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<mods:title id="EB4F4B03AD5B094F45A77E4AA20C109D">Reinstatement of species belonging Marphysa sanguinea complex (Annelida Eunicidae) and description of new species from the mid-Pacific Ocean and the Adriatic Sea</mods:title>
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<mods:namePart id="7B9B2CABF3763AB33C4A95A52BF4BBAE">Idris, Izwandy</mods:namePart>
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<heading id="3058967FFFAAFFDE46D6F8C67E38F890" bold="true" box="[151,576,1828,1855]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="ACAF5A90FFAAFFDE46D6F8C67E38F890" ID-CoL="3Y896" authority="Moore, 1909" authorityName="Moore" authorityYear="1909" box="[151,576,1828,1855]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="californica">
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<emphasis id="59DBFD01FFAAFFDE46D6F8C67DE5F891" bold="true" box="[151,413,1828,1855]" italics="true" pageId="14" pageNumber="15">Marphysa californica</emphasis>
<bibRefCitation id="0F3E5CE2FFAAFFDE47E5F8C67E38F890" author="Moore, J. P." box="[420,576,1828,1854]" pageId="14" pageNumber="15" pagination="235 - 295" refId="ref31817" refString="Moore, J. P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235 - 295." type="journal article" year="1909">Moore, 1909</bibRefCitation>
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<paragraph id="6B102113FFAAFFDE46D6F8AA7DF6F8CC" blockId="14.[151,576,1828,1890]" box="[151,398,1863,1890]" pageId="14" pageNumber="15">
<figureCitation id="F3943D96FFAAFFDE46D6F8AA7D7AF8CC" box="[151,258,1863,1890]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="14" pageNumber="15">Figures 5</figureCitation>
,
<figureCitation id="F3943D96FFAAFFDE474FF8AA7D57F8CC" box="[270,303,1864,1890]" captionStart="FIGURE 9" captionStartId="26.[151,250,1874,1899]" captionTargetBox="[162,1431,194,1836]" captionTargetId="figure@26.[151,1436,181,1849]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of branchial filaments throughout the body in A. Marphysa americana Monro, 1933 n. status.; B. Marphysa baileybrockae n. sp.; C. Marphysa birgeri n. sp.; D. Marphysa brevibranchiata Treadwell, 1921 n. status.; E. Marphysa californica (Moore, 1909); F. Marphysa leidii de Quatrefages, 1866; G. Marphysa parishii Baird, 1869; H. Marphysa sanguinea (Montagu, 1813)." figureDoi="http://doi.org/10.5281/zenodo.3954065" httpUri="https://zenodo.org/record/3954065/files/figure.png" pageId="14" pageNumber="15">9E</figureCitation>
,
<tableCitation id="262D14A8FFAAFFDE477AF8AA7DF6F8CC" box="[315,398,1864,1890]" captionStart="TABLE 1" captionStartId="30.[151,235,468,493]" captionText="TABLE 1. (Continued)" pageId="14" pageNumber="15">Table 1</tableCitation>
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<paragraph id="6B102113FFAAFFDE46D6F8657EF3F810" blockId="14.[151,1437,1926,2043]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="4BBF063DFFAAFFDE46D6F8657EF3F810" pageId="14" pageNumber="15">
<treatmentCitation id="EA0E0702FFAAFFDE46D6F8657E1BF831" author="Moore, J. P." box="[151,611,1926,1951]" page="253" pageId="14" pageNumber="15" year="1909">
<taxonomicName id="ACAF5A90FFAAFFDE46D6F8657E1BF831" ID-CoL="3Y896" authority="Moore, 1909: 251 - 253" authorityName="Moore" authorityPageNumber="251 - 253" authorityYear="1909" box="[151,611,1926,1951]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFAAFFDE46D6F8657D0BF831" box="[151,371,1927,1951]" italics="true" pageId="14" pageNumber="15">Marphysa californica</emphasis>
<bibRefCitation id="0F3E5CE2FFAAFFDE4738F86A7E1BF831" author="Moore, J. P." box="[377,611,1926,1951]" pageId="14" pageNumber="15" pagination="235 - 295" refId="ref31817" refString="Moore, J. P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235 - 295." type="journal article" year="1909">Moore, 1909: 251–253</bibRefCitation>
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, Pls. 7–8, Figs. 13–20.—
<treatmentCitation id="EA0E0702FFAAFFDE4523F865782FF831" author="Carrera-Parra, L. F." box="[866,1111,1927,1951]" page="179" pageId="14" pageNumber="15" year="2009">
<bibRefCitation id="0F3E5CE2FFAAFFDE4523F865782FF831" author="Carrera-Parra, L. F." box="[866,1111,1927,1951]" pageId="14" pageNumber="15" pagination="165 - 181" refId="ref28848" refString="Carrera-Parra, L. F. (2009) Eunicidae Berthold, 1827. In: de Leon-Gonzalez, J. A., Bastida-Zavala, J. R., Carrera-Parra, L. F., Garcia-Garza, M. E., Pena-Rivera, A., Salazar-Vallejo, S. I. &amp; Solis-Weiss, V. (Eds.), Poliquetos (Annelida: Polychaeta) de Mexico y America Tropical. Universidad Autonoma de Nuevo Leon, Monterrey, pp. 165 - 181." type="book chapter" year="2009">Carrera-Parra 2009: 179</bibRefCitation>
</treatmentCitation>
.—
<bibRefCitation id="0F3E5CE2FFAAFFDE4236F864795FF831" author="Zanol, J. &amp; Halanych, K. M. &amp; Struck, T. H. &amp; Fauchald, K." box="[1143,1319,1926,1951]" pageId="14" pageNumber="15" pagination="660 - 676" refId="ref32780" refString="Zanol, J., Halanych, K. M., Struck, T. H. &amp; Fauchald, K. (2010) Phylogeny of the bristle worm family Eunicidae (Eunicida, Annelida) and the phylogenetic utility of noncongruent 16 S, COI and 18 S in combined analyses. Molecular Phylogenetics and Evolution, 55 (2), 660 - 676. https: // doi. org / 10.1016 / j. ympev. 2009.12.024" type="journal article" year="2010">
Zanol
<emphasis id="59DBFD01FFAAFFDE42F9F86A7892F831" box="[1208,1258,1927,1951]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2010
</bibRefCitation>
:
<figureCitation id="F3943D96FFAAFFDE436EF86479FDF831" box="[1327,1413,1926,1951]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="5.[151,250,1760,1785]" captionStartId-1="8.[151,250,1749,1774]" captionStartId-2="10.[151,250,1752,1777]" captionTargetBox-0="[266,1322,179,1732]" captionTargetBox-1="[269,1316,181,1723]" captionTargetBox-2="[268,1319,181,1726]" captionTargetId-0="figure@5.[266,1322,179,1732]" captionTargetId-1="figure@8.[269,1318,181,1723]" captionTargetId-2="figure@10.[268,1319,181,1726]" captionTargetPageId-0="5" captionTargetPageId-1="8" captionTargetPageId-2="10" captionText-0="FIGURE 1. Marphysa americana. Holotype BNHM 1932.12.24.554-556. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 247; K. Parapodium 377; L. Compound spinigers, chaetiger 6; M. Thin, isodont narrow, asymmetric, with long and slender teeth, chaetiger 6; N. Thick, isodont wide, symmetric, with short and slender teeth, chaetiger 325; O. Thick, isodont wide, asymmetric, with long and thick teeth, chaetiger 325; P. Thick, anodont wide, asymmetric, with long and thick teeth, chaetiger 377; Q. Subacicular hook, chaetiger 367.All chaetigers in anterior view; al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV. Scale bars: A–C, 3 mm; D, 1.4 mm; E. 0.9 mm; F. 1.3 mm; G–K, 0.2 mm; L, 0.1 mm; M–Q, 30 µm." captionText-1="FIGURE 2. Marphysa baileybrockae n. sp. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 8; I. Parapodium 16; J. Parapodium 250; K. Parapodium 308; L. Thin, isodont narrow, symmetric, with short and slender teeth, chaetiger 8; M. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 295; N. Thick, isodont wide, symmetric, with long and slender teeth, chaetiger 358; O. Compound spinigers, chaetiger 8; P. Subacicular hook unidentate, chaetiger 122; Q. Subacicular hook bidentate, chaetiger 152. A–O from holotype USNM 5444, P–Q from paratype AMNH 367. All chaetigers in anterior view; al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV. Scale bars: A–C, 2.9 mm; D–E, 1 mm; F, 0.8 mm; G–K, 0.1 mm; L–Q, 30 µm." captionText-2="FIGURE 3. Marphysa birgeri n. sp. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior end, lateral view; D. Maxillary apparatus, dorsal view; E. Mandible; F. Parapodium 3; G. Parapodium 7; H. Parapodium 14; I. Parapodium 67; J. Parapodium 103; K. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 14; L. Thin, isodont wide, asymmetric, with short and slender teeth, chaetiger 85; M. Thick, anodont wide, symmetric, with long and slender teeth, chaetiger 85; N. Thick, anodont wide, asymmetric, with long and thick teeth, chaetiger 105; O. Compound spinigers, chaetiger 7; P. Subacicular hook, chaetiger 85. A–O from holotype F.2021, P from paratype 12273. All chaetigers in anterior view; al-MIII: attachment lamella MIII; al-MIV: attachment lamella MIV. Scale bars: A–C, 1.4 mm; D–E, 0.5 mm; F–J, 0.2 mm; K–N, 30 µm; O–P, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3954049" figureDoi-1="http://doi.org/10.5281/zenodo.3954051" figureDoi-2="http://doi.org/10.5281/zenodo.3954053" httpUri-0="https://zenodo.org/record/3954049/files/figure.png" httpUri-1="https://zenodo.org/record/3954051/files/figure.png" httpUri-2="https://zenodo.org/record/3954053/files/figure.png" pageId="14" pageNumber="15">Fig. 1–3</figureCitation>
,
<figureCitation id="F3943D96FFAAFFDE43D1F86479E5F831" box="[1424,1437,1926,1951]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="14" pageNumber="15">5</figureCitation>
(phylogenetic trees).—
<treatmentCitation id="EA0E0702FFAAFFDE47F0F8477EFCF810" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[433,644,1957,1982]" page="93" pageId="14" pageNumber="15" year="2014">
<bibRefCitation id="0F3E5CE2FFAAFFDE47F0F8477EFCF810" author="Zanol, J. &amp; Halanych, K. M. &amp; Fauchald, K." box="[433,644,1957,1982]" pageId="14" pageNumber="15" pagination="79 - 100" refId="ref32863" refString="Zanol, J., Halanych, K. M. &amp; Fauchald, K. (2014) Reconciling taxonomy and phylogeny in the bristleworm family Eunicidae (Polychaete: Annelida). Zoologica Scripta, 43, 79 - 100. https: // doi. org / 10.1111 / zsc. 12034" type="journal article" year="2014">
Zanol
<emphasis id="59DBFD01FFAAFFDE47B3F8457E67F810" box="[498,543,1958,1982]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2014: 93
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="6B102113FFAAFFDE46D6F8267D71F855" blockId="14.[151,1437,1926,2043]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="4BBF063DFFAAFFDE46D6F8267D71F855" pageId="14" pageNumber="15">
<emphasis id="59DBFD01FFAAFFDE46D6F8267D00F872" box="[151,376,1988,2012]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="ACAF5A90FFAAFFDE46D6F8267D0CF872" ID-CoL="3Y8B5" baseAuthorityName="Montagu" baseAuthorityYear="1813" box="[151,372,1988,2012]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="14" pageNumber="15" phylum="Annelida" rank="species" species="sanguinea">Marphysa sanguinea</taxonomicName>
.
</emphasis>
—
<treatmentCitation id="EA0E0702FFAAFFDE47D0F8277E45F873" author="Monro, C. C. A." box="[401,573,1988,2013]" page="69" pageId="14" pageNumber="15" year="1933">
<bibRefCitation id="0F3E5CE2FFAAFFDE47D0F8277E45F873" author="Monro, C. C. A." box="[401,573,1988,2013]" pageId="14" pageNumber="15" pagination="1 - 196" refId="ref31679" refString="Monro, C. C. A. (1933) The Polychaeta Errantia collected by Dr. C. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the expedition of the S. Y. ' St. George'. Proceedings of the Zoological Society of London, 1933, 1 - 196. https: // doi. org / 10.1111 / j. 1096 - 3642.1933. tb 01578. x" type="journal article" year="1933">Monro 1933: 69</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="EA0E0702FFAAFFDE441FF8277F13F872" author="Hartman, O." box="[606,875,1988,2012]" page="127" pageId="14" pageNumber="15" year="1944">
<bibRefCitation id="0F3E5CE2FFAAFFDE441FF8277F13F872" author="Hartman, O." box="[606,875,1988,2012]" pageId="14" pageNumber="15" pagination="1 - 237" refId="ref30209" refString="Hartman, O. (1944 a) Polychaetous Annelids. Part V. Eunicea. Allan Hancock Pacific Expedition, 10 (1), 1 - 237." type="journal article" year="1944">Hartman 1944a: 127–128</bibRefCitation>
</treatmentCitation>
, Pl. 8, Figs. 179–183.—1961: 84–85 (
<emphasis id="59DBFD01FFAAFFDE434BF8277948F872" box="[1290,1328,1989,2012]" italics="true" pageId="14" pageNumber="15">non</emphasis>
<bibRefCitation id="0F3E5CE2FFAAFFDE437BF8277C85F855" author="Montagu, G." pageId="14" pageNumber="15" pagination="18 - 21" refId="ref31759" refString="Montagu, G. (1813) Descriptions of several new or rare animals principally marine, found on the south coast of Devonshire. Transactions of the Linnean Society of London, 11, 18 - 21. https: // doi. org / 10.1111 / j. 1096 - 3642.1813. tb 00035. x" type="journal article" year="1813">Montagu, 1813</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="23B57298FFABFFDF46D6FF7A78CFFF55" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="6B102113FFABFFDF46D6FF7A78CFFF55" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="59DBFD01FFABFFDF46D6FF7A7E5FFF1D" box="[151,551,152,179]" italics="true" pageId="15" pageNumber="16">
<emphasis id="59DBFD01FFABFFDF46D6FF7A7D0BFF1D" bold="true" box="[151,371,152,179]" italics="true" pageId="15" pageNumber="16">Material examined</emphasis>
. Type material:
</emphasis>
<materialsCitation id="DBC72B4EFFABFFDF4471FF7B7C9DFF79" collectionCode="CAS" collectorName="E. C. Starks." country="United States of America" county="San Diego County" location="San Diego County" pageId="15" pageNumber="16" specimenCode="CAS 020357" specimenCount="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="B4149FB1FFABFFDF4471FF7B7EE0FF1D" box="[560,664,153,179]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
<specimenCode id="3B098968FFABFFDF44E1FF7B7F4EFF1D" box="[672,822,152,179]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="16" type="Museum">CAS 020357</specimenCode>
,
<collectingCounty id="8271599FFFABFFDF4503FF7B786EFF1D" box="[834,1046,153,179]" pageId="15" pageNumber="16">San Diego County</collectingCounty>
,
<collectingRegion id="A96BEFF1FFABFFDF4260FF7B78EDFF1D" box="[1057,1173,153,179]" country="United States of America" name="California" pageId="15" pageNumber="16">California</collectingRegion>
,
<collectingCountry id="13B86183FFABFFDF42E3FF7B78A5FF1D" box="[1186,1245,153,179]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
, 1902, coll.
<collectorName id="C65A44C5FFABFFDF432BFF7B7C9DFF79" pageId="15" pageNumber="16">E.C. Starks.</collectorName>
</materialsCitation>
<emphasis id="59DBFD01FFABFFDF46B1FF5F7DA2FF79" box="[240,474,189,215]" italics="true" pageId="15" pageNumber="16">Additional material:</emphasis>
<materialsCitation id="DBC72B4EFFABFFDF47A7FF5F79E0FF79" box="[486,1432,188,216]" collectingDate="1911-04-10" collectionCode="LACM-AHF" country="United States of America" location="Point Fermin" pageId="15" pageNumber="16" specimenCode="POLY 52710" specimenCount="1" stateProvince="California">
<collectionCode id="0DBEB9D6FFABFFDF47A7FF5F7E00FF79" box="[486,632,189,215]" pageId="15" pageNumber="16">LACM-AHF</collectionCode>
<specimenCode id="3B098968FFABFFDF44C4FF5F7F64FF79" box="[645,796,188,215]" collectionCode="POLY" pageId="15" pageNumber="16">POLY 52710</specimenCode>
(
<specimenCount id="7DA9EA9AFFABFFDF4572FF5F7FC1FF76" box="[819,953,189,216]" pageId="15" pageNumber="16" type="generic">1 specimen</specimenCount>
),
<location id="6E7077C8FFABFFDF4590FF5F7814FF79" LSID="urn:lsid:plazi:treatment:E3069005FFAAFFC146D6F8C6789DFAF9:6E7077C8FFABFFDF4590FF5F7814FF79" box="[977,1132,189,215]" country="United States of America" name="Point Fermin" pageId="15" pageNumber="16" stateProvince="California">Point Fermin</location>
,
<collectingRegion id="A96BEFF1FFABFFDF423DFF5F7888FF79" box="[1148,1264,189,215]" country="United States of America" name="California" pageId="15" pageNumber="16">California</collectingRegion>
,
<date id="1F1107D3FFABFFDF4341FF5F79E0FF79" box="[1280,1432,189,215]" pageId="15" pageNumber="16" value="1911-04-10">
<collectingDate id="0F55FE3BFFABFFDF4341FF5F79E0FF79" box="[1280,1432,189,215]" pageId="15" pageNumber="16" value="1911-04-10">10 Apr 1911</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="DBC72B4EFFABFFDF46D6FF0378CBFF55" box="[151,1203,224,252]" collectingDate="1935-10-26" collectionCode="LACM-AHF, POLY" country="United States of America" location="Redondo Beach" municipality="Bluff Cove" pageId="15" pageNumber="16" specimenCount="1" stateProvince="California">
<collectionCode id="0DBEB9D6FFABFFDF46D6FF037D0DFF55" box="[151,373,225,251]" pageId="15" pageNumber="16">LACM-AHF POLY</collectionCode>
0000 (
<specimenCount id="7DA9EA9AFFABFFDF4786FF037E30FF52" box="[455,584,225,252]" pageId="15" pageNumber="16" type="generic">1 specimen</specimenCount>
),
<collectingMunicipality id="8B74BB69FFABFFDF441AFF037EA3FF55" box="[603,731,225,251]" pageId="15" pageNumber="16">Bluff Cove</collectingMunicipality>
,
<location id="6E7077C8FFABFFDF44A7FF037FE3FF55" LSID="urn:lsid:plazi:treatment:E3069005FFAAFFC146D6F8C6789DFAF9:6E7077C8FFABFFDF44A7FF037FE3FF55" box="[742,923,225,251]" country="United States of America" municipality="Bluff Cove" name="Redondo Beach" pageId="15" pageNumber="16" stateProvince="California">Redondo Beach</location>
,
<collectingRegion id="A96BEFF1FFABFFDF45E7FF037862FF55" box="[934,1050,225,251]" country="United States of America" name="California" pageId="15" pageNumber="16">California</collectingRegion>
,
<date id="1F1107D3FFABFFDF4264FF0378CBFF55" box="[1061,1203,224,251]" pageId="15" pageNumber="16" value="1935-10-26">
<collectingDate id="0F55FE3BFFABFFDF4264FF0378CBFF55" box="[1061,1203,224,251]" pageId="15" pageNumber="16" value="1935-10-26">26 Oct 1935</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="23B57298FFABFFC14686FEE77E53FEB1" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="6B102113FFABFFDF4686FEE77E00FEC9" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
<emphasis id="59DBFD01FFABFFDF4686FEE77D2CFEB1" bold="true" box="[199,340,261,287]" italics="true" pageId="15" pageNumber="16">Description.</emphasis>
<typeStatus id="B4149FB1FFABFFDF471BFEE77DBAFEB1" box="[346,450,261,287]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
incomplete, ventrally dissected, with 92 chaetigers, L10 =
<quantity id="AC578CF6FFABFFDF420DFEE678C9FEB1" box="[1100,1201,260,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="15" pageNumber="16" unit="mm" value="15.6">15.6 mm</quantity>
, W10 =
<quantity id="AC578CF6FFABFFDF434DFEE77919FEB1" box="[1292,1377,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="15" pageNumber="16" unit="mm" value="8.3">8.3 mm</quantity>
TL =
<quantity id="AC578CF6FFABFFDF46D6FECB7C90FEED" box="[151,232,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="15" pageNumber="16" unit="mm" value="86.0">86 mm</quantity>
. Anterior region of the body with dorsum convex and flat ventrum; body depressed from chaetiger 11, widest at chaetiger 30, tapering after chaetiger 43.
</paragraph>
<paragraph id="6B102113FFABFFDF4686FE937EFAFDB5" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Prostomium bilobed,
<quantity id="AC578CF6FFABFFDF47FBFE937E68FE25" box="[442,528,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="15" pageNumber="16" unit="mm" value="3.1">3.1 mm</quantity>
long,
<quantity id="AC578CF6FFABFFDF4416FE927EE0FE25" box="[599,664,368,395]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="mm" value="5.0">5 mm</quantity>
wide; lobes frontally rounded; median sulcus shallow anteriorly and deep ventrally (
<figureCitation id="F3943D96FFABFFDF4707FE777DFFFE01" box="[326,391,404,431]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
A–B). Prostomial appendages in a semicircle, median antenna isolated by a gap. Palps reaching first chaetiger; lateral antennae reaching middle of first chaetiger; median antenna reaching second chaetiger. Palpophores and ceratophores ring-shaped, short, thick; palpostyles and ceratostyles tapering, slender. Eyes ovate, brown, between palps, and lateral antennae.
</paragraph>
<paragraph id="6B102113FFABFFDF4686FDC77EB7FD29" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Peristomium (
<quantity id="AC578CF6FFABFFDF4726FDC77DB9FD91" box="[359,449,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="16" unit="mm" value="3.8">3.8 mm</quantity>
long,
<quantity id="AC578CF6FFABFFDF444AFDC77E37FD91" box="[523,591,549,575]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="16" unit="mm" value="9.0">9 mm</quantity>
wide) larger and wider than prostomium; first ring two times longer than second ring, separation between rings distinct on all sides (
<figureCitation id="F3943D96FFABFFDF4503FDAB7FFEFDCD" box="[834,902,584,611]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
A–C). Ventral lip with a slight central anterior depression and several shallow wrinkles (
<figureCitation id="F3943D96FFABFFDF442BFD8F7EBAFD29" box="[618,706,620,647]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="6B102113FFABFFDF4686FD7378D2FC41" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Maxillary apparatus with MF = 1 + 1, 4 + 4, 5 + 0, 4 + 8, 1 + 1 (
<figureCitation id="F3943D96FFABFFDF45D9FD737F89FD05" box="[920,1009,656,683]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5D</figureCitation>
). MI three times longer than length of maxillary carriers. MI forceps-like, MI 4.4 times longer than length of closing system; ligament between MI and
<collectingRegion id="A96BEFF1FFABFFDF46D6FD3B7CBDFD5D" box="[151,197,729,755]" country="Japan" name="Mie" pageId="15" pageNumber="16">MII</collectingRegion>
, strongly sclerotized.
<collectingRegion id="A96BEFF1FFABFFDF4781FD3B7D94FD5D" box="[448,492,729,755]" country="Japan" name="Mie" pageId="15" pageNumber="16">MII</collectingRegion>
with recurved triangular teeth;
<collectingRegion id="A96BEFF1FFABFFDF4516FD3B7FFBFD5D" box="[855,899,729,755]" country="Japan" name="Mie" pageId="15" pageNumber="16">MII</collectingRegion>
4.5 times longer than length of cavity opening (
<figureCitation id="F3943D96FFABFFDF46E1FD1F7C9AFCB9" box="[160,226,764,791]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
D–E); ligament between
<collectingRegion id="A96BEFF1FFABFFDF47BFFD1F7E52FCB9" box="[510,554,765,791]" country="Japan" name="Mie" pageId="15" pageNumber="16">MII</collectingRegion>
and MIII and right MIV sclerotized. MIII with triangular teeth; with irregular attachment lamella, situated only in center of right edge of the plate, sclerotized (
<figureCitation id="F3943D96FFABFFDF4268FCC37813FC95" box="[1065,1131,800,827]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
D–E). Left MIV with three lateral teeth smaller than rest; attachment lamella semicircle, slender, better developed on right side, situated 2/3 of anterior edge of maxilla. Right MIV with three lateral teeth larger than rest; attachment lamella semicircle, wide, better developed in central side, situated 3/4 of anterior edge of maxilla, sclerotized (
<figureCitation id="F3943D96FFABFFDF4210FC6F78EBFC09" box="[1105,1171,908,935]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
D–E). MV square, with a short, rounded tooth. Mandibles dark, becoming light amber toward the outer edge; with calcareous cutting plates present and sclerotized cutting plates brown (
<figureCitation id="F3943D96FFABFFDF44D2FC377E90FC41" box="[659,744,980,1007]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5F</figureCitation>
). Number of growth rings not observed.
</paragraph>
<paragraph id="6B102113FFABFFDF4686FC1B7D74FBD1" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Pectinate branchiae with up to six long filaments, present from chaetigers 31L–34R to last chaetiger of the fragment (
<figureCitation id="F3943D96FFABFFDF469FFBFF7D67FB99" box="[222,287,1052,1079]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
J–K). Two filaments in first chaetiger; reaching the maximum five or six filaments from chaetigers 58L to the last chaetiger of the fragment (
<figureCitation id="F3943D96FFABFFDF447AFBA37EEBFBF5" box="[571,659,1089,1115]" captionStart="FIGURE 9" captionStartId="26.[151,250,1874,1899]" captionTargetBox="[162,1431,194,1836]" captionTargetId="figure@26.[151,1436,181,1849]" captionTargetPageId="26" captionText="FIGURE 9. Distribution of branchial filaments throughout the body in A. Marphysa americana Monro, 1933 n. status.; B. Marphysa baileybrockae n. sp.; C. Marphysa birgeri n. sp.; D. Marphysa brevibranchiata Treadwell, 1921 n. status.; E. Marphysa californica (Moore, 1909); F. Marphysa leidii de Quatrefages, 1866; G. Marphysa parishii Baird, 1869; H. Marphysa sanguinea (Montagu, 1813)." figureDoi="http://doi.org/10.5281/zenodo.3954065" httpUri="https://zenodo.org/record/3954065/files/figure.png" pageId="15" pageNumber="16">Fig. 9E</figureCitation>
). Branchial filaments longer than dorsal cirri except in the first three branchiae.
</paragraph>
<paragraph id="6B102113FFABFFDF4686FB6B7F25FA6D" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
First two parapodia smaller, best developed in chaetigers 5–32, following ones becoming gradually smaller. Dorsal cirri conical; longer than ventral cirri in anterior region, following ones of similar size; best developed in chaetigers 4–25, following ones gradually smaller (
<figureCitation id="F3943D96FFABFFDF44A4FB337F50FB45" box="[741,808,1232,1259]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K). Prechaetal lobes short, as transverse folds in all chaetigers (
<figureCitation id="F3943D96FFABFFDF475AFB177D18FAA1" box="[283,352,1268,1295]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K). Chaetal lobes in the first 46 chaetigers rectangular, shorter than postchaetal lobe, with aciculae emerging dorsal to midline; from chaetiger 47 triangular, longer than other lobes (
<figureCitation id="F3943D96FFABFFDF42C7FAFB78BFFA9D" box="[1158,1223,1304,1331]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K), with acicula emerging in midline. Postchaetal lobes well developed in the first 92 chaetigers; digitiform in the first three chaetigers, ovoid in chaetigers 4–10, rounded from chaetiger 11, progressively smaller from chaetiger 22; from chaetiger 93 inconspicuous (
<figureCitation id="F3943D96FFABFFDF4728FA677DD1FA31" box="[361,425,1412,1439]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K). Ventral cirri digitiform in first 12 chaetigers; in chaetigers 13 to last part of the fragment with an oval swollen base and digitiform tip (
<figureCitation id="F3943D96FFABFFDF4492FA4B7F6DFA6D" box="[723,789,1448,1475]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K).
</paragraph>
<paragraph id="6B102113FFABFFDF4686FA2F7DFFF981" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Aciculae blunt, reddish along most of its length, amber on the distal tip (
<figureCitation id="F3943D96FFABFFDF45ACFA2F7856FA49" box="[1005,1070,1484,1511]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
G–K). First 2 chaetigers with six aciculae; in chaetigers 3–9 with five; in chaetigers 10–54 with four; in chaetigers 55–61 with three; from chaetiger 62 with two aciculae.
</paragraph>
<paragraph id="6B102113FFABFFDF4686F9DB796DF839" blockId="15.[151,1437,152,2015]" pageId="15" pageNumber="16">
Limbate chaetae of two lengths in same chaetiger: long and short, long blades are in dorsal position, short blades in ventral position; limbate chaetae reduced in number around chaetiger 28, and then maintained a similar number until the posterior end. Four
<typeStatus id="B4149FB1FFABFFDF4794F9637E68F935" box="[469,528,1665,1691]" pageId="15" pageNumber="16">types</typeStatus>
of pectinate chaetae; in anterior chaetigers, thin, isodont narrow, symmetric, with short, and slender teeth, 2–3 chaetae, up to 15 teeth (
<figureCitation id="F3943D96FFABFFDF44B2F9477F33F911" box="[755,843,1700,1727]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5L</figureCitation>
). In median-posterior chaetigers, thin, isodont wide, symmetric, with short, and slender teeth, 8–9 chaetae, up to 26 teeth (
<figureCitation id="F3943D96FFABFFDF45DDF92B7F81F94D" box="[924,1017,1736,1763]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5M</figureCitation>
), and thick, isodont wide, symmetric, with long and thick teeth, 1–2 pectinate, and up to 18 teeth. In posterior chaetigers, thick, pectinate anodont, symmetric, wide with long and thick teeth, 1–2 chaetae, up to 18 teeth (
<figureCitation id="F3943D96FFABFFDF45C6F8F37F9AF885" box="[903,994,1808,1835]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5N</figureCitation>
). Compound spinigers present throughout, in anterior chaetiger with blades of two lengths: shorter blades slightly more abundant than longer blades (
<figureCitation id="F3943D96FFABFFDF432EF8D77CC5F8DD" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5O</figureCitation>
). Subacicular hook absent; in additional material LACM-AHF POLY 52710 (L10 =
<quantity id="AC578CF6FFABFFDF422AF8BA78B7F8DD" box="[1131,1231,1880,1907]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="15" pageNumber="16" unit="mm" value="13.5">13.5 mm</quantity>
) subacicular hook unidentate, translucent; starting from chaetiger 204, present discontinuously in all chaetigers (
<figureCitation id="F3943D96FFABFFDF42F3F89F7970F839" box="[1202,1288,1916,1943]" captionStart="FIGURE 5" captionStartId="16.[151,250,1761,1786]" captionTargetBox="[265,1322,181,1736]" captionTargetId="figure@16.[265,1322,181,1736]" captionTargetPageId="16" captionText="FIGURE 5. Marphysa californica Moore, 1909. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars:A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="15" pageNumber="16">Fig. 5P</figureCitation>
).
</paragraph>
<paragraph id="6B102113FFABFFC14686F8437E53FEB1" blockId="15.[151,1437,152,2015]" lastBlockId="17.[151,1437,152,1367]" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
<emphasis id="59DBFD01FFABFFDF4686F8437D41F815" bold="true" box="[199,313,1953,1979]" italics="true" pageId="15" pageNumber="16">Variation.</emphasis>
Material examined with L10 =
<quantity id="AC578CF6FFABFFDF44DFF8427F56F815" box="[670,814,1952,1979]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.9" metricValueMin="1.35" pageId="15" pageNumber="16" unit="mm" value="16.25" valueMax="19.0" valueMin="13.5">13.5–19 mm</quantity>
, W10 =
<quantity id="AC578CF6FFABFFDF45CCF842785EF815" box="[909,1062,1952,1979]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" metricValueMax="10.3" metricValueMin="6.5" pageId="15" pageNumber="16" unit="mm" value="8.4" valueMax="10.3" valueMin="6.5">6.5–10.3 mm</quantity>
. All specimens incomplete. Palps reaching second peristomial ring or first chaetiger; lateral antennae reaching middle of first chaetiger; median antenna reaching second chaetiger. Maxillary formula is variable:
<collectingRegion id="A96BEFF1FFB5FFC145CBFF7B7FCEFF1D" box="[906,950,153,179]" country="Japan" name="Mie" pageId="17" pageNumber="18">MII</collectingRegion>
3–4 + 5, MIII 5, MIV 4 + 7–8. Branchiae from chaetigers 28–40. Maximum number of branchial filaments varied from four to eight and postchaetal lobes were conspicuous in first 41–92 chaetigers. Ventral cirri with a swollen base from chaetigers 13–14. Subacicular hook from chaetigers 204 or absent.
</paragraph>
</subSubSection>
<caption id="3FD0719BFFB4FFC046D6F9037E7DF85B" ID-DOI="http://doi.org/10.5281/zenodo.3954057" ID-Zenodo-Dep="3954057" httpUri="https://zenodo.org/record/3954057/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1761,1786]" targetBox="[265,1322,181,1736]" targetPageId="16">
<paragraph id="6B102113FFB4FFC046D6F9037E7DF85B" blockId="16.[151,1437,1761,2038]" pageId="16" pageNumber="17">
<emphasis id="59DBFD01FFB4FFC046D6F9037D6BF954" bold="true" box="[151,275,1761,1786]" pageId="16" pageNumber="17">FIGURE 5.</emphasis>
<taxonomicName id="ACAF5A90FFB4FFC0475BF9007EFAF954" authority="Moore, 1909" authorityName="Moore" authorityYear="1909" box="[282,642,1762,1786]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="16" pageNumber="17" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB4FFC0475BF9007D8EF954" box="[282,502,1762,1786]" italics="true" pageId="16" pageNumber="17">Marphysa californica</emphasis>
<bibRefCitation id="0F3E5CE2FFB4FFC047BDF9007EFAF954" author="Moore, J. P." box="[508,642,1762,1786]" pageId="16" pageNumber="17" pagination="235 - 295" refId="ref31817" refString="Moore, J. P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235 - 295." type="journal article" year="1909">Moore, 1909</bibRefCitation>
</taxonomicName>
. A. Anterior end, dorsal view; B. Anterior end, ventral view; C. Anterior view, lateral view; D. Maxillary apparatus, dorsal view; E. Left MI-II-III-IV-V, lateral view; F. Mandible; G. Parapodium 3; H. Parapodium 6; I. Parapodium 12; J. Parapodium 86; K. Parapodium 212; L. Thin, isodont narrow, symmetric, with long and slender teeth, chaetiger 17; M. Thin, isodont wide, symmetric, with short and slender teeth, chaetiger 86; N. Thick, anodont wide, symmetric, with long and thick teeth, chaetiger 212; O. Compound spinigers, chaetiger 12; P. Subacicular hook, chaetiger 212. A–J, O from holotype CAS 020357; K, M–N, P from LACM-AHF POLY 52710. All chaetigers in anterior view; LMI-II: Ligament between MI and MII; LMII-III: Ligament between MII and MIII. Scale bars: A–C, 3.9 mm; D–E, 1.6 mm; F, 1.5 mm; G–K, 0.2 mm; L–M, N, P, 30 µm; O, 0.1 mm.
</paragraph>
</caption>
<subSubSection id="23B57298FFB5FFC14686FECA7E13FE25" pageId="17" pageNumber="18" type="distribution">
<paragraph id="6B102113FFB5FFC14686FECA7E58FEED" blockId="17.[151,1437,152,1367]" box="[199,544,296,323]" pageId="17" pageNumber="18">
<emphasis id="59DBFD01FFB5FFC14686FECA7D22FEED" bold="true" box="[199,346,296,323]" italics="true" pageId="17" pageNumber="18">Distribution.</emphasis>
<collectingRegion id="A96BEFF1FFB5FFC14720FECB7DADFEED" box="[353,469,297,323]" country="United States of America" name="California" pageId="17" pageNumber="18">California</collectingRegion>
,
<collectingCountry id="13B86183FFB5FFC147A0FECB7E65FEED" box="[481,541,297,323]" name="United States of America" pageId="17" pageNumber="18">USA</collectingCountry>
.
</paragraph>
<paragraph id="6B102113FFB5FFC14686FEAE7E13FE25" blockId="17.[151,1437,152,1367]" pageId="17" pageNumber="18">
<emphasis id="59DBFD01FFB5FFC14686FEAE7D5DFEC9" bold="true" box="[199,293,332,359]" italics="true" pageId="17" pageNumber="18">Habitat.</emphasis>
According to
<bibRefCitation id="0F3E5CE2FFB5FFC14785FEAF7E18FEC9" author="Moore, J. P." box="[452,608,333,359]" pageId="17" pageNumber="18" pagination="235 - 295" refId="ref31817" refString="Moore, J. P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235 - 295." type="journal article" year="1909">Moore (1909)</bibRefCitation>
the
<typeStatus id="B4149FB1FFB5FFC144D1FEAF7E8AFEC9" box="[656,754,333,359]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
was collected by Profesor E.C. Starks from San Diego Bays, in the sand-bar, within the intertidal zone.
</paragraph>
</subSubSection>
<subSubSection id="23B57298FFB5FFC14686FE76789DFAF9" pageId="17" pageNumber="18" type="discussion">
<paragraph id="6B102113FFB5FFC14686FE767E83FCF1" blockId="17.[151,1437,152,1367]" pageId="17" pageNumber="18">
<emphasis id="59DBFD01FFB5FFC14686FE767E56FE01" box="[199,558,404,431]" italics="true" pageId="17" pageNumber="18">
<emphasis id="59DBFD01FFB5FFC14686FE767D4DFE01" bold="true" box="[199,309,404,431]" italics="true" pageId="17" pageNumber="18">Remarks.</emphasis>
<taxonomicName id="ACAF5A90FFB5FFC1477AFE777E56FE01" authorityName="Moore" authorityYear="1909" box="[315,558,405,431]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">Marphysa californica</taxonomicName>
</emphasis>
, described by
<bibRefCitation id="0F3E5CE2FFB5FFC14491FE777F14FE01" author="Moore, J. P." box="[720,876,405,431]" pageId="17" pageNumber="18" pagination="235 - 295" refId="ref31817" refString="Moore, J. P. (1909) Polychaetous annelids from Monterey Bay and San Diego, California. Proceedings of the Academy of Natural Sciences of Philadelphia, 61, 235 - 295." type="journal article" year="1909">Moore (1909)</bibRefCitation>
from San Diego (
<collectingRegion id="A96BEFF1FFB5FFC14272FE7778D3FE1E" box="[1075,1195,405,432]" country="United States of America" name="California" pageId="17" pageNumber="18">California</collectingRegion>
), it was later consider as a junior synonym of
<taxonomicName id="ACAF5A90FFB5FFC147E8FE587E3CFE7D" baseAuthorityName="Montagu" baseAuthorityYear="1813" box="[425,580,442,467]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="sanguinea">
<emphasis id="59DBFD01FFB5FFC147E8FE587E3CFE7D" box="[425,580,442,467]" italics="true" pageId="17" pageNumber="18">M. sanguinea</emphasis>
</taxonomicName>
by
<bibRefCitation id="0F3E5CE2FFB5FFC14433FE5B7F6DFE7D" author="Monro, C. C. A." box="[626,789,441,467]" pageId="17" pageNumber="18" pagination="1 - 196" refId="ref31679" refString="Monro, C. C. A. (1933) The Polychaeta Errantia collected by Dr. C. Crossland at Colon, in the Panama Region, and the Galapagos Islands during the expedition of the S. Y. ' St. George'. Proceedings of the Zoological Society of London, 1933, 1 - 196. https: // doi. org / 10.1111 / j. 1096 - 3642.1933. tb 01578. x" type="journal article" year="1933">Monro (1933)</bibRefCitation>
. Based on the distance between the
<typeStatus id="B4149FB1FFB5FFC142F7FE5B789EFE7D" box="[1206,1254,441,467]" pageId="17" pageNumber="18">type</typeStatus>
localities of
<taxonomicName id="ACAF5A90FFB5FFC1433FFE587D72FE59" baseAuthorityName="Montagu" baseAuthorityYear="1813" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="sanguinea">
<emphasis id="59DBFD01FFB5FFC1433FFE587D72FE59" italics="true" pageId="17" pageNumber="18">M. sanguinea</emphasis>
</taxonomicName>
and
<taxonomicName id="ACAF5A90FFB5FFC1477EFE3C7EADFE59" authority=", Carrera-Parra (2009)" authorityName=", Carrera-Parra" authorityYear="2009" box="[319,725,477,503]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC1477EFE3C7DA6FE59" box="[319,478,477,503]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
,
<bibRefCitation id="0F3E5CE2FFB5FFC147AAFE3F7EADFE59" author="Carrera-Parra, L. F." box="[491,725,477,503]" pageId="17" pageNumber="18" pagination="165 - 181" refId="ref28848" refString="Carrera-Parra, L. F. (2009) Eunicidae Berthold, 1827. In: de Leon-Gonzalez, J. A., Bastida-Zavala, J. R., Carrera-Parra, L. F., Garcia-Garza, M. E., Pena-Rivera, A., Salazar-Vallejo, S. I. &amp; Solis-Weiss, V. (Eds.), Poliquetos (Annelida: Polychaeta) de Mexico y America Tropical. Universidad Autonoma de Nuevo Leon, Monterrey, pp. 165 - 181." type="book chapter" year="2009">Carrera-Parra (2009)</bibRefCitation>
</taxonomicName>
suggested a review on
<taxonomicName id="ACAF5A90FFB5FFC14597FE3C780DFE59" authorityName=", Carrera-Parra" authorityYear="2009" box="[982,1141,477,503]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14597FE3C780DFE59" box="[982,1141,477,503]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
to solve the validity of the species. Later, according to molecular evidence taken from the
<typeStatus id="B4149FB1FFB5FFC14517FDE37FFEFDB5" box="[854,902,513,539]" pageId="17" pageNumber="18">type</typeStatus>
locality,
<bibRefCitation id="0F3E5CE2FFB5FFC145ADFDE378C5FDB5" author="Zanol, J. &amp; Halanych, K. M. &amp; Struck, T. H. &amp; Fauchald, K." box="[1004,1213,513,539]" pageId="17" pageNumber="18" pagination="660 - 676" refId="ref32780" refString="Zanol, J., Halanych, K. M., Struck, T. H. &amp; Fauchald, K. (2010) Phylogeny of the bristle worm family Eunicidae (Eunicida, Annelida) and the phylogenetic utility of noncongruent 16 S, COI and 18 S in combined analyses. Molecular Phylogenetics and Evolution, 55 (2), 660 - 676. https: // doi. org / 10.1016 / j. ympev. 2009.12.024" type="journal article" year="2010">
Zanol
<emphasis id="59DBFD01FFB5FFC14275FDE0781DFDB5" box="[1076,1125,513,539]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2010)
</bibRefCitation>
recognized
<taxonomicName id="ACAF5A90FFB5FFC14305FDE07C90FD91" authorityName=", Carrera-Parra" authorityYear="2009" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14305FDE07C90FD91" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
as valid species and different from
<taxonomicName id="ACAF5A90FFB5FFC14434FDC47F75FD91" baseAuthorityName="Montagu" baseAuthorityYear="1813" box="[629,781,550,575]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="sanguinea">
<emphasis id="59DBFD01FFB5FFC14434FDC47F75FD91" box="[629,781,550,575]" italics="true" pageId="17" pageNumber="18">M. sanguinea</emphasis>
</taxonomicName>
, although they did not clarify its taxonomic status. Herein, after a morphological comparison between both
<typeStatus id="B4149FB1FFB5FFC144EFFDAB7EA6FDCD" box="[686,734,585,611]" pageId="17" pageNumber="18">type</typeStatus>
and additional materials, the following differences were found. In
<taxonomicName id="ACAF5A90FFB5FFC146F7FD8C7D2EFD29" authorityName=", Carrera-Parra" authorityYear="2009" box="[182,342,621,647]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC146F7FD8C7D2EFD29" box="[182,342,621,647]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
(L10 =
<quantity id="AC578CF6FFB5FFC147F3FD8E7E39FD29" box="[434,577,620,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.9" metricValueMin="1.35" pageId="17" pageNumber="18" unit="mm" value="16.25" valueMax="19.0" valueMin="13.5">13.5–19 mm</quantity>
) the branchiae start from chaetigers 28–40, the chaetal lobe is rectangular in anterior region, the ventral cirri with swollen base start from chaetigers 13–14, there are two
<typeStatus id="B4149FB1FFB5FFC142E5FD7378A7FD05" box="[1188,1247,657,683]" pageId="17" pageNumber="18">types</typeStatus>
of isodont pectinate chaetae in median-posterior region, and the subacicular hook is translucent. In
<taxonomicName id="ACAF5A90FFB5FFC1427AFD5478ABFD61" baseAuthorityName="Montagu" baseAuthorityYear="1813" box="[1083,1235,694,719]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="sanguinea">
<emphasis id="59DBFD01FFB5FFC1427AFD5478ABFD61" box="[1083,1235,694,719]" italics="true" pageId="17" pageNumber="18">M. sanguinea</emphasis>
</taxonomicName>
(L10 =
<quantity id="AC578CF6FFB5FFC1436CFD567CBCFD5D" metricMagnitude="-2" metricUnit="m" metricValue="1.595" metricValueMax="2.04" metricValueMin="1.15" pageId="17" pageNumber="18" unit="mm" value="15.95" valueMax="20.4" valueMin="11.5">11.5–20.4 mm</quantity>
) the branchiae start from chaetigers 21–25, the chaetal lobe is rounded in anterior region, the ventral cirri with swollen base start from chaetigers 5–8, there is only one
<typeStatus id="B4149FB1FFB5FFC1455AFD1F7F33FCB9" box="[795,843,765,791]" pageId="17" pageNumber="18">type</typeStatus>
of isodont pectinate chaetae in posterior region, and the subacicular hook is reddish. Because of the above and the molecular evidence presented by
<bibRefCitation id="0F3E5CE2FFB5FFC14285FCC379E0FC95" author="Zanol, J. &amp; Halanych, K. M. &amp; Struck, T. H. &amp; Fauchald, K." box="[1220,1432,801,827]" pageId="17" pageNumber="18" pagination="660 - 676" refId="ref32780" refString="Zanol, J., Halanych, K. M., Struck, T. H. &amp; Fauchald, K. (2010) Phylogeny of the bristle worm family Eunicidae (Eunicida, Annelida) and the phylogenetic utility of noncongruent 16 S, COI and 18 S in combined analyses. Molecular Phylogenetics and Evolution, 55 (2), 660 - 676. https: // doi. org / 10.1016 / j. ympev. 2009.12.024" type="journal article" year="2010">
Zanol
<emphasis id="59DBFD01FFB5FFC1434DFCC07945FC95" box="[1292,1341,801,827]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
. (2010)
</bibRefCitation>
,
<taxonomicName id="ACAF5A90FFB5FFC146D6FCA47D40FCF1" authorityName=", Carrera-Parra" authorityYear="2009" box="[151,312,837,863]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC146D6FCA47D40FCF1" box="[151,312,837,863]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
should be considered as a valid species.
</paragraph>
<paragraph id="6B102113FFB5FFC14686FC8B789DFAF9" blockId="17.[151,1437,152,1367]" pageId="17" pageNumber="18">
<taxonomicName id="ACAF5A90FFB5FFC14686FC8B7DC3FC2D" authorityName=", Carrera-Parra" authorityYear="2009" box="[199,443,873,899]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14686FC8B7DC3FC2D" box="[199,443,873,899]" italics="true" pageId="17" pageNumber="18">Marphysa californica</emphasis>
</taxonomicName>
resembles
<taxonomicName id="ACAF5A90FFB5FFC1447AFC887F1CFC2A" authority="(Bermuda)" baseAuthorityName="Bermuda" box="[571,868,873,900]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="acicularum">
<emphasis id="59DBFD01FFB5FFC1447AFC887E99FC2D" box="[571,737,873,899]" italics="true" pageId="17" pageNumber="18">M. acicularum</emphasis>
(
<collectingCountry id="13B86183FFB5FFC144B1FC8B7F25FC2A" box="[752,861,873,900]" name="Bermuda" pageId="17" pageNumber="18">Bermuda</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="ACAF5A90FFB5FFC14531FC88784CFC2D" authority="Molina-Acevedo &amp; Idris, 2020" authorityName="Molina-Acevedo &amp; Idris" authorityYear="2020" box="[880,1076,873,899]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="baileybrockae" status="sp. nov.">
<emphasis id="59DBFD01FFB5FFC14531FC88784CFC2D" box="[880,1076,873,899]" italics="true" pageId="17" pageNumber="18">M. baileybrockae</emphasis>
</taxonomicName>
<emphasis id="59DBFD01FFB5FFC1427AFC8B7803FC2D" bold="true" box="[1083,1147,873,899]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="42E8407AFFB5FFC1427AFC8B7803FC2D" box="[1083,1147,873,899]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(Hawaii),
<taxonomicName id="ACAF5A90FFB5FFC142B4FC887C89FC06" authority="(Brazil)" baseAuthorityName="Brazil" baseAuthorityYear="1882" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="brasiliensis">
<emphasis id="59DBFD01FFB5FFC142B4FC8879E4FC2D" box="[1269,1436,873,899]" italics="true" pageId="17" pageNumber="18">M. brasiliensis</emphasis>
(
<collectingCountry id="13B86183FFB5FFC146DEFC6F7C91FC06" box="[159,233,909,936]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
)
</taxonomicName>
,
<taxonomicName id="ACAF5A90FFB5FFC146BDFC6C7D23FC09" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[252,347,909,935]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="bulla">
<emphasis id="59DBFD01FFB5FFC146BDFC6C7D23FC09" box="[252,347,909,935]" italics="true" pageId="17" pageNumber="18">M. bulla</emphasis>
</taxonomicName>
(Yellow Sea,
<collectingCountry id="13B86183FFB5FFC147B7FC6F7E45FC06" box="[502,573,909,936]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
),
<taxonomicName id="ACAF5A90FFB5FFC14411FC6C7F51FC09" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[592,809,909,935]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="maxidenticulata">
<emphasis id="59DBFD01FFB5FFC14411FC6C7F51FC09" box="[592,809,909,935]" italics="true" pageId="17" pageNumber="18">M. maxidenticulata</emphasis>
</taxonomicName>
(Yellow Sea,
<collectingCountry id="13B86183FFB5FFC14584FC6F7874FC06" box="[965,1036,909,936]" name="China" pageId="17" pageNumber="18">China</collectingCountry>
), and
<taxonomicName id="ACAF5A90FFB5FFC1420CFC6C795FFC06" authority="(Brazil)" authorityName="Baird" authorityYear="1869" baseAuthorityName="Brazil" box="[1101,1319,909,936]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="parishii">
<emphasis id="59DBFD01FFB5FFC1420CFC6C78B1FC09" box="[1101,1225,909,935]" italics="true" pageId="17" pageNumber="18">M. parishii</emphasis>
(
<collectingCountry id="13B86183FFB5FFC14299FC6F7967FC06" box="[1240,1311,909,936]" name="Brazil" pageId="17" pageNumber="18">Brazil</collectingCountry>
)
</taxonomicName>
by having branchiae pectinate with long branchial filaments, the absence of the subacicular limbate chaetae, and the subacicular hook translucent. However,
<taxonomicName id="ACAF5A90FFB5FFC147B3FC347EE9FC41" authorityName=", Carrera-Parra" authorityYear="2009" box="[498,657,981,1007]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC147B3FC347EE9FC41" box="[498,657,981,1007]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
has eyes, in contrast of
<taxonomicName id="ACAF5A90FFB5FFC145D9FC347F8EFC41" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[920,1014,981,1007]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="bulla">
<emphasis id="59DBFD01FFB5FFC145D9FC347F8EFC41" box="[920,1014,981,1007]" italics="true" pageId="17" pageNumber="18">M. bulla</emphasis>
</taxonomicName>
and
<taxonomicName id="ACAF5A90FFB5FFC14268FC34797AFC41" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[1065,1282,981,1007]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="maxidenticulata">
<emphasis id="59DBFD01FFB5FFC14268FC34797AFC41" box="[1065,1282,981,1007]" italics="true" pageId="17" pageNumber="18">M. maxidenticulata</emphasis>
</taxonomicName>
in which lack eyes. Moreover,
<taxonomicName id="ACAF5A90FFB5FFC14712FC187D8CFBBD" authorityName=", Carrera-Parra" authorityYear="2009" box="[339,500,1017,1043]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14712FC187D8CFBBD" box="[339,500,1017,1043]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
has conical dorsal cirri in all chaetigers, whereas in
<taxonomicName id="ACAF5A90FFB5FFC14205FC187892FBBD" box="[1092,1258,1017,1043]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="acicularum">
<emphasis id="59DBFD01FFB5FFC14205FC187892FBBD" box="[1092,1258,1017,1043]" italics="true" pageId="17" pageNumber="18">M. acicularum</emphasis>
</taxonomicName>
, the dorsal cirri are conical with a broad base.
<taxonomicName id="ACAF5A90FFB5FFC147AAFBFF7EA7FB99" authorityName=", Carrera-Parra" authorityYear="2009" box="[491,735,1053,1079]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC147AAFBFF7EA7FB99" box="[491,735,1053,1079]" italics="true" pageId="17" pageNumber="18">Marphysa californica</emphasis>
</taxonomicName>
has the prechaetal lobe as a transversal fold in anterior region, instaed of
<taxonomicName id="ACAF5A90FFB5FFC14750FBA07DC1FBF5" box="[273,441,1089,1115]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="acicularum">
<emphasis id="59DBFD01FFB5FFC14750FBA07DC1FBF5" box="[273,441,1089,1115]" italics="true" pageId="17" pageNumber="18">M. acicularum</emphasis>
</taxonomicName>
and
<taxonomicName id="ACAF5A90FFB5FFC147B5FBA07EC3FBF5" authority="Molina-Acevedo &amp; Idris, 2020" authorityName="Molina-Acevedo &amp; Idris" authorityYear="2020" box="[500,699,1089,1115]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="baileybrockae" status="sp. nov.">
<emphasis id="59DBFD01FFB5FFC147B5FBA07EC3FBF5" box="[500,699,1089,1115]" italics="true" pageId="17" pageNumber="18">M. baileybrockae</emphasis>
</taxonomicName>
<emphasis id="59DBFD01FFB5FFC14485FBA37F7DFBF5" bold="true" box="[708,773,1089,1115]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="42E8407AFFB5FFC14485FBA37F7DFBF5" box="[708,773,1089,1115]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in which the prechaetal lobe has dorsal edge longer than ventral in the same region. In addition,
<taxonomicName id="ACAF5A90FFB5FFC1440AFB847E93FBD1" authorityName=", Carrera-Parra" authorityYear="2009" box="[587,747,1125,1151]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC1440AFB847E93FBD1" box="[587,747,1125,1151]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
has the chaetal lobe rectangular in first chaetigers, whereas in
<taxonomicName id="ACAF5A90FFB5FFC146D6FB687D45FB0D" box="[151,317,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="acicularum">
<emphasis id="59DBFD01FFB5FFC146D6FB687D45FB0D" box="[151,317,1161,1187]" italics="true" pageId="17" pageNumber="18">M. acicularum</emphasis>
</taxonomicName>
,
<taxonomicName id="ACAF5A90FFB5FFC1470BFB687E75FB0D" authority="Molina-Acevedo &amp; Idris, 2020" authorityName="Molina-Acevedo &amp; Idris" authorityYear="2020" box="[330,525,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="baileybrockae" status="sp. nov.">
<emphasis id="59DBFD01FFB5FFC1470BFB687E75FB0D" box="[330,525,1161,1187]" italics="true" pageId="17" pageNumber="18">M. baileybrockae</emphasis>
</taxonomicName>
<emphasis id="59DBFD01FFB5FFC14455FB6B7E2BFB0D" bold="true" box="[532,595,1161,1187]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="42E8407AFFB5FFC14455FB6B7E2BFB0D" box="[532,595,1161,1187]" pageId="17" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="ACAF5A90FFB5FFC14420FB687F7FFB0D" baseAuthorityName="Hansen" baseAuthorityYear="1882" box="[609,775,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="brasiliensis">
<emphasis id="59DBFD01FFB5FFC14420FB687F7FFB0D" box="[609,775,1161,1187]" italics="true" pageId="17" pageNumber="18">M. brasiliensis</emphasis>
</taxonomicName>
,
<taxonomicName id="ACAF5A90FFB5FFC14554FB687F0DFB0D" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[789,885,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="bulla">
<emphasis id="59DBFD01FFB5FFC14554FB687F0DFB0D" box="[789,885,1161,1187]" italics="true" pageId="17" pageNumber="18">M. bulla</emphasis>
</taxonomicName>
,
<taxonomicName id="ACAF5A90FFB5FFC145C3FB687824FB0D" authorityName="Liu, Hutchings &amp; Kupriyanova" authorityYear="2018" box="[898,1116,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="maxidenticulata">
<emphasis id="59DBFD01FFB5FFC145C3FB687824FB0D" box="[898,1116,1161,1187]" italics="true" pageId="17" pageNumber="18">M. maxidenticulata</emphasis>
</taxonomicName>
, and
<taxonomicName id="ACAF5A90FFB5FFC142D8FB68796EFB0D" authorityName="Baird" authorityYear="1869" box="[1177,1302,1161,1187]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="parishii">
<emphasis id="59DBFD01FFB5FFC142D8FB68796EFB0D" box="[1177,1302,1161,1187]" italics="true" pageId="17" pageNumber="18">M. parishii</emphasis>
</taxonomicName>
, the chaetal lobe is rounded in first chaetigers. Also,
<taxonomicName id="ACAF5A90FFB5FFC14422FB4C7F7CFB69" authorityName=", Carrera-Parra" authorityYear="2009" box="[611,772,1197,1223]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14422FB4C7F7CFB69" box="[611,772,1197,1223]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
(
<typeStatus id="B4149FB1FFB5FFC14555FB4F7F3EFB69" box="[788,838,1197,1223]" pageId="17" pageNumber="18">type</typeStatus>
and additional material, L10 =
<quantity id="AC578CF6FFB5FFC142ECFB4E7945FB69" box="[1197,1341,1196,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.625" metricValueMax="1.9" metricValueMin="1.35" pageId="17" pageNumber="18" unit="mm" value="16.25" valueMax="19.0" valueMin="13.5">13.5–19 mm</quantity>
) has the postchaetal lobe developed up to chaetigers 41–92, different from
<taxonomicName id="ACAF5A90FFB5FFC145C6FB307857FB45" baseAuthorityName="Hansen" baseAuthorityYear="1882" box="[903,1071,1233,1259]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="brasiliensis">
<emphasis id="59DBFD01FFB5FFC145C6FB307857FB45" box="[903,1071,1233,1259]" italics="true" pageId="17" pageNumber="18">M. brasiliensis</emphasis>
</taxonomicName>
(L10 =
<quantity id="AC578CF6FFB5FFC142CCFB3378A6FB45" box="[1165,1246,1233,1259]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="18" unit="mm" value="17.0">17 mm</quantity>
) and
<taxonomicName id="ACAF5A90FFB5FFC1435EFB3079E4FB45" authorityName="Baird" authorityYear="1869" box="[1311,1436,1233,1259]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="parishii">
<emphasis id="59DBFD01FFB5FFC1435EFB3079E4FB45" box="[1311,1436,1233,1259]" italics="true" pageId="17" pageNumber="18">M. parishii</emphasis>
</taxonomicName>
(L10 =
<quantity id="AC578CF6FFB5FFC146ABFB177D2AFAA1" box="[234,338,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="17" pageNumber="18" unit="mm" value="12.2">12.2 mm</quantity>
), in which the postchaetal lobe is developed in more than 200 chaetigers. Finally, in
<taxonomicName id="ACAF5A90FFB5FFC142BDFB1479E4FAA1" authorityName=", Carrera-Parra" authorityYear="2009" box="[1276,1436,1269,1295]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC142BDFB1479E4FAA1" box="[1276,1436,1269,1295]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
the ventral cirri with swollen base start from chaetigers 13–14, whereas in
<taxonomicName id="ACAF5A90FFB5FFC1459DFAF878F9FA9D" box="[988,1153,1305,1331]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="acicularum">
<emphasis id="59DBFD01FFB5FFC1459DFAF878F9FA9D" box="[988,1153,1305,1331]" italics="true" pageId="17" pageNumber="18">M. acicularum</emphasis>
</taxonomicName>
and
<taxonomicName id="ACAF5A90FFB5FFC142F6FAF87926FA9D" baseAuthorityName="Hansen" baseAuthorityYear="1882" box="[1207,1374,1305,1331]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="brasiliensis">
<emphasis id="59DBFD01FFB5FFC142F6FAF87926FA9D" box="[1207,1374,1305,1331]" italics="true" pageId="17" pageNumber="18">M. brasiliensis</emphasis>
</taxonomicName>
, start from chaetigers 3–5. The comparison of
<taxonomicName id="ACAF5A90FFB5FFC14421FADC7F78FAF9" authorityName=", Carrera-Parra" authorityYear="2009" box="[608,768,1341,1367]" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" order="Eunicida" pageId="17" pageNumber="18" phylum="Annelida" rank="species" species="californica">
<emphasis id="59DBFD01FFB5FFC14421FADC7F78FAF9" box="[608,768,1341,1367]" italics="true" pageId="17" pageNumber="18">M. californica</emphasis>
</taxonomicName>
with related species is provided in
<tableCitation id="262D14A8FFB5FFC142CAFADF78A7FAF9" box="[1163,1247,1341,1367]" captionStart="TABLE 1" captionStartId="30.[151,235,468,493]" captionText="TABLE 1. (Continued)" pageId="17" pageNumber="18">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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