1185" longitude="54.5" municipality="Berlese" pageId="5" pageNumber="237" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Golestan" typeStatus="allotype">
<typeStatus id="54AC88554367FFD87BF0F9DDFDDA8D0A" box="[423,526,1627,1651]" pageId="5" pageNumber="237" type="allotype">Allotype</typeStatus>
<specimenCount id="9D11FD7E4367FFD87842F9DAFDFE8D0A" box="[533,554,1628,1651]" pageId="5" pageNumber="237" type="male"></specimenCount>
, MNHN-Ac1153,
<collectingCountry id="F30076674367FFD87AC8F9F9FF058DEF" box="[159,209,1663,1686]" name="Iran" pageId="5" pageNumber="237">Iran</collectingCountry>
,
<collectingRegion id="49D3F8154367FFD87A8BF9F8FE618DEF" box="[220,437,1662,1686]" country="Iran" name="Golestan" pageId="5" pageNumber="237">Golestan Province</collectingRegion>
,
<collectingCounty id="62C94E7B4367FFD87B97F9F9FDCC8DEE" box="[448,536,1663,1687]" pageId="5" pageNumber="237">Gorgan</collectingCounty>
(
<geoCoordinate id="EE2350304367FFD87870F9F8FD538DEF" box="[551,647,1662,1686]" degrees="36" direction="north" minutes="50" orientation="latitude" pageId="5" pageNumber="237" precision="925" value="36.833332">36°50N</geoCoordinate>
,
<geoCoordinate id="EE2350304367FFD878C4F9F8FD398DEF" box="[659,749,1662,1686]" degrees="54" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="237" precision="925" value="54.5">54°30E</geoCoordinate>
),
<collectingMunicipality id="6BCCAC8D4367FFD87AC8F927FF278DC0" box="[159,243,1697,1721]" pageId="5" pageNumber="237">Berlese</collectingMunicipality>
extraction of soil. A reference collection is kept at the
<location id="8EC8602C4367FFD87B70F943FE2F8DA5" LSID="urn:lsid:plazi:treatment:03BE87E14367FFD57AECFA5BFDF68889:8EC8602C4367FFD87B70F943FE2F8DA5" box="[295,507,1733,1757]" country="Iran" county="Gorgan" latitude="36.833332" longLatPrecision="1185" longitude="54.5" municipality="Berlese" name="Zoology Museum" pageId="5" pageNumber="237" stateProvince="Golestan">Zoology Museum</location>
of
<collectingRegion id="49D3F8154367FFD87874F943FDA08DA4" box="[547,628,1733,1757]" country="Iran" name="Tehran" pageId="5" pageNumber="237">Tehran</collectingRegion>
<location id="8EC8602C4367FFD8782AF943FD2E8DA4" LSID="urn:lsid:plazi:treatment:03BE87E14367FFD57AECFA5BFDF68889:8EC8602C4367FFD8782AF943FD2E8DA4" box="[637,762,1733,1757]" country="Iran" county="Gorgan" latitude="36.833332" longLatPrecision="1185" longitude="54.5" municipality="Berlese" name="University" pageId="5" pageNumber="237" stateProvince="Golestan">University</location>
(three females)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D657C4367FFD57A92F895FDF68889" lastPageId="8" lastPageNumber="240" pageId="5" pageNumber="237" type="description">
<paragraph id="8BA836F74367FFD87A92F895FE7B8C37" blockId="5.[159,762,1557,1948]" pageId="5" pageNumber="237">
Female — Dorsal shield
<quantity id="4CEF9B124367FFD87BB6F895FDA38C52" box="[481,631,1811,1835]" metricMagnitude="-1" metricUnit="m" metricValue="6.2" metricValueMax="6.7" metricValueMin="5.7" pageId="5" pageNumber="237" unit="mm" value="620.0" valueMax="670.0" valueMin="570.0">570 670 µm</quantity>
long, 260 302 wide. Male similar.
</paragraph>
<paragraph id="8BA836F74367FFDB7A92F8E7FE878AA2" blockId="5.[159,762,1557,1948]" lastBlockId="6.[159,762,311,476]" lastPageId="6" lastPageNumber="238" pageId="5" pageNumber="237">
Dorsal shield — (
<figureCitation id="132C2A724367FFD87BF7F8E7FD258C03" box="[416,753,1889,1914]" captionStart="FIGURE" captionStartId="4.[159,171,1766,1785]" captionTargetBox="[159,1429,416,1813]" captionTargetId="figure-4@4.[245,1349,416,1809]" captionTargetPageId="4" captionText="FIGURE 1: Labidostoma (Labidostoma) intermedia n. sp., female: A dorsal shield; B epimera (p: epimeral and post epimeral pores; C-D dorsal trichobotria; E detail of dorso-lateral line of pores." figureDoi="http://doi.org/10.5281/zenodo.4639920" httpUri="https://zenodo.org/record/4639920/files/figure.png" pageId="5" pageNumber="237">Figures 1A, 1C, 1D and 1E</figureCitation>
) Body elongated, covered by a reticular pattern even in the central zone. As in many species the polygons become less regular and less distinct and increasingly granulate dorsally as well as in the posterior part. Two pairs of long trichobothria;
<emphasis id="B963EAE54367FFD87F5DFA9CFAFB8E4B" box="[1290,1327,1306,1330]" italics="true" pageId="5" pageNumber="237">bop</emphasis>
longest. Aspidosomal setae:
<emphasis id="B963EAE54367FFD87E66FAB8FB9D8E2C" box="[1073,1097,1342,1365]" italics="true" pageId="5" pageNumber="237">ge</emphasis>
&gt;
<emphasis id="B963EAE54367FFD87E3BFAB8FB518E2C" box="[1132,1157,1342,1365]" italics="true" pageId="5" pageNumber="237">ga</emphasis>
=
<emphasis id="B963EAE54367FFD87EFEFAB8FB158E2C" box="[1193,1217,1342,1365]" italics="true" pageId="5" pageNumber="237">gr</emphasis>
&lt;
<emphasis id="B963EAE54367FFD87EB2FAB8FAD98E2C" box="[1253,1293,1342,1365]" italics="true" pageId="5" pageNumber="237">gm,</emphasis>
all simple. Usual paired dorsal and lateral setae, all simple, posterior setae (
<emphasis id="B963EAE54367FFD879A1FA05FBC58EE2" box="[1014,1041,1411,1435]" italics="true" pageId="5" pageNumber="237">dd</emphasis>
,
<emphasis id="B963EAE54367FFD87E49FA05FBE28EE2" box="[1054,1078,1411,1435]" italics="true" pageId="5" pageNumber="237">de</emphasis>
, and
<emphasis id="B963EAE54367FFD87E2FFA05FB5E8EE2" box="[1144,1162,1411,1435]" italics="true" pageId="5" pageNumber="237">le</emphasis>
) longest. Dorsal pores visible around
<emphasis id="B963EAE54367FFD879A5FA20FBC38EC7" box="[1010,1047,1446,1470]" italics="true" pageId="5" pageNumber="237">bop</emphasis>
and grouped in area posteriad to setae
<emphasis id="B963EAE54367FFD879CCFA4CFC698E98" box="[923,957,1482,1505]" italics="true" pageId="5" pageNumber="237">gm</emphasis>
and latero-posterior to
<emphasis id="B963EAE54367FFD87E82FA4FFB3B8E98" box="[1237,1263,1481,1505]" italics="true" pageId="5" pageNumber="237">db</emphasis>
. Anterior eye present,
<quantity id="4CEF9B124367FFD879C9FA6BFC318D7D" box="[926,997,1516,1541]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="237" unit="mm" value="23.0">23 µm</quantity>
in diameter, in subterminal position. One pair of large lateral pustules (diameter
<quantity id="4CEF9B124367FFD87F14F996FA588D5E" box="[1347,1420,1551,1576]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="237" unit="mm" value="34.0">34 µm</quantity>
) each close to the small lateral eyes. Lateral lyriform organ present, extending over three to five cuticular cells. The pustule cuts the lateral line of pores that are connected by a sclerotized
<emphasis id="B963EAE54367FFD87E8CF91AFB3C8DCA" box="[1243,1256,1692,1715]" italics="true" pageId="5" pageNumber="237">&quot;</emphasis>
ridge
<emphasis id="B963EAE54367FFD87F71F91AFAE78DCA" box="[1318,1331,1692,1715]" italics="true" pageId="5" pageNumber="237">&quot;</emphasis>
(figures 3A and 3B). The posterior part of this ridge (
<emphasis id="B963EAE54367FFD87F14F939FAA38DAF" box="[1347,1399,1727,1750]" italics="true" pageId="5" pageNumber="237">arOEte</emphasis>
of Grandjean,
<emphasis id="B963EAE54367FFD87993F965FBD88D83" box="[964,1036,1763,1786]" italics="true" pageId="5" pageNumber="237">op. cit</emphasis>
.) follows the margin on the dorsal shield backward and continues from the right pustule to the symmetric pustule. From the oculopustular zone, the anterior branches are inclined toward the axis of the body and are long enough to reach the aspidosomal setae
<emphasis id="B963EAE54364FFDB7BA1FEBEFDDA8A36" box="[502,526,312,335]" italics="true" pageId="6" pageNumber="238">ge</emphasis>
. In some individuals the anterior ridge ends below the lateral ocular lenses, the posterior branch ends at the level of the pustule and sometimes tends to curve below this organ (figure 3B).
</paragraph>
<caption id="DF68667F4364FFDB796DFC71FACB8F5C" ID-DOI="http://doi.org/10.5281/zenodo.4639926" ID-Zenodo-Dep="4639926" httpUri="https://zenodo.org/record/4639926/files/figure.png" pageId="6" pageNumber="238" startId="6.[826,838,1015,1034]" targetBox="[828,1505,299,1000]" targetPageId="6">
<paragraph id="8BA836F74364FFDB796DFC71FACB8F5C" blockId="6.[826,1429,1014,1061]" pageId="6" pageNumber="238">
FIGURE 4:
<taxonomicName id="4C174D744364FFDB79F7FC70FBB68F70" authorityName="Berlese" authorityYear="1911" box="[928,1122,1014,1033]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54364FFDB79F7FC70FBB68F70" box="[928,1122,1014,1033]" italics="true" pageId="6" pageNumber="238">Labidostoma integrum</emphasis>
</taxonomicName>
from Kazaksthan: A Chelicera antiaxial view; B infracapitulum, ventral view.
</paragraph>
</caption>
<caption id="DF68667F4364FFDB7AC8FACFFDBE8EEA" ID-DOI="http://doi.org/10.5281/zenodo.4639928" ID-Zenodo-Dep="4639928" httpUri="https://zenodo.org/record/4639928/files/figure.png" pageId="6" pageNumber="238" startId="6.[159,171,1353,1372]" targetBox="[163,785,568,1361]" targetPageId="6">
<paragraph id="8BA836F74364FFDB7AC8FACFFDBE8EEA" blockId="6.[159,762,1352,1427]" pageId="6" pageNumber="238">
FIGURE 3:
<taxonomicName id="4C174D744364FFDB7B47FACEFDB08E22" authority="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori, 2012" authorityName="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori" authorityYear="2012" box="[272,612,1352,1371]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov." subGenus="Labidostoma">
<emphasis id="B963EAE54364FFDB7B47FACEFEA88E22" box="[272,380,1352,1371]" italics="true" pageId="6" pageNumber="238">Labidostoma</emphasis>
(
<emphasis id="B963EAE54364FFDB7BDBFACEFE2C8E22" box="[396,504,1352,1371]" italics="true" pageId="6" pageNumber="238">Labidostoma</emphasis>
)
<emphasis id="B963EAE54364FFDB785FFACEFDB08E22" box="[520,612,1352,1371]" italics="true" pageId="6" pageNumber="238">intermedia</emphasis>
</taxonomicName>
<emphasis id="B963EAE54364FFDB783AFACEFD798E22" bold="true" box="[621,685,1352,1371]" pageId="6" pageNumber="238">
<taxonomicNameLabel id="A250579E4364FFDB783AFACEFD798E22" box="[621,685,1352,1371]" pageId="6" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A-B oculo-pustular zone; C-D chelicera, lateral view and detail of terminal teeth of fixed digit (ventral view).
</paragraph>
</caption>
<paragraph id="8BA836F74364FFDB7A92F9CFFED08CE4" blockId="6.[159,762,1609,1949]" pageId="6" pageNumber="238">
Ventral shield — (figure 1B) Entirely covered by ornamentation. Epimeral setae short ((18-24)-(14)- (9)-(12)). Usual coxal pore on epimera I. Fourteen pores on cuticle behind the fourth epimeral plate and a transverse line of eight setae (the number of pores differs between
<taxonomicName id="4C174D744364FFDB7BF5F97FFDD28C69" authorityName="Berlese" authorityYear="1911" box="[418,518,1785,1808]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54364FFDB7BF5F97FFDD28C69" box="[418,518,1785,1808]" italics="true" pageId="6" pageNumber="238">integrum</emphasis>
</taxonomicName>
and
<emphasis id="B963EAE54364FFDB7816F97FFD698C69" box="[577,701,1785,1808]" italics="true" pageId="6" pageNumber="238">caucasicum</emphasis>
with 10 and 12 pores respectively). Anogenital ring surrounding anal and genital shields in the female, distinct genital and anal rings in the male (figures 2C and 2D).
</paragraph>
<caption id="DF68667F4364FFDB796DF8E2FC248CEA" ID-DOI="http://doi.org/10.5281/zenodo.4639930" ID-Zenodo-Dep="4639930" httpUri="https://zenodo.org/record/4639930/files/figure.png" pageId="6" pageNumber="238" startId="6.[826,838,1892,1911]" targetBox="[832,1399,1180,1839]" targetPageId="6">
<paragraph id="8BA836F74364FFDB796DF8E2FC248CEA" blockId="6.[826,1429,1892,1939]" pageId="6" pageNumber="238">
FIGURE 5:
<taxonomicName id="4C174D744364FFDB79F6F8E2FBBA8C0E" authority="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori, 2012" authorityName="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori" authorityYear="2012" box="[929,1134,1892,1911]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="6" pageNumber="238" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B963EAE54364FFDB79F6F8E2FBBA8C0E" box="[929,1134,1892,1911]" italics="true" pageId="6" pageNumber="238">Labidostoma intermedia</emphasis>
</taxonomicName>
<emphasis id="B963EAE54364FFDB7E23F8E2FB7D8C0E" bold="true" box="[1140,1193,1892,1911]" pageId="6" pageNumber="238">
<taxonomicNameLabel id="A250579E4364FFDB7E23F8E2FB7D8C0E" box="[1140,1193,1892,1911]" pageId="6" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A Tarsus I, external; B paraxial view.
</paragraph>
</caption>
<caption id="DF68667F4365FFDA7AC8FA49FC508D60" ID-DOI="http://doi.org/10.5281/zenodo.4639932" ID-Zenodo-Dep="4639932" httpUri="https://zenodo.org/record/4639932/files/figure.png" pageId="7" pageNumber="239" startId="7.[159,171,1487,1506]" targetBox="[244,1286,356,1337]" targetPageId="7">
<paragraph id="8BA836F74365FFDA7AC8FA49FC508D60" blockId="7.[159,1429,1486,1561]" pageId="7" pageNumber="239">
FIGURE 6: Compared cheliceral morphology in the
<taxonomicName id="4C174D744365FFDA78C5FA49FD368E98" authorityName="Berlese" authorityYear="1911" box="[658,738,1487,1505]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA78C5FA49FD368E98" box="[658,738,1487,1505]" italics="true" pageId="7" pageNumber="239">integrum</emphasis>
</taxonomicName>
-species group: A
<taxonomicName id="4C174D744365FFDA79F0FA48FBBE8E98" authorityName="Berlese" authorityYear="1911" box="[935,1130,1486,1505]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA79F0FA48FBBE8E98" box="[935,1130,1486,1505]" italics="true" pageId="7" pageNumber="239">Labidostoma integrum</emphasis>
</taxonomicName>
(Aude, France, Bertrand coll.); B-C
<taxonomicName id="4C174D744365FFDA7AADFA6CFE6F8E85" authorityName="Berlese" authorityYear="1911" box="[250,443,1514,1533]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA7AADFA6CFE6F8E85" box="[250,443,1514,1533]" italics="true" pageId="7" pageNumber="239">Labidostoma integrum</emphasis>
</taxonomicName>
from Grandjean 1942 Modified (possible Algerian specimen); D-E
<taxonomicName id="4C174D744365FFDA7E16FA6CFABF8E84" authority="(Romania)" authorityName="Berlese" authorityYear="1911" baseAuthorityName="Romania" box="[1089,1387,1514,1533]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA7E16FA6CFAD68E85" box="[1089,1282,1514,1533]" italics="true" pageId="7" pageNumber="239">Labidostoma integrum</emphasis>
(Romania)
</taxonomicName>
and
<emphasis id="B963EAE54365FFDA7A96F980FE418D61" box="[193,405,1542,1561]" italics="true" pageId="7" pageNumber="239">
<taxonomicName id="4C174D744365FFDA7A96F980FEF98D60" authorityName=", Kramer" authorityYear="1879" box="[193,301,1542,1561]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="genus">Labidostoma</taxonomicName>
caucasicum
</emphasis>
(Romania) (from
<bibRefCitation id="EF864B064365FFDA786BF980FCCA8D60" author="Feider Z. &amp; Vasiliu N." box="[572,798,1542,1561]" pageId="7" pageNumber="239" pagination="1868 - 1968" refId="ref6845" refString="Feider Z., Vasiliu N. 1970 - EspLces de Nicoletiellidae (Acariformes) de Roumanie - AcadOmie de la ROpublique socialiste de Roumanie. Livre du Centenaire Emile G. Racovitza 1868 - 1968. 371 - 391." type="book chapter" year="1970">Feider and Vasiliu, 1970</bibRefCitation>
, redrawn).
</paragraph>
</caption>
<paragraph id="8BA836F74365FFDA7A92F9DCFB7C8DEC" blockId="7.[159,762,1626,1931]" lastBlockId="7.[826,1429,1626,1685]" pageId="7" pageNumber="239">
Infracapitulum — The labrum (dorsal lip) is shorter than the lateral lips, which are rather large, each with a minute seta. Setae
<emphasis id="B963EAE54365FFDA785EF927FDFD8DC1" box="[521,553,1697,1720]" italics="true" pageId="7" pageNumber="239">ma</emphasis>
and
<emphasis id="B963EAE54365FFDA7833F926FD518DC1" box="[612,645,1696,1720]" italics="true" pageId="7" pageNumber="239">mb</emphasis>
, plus two additional setae near insertion of the palp. Palps with usual chaetotaxy, dorsal solenidion. The presence of additional setae was noted on Grandjeans drawings for
<taxonomicName id="4C174D744365FFDA7B68F8ABFE778C3D" authorityName="Berlese" authorityYear="1911" box="[319,419,1837,1860]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA7B68F8ABFE778C3D" box="[319,419,1837,1860]" italics="true" pageId="7" pageNumber="239">integrum</emphasis>
</taxonomicName>
; Feider and Vasiliu recorded two setae in Romanian specimens of
<taxonomicName id="4C174D744365FFDA7839F8D6FD268C1E" authorityName="Berlese" authorityYear="1911" box="[622,754,1872,1895]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA7839F8D6FD268C1E" box="[622,754,1872,1895]" italics="true" pageId="7" pageNumber="239">L. integrum</emphasis>
</taxonomicName>
, whereas they have drawn three setae in
<emphasis id="B963EAE54365FFDA7821F8F5FD268CF3" box="[630,754,1907,1930]" italics="true" pageId="7" pageNumber="239">caucasicum</emphasis>
. Such additional setae are absent on Kazakhstani specimens of
<taxonomicName id="4C174D744365FFDA798EF9F8FBE98DEC" authorityName="Berlese" authorityYear="1911" box="[985,1085,1662,1685]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA798EF9F8FBE98DEC" box="[985,1085,1662,1685]" italics="true" pageId="7" pageNumber="239">integrum</emphasis>
</taxonomicName>
(fig. 5B).
</paragraph>
<paragraph id="8BA836F74365FFD57937F934FDCC891E" blockId="7.[826,1429,1714,1948]" lastBlockId="8.[159,762,311,1008]" lastPageId="8" lastPageNumber="240" pageId="7" pageNumber="239">
Chelicerae — (figures 3C and 3D) The chelicerae differ essentially from those of the other species of the group by the shape of the paraxial tooth, which is less developed and not smooth. It differs clearly from the French (figure 6A), Algerian (figures 6B and 6C), Romanian (figure 6E) and Kazakhstani (figure 5A) specimens of
<taxonomicName id="4C174D744365FFDA7E81F803FA888CE5" authorityName="Berlese" authorityYear="1911" box="[1238,1372,1925,1948]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="7" pageNumber="239" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE54365FFDA7E81F803FA888CE5" box="[1238,1372,1925,1948]" italics="true" pageId="7" pageNumber="239">L. integrum</emphasis>
</taxonomicName>
and from
<emphasis id="B963EAE5436AFFD57A89FEBEFEA28A37" box="[222,374,311,335]" italics="true" pageId="8" pageNumber="240">L. caucasicum</emphasis>
(figure 6D) by the teeth and denticles of the mobile digit. Note that the chelicerae of French specimens of
<taxonomicName id="4C174D74436AFFD57B94FEF8FD938AEC" authorityName="Berlese" authorityYear="1911" box="[451,583,382,405]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="240" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE5436AFFD57B94FEF8FD938AEC" box="[451,583,382,405]" italics="true" pageId="8" pageNumber="240">L. integrum</emphasis>
</taxonomicName>
show an inferior tooth very similar to the drawings of
<bibRefCitation id="EF864B06436AFFD5782BFE26FE908AA2" author="Feider Z. &amp; Vasiliu N." pageId="8" pageNumber="240" pagination="1868 - 1968" refId="ref6845" refString="Feider Z., Vasiliu N. 1970 - EspLces de Nicoletiellidae (Acariformes) de Roumanie - AcadOmie de la ROpublique socialiste de Roumanie. Livre du Centenaire Emile G. Racovitza 1868 - 1968. 371 - 391." type="book chapter" year="1970">Feider and Vasiliu (1970)</bibRefCitation>
, but differ from the
<taxonomicName id="4C174D74436AFFD57811FE42FD7E8AA2" authorityName="Berlese" authorityYear="1911" box="[582,682,452,475]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="240" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE5436AFFD57811FE42FD7E8AA2" box="[582,682,452,475]" italics="true" pageId="8" pageNumber="240">integrum</emphasis>
</taxonomicName>
s chelicerae drawn by Grandjean (
<figureCitation id="132C2A72436AFFD57BB8FE60FD8F8A87" box="[495,603,486,510]" captionStart="FIGURE" captionStartId="7.[159,171,1487,1506]" captionTargetBox="[244,1286,356,1337]" captionTargetId="figure-4@7.[242,1346,302,1454]" captionTargetPageId="7" captionText="FIGURE 6: Compared cheliceral morphology in the integrum-species group: A Labidostoma integrum (Aude, France, Bertrand coll.); B-C Labidostoma integrum from Grandjean 1942 Modified (possible Algerian specimen); D-E Labidostoma integrum (Romania) and Labidostoma caucasicum (Romania) (from Feider and Vasiliu, 1970, redrawn)." figureDoi="http://doi.org/10.5281/zenodo.4639932" httpUri="https://zenodo.org/record/4639932/files/figure.png" pageId="8" pageNumber="240">Figures 6</figureCitation>
A-6D). It may be supposed that Grandjean chose North African specimens because they were of a bigger size than French ones (
<bibRefCitation id="EF864B06436AFFD57B6DFDC9FDDF891E" author="Grandjean F." box="[314,523,591,615]" pageId="8" pageNumber="240" pagination="185 - 192" refId="ref7030" refString="Grandjean F. 1942 b - Observations sur les Labidostommidae. 3 e SOrie - Bull. Mus., Paris, 2 e SOrie, 14 (3): 185 - 192." type="journal article" year="1942">Grandjean, 1942b</bibRefCitation>
).
</paragraph>
<paragraph id="8BA836F7436AFFD57A92FDFCFDF68889" blockId="8.[159,762,311,1008]" pageId="8" pageNumber="240">
Legs — Among the species-group, the first pair of legs is remarkable for its lengthened tibia, genu and mesofemur.
<taxonomicName id="4C174D74436AFFD57B3FFD47FE2389A1" authority="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori, 2012" authorityName="Bertrand &amp; Bagheri &amp; Akbari &amp; Yazdanian &amp; Irani-Nejad &amp; Mohajer &amp; Saboori" authorityYear="2012" box="[360,503,704,728]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="240" phylum="Arthropoda" rank="species" species="intermedia" status="sp. nov.">
<emphasis id="B963EAE5436AFFD57B3FFD47FE2389A1" box="[360,503,704,728]" italics="true" pageId="8" pageNumber="240">L. intermedia</emphasis>
</taxonomicName>
<emphasis id="B963EAE5436AFFD57BA8FD46FD9789A1" bold="true" box="[511,579,704,728]" pageId="8" pageNumber="240">
<taxonomicNameLabel id="A250579E436AFFD57BA8FD46FD9789A1" box="[511,579,704,728]" pageId="8" pageNumber="240" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
exhibits a relatively short genu, subequal in length to the mesofemur, the tibia being the longest article (ratio genual/tibia = 0.7 vs.
<quantity id="4CEF9B12436AFFD57B2FFCAFFE6C8838" box="[376,440,809,833]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="240" unit="in" value="0.8">0.8 in</quantity>
<taxonomicName id="4C174D74436AFFD57BE8FCACFDEB8838" authorityName="Berlese" authorityYear="1911" box="[447,575,810,833]" class="Arachnida" family="Labidostommatidae" genus="Labidostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="8" pageNumber="240" phylum="Arthropoda" rank="species" species="integrum">
<emphasis id="B963EAE5436AFFD57BE8FCACFDEB8838" box="[447,575,810,833]" italics="true" pageId="8" pageNumber="240">L. integrum</emphasis>
</taxonomicName>
) (
<tableCitation id="C695034C436AFFD57800FCAFFD7A8838" box="[599,686,809,833]" captionStart="TABLE" captionStartId="8.[159,172,1453,1472]" captionTargetPageId="8" captionText="TABLE 2: L. intermedia n. sp. Length of leg articles (4 individuals) and contribution of tibia, genu and mesofemur to the total length of PI (2nd column)." httpUri="http://table.plazi.org/id/DF68667F436AFFD57AC8FA2BFE958EA2" pageId="8" pageNumber="240" tableUuid="DF68667F436AFFD57AC8FA2BFE958EA2">Table 2</tableCitation>
). Simple setae on leg I. Tarsi of PII, PIII and PIV with subterminal ventral setae plumose (= &quot;
<emphasis id="B963EAE5436AFFD5781AFCE9FD7688FE" box="[589,674,879,903]" italics="true" pageId="8" pageNumber="240">scobales</emphasis>
&quot; sensu Feider and Vasiliu), other setae simple. Tarsus I with usual solenidia
<emphasis id="B963EAE5436AFFD57BCCFC33FE6E88B7" box="[411,442,949,974]" italics="true" pageId="8" pageNumber="240">ω1</emphasis>
and
<emphasis id="B963EAE5436AFFD57BAEFC33FDCC88B7" box="[505,536,949,974]" italics="true" pageId="8" pageNumber="240">ω2</emphasis>
, and spine-like famulus, tarsal eupathidia present.
</paragraph>
</subSubSection>
</treatment>
</document>