treatments-xml/data/DF/18/F9/DF18F971FFCB0B3AFF4BF9BD68D5BE1B.xml

<document id="3FB96EE5D1FB122C784219D3F0B67139" ID-CLB-Dataset="27575" ID-DOI="10.11646/zootaxa.4554.1.9" ID-GBIF-Dataset="ff7707f0-058f-4f40-8eb8-2b6c9c75a173" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623565" ID-ZooBank="A804E949-109A-468D-B58B-CF7C8BCB3059" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" checkinTime="1554271225461" checkinUser="plazi" docAuthor="Vandenberg, Natalia J. &amp; Hanson, Paul E." docDate="2019" docId="DF18F971FFCB0B3AFF4BF9BD68D5BE1B" docLanguage="en" docName="zootaxa.4554.1.9.pdf" docOrigin="Zootaxa 4554 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Moiradiomus Vandenberg &amp; Hanson, 2019, gen. nov." docType="treatment" docVersion="10" lastPageNumber="271" masterDocId="23218109FFC20B2BFFDCFFF96C54BC27" masterDocTitle="Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)" masterLastPageNumber="285" masterPageNumber="255" pageNumber="264" updateTime="1728643685052" updateUser="ExternalLinkService">
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<mods:title id="853B5E64C8634CE83CCC2A0F5F9F2C1D">Overview of the lady beetle tribe Diomini (Coleoptera: Coccinellidae) and description of a new phytophagous, silk-spinning genus from Costa Rica that induces food bodies on leaves of Piper (Piperaceae)</mods:title>
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<mods:namePart id="E0DE692351A92DBC1FF54DF1D94CCA30">Vandenberg, Natalia J.</mods:namePart>
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<mods:namePart id="8291683CA11A00126D2C5E1015AB1F5D">Hanson, Paul E.</mods:namePart>
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<treatment id="DF18F971FFCB0B3AFF4BF9BD68D5BE1B" ID-DOI="http://doi.org/10.5281/zenodo.5921882" ID-GBIF-Taxon="154535012" ID-Zenodo-Dep="5921882" LSID="urn:lsid:plazi:treatment:DF18F971FFCB0B3AFF4BF9BD68D5BE1B" httpUri="http://treatment.plazi.org/id/DF18F971FFCB0B3AFF4BF9BD68D5BE1B" lastPageId="17" lastPageNumber="271" pageId="9" pageNumber="264">
<subSubSection id="1FAB1BECFFCB0B22FF4BF9BD6DFBBA79" box="[151,431,1604,1630]" pageId="9" pageNumber="264" type="nomenclature">
<paragraph id="570E4867FFCB0B22FF4BF9BD6DFBBA79" blockId="9.[151,525,1604,1663]" box="[151,431,1604,1630]" pageId="9" pageNumber="264">
<heading id="0C46FF0BFFCB0B22FF4BF9BD6DFBBA79" bold="true" box="[151,431,1604,1630]" fontSize="11" level="1" pageId="9" pageNumber="264" reason="1">
<emphasis id="65C59475FFCB0B22FF4BF9BD6DFBBA79" bold="true" box="[151,431,1604,1630]" pageId="9" pageNumber="264">
<taxonomicName id="90B133E4FFCB0B22FF4BF9BD6D6EBA79" ID-CoL="CG2R4" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[151,314,1604,1630]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="65C59475FFCB0B22FF4BF9BD6D6EBA79" bold="true" box="[151,314,1604,1630]" italics="true" pageId="9" pageNumber="264">Moiradiomus</emphasis>
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<taxonomicNameLabel id="7EF6290EFFCB0B22FE9DF9BD6DFBBA79" box="[321,431,1604,1630]" pageId="9" pageNumber="264" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="1FAB1BECFFCB0B22FF4BF99F6E1DBAE3" pageId="9" pageNumber="264" type="description">
<paragraph id="570E4867FFCB0B22FF4BF99F6E59BA58" blockId="9.[151,525,1604,1663]" box="[151,525,1638,1663]" pageId="9" pageNumber="264">
(
<figureCitation id="CF8A54E2FFCB0B22FF43F99F6D60BA58" box="[159,308,1638,1663]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon &amp; Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="9" pageNumber="264">Figs. 6, 8, 10</figureCitation>
, 
<figureCitation id="CF8A54E2FFCB0B22FE9CF99E6DDDBA58" box="[320,393,1639,1663]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="9" pageNumber="264">13–20</figureCitation>
, 
<figureCitation id="CF8A54E2FFCB0B22FE48F99E6DE6BA58" box="[404,434,1639,1663]" captionStart="FIGURES 21–24" captionStartId="15.[151,264,1269,1292]" captionTargetBox="[157,1430,215,1228]" captionTargetId="figure@15.[151,1436,213,1234]" captionTargetPageId="15" captionText="FIGURES 21–24. Dorsal view of right mandible: 21, Subcoccinella vigintiquatuorpunctata (L.) (folivorous) (after Kovář 1996); 22, Bulaea sp. (omnivorous with emphasis on pollinivory/phytophagy); 23, Moiradiomus lachesis (enlarged detail of apical portion of incisor blade shown to left of main structure) (specialized on Piper food bodies); 24, Adalia bipunctata (entomophagous with emphasis on aphids) (after Kovář 1996)." figureDoi="http://doi.org/10.5281/zenodo.2623575" httpUri="https://zenodo.org/record/2623575/files/figure.png" pageId="9" pageNumber="264">23</figureCitation>
, 
<figureCitation id="CF8A54E2FFCB0B22FE62F99F6E52BA58" box="[446,518,1638,1663]" captionStart-0="FIGURES 25–28" captionStart-1="FIGURES 29–32" captionStart-2="FIGURES 33–39" captionStart-3="FIGURES 40–42" captionStartId-0="15.[151,264,1975,1998]" captionStartId-1="16.[151,264,1700,1723]" captionStartId-2="19.[151,264,1894,1917]" captionStartId-3="20.[151,264,1957,1980]" captionTargetBox-0="[151,1436,1466,1918]" captionTargetBox-1="[154,1436,190,1666]" captionTargetBox-2="[151,1435,237,1813]" captionTargetBox-3="[151,1436,193,1936]" captionTargetId-0="figure@15.[151,1436,1461,1924]" captionTargetId-1="figure@16.[151,1436,189,1667]" captionTargetId-2="figure@19.[151,1436,235,1848]" captionTargetId-3="figure@20.[151,1436,193,1936]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." captionText-1="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." captionText-2="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." captionText-3="FIGURES 40–42. Tents of Moiradiomus spp.: 40, M. lachesis on Piper lanosibracteum; 41, M. nanita on P. reticulatum; 42, M. atropos on P. friedrichsthalii." figureDoi-0="http://doi.org/10.5281/zenodo.2623578" figureDoi-1="http://doi.org/10.5281/zenodo.2623580" figureDoi-2="http://doi.org/10.5281/zenodo.2623582" figureDoi-3="http://doi.org/10.5281/zenodo.2623584" httpUri-0="https://zenodo.org/record/2623578/files/figure.png" httpUri-1="https://zenodo.org/record/2623580/files/figure.png" httpUri-2="https://zenodo.org/record/2623582/files/figure.png" httpUri-3="https://zenodo.org/record/2623584/files/figure.png" pageId="9" pageNumber="264">25–42</figureCitation>
)
</paragraph>
<paragraph id="570E4867FFCB0B22FF4BF9566E1DBAE3" blockId="9.[151,585,1710,1732]" box="[151,585,1710,1732]" pageId="9" pageNumber="264">
<typeStatus id="880AF6C5FFCB0B22FF4BF9566C9DBAE3" box="[151,201,1711,1732]" pageId="9" pageNumber="264">Type</typeStatus>
species: 
<taxonomicName id="90B133E4FFCB0B22FEFAF9576DA0BAE3" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[294,500,1710,1732]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="clotho" status="sp. nov.">
<emphasis id="65C59475FFCB0B22FEFAF9576DA0BAE3" box="[294,500,1710,1732]" italics="true" pageId="9" pageNumber="264">Moiradiomus clotho</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7EF6290EFFCB0B22FE26F9566E1DBAE3" box="[506,585,1711,1732]" pageId="9" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFCB0B21FF4BF9086E04BE1A" lastPageId="10" lastPageNumber="265" pageId="9" pageNumber="264" type="diagnosis">
<paragraph id="570E4867FFCB0B21FF4BF9086E29BCDF" blockId="9.[151,1436,1777,2018]" lastBlockId="10.[151,1437,151,1979]" lastPageId="10" lastPageNumber="265" pageId="9" pageNumber="264">
<emphasis id="65C59475FFCB0B22FF4BF9086D45BB2D" bold="true" box="[151,273,1777,1802]" pageId="9" pageNumber="264">Diagnosis.</emphasis>
Distinguished from other members of the tribe 
<taxonomicName id="90B133E4FFCB0B22FCB4F9086F91BB2D" authorityName="Gordon" authorityYear="1999" box="[872,965,1777,1802]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
by the combination of morphological, anatomical, and behavioral traits: antenna consistently comprised of 10 antennomeres, distal antennomere not greatly enlarged or elongated relative to penultimate, pronotal anterolateral angles lacking submarginal carinae (
<figureCitation id="CF8A54E2FFCB0B22FF43F8A46D7EBB51" box="[159,298,1885,1910]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="9" pageNumber="264">Figs. 13–20</figureCitation>
), prosternum (
<figureCitation id="CF8A54E2FFCB0B22FE0CF8A46E0CBB51" box="[464,600,1885,1910]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="9" pageNumber="264">Figs. 25–28</figureCitation>
) shaped like a short stemmed Y, short anterior to coxal cavities, about ½ diameter of cavity, with carinae of intercoxal process extending to anterior margin, male genitalia (
<figureCitation id="CF8A54E2FFCB0B22FAD7F87869C0BBBD" box="[1291,1428,1921,1946]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="9" pageNumber="264">Figs. 29–32</figureCitation>
) with basal lobe (=penis guide) asymmetrical (
<figureCitation id="CF8A54E2FFCB0B22FD7EF85C6FEABB99" box="[674,958,1957,1982]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="9" pageNumber="264">Figs. 29b, 30b, 31b, 32b</figureCitation>
), penis (=siphonal) capsule (
<figureCitation id="CF8A54E2FFCB0B22FAD4F85C690FBB99" box="[1288,1371,1957,1982]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon &amp; Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="9" pageNumber="264">Fig. 10</figureCitation>
) well developed, with inner and outer arms broadly joined into single wedge-shaped structure with sinuous distal margin, capsule enclosing ejaculatory duct prior to its entry into the main penis tube, penis apex lacking flagellum (
<figureCitation id="CF8A54E2FFC80B21FA94FF6E69C3BC97" box="[1352,1431,151,176]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon &amp; Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="10" pageNumber="265">Figs. 8</figureCitation>
, 
<figureCitation id="CF8A54E2FFC80B21FF4BFF456D27BCF3" box="[151,371,188,212]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="10" pageNumber="265">29c, 30c, 31c, 32c</figureCitation>
), larva silk spinning, adult and larva phytophagous, parasitic on plants in the genus 
<taxonomicName id="90B133E4FFC80B21FA84FF4469C1BCF3" box="[1368,1429,189,212]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="10" pageNumber="265" phylum="Tracheophyta" rank="genus">
<emphasis id="65C59475FFC80B21FA84FF4469C1BCF3" box="[1368,1429,189,212]" italics="true" pageId="10" pageNumber="265">Piper</emphasis>
</taxonomicName>
, inducing and feeding on 
<taxonomicName id="90B133E4FFC80B21FE71FF186DBFBCDF" box="[429,491,225,248]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="10" pageNumber="265" phylum="Tracheophyta" rank="genus">
<emphasis id="65C59475FFC80B21FE71FF186DBFBCDF" box="[429,491,225,248]" italics="true" pageId="10" pageNumber="265">Piper</emphasis>
</taxonomicName>
food bodies.
</paragraph>
<paragraph id="570E4867FFC80B21FF1BFEFD6E04BE1A" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
The reduced number of antennomeres (10 as opposed to 11) will distinguished the new genus from all other 
<taxonomicName id="90B133E4FFC80B21FF4BFEDE6CA0BD67" authorityName="Gordon" authorityYear="1999" box="[151,244,295,320]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
except 
<taxonomicName id="90B133E4FFC80B21FE97FEDE6D83BD67" authorityName="Chapin" authorityYear="1933" box="[331,471,295,320]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FE97FEDE6D83BD67" box="[331,471,295,320]" italics="true" pageId="10" pageNumber="265">Decadiomus</emphasis>
</taxonomicName>
and a few species or individuals presently classified in 
<taxonomicName id="90B133E4FFC80B21FB94FED068CBBD67" authorityName="Mulsant" authorityYear="1850" box="[1096,1183,297,320]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FB94FED068CBBD67" box="[1096,1183,297,320]" italics="true" pageId="10" pageNumber="265">Diomus</emphasis>
</taxonomicName>
. From these latter two genera, 
<taxonomicName id="90B133E4FFC80B21FF2CFEB56DD3BD42" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[240,391,332,357]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FF2CFEB56DD3BD42" box="[240,391,332,357]" italics="true" pageId="10" pageNumber="265">Moiradiomus</emphasis>
</taxonomicName>
can be distinguished by the highly consistent configuration of the penis of the male genitalia (as described above), and by the unusual life history and trophic relations shared by its members. Thus far the known 
<taxonomicName id="90B133E4FFC80B21FF37FE6D6D88BD8A" box="[235,476,404,429]" pageId="10" pageNumber="265">
<emphasis id="65C59475FFC80B21FF37FE6D6DD6BD8A" box="[235,386,404,429]" italics="true" pageId="10" pageNumber="265">Moiradiomus</emphasis>
species
</taxonomicName>
all have a similar elytral color pattern (
<figureCitation id="CF8A54E2FFC80B21FC7DFE6D687DBD8B" box="[929,1065,404,429]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="10" pageNumber="265">Figs. 13–20</figureCitation>
) consisting of a medium to dark brown background with apex suffusely lightened or bearing a distinct band of pale yellow. The elytral color patterns in 
<taxonomicName id="90B133E4FFC80B21FEC1FE256DFDBDD2" authorityName="Chapin" authorityYear="1933" box="[285,425,476,501]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FEC1FE256DFDBDD2" box="[285,425,476,501]" italics="true" pageId="10" pageNumber="265">Decadiomus</emphasis>
</taxonomicName>
are more variable, with either a dark or light background and often with contrasting maculae. The tarsal claw is sexually dimorphic in the known species 
<taxonomicName id="90B133E4FFC80B21FC40FE066867BE3F" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[924,1075,511,536]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FC40FE066867BE3F" box="[924,1075,511,536]" italics="true" pageId="10" pageNumber="265">Moiradiomus</emphasis>
</taxonomicName>
, but this characteristic occurs in some 
<taxonomicName id="90B133E4FFC80B21FF05FDDC6D64BE1B" authorityName="Mulsant" authorityYear="1850" box="[217,304,549,572]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FF05FDDC6D64BE1B" box="[217,304,549,572]" italics="true" pageId="10" pageNumber="265">Diomus</emphasis>
</taxonomicName>
and 
<taxonomicName id="90B133E4FFC80B21FEBAFDDD6DA6BE1A" authorityName="Chapin" authorityYear="1933" box="[358,498,548,573]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFC80B21FEBAFDDD6DA6BE1A" box="[358,498,548,573]" italics="true" pageId="10" pageNumber="265">Decadiomus</emphasis>
</taxonomicName>
as well.
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFC80B21FF1BFDBE68C4BB9C" pageId="10" pageNumber="265" type="description">
<paragraph id="570E4867FFC80B21FF1BFDBE6E90BF82" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
<emphasis id="65C59475FFC80B21FF1BFDBE6D0DBE47" bold="true" box="[199,345,583,608]" pageId="10" pageNumber="265">Description.</emphasis>
Form (
<figureCitation id="CF8A54E2FFC80B21FE71FDBE6E60BE47" box="[429,564,583,608]" captionStart-0="FIGURES 13–16" captionStart-1="FIGURES 17–20" captionStartId-0="13.[151,264,1957,1980]" captionStartId-1="14.[151,264,1965,1988]" captionTargetBox-0="[151,1436,245,1896]" captionTargetBox-1="[151,1436,260,1911]" captionTargetId-0="figure@13.[151,1436,245,1896]" captionTargetId-1="figure@14.[151,1436,260,1911]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURES 13–16. Moiradiomus species habitus illustrations. 13, M. clotho: holotype, male; 14, M. clotho: paratype, female. 15, M. lachesis: holotype, male; 16, M. lachesis: paratype, female. Scale bar = 0.5 mm." captionText-1="FIGURES 17–20. Moiradiomus species habitus illustrations. 17, M. atropos: holotype, male; 18, M. atropos: paratype, female. 19, M. nanita: holotype, male; 20, M. nanita: paratype, female. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.2623571" figureDoi-1="http://doi.org/10.5281/zenodo.2623573" httpUri-0="https://zenodo.org/record/2623571/files/figure.png" httpUri-1="https://zenodo.org/record/2623573/files/figure.png" pageId="10" pageNumber="265">Figs. 13–20</figureCitation>
) broadly oval to elongate, length 1.1–2.0 mm, weakly to moderately convex, pubescent. Color pattern simple, unsaturated; predominantly brown with yellow; female pronotum generally darker than in male. Head transverse; eyes finely facetted, well separated dorsally, bearing interfacetal setae, with interocular distance greater than eye width in frontal view, weakly emarginated near antennal insertions by small rounded eye canthus; antenna short, approximately ½ head width, clavate, composed of 10 antennomeres with last 4 or 5 forming irregular club; scape normal; pedicel bead like, slightly narrower than scape; antennomere 3 elongate, longer than 4, with distal antennomere subequal to or only slightly longer than penultimate. Mandible (
<figureCitation id="CF8A54E2FFC80B21FF43FCBD6CA5BF7B" box="[159,241,836,861]" captionStart="FIGURES 21–24" captionStartId="15.[151,264,1269,1292]" captionTargetBox="[157,1430,215,1228]" captionTargetId="figure@15.[151,1436,213,1234]" captionTargetPageId="15" captionText="FIGURES 21–24. Dorsal view of right mandible: 21, Subcoccinella vigintiquatuorpunctata (L.) (folivorous) (after Kovář 1996); 22, Bulaea sp. (omnivorous with emphasis on pollinivory/phytophagy); 23, Moiradiomus lachesis (enlarged detail of apical portion of incisor blade shown to left of main structure) (specialized on Piper food bodies); 24, Adalia bipunctata (entomophagous with emphasis on aphids) (after Kovář 1996)." figureDoi="http://doi.org/10.5281/zenodo.2623575" httpUri="https://zenodo.org/record/2623575/files/figure.png" pageId="10" pageNumber="265">Fig. 23</figureCitation>
) with bifid apex, with about 40 minute blunt teeth on incisor blade; molar part with curved tooth near top (=basal tooth sens. 
<bibRefCitation id="33203596FFC80B21FEADFC9E6E4EBFA7" author="Slipinski, S. A." box="[369,538,871,896]" pageId="10" pageNumber="265" refId="ref22567" refString="Slipinski, S. A. (2007) Australian ladybird beetles (Coleoptera: Coccinellidae). Their biology and classification. ABRS, Canberra, 288 pp." type="book" year="2007">Ślipiński 2007</bibRefCitation>
). Maxillary palp with 4 palpomeres; terminal palpomere moderately to strongly expanded distally. Labial palp with 3 palpomeres.
</paragraph>
<paragraph id="570E4867FFC80B21FF1BFC4968E4B95F" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
Pronotum transverse, evenly convex, base with raised border; pronotal anterolateral angle lacking submarginal carina. Elytron with raised lateral ridge; epipleuron narrow, oblique, not excavated to receive femoral apices, not reaching elytral apex. Prosternum (
<figureCitation id="CF8A54E2FFC80B21FDC2FC0E6EF0B837" box="[542,676,1015,1040]" captionStart="FIGURES 25–28" captionStartId="15.[151,264,1975,1998]" captionTargetBox="[151,1436,1466,1918]" captionTargetId="figure@15.[151,1436,1461,1924]" captionTargetPageId="15" captionText="FIGURES 25–28. Diagrammatic ventral view of Moiradiomus prosterna (setae, color patterns, and surface punctation not indicated): 25, M. clotho; 26, M. lachesis; 27, M. atopos; 28, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="10" pageNumber="265">Figs. 25–28</figureCitation>
) shaped like short stemmed Y, short anterior to coxal cavities, about ½ diameter of cavity, with carinae of intercoxal process extending to anterior margin or nearly so. Mesoventrite with raised anterior border, with anterior face excavated to receive apex of prosternal intercoxal process. Tibial spurs lacking; tarsal claw sexually dimorphic in species thus far known, with short triangular tooth in female, longer scythe-like tooth in male. Abdomen with 6 ventrites in both sexes; abdominal postcoxal line curving posterolaterally, merging with posterior margin of ventrite. Male genitalia (
<figureCitation id="CF8A54E2FFC80B21FBD5FB5568C0B8E3" box="[1033,1172,1196,1221]" captionStart="FIGURES 29–32" captionStartId="16.[151,264,1700,1723]" captionTargetBox="[154,1436,190,1666]" captionTargetId="figure@16.[151,1436,189,1667]" captionTargetPageId="16" captionText="FIGURES 29–32. Male genitalia of Moiradiomus species (a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, M. clotho; 30, M. lachesis; 31, M. atopos; 32, M. nanita." figureDoi="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="10" pageNumber="265">Figs. 29–32</figureCitation>
) with outer margin of paramere densely setiferous; setae long, slender, flexible; basal lobe (=penis guide) asymmetrical, penis capsule (
<figureCitation id="CF8A54E2FFC80B21FF43FB0D6CA4B92B" box="[159,240,1268,1293]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon &amp; Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="10" pageNumber="265">Fig. 10</figureCitation>
) well developed, with inner and outer arms broadly joined into single wedge-shaped structure with sinuous distal margin, capsule enclosing ejaculatory duct prior to its entry into main penis tube; penis apex lacking flagellum. Female genitalia (
<figureCitation id="CF8A54E2FFC80B21FE38FAC56E7FB972" box="[484,555,1340,1365]" captionStart="FIGURES 3–12" captionStartId="12.[151,264,1784,1807]" captionTargetBox="[151,1433,212,1735]" captionTargetId="figure@12.[151,1436,193,1763]" captionTargetPageId="12" captionText="FIGURES 3–12. Genitalia of Diomini and Selvadiina (Hyperaspidini): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, Selvadius nunenmacheri Gordon 1970 (Hyperaspidini: Selvadiina), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, Erratodiomus brindisi Gordon 1999 (Hyperaspidini: Selvadiina), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, Decadiomus hughesi Gordon &amp; Hilburn 1990 (Diomini), bursa through spermatheca (note long convoluted sperm duct); 6, M. lachesis (Diomini) (A.g.= accessory gland); 7–12, male genitalia of Diomini, left lateral view of penis: 7, D. hughesi; 8, M. lachesis; 9, Diomus donatus Gordon (capsule); 10, M. lachesis (capsule); 11, Decadiomus bigemmeus (capsule); 12, Decadiomus liebecki (capsule)" figureDoi="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="10" pageNumber="265">Fig. 6</figureCitation>
) with spermathecal capsule fully developed, bent in basal 1/2 with bulbous base; sperm duct moderately long with sclerotized distally tapered sheath enclosing basal 1/2.
</paragraph>
<paragraph id="570E4867FFC80B21FF08FA7F6DE7BA74" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
<emphasis id="65C59475FFC80B21FF08FA7F6F93B9B8" bold="true" box="[212,967,1411,1439]" pageId="10" pageNumber="265">
Larva, final (4 
<superScript id="A0C4E52FFFC80B21FE52FA7A6DCFB9B6" attach="none" box="[398,411,1411,1425]" fontSize="6" pageId="10" pageNumber="265">th</superScript>
) instar (
<figureCitation id="CF8A54E2FFC80B21FDD2FA7F6EC9B9B8" box="[526,669,1414,1439]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Figs. 33–39</figureCitation>
, based on 
<taxonomicName id="90B133E4FFC80B21FCF2FA7E6FECB9B8" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[814,952,1414,1439]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="265" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFC80B21FCF2FA7E6FECB9B8" bold="true" box="[814,952,1414,1439]" italics="true" pageId="10" pageNumber="265">M. lachesis</emphasis>
</taxonomicName>
).
</emphasis>
Body off white, pruinose, soft-bodied, subovate, weakly convex dorsally, setiferous. Setae (
<figureCitation id="CF8A54E2FFC80B21FD28FA536F1CB9E4" box="[756,840,1450,1475]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 37</figureCitation>
) minutely barbed, of variable form, some flattened, scale-like, falcate, spatulate, clavate or frayed. Head tapered anteriorly, bearing three stemmata on each side behind antenna; frontal arms of epicranial suture indistinct; epicranial stem absent; mandible (
<figureCitation id="CF8A54E2FFC80B21FB55FA0B688ABA2C" box="[1161,1246,1522,1547]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 36</figureCitation>
) falciform, with unidentate apex, shallow incisor groove, triangular plate-like angulation near middle of incisor edge, with simple rectangular molar region.
</paragraph>
<paragraph id="570E4867FFC80B21FF1BF9A668A3BB68" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
Antenna (
<figureCitation id="CF8A54E2FFC80B21FEE9F9A76DD2BA50" box="[309,390,1630,1655]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 35</figureCitation>
) 3-segmented, short; scape annular; pedicle narrower, nearly as long as wide, bearing conical sensorium, pair of preapical setae present; flagellum with long apical seta. Mala small, apically rounded. Maxilla (
<figureCitation id="CF8A54E2FFC80B21FF43F95F6CA6BA98" box="[159,242,1702,1727]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 34</figureCitation>
) with two conspicuous long setae on fused cardo/stipes; maxillary palp 2-segmented; maxillary palpifer incompletely annular. Labial palp 2-segmented; palpifer unsclerotized, indistinct. Fused submentum/mentum with several long conspicuous setae. Leg well developed, visible in dorsal view; tibiotarsus (
<figureCitation id="CF8A54E2FFC80B21FB52F91768B6BB20" box="[1166,1250,1774,1799]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 38</figureCitation>
) with cluster of capitate setae near apex; tarsungulus (
<figureCitation id="CF8A54E2FFC80B21FD95F8EB6EC9BB0C" box="[585,669,1810,1835]" captionStart="FIGURES 33–39" captionStartId="19.[151,264,1894,1917]" captionTargetBox="[151,1435,237,1813]" captionTargetId="figure@19.[151,1436,235,1848]" captionTargetPageId="19" captionText="FIGURES 33–39. Fourth instar larva of Moiradiomus lachesis: 33, dorsal habitus view, scale bar = 0.5 mm; 34, ventral view of mouthparts, enlarged; 35, right antenna, enlarged; 36, dorsal view of right mandible, enlarged; 37, diverse dorsal setae, enlarged; 38, tibiotarsus showing distribution of clavate setae, enlarged; 39, apex of tibiotarsus (setae removed) showing shape of tarsal claw, enlarged." figureDoi="http://doi.org/10.5281/zenodo.2623582" httpUri="https://zenodo.org/record/2623582/files/figure.png" pageId="10" pageNumber="265">Fig. 39</figureCitation>
) with broad base, pointed apex, bearing single short simple seta at base of concave inner side. Paired dorsal abdominal glands not evident, possibly vestigial or absent.
</paragraph>
<paragraph id="570E4867FFC80B21FF1BF8A368C4BB9C" blockId="10.[151,1437,151,1979]" pageId="10" pageNumber="265">
<emphasis id="65C59475FFC80B21FF1BF8A36DCEBB54" bold="true" box="[199,410,1882,1907]" pageId="10" pageNumber="265">Trophic relations.</emphasis>
Larvae develop on food bodies of 
<taxonomicName id="90B133E4FFC80B21FCC1F8A56FEFBB54" box="[797,955,1882,1907]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="10" pageNumber="265" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="65C59475FFC80B21FCC1F8A56F0FBB54" box="[797,859,1884,1907]" italics="true" pageId="10" pageNumber="265">Piper</emphasis>
species.
</taxonomicName>
Although the four species described below are all from 
<collectingCountry id="2FA608F7FFC80B21FEC2F8866DCFBBB0" box="[286,411,1918,1943]" name="Costa Rica" pageId="10" pageNumber="265">Costa Rica</collectingCountry>
, the distinctive larval tents have been observed on leaves of 
<taxonomicName id="90B133E4FFC80B21FBE4F8796823BBB0" box="[1080,1143,1920,1943]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="10" pageNumber="265" phylum="Tracheophyta" rank="genus">
<emphasis id="65C59475FFC80B21FBE4F8796823BBB0" box="[1080,1143,1920,1943]" italics="true" pageId="10" pageNumber="265">Piper</emphasis>
</taxonomicName>
as far north as San Martin Tuxtla, 
<collectingRegion id="95758685FFC80B21FF31F85A6D00BB9D" box="[237,340,1955,1978]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="265">Veracruz</collectingRegion>
, 
<collectingCountry id="2FA608F7FFC80B21FE82F85B6DE2BB9C" box="[350,438,1954,1979]" name="Mexico" pageId="10" pageNumber="265">Mexico</collectingCountry>
, and as far south as Itacolomi State Park in 
<collectingRegion id="95758685FFC80B21FC7CF85B686DBB9C" box="[928,1081,1954,1979]" country="Brazil" name="Minas Gerais" pageId="10" pageNumber="265">Minas Gerais</collectingRegion>
, 
<collectingCountry id="2FA608F7FFC80B21FB98F85B68D8BB9C" box="[1092,1164,1954,1979]" name="Brazil" pageId="10" pageNumber="265">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="03CE18EFFFC90B20FF4BF9156D04BBFE" httpUri="https://zenodo.org/record/2623567/files/figure.png" pageId="11" pageNumber="266" startId="11.[151,264,1772,1795]" targetBox="[174,1423,193,1748]" targetPageId="11">
<paragraph id="570E4867FFC90B20FF4BF9156D04BBFE" blockId="11.[151,1436,1772,2009]" pageId="11" pageNumber="266">
<emphasis id="65C59475FFC90B20FF4BF9156D6BBB26" bold="true" box="[151,319,1772,1795]" pageId="11" pageNumber="266">FIGURES 1–2.</emphasis>
Diagrammatic representation of major diagnostic features of 
<taxonomicName id="90B133E4FFC90B20FC09F915687DBB25" authorityName="Gordon" authorityYear="1999" box="[981,1065,1772,1794]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
and 
<taxonomicName id="90B133E4FFC90B20FBB8F91568A3BB25" authorityName="Mulsant" authorityYear="1846" box="[1124,1271,1772,1794]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
: 1, 
<taxonomicName id="90B133E4FFC90B20FAF8F9156929BB25" authorityName="Gordon" authorityYear="1999" box="[1316,1405,1772,1794]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
: a, ventral view of head capsule showing position of maxillary palp when retracted; b, left antennal club from dorsal view when extended; c, apex of mesothoracic leg showing trimerous tarsus; d, left half of first abdominal ventrite showing configuration of postcoxal line. 2, 
<taxonomicName id="90B133E4FFC90B20FE95F8B16D89BB79" authorityName="Mulsant" authorityYear="1846" box="[329,477,1864,1886]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="266" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
: a, ventral view of head capsule showing position of maxillary palp when retracted (left side of image) or extended (right side of image); b, left antennal club from dorsal view when extended (left image), same rotated 90 degrees counter clockwise to show membranous sensory patches of last two antennomeres (right image); c, apex of mesothoracic leg showing cryptotetramerous tarsus; d, left half of first abdominal ventrite showing various configurations of the postcoxal line.
</paragraph>
</caption>
<caption id="03CE18EFFFCE0B27FF4BF9016913BBE1" ID-DOI="http://doi.org/10.5281/zenodo.2623569" httpUri="https://zenodo.org/record/2623569/files/figure.png" pageId="12" pageNumber="267" startId="12.[151,264,1784,1807]" targetBox="[151,1433,212,1735]" targetPageId="12">
<paragraph id="570E4867FFCE0B27FF4BF9016913BBE1" blockId="12.[151,1437,1784,1991]" pageId="12" pageNumber="267">
<emphasis id="65C59475FFCE0B27FF4BF9016D12BB2A" bold="true" box="[151,326,1784,1807]" pageId="12" pageNumber="267">FIGURES 3–12.</emphasis>
Genitalia of 
<taxonomicName id="90B133E4FFCE0B27FE16F9016E4ABB29" authorityName="Gordon" authorityYear="1999" box="[458,542,1784,1806]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
and 
<taxonomicName id="90B133E4FFCE0B27FD93F9016EEEBB29" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[591,698,1784,1806]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="261" phylum="Arthropoda" rank="subTribe" subTribe="Selvadiina">Selvadiina</taxonomicName>
(
<taxonomicName id="90B133E4FFCE0B27FD1BF9016F0ABB29" authorityName="Mulsant" authorityYear="1846" box="[711,862,1784,1806]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
): 3–6, female genitalia, dorsal view (spermatheca shaded gray): 3, 
<taxonomicName id="90B133E4FFCE0B27FED9F8EE6EF1BB0A" authority="Gordon 1970" authorityName="Gordon" authorityYear="1970" box="[261,677,1815,1837]" class="Insecta" family="Coccinellidae" genus="Selvadius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="nunenmacheri">
<emphasis id="65C59475FFCE0B27FED9F8EE6E52BB0A" box="[261,518,1815,1837]" italics="true" pageId="12" pageNumber="267">Selvadius nunenmacheri</emphasis>
Gordon 1970
</taxonomicName>
(
<taxonomicName id="90B133E4FFCE0B27FD67F8EE6F04BB0A" authorityName="Mulsant" authorityYear="1846" box="[699,848,1815,1837]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
: 
<taxonomicName id="90B133E4FFCE0B27FCBFF8EE6F86BB0A" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[867,978,1815,1837]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="261" phylum="Arthropoda" rank="subTribe" subTribe="Selvadiina">Selvadiina</taxonomicName>
), showing distal end of sperm duct and spermatheca, accessory gland not shown (after Gordon 1985); 4, 
<taxonomicName id="90B133E4FFCE0B27FCB0F8CC68A4BB6C" authority="Gordon 1999" authorityName="Gordon" authorityYear="1999" box="[876,1264,1845,1867]" class="Insecta" family="Coccinellidae" genus="Erratodiomus" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="brindisi">
<emphasis id="65C59475FFCE0B27FCB0F8CC6801BB6C" box="[876,1109,1845,1867]" italics="true" pageId="12" pageNumber="267">Erratodiomus brindisi</emphasis>
Gordon 1999
</taxonomicName>
(
<taxonomicName id="90B133E4FFCE0B27FAD8F8CC69CDBB6C" authorityName="Mulsant" authorityYear="1846" box="[1284,1433,1845,1867]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Hyperaspidini">Hyperaspidini</taxonomicName>
: 
<taxonomicName id="90B133E4FFCE0B27FF4BF8AD6D52BB4D" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[151,262,1876,1898]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="261" phylum="Arthropoda" rank="subTribe" subTribe="Selvadiina">Selvadiina</taxonomicName>
), bursa through spermatheca, accessory gland not shown (after Gordon 1999); 5, 
<taxonomicName id="90B133E4FFCE0B27FB88F8AD6D70BBAE" authority="Gordon &amp; Hilburn 1990" authorityName="Gordon &amp; Hilburn" authorityYear="1990" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="hughesi">
<emphasis id="65C59475FFCE0B27FB88F8AD697FBB4D" box="[1108,1323,1876,1898]" italics="true" pageId="12" pageNumber="267">Decadiomus hughesi</emphasis>
Gordon &amp; Hilburn 1990
</taxonomicName>
(
<taxonomicName id="90B133E4FFCE0B27FEE9F88A6DDBBBAE" authorityName="Gordon" authorityYear="1999" box="[309,399,1907,1929]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
), bursa through spermatheca (note long convoluted sperm duct); 6, 
<taxonomicName id="90B133E4FFCE0B27FBB5F88D68B6BBAE" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1129,1250,1907,1929]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCE0B27FBB5F88D682ABBAE" box="[1129,1150,1908,1929]" italics="true" pageId="12" pageNumber="267">M</emphasis>
. 
<emphasis id="65C59475FFCE0B27FB4CF88A68B6BBAE" box="[1168,1250,1907,1929]" italics="true" pageId="12" pageNumber="267">lachesis</emphasis>
</taxonomicName>
(
<taxonomicName id="90B133E4FFCE0B27FB2FF88A691EBBAE" authorityName="Gordon" authorityYear="1999" box="[1267,1354,1907,1929]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
) (
<emphasis id="65C59475FFCE0B27FAB9F88D69DABBAE" box="[1381,1422,1908,1929]" italics="true" pageId="12" pageNumber="267">A.g.</emphasis>
= accessory gland); 7–12, male genitalia of 
<taxonomicName id="90B133E4FFCE0B27FD9DF86B6ECCBB8F" authorityName="Gordon" authorityYear="1999" box="[577,664,1938,1960]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
, left lateral view of penis: 7, 
<taxonomicName id="90B133E4FFCE0B27FC18F86A6866BB8F" authorityName="Gordon &amp; Hilburn" authorityYear="1990" box="[964,1074,1938,1960]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="hughesi">
<emphasis id="65C59475FFCE0B27FC18F86A6866BB8F" box="[964,1074,1938,1960]" italics="true" pageId="12" pageNumber="267">D. hughesi</emphasis>
</taxonomicName>
; 8, 
<taxonomicName id="90B133E4FFCE0B27FB86F86A6899BB8F" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1114,1229,1938,1960]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCE0B27FB86F86A6899BB8F" box="[1114,1229,1938,1960]" italics="true" pageId="12" pageNumber="267">M. lachesis</emphasis>
</taxonomicName>
; 9, 
<taxonomicName id="90B133E4FFCE0B27FB2AF86A6CB0BBE1" authority="Gordon" authorityName="Gordon" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="donatus">
<emphasis id="65C59475FFCE0B27FB2AF86A69C8BB8F" box="[1270,1436,1938,1960]" italics="true" pageId="12" pageNumber="267">Diomus donatus</emphasis>
Gordon
</taxonomicName>
(capsule); 10, 
<taxonomicName id="90B133E4FFCE0B27FEA7F84B6DBABBE1" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[379,494,1968,1991]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCE0B27FEA7F84B6DBABBE1" box="[379,494,1968,1991]" italics="true" pageId="12" pageNumber="267">M. lachesis</emphasis>
</taxonomicName>
(capsule); 11, 
<taxonomicName id="90B133E4FFCE0B27FD5FF8496F2EBBE1" baseAuthorityName="Horn" baseAuthorityYear="1895" box="[643,890,1968,1990]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="bigemmeus">
<emphasis id="65C59475FFCE0B27FD5FF8496F2EBBE1" box="[643,890,1968,1990]" italics="true" pageId="12" pageNumber="267">Decadiomus bigemmeus</emphasis>
</taxonomicName>
(capsule); 12, 
<taxonomicName id="90B133E4FFCE0B27FBCCF84968B0BBE1" baseAuthorityName="Horn" baseAuthorityYear="1895" box="[1040,1252,1968,1990]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="267" phylum="Arthropoda" rank="species" species="liebecki">
<emphasis id="65C59475FFCE0B27FBCCF84968B0BBE1" box="[1040,1252,1968,1990]" italics="true" pageId="12" pageNumber="267">Decadiomus liebecki</emphasis>
</taxonomicName>
(capsule)
</paragraph>
</caption>
<caption id="03CE18EFFFCF0B26FF4BF85C685EBBFC" ID-DOI="http://doi.org/10.5281/zenodo.2623571" httpUri="https://zenodo.org/record/2623571/files/figure.png" pageId="13" pageNumber="268" startId="13.[151,264,1957,1980]" targetBox="[151,1436,245,1896]" targetPageId="13">
<paragraph id="570E4867FFCF0B26FF4BF85C685EBBFC" blockId="13.[151,1437,1957,2011]" pageId="13" pageNumber="268">
<emphasis id="65C59475FFCF0B26FF4BF85C6D01BB9C" bold="true" box="[151,341,1957,1980]" pageId="13" pageNumber="268">FIGURES 13–16.</emphasis>
<taxonomicName id="90B133E4FFCF0B26FE80F85F6E61BB9B" box="[348,565,1958,1980]" pageId="13" pageNumber="268">
<emphasis id="65C59475FFCF0B26FE80F85F6DB0BB9B" box="[348,484,1958,1980]" italics="true" pageId="13" pageNumber="268">Moiradiomus</emphasis>
species
</taxonomicName>
habitus illustrations. 13, 
<taxonomicName id="90B133E4FFCF0B26FCE7F85E6FC9BB9B" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[827,925,1958,1980]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="clotho">
<emphasis id="65C59475FFCF0B26FCE7F85E6FC9BB9B" box="[827,925,1958,1980]" italics="true" pageId="13" pageNumber="268">M. clotho</emphasis>
</taxonomicName>
: holotype, male; 14, 
<taxonomicName id="90B133E4FFCF0B26FBA6F85E6888BB9B" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1146,1244,1958,1980]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="clotho">
<emphasis id="65C59475FFCF0B26FBA6F85E6888BB9B" box="[1146,1244,1958,1980]" italics="true" pageId="13" pageNumber="268">M. clotho</emphasis>
</taxonomicName>
: paratype, female. 15, 
<taxonomicName id="90B133E4FFCF0B26FF61F83F6D64BBFC" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[189,304,1989,2011]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCF0B26FF61F83F6D64BBFC" box="[189,304,1989,2011]" italics="true" pageId="13" pageNumber="268">M. lachesis</emphasis>
</taxonomicName>
: holotype, male; 16, 
<taxonomicName id="90B133E4FFCF0B26FDDBF83F6E2FBBFC" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[519,635,1989,2011]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCF0B26FDDBF83F6E2FBBFC" box="[519,635,1989,2011]" italics="true" pageId="13" pageNumber="268">M. lachesis</emphasis>
</taxonomicName>
: paratype, female. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="03CE18EFFFCC0B25FF4BF8546FBCBBC5" ID-DOI="http://doi.org/10.5281/zenodo.2623573" httpUri="https://zenodo.org/record/2623573/files/figure.png" pageId="14" pageNumber="269" startId="14.[151,264,1965,1988]" targetBox="[151,1436,260,1911]" targetPageId="14">
<paragraph id="570E4867FFCC0B25FF4BF8546FBCBBC5" blockId="14.[151,1437,1965,2018]" pageId="14" pageNumber="269">
<emphasis id="65C59475FFCC0B25FF4BF8546D07BBE4" bold="true" box="[151,339,1965,1988]" pageId="14" pageNumber="269">FIGURES 17–20.</emphasis>
<taxonomicName id="90B133E4FFCC0B25FE85F8546E64BBE4" box="[345,560,1965,1987]" pageId="14" pageNumber="269">
<emphasis id="65C59475FFCC0B25FE85F8546DB5BBE4" box="[345,481,1965,1987]" italics="true" pageId="14" pageNumber="269">Moiradiomus</emphasis>
species
</taxonomicName>
habitus illustrations. 17, 
<taxonomicName id="90B133E4FFCC0B25FCF3F8566FC8BBE3" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[815,924,1967,1988]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="atropos">
<emphasis id="65C59475FFCC0B25FCF3F8566FC8BBE3" box="[815,924,1967,1988]" italics="true" pageId="14" pageNumber="269">M. atropos</emphasis>
</taxonomicName>
: holotype, male; 18, 
<taxonomicName id="90B133E4FFCC0B25FBAEF85668B4BBE3" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1138,1248,1967,1988]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="atropos">
<emphasis id="65C59475FFCC0B25FBAEF85668B4BBE3" box="[1138,1248,1967,1988]" italics="true" pageId="14" pageNumber="269">M. atropos</emphasis>
</taxonomicName>
: paratype, female. 19, 
<taxonomicName id="90B133E4FFCC0B25FF61F8346D4BBBC5" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[189,287,1997,2018]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="nanita">
<emphasis id="65C59475FFCC0B25FF61F8346D4BBBC5" box="[189,287,1997,2018]" italics="true" pageId="14" pageNumber="269">M. nanita</emphasis>
</taxonomicName>
: holotype, male; 20, 
<taxonomicName id="90B133E4FFCC0B25FE2AF8346E0DBBC5" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[502,601,1997,2018]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="nanita">
<emphasis id="65C59475FFCC0B25FE2AF8346E0DBBC5" box="[502,601,1997,2018]" italics="true" pageId="14" pageNumber="269">M. nanita</emphasis>
</taxonomicName>
: paratype, female. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="03CE18EFFFCD0B24FF4BFB0C6F47B94F" ID-DOI="http://doi.org/10.5281/zenodo.2623575" httpUri="https://zenodo.org/record/2623575/files/figure.png" pageId="15" pageNumber="270" startId="15.[151,264,1269,1292]" targetBox="[157,1430,215,1228]" targetPageId="15">
<paragraph id="570E4867FFCD0B24FF4BFB0C6F47B94F" blockId="15.[151,1436,1269,1384]" pageId="15" pageNumber="270">
<emphasis id="65C59475FFCD0B24FF4BFB0C6D03B92C" bold="true" box="[151,343,1269,1292]" pageId="15" pageNumber="270">FIGURES 21–24.</emphasis>
Dorsal view of right mandible: 21, 
<taxonomicName id="90B133E4FFCD0B24FD00FB0F680AB92B" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[732,1118,1270,1292]" class="Insecta" family="Coccinellidae" genus="Subcoccinella" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="vigintiquatuorpunctata">
<emphasis id="65C59475FFCD0B24FD00FB0F680AB92B" box="[732,1118,1270,1292]" italics="true" pageId="15" pageNumber="270">Subcoccinella vigintiquatuorpunctata</emphasis>
</taxonomicName>
(L.) (folivorous) (after Kovář 1996); 22, 
<taxonomicName id="90B133E4FFCD0B24FED5FAEC6D22B90C" box="[265,374,1301,1323]" class="Insecta" family="Coccinellidae" genus="Bulaea" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="65C59475FFCD0B24FED5FAEC6D05B90C" box="[265,337,1301,1323]" italics="true" pageId="15" pageNumber="270">Bulaea</emphasis>
sp.
</taxonomicName>
(omnivorous with emphasis on pollinivory/phytophagy); 23, 
<taxonomicName id="90B133E4FFCD0B24FC28FAEC6883B90C" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1012,1239,1301,1323]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCD0B24FC28FAEC6883B90C" box="[1012,1239,1301,1323]" italics="true" pageId="15" pageNumber="270">Moiradiomus lachesis</emphasis>
</taxonomicName>
(enlarged detail of apical portion of incisor blade shown to left of main structure) (specialized on 
<taxonomicName id="90B133E4FFCD0B24FC3EFACC684FB96D" box="[994,1051,1333,1354]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="15" pageNumber="270" phylum="Tracheophyta" rank="genus">
<emphasis id="65C59475FFCD0B24FC3EFACC684FB96D" box="[994,1051,1333,1354]" italics="true" pageId="15" pageNumber="270">Piper</emphasis>
</taxonomicName>
food bodies); 24, 
<taxonomicName id="90B133E4FFCD0B24FB3EFACD69C8B96D" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1250,1436,1332,1354]" class="Insecta" family="Coccinellidae" genus="Adalia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="65C59475FFCD0B24FB3EFACD69C8B96D" box="[1250,1436,1332,1354]" italics="true" pageId="15" pageNumber="270">Adalia bipunctata</emphasis>
</taxonomicName>
(entomophagous with emphasis on aphids) (after Kovář 1996).
</paragraph>
</caption>
<caption id="03CE18EFFFCD0B24FF4BF84E6F39BBCA" ID-DOI="http://doi.org/10.5281/zenodo.2623578" httpUri="https://zenodo.org/record/2623578/files/figure.png" pageId="15" pageNumber="270" startId="15.[151,264,1975,1998]" targetBox="[151,1436,1466,1918]" targetPageId="15">
<paragraph id="570E4867FFCD0B24FF4BF84E6F39BBCA" blockId="15.[151,1435,1975,2029]" pageId="15" pageNumber="270">
<emphasis id="65C59475FFCD0B24FF4BF84E6D03BBEA" bold="true" box="[151,343,1975,1998]" pageId="15" pageNumber="270">FIGURES 25–28.</emphasis>
Diagrammatic ventral view of 
<taxonomicName id="90B133E4FFCD0B24FD78F8416F79BBE9" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[676,813,1976,1998]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFCD0B24FD78F8416F79BBE9" box="[676,813,1976,1998]" italics="true" pageId="15" pageNumber="270">Moiradiomus</emphasis>
</taxonomicName>
prosterna (setae, color patterns, and surface punctation not indicated): 25, 
<taxonomicName id="90B133E4FFCD0B24FEF3F8216DC4BBCA" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[303,400,2007,2029]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="clotho">
<emphasis id="65C59475FFCD0B24FEF3F8216DC4BBCA" box="[303,400,2007,2029]" italics="true" pageId="15" pageNumber="270">M. clotho</emphasis>
</taxonomicName>
; 26, 
<taxonomicName id="90B133E4FFCD0B24FE18F8216E63BBCA" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[452,567,2007,2029]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFCD0B24FE18F8216D8DBBCA" box="[452,473,2008,2029]" italics="true" pageId="15" pageNumber="270">M</emphasis>
. 
<emphasis id="65C59475FFCD0B24FE39F82E6E63BBCA" box="[485,567,2007,2029]" italics="true" pageId="15" pageNumber="270">lachesis</emphasis>
</taxonomicName>
; 27, 
<taxonomicName id="90B133E4FFCD0B24FDB6F8216E9BBBCA" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[618,719,2008,2029]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="atopos">
<emphasis id="65C59475FFCD0B24FDB6F8216E9BBBCA" box="[618,719,2008,2029]" italics="true" pageId="15" pageNumber="270">M. atopos</emphasis>
</taxonomicName>
; 28, 
<taxonomicName id="90B133E4FFCD0B24FCDFF8216F3CBBCA" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[771,872,2008,2029]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="270" phylum="Arthropoda" rank="species" species="nanita">M. nanita</taxonomicName>
.
</paragraph>
</caption>
<caption id="03CE18EFFFD20B3BFF4BF95D6913BAFD" ID-DOI="http://doi.org/10.5281/zenodo.2623580" httpUri="https://zenodo.org/record/2623580/files/figure.png" pageId="16" pageNumber="271" startId="16.[151,264,1700,1723]" targetBox="[154,1436,190,1666]" targetPageId="16">
<paragraph id="570E4867FFD20B3BFF4BF95D6913BAFD" blockId="16.[151,1436,1700,1754]" pageId="16" pageNumber="271">
<emphasis id="65C59475FFD20B3BFF4BF95D6D01BA9D" bold="true" box="[151,341,1700,1723]" pageId="16" pageNumber="271">FIGURES 29–32.</emphasis>
Male genitalia of 
<taxonomicName id="90B133E4FFD20B3BFDCEF95C6EBFBA9C" box="[530,747,1701,1723]" pageId="16" pageNumber="271">
<emphasis id="65C59475FFD20B3BFDCEF95C6ECEBA9C" box="[530,666,1701,1723]" italics="true" pageId="16" pageNumber="271">Moiradiomus</emphasis>
species
</taxonomicName>
(a, left lateral view of phallobase; b, ventral view of phallobase; c, left lateral view of penis; d, apex of basal lobe, enlarged): 29, 
<taxonomicName id="90B133E4FFD20B3BFCD6F93C6F3FBAFD" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[778,875,1732,1754]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="species" species="clotho">
<emphasis id="65C59475FFD20B3BFCD6F93C6F3FBAFD" box="[778,875,1732,1754]" italics="true" pageId="16" pageNumber="271">M. clotho</emphasis>
</taxonomicName>
; 30, 
<taxonomicName id="90B133E4FFD20B3BFC42F93C6846BAFD" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[926,1042,1732,1754]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="species" species="lachesis">
<emphasis id="65C59475FFD20B3BFC42F93C6846BAFD" box="[926,1042,1732,1754]" italics="true" pageId="16" pageNumber="271">M. lachesis</emphasis>
</taxonomicName>
; 31, 
<taxonomicName id="90B133E4FFD20B3BFB99F93C68FEBAFD" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1093,1194,1733,1754]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="species" species="atopos">
<emphasis id="65C59475FFD20B3BFB99F93C68FEBAFD" box="[1093,1194,1733,1754]" italics="true" pageId="16" pageNumber="271">M. atopos</emphasis>
</taxonomicName>
; 32, 
<taxonomicName id="90B133E4FFD20B3BFB01F93C6917BAFD" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1245,1347,1733,1754]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="species" species="nanita">M. nanita</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="1FAB1BECFFD20B3BFF1BF8F86C90BB8D" pageId="16" pageNumber="271" type="etymology">
<paragraph id="570E4867FFD20B3BFF1BF8F86C90BB8D" blockId="16.[151,1437,1793,2034]" pageId="16" pageNumber="271">
<emphasis id="65C59475FFD20B3BFF1BF8F86D1ABB3D" bold="true" box="[199,334,1793,1818]" pageId="16" pageNumber="271">Etymology.</emphasis>
<taxonomicName id="90B133E4FFD20B3BFE88F8F86DBFBB3D" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[340,491,1793,1818]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD20B3BFE88F8F86DBFBB3D" box="[340,491,1793,1818]" italics="true" pageId="16" pageNumber="271">Moiradiomus</emphasis>
</taxonomicName>
(gender Masculine) formed from a combination of Moira (=fate) + 
<taxonomicName id="90B133E4FFD20B3BFB3CF8FB6963BB3E" authorityName="Mulsant" authorityYear="1850" box="[1248,1335,1794,1817]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD20B3BFB3CF8FB6963BB3E" box="[1248,1335,1794,1817]" italics="true" pageId="16" pageNumber="271">Diomus</emphasis>
</taxonomicName>
(a genus of lady beetle, possibly in reference to Greek mythology where 
<taxonomicName id="90B133E4FFD20B3BFC81F8DC6FE2BB19" authorityName="Mulsant" authorityYear="1850" box="[861,950,1829,1854]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="285" phylum="Arthropoda" rank="genus">Diomus</taxonomicName>
was an Athenian hero, son of Colyttus, and a companion of Heracles). Moira is an Ancient Greek word (µoῖρα) meaning a portion or lot of the whole. In Greek mythology the Moirae are the three fates, daughters of Zeus and Themis, who spin, measure, and cut the thread of life.
</paragraph>
</subSubSection>
<subSubSection id="1FAB1BECFFD20B3AFF1BF84C68D5BE1B" lastPageId="17" lastPageNumber="272" pageId="16" pageNumber="271" type="discussion">
<paragraph id="570E4867FFD20B3AFF1BF84C68D5BE1B" blockId="16.[151,1437,1793,2034]" lastBlockId="17.[151,1436,151,573]" lastPageId="17" lastPageNumber="272" pageId="16" pageNumber="271">
<emphasis id="65C59475FFD20B3BFF1BF84C6D6FBBE9" bold="true" box="[199,315,1973,1998]" pageId="16" pageNumber="271">Remarks.</emphasis>
The relationships between the new genus and other genera of 
<taxonomicName id="90B133E4FFD20B3BFBF3F84C68D8BBE9" authorityName="Gordon" authorityYear="1999" box="[1071,1164,1973,1998]" class="Insecta" family="Coccinellidae" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="tribe" tribe="Diomini">Diomini</taxonomicName>
(sens. nov.) remains a mystery. Adult members of 
<taxonomicName id="90B133E4FFD20B3BFE08F8206E3FBBD5" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[468,619,2009,2034]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD20B3BFE08F8206E3FBBD5" box="[468,619,2009,2034]" italics="true" pageId="16" pageNumber="271">Moiradiomus</emphasis>
</taxonomicName>
key to 
<taxonomicName id="90B133E4FFD20B3BFD1EF8206F1ABBD5" authorityName="Chapin" authorityYear="1933" box="[706,846,2009,2034]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD20B3BFD1EF8206F1ABBD5" box="[706,846,2009,2034]" italics="true" pageId="16" pageNumber="271">Decadiomus</emphasis>
</taxonomicName>
in 
<bibRefCitation id="33203596FFD20B3BFCA8F8206848BBD5" author="Gordon, R. D." box="[884,1052,2009,2034]" pageId="16" pageNumber="271" pagination="1 - 219" refId="ref20207" refString="Gordon, R. D. (1999) South American Coccinellidae (Coleoptera), Part VI: A systematic revision of the South American Diomini, new tribe (Scymninae). Annales Zoologici, 49 (1), 1 - 219." type="journal article" year="1999">Gordon (1999)</bibRefCitation>
because the antenna is composed of 10 antennomeres. However, individuals with only 10 antennomeres were also noted to randomly occur in certain Australian 
<taxonomicName id="90B133E4FFD30B3AFECFFF446D96BCF2" box="[275,450,188,213]" pageId="17" pageNumber="272">
<emphasis id="65C59475FFD30B3AFECFFF446D3EBCF3" box="[275,362,189,212]" italics="true" pageId="17" pageNumber="272">Diomus</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="33203596FFD30B3AFE0DFF456E83BCF3" author="Pang, H. &amp; Slipinski, A." box="[465,727,188,213]" pageId="17" pageNumber="272" pagination="641 - 698" refId="ref21604" refString="Pang, H. &amp; Slipinski, A. (2009) Revision of the Australian Coccinellidae (Coleoptera). genus Diomus Mulsant. Part 1. Annales Zoologici, 59 (4), 641 - 698. https: // doi. org / 10.3161 / 000345409 x 485008" type="journal article" year="2009">Pang &amp; Ślipiński 2009</bibRefCitation>
). The female genitalia of 
<taxonomicName id="90B133E4FFD30B3AFC2BFF4568DABCF2" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[1015,1166,188,213]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFC2BFF4568DABCF2" box="[1015,1166,188,213]" italics="true" pageId="17" pageNumber="272">Moiradiomus</emphasis>
</taxonomicName>
are quite different from members of 
<taxonomicName id="90B133E4FFD30B3AFEFEFF266DFABCDF" authorityName="Chapin" authorityYear="1933" box="[290,430,223,248]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFEFEFF266DFABCDF" box="[290,430,223,248]" italics="true" pageId="17" pageNumber="272">Decadiomus</emphasis>
</taxonomicName>
and instead match those of 
<taxonomicName id="90B133E4FFD30B3AFD39FF266FECBCD0" box="[741,952,223,248]" class="Insecta" family="Coccinellidae" genus="Heterodiomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="65C59475FFD30B3AFD39FF266FD7BCDF" box="[741,899,223,248]" italics="true" pageId="17" pageNumber="272">Heterodiomus</emphasis>
spp.
</taxonomicName>
(
<bibRefCitation id="33203596FFD30B3AFC1AFF26680BBCDF" author="Gordon, R. D." box="[966,1119,223,248]" pageId="17" pageNumber="272" pagination="1 - 219" refId="ref20207" refString="Gordon, R. D. (1999) South American Coccinellidae (Coleoptera), Part VI: A systematic revision of the South American Diomini, new tribe (Scymninae). Annales Zoologici, 49 (1), 1 - 219." type="journal article" year="1999">Gordon 1999</bibRefCitation>
; 
<bibRefCitation id="33203596FFD30B3AFBB6FF2669C5BCDF" author="Gordon, R. D. &amp; Gonzalez, G." box="[1130,1425,223,248]" pageId="17" pageNumber="272" pagination="237 - 239" refId="ref20305" refString="Gordon, R. D. &amp; Gonzalez, G. (2003) Descriptions of the male of Heterodiomus marchali Brethes and a new species of Chilean Heterodiomus Brethes (Coleoptera: Coccinellidae: Scymninae: Diomini). Insecta Mundi, 17 (3 - 4), 237 - 239." type="journal article" year="2003">Gordon &amp; González 2003</bibRefCitation>
). The larva of the new genus differs greatly from the unusual onisciform larva of two myrmecophilous species, 
<taxonomicName id="90B133E4FFD30B3AFF4BFED06D3BBD67" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[151,367,295,320]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="thoracicus">
<emphasis id="65C59475FFD30B3AFF4BFED06D3BBD67" box="[151,367,295,320]" italics="true" pageId="17" pageNumber="272">Diomus thoracicus</emphasis>
</taxonomicName>
(
<typeStatus id="880AF6C5FFD30B3AFE5EFED16DE0BD67" box="[386,436,296,320]" pageId="17" pageNumber="272">type</typeStatus>
species of 
<taxonomicName id="90B133E4FFD30B3AFDE1FED06EC0BD67" authorityName="Mulsant" authorityYear="1850" box="[573,660,297,320]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFDE1FED06EC0BD67" box="[573,660,297,320]" italics="true" pageId="17" pageNumber="272">Diomus</emphasis>
</taxonomicName>
) and 
<taxonomicName id="90B133E4FFD30B3AFD00FED06C8FBD43" authority="Vandenberg et al. (Vandenberg et al. 2018)" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="lupusapudoves">
<emphasis id="65C59475FFD30B3AFD00FED06FB0BD67" box="[732,996,295,320]" italics="true" pageId="17" pageNumber="272">Diomus lupusapudoves</emphasis>
<bibRefCitation id="33203596FFD30B3AFC33FEDE6C8FBD43" author="Vandenberg, N. J. &amp; Iverson, A. &amp; Liere, H." pageId="17" pageNumber="272" pagination="113 - 122" refId="ref22928" refString="Vandenberg, N. J., Iverson, A. &amp; Liere, H. (2018) A new species of myrmecophilous lady beetle in the genus Diomus (Coleoptera: Coccinellidae: Diomini) from Chiapas, Mexico that feeds on green coffee scale, Coccus viridis (Green) (Hemiptera: Coccidae). Zootaxa, 4420 (1), 113 - 122. https: // doi. org / 10.11646 / zootaxa. 4420.1.6" type="journal article" year="2018">
Vandenberg 
<emphasis id="65C59475FFD30B3AFBA3FED068E0BD67" box="[1151,1204,295,320]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. (Vandenberg 
<emphasis id="65C59475FFD30B3AFABCFED069C0BD67" box="[1376,1428,295,320]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. 2018)
</bibRefCitation>
</taxonomicName>
, but is similar to the setiferous, spindle-shaped forms found in 
<taxonomicName id="90B133E4FFD30B3AFC63FEB46972BD42" authority="(Say, 1835)" baseAuthorityName="Say" baseAuthorityYear="1835" box="[959,1318,332,357]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="terminatus">
<emphasis id="65C59475FFD30B3AFC63FEB468CCBD43" box="[959,1176,333,356]" italics="true" pageId="17" pageNumber="272">Diomus terminatus</emphasis>
(Say, 1835)
</taxonomicName>
(
<bibRefCitation id="33203596FFD30B3AFAEBFEB56D62BDAF" author="Hentz, M. G. &amp; Nuessly, G. S." pageId="17" pageNumber="272" pagination="276 - 278" refId="ref20409" refString="Hentz, M. G. &amp; Nuessly, G. S. (2002) Morphology and biology of Diomus terminatus (Coleoptera: Coccinellidae), a predator of Sipha flava (Homoptera: Aphidae). Florida Entomologist, 85, 276 - 278. https: // doi. org / 10.1653 / 0015 - 4040 (2002) 085 [0276: MABODT] 2.0. CO; 2" type="journal article" year="2002">
Hentz 
<emphasis id="65C59475FFD30B3AFA5AFEB469C8BD43" box="[1414,1436,333,356]" italics="true" pageId="17" pageNumber="272">&amp;</emphasis>
Nuessly 2002
</bibRefCitation>
, 
<bibRefCitation id="33203596FFD30B3AFE9FFE966E44BDAF" author="Akbar, W. &amp; Carlton, C. &amp; Reagan, T. E." box="[323,528,367,392]" pageId="17" pageNumber="272" pagination="96 - 103" refId="ref18759" refString="Akbar, W., Carlton, C. &amp; Reagan, T. E. (2009) Life cycle and larval morphology of Diomus terminatus (Coleoptera: Coccinellidae) and its potential as a biological control agent of Melanaphis sacchari (Hemiptera: Aphididae). Annals of the Entomological Society of America, 102, 96 - 103. https: // doi. org / 10.1603 / 008.102.0110" type="journal article" year="2009">
Akbar 
<emphasis id="65C59475FFD30B3AFE4FFE886D91BDAF" box="[403,453,367,392]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. 2009
</bibRefCitation>
) and 
<taxonomicName id="90B133E4FFD30B3AFD8FFE8868D3BDAF" authority="(Mulsant, 1853) (Rees et al. 1994)" authorityName="Rees et al." authorityYear="1994" baseAuthorityName="Mulsant" baseAuthorityYear="1853" box="[595,1159,367,392]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="roseicollis">
<emphasis id="65C59475FFD30B3AFD8FFE886EB8BDAF" box="[595,748,367,392]" italics="true" pageId="17" pageNumber="272">D. roseicollis</emphasis>
(Mulsant, 1853) (
<bibRefCitation id="33203596FFD30B3AFC61FE89682BBDAF" author="Rees, B. E. &amp; Anderson, D. M. &amp; Bouk, D. &amp; Gordon, R. D." box="[957,1151,367,392]" pageId="17" pageNumber="272" pagination="387 - 412" refId="ref21900" refString="Rees, B. E., Anderson, D. M., Bouk, D. &amp; Gordon, R. D. (1994) Larval key to genera and selected species of North American Coccinellidae (Coleoptera). Proceedings of the Entomological Society of Washington, 96, 387 - 412." type="journal article" year="1994">
Rees 
<emphasis id="65C59475FFD30B3AFC23FE886867BDAF" box="[1023,1075,367,392]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. 1994
</bibRefCitation>
)
</taxonomicName>
, as well as 
<taxonomicName id="90B133E4FFD30B3AFACCFE966DD5BD8B" authority="Chapin, 1933" authorityName="Chapin" authorityYear="1933" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="65C59475FFD30B3AFACCFE966C8EBD8B" italics="true" pageId="17" pageNumber="272">Decadiomus pictus</emphasis>
<bibRefCitation id="33203596FFD30B3AFF39FE6D6DD5BD8B" author="Chapin, E. A." box="[229,385,404,429]" pageId="17" pageNumber="272" pagination="95 - 99" refId="ref19138" refString="Chapin, E. A. (1933) A new genus of West Indian Coccinellidae (Coleoptera). Proceedings of the Biological Society of Washington, 46, 95 - 99." type="journal article" year="1933">Chapin, 1933</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="33203596FFD30B3AFE48FE6D6E64BD8B" author="Boving, A. G." box="[404,560,404,429]" pageId="17" pageNumber="272" refId="ref18892" refString="Boving, A. G. (1933) Description of the larvae of Decadiomus pictus Chapin (Scymnini, Coccinellidae). Proceedings of the Entomological Society of Washington, 46, 101 - 04." type="book" year="1933">Böving 1933</bibRefCitation>
). 
<taxonomicName id="90B133E4FFD30B3AFD94FE6D6E8BBD8A" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[584,735,404,429]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFD94FE6D6E8BBD8A" box="[584,735,404,429]" italics="true" pageId="17" pageNumber="272">Moiradiomus</emphasis>
</taxonomicName>
larvae also appear to lack the dorsal intersegmental glands reported in 
<taxonomicName id="90B133E4FFD30B3AFEC0FE406D27BDF7" authorityName="Mulsant" authorityYear="1850" box="[284,371,441,464]" class="Insecta" family="Coccinellidae" genus="Diomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFEC0FE406D27BDF7" box="[284,371,441,464]" italics="true" pageId="17" pageNumber="272">Diomus</emphasis>
</taxonomicName>
and 
<taxonomicName id="90B133E4FFD30B3AFE72FE4E6E6EBDF7" authorityName="Chapin" authorityYear="1933" box="[430,570,439,464]" class="Insecta" family="Coccinellidae" genus="Decadiomus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFE72FE4E6E6EBDF7" box="[430,570,439,464]" italics="true" pageId="17" pageNumber="272">Decadiomus</emphasis>
</taxonomicName>
larvae (
<bibRefCitation id="33203596FFD30B3AFD45FE416F0DBDF7" author="Rees, B. E. &amp; Anderson, D. M. &amp; Bouk, D. &amp; Gordon, R. D." box="[665,857,439,464]" pageId="17" pageNumber="272" pagination="387 - 412" refId="ref21900" refString="Rees, B. E., Anderson, D. M., Bouk, D. &amp; Gordon, R. D. (1994) Larval key to genera and selected species of North American Coccinellidae (Coleoptera). Proceedings of the Entomological Society of Washington, 96, 387 - 412." type="journal article" year="1994">
Rees 
<emphasis id="65C59475FFD30B3AFD06FE406F41BDF7" box="[730,789,439,464]" italics="true" pageId="17" pageNumber="272">et al.</emphasis>
1994
</bibRefCitation>
; 
<bibRefCitation id="33203596FFD30B3AFCBBFE4E6822BDF7" author="Vandenberg, N. J. &amp; Iverson, A. &amp; Liere, H." box="[871,1142,439,464]" pageId="17" pageNumber="272" pagination="113 - 122" refId="ref22928" refString="Vandenberg, N. J., Iverson, A. &amp; Liere, H. (2018) A new species of myrmecophilous lady beetle in the genus Diomus (Coleoptera: Coccinellidae: Diomini) from Chiapas, Mexico that feeds on green coffee scale, Coccus viridis (Green) (Hemiptera: Coccidae). Zootaxa, 4420 (1), 113 - 122. https: // doi. org / 10.11646 / zootaxa. 4420.1.6" type="journal article" year="2018">
Vandenberg 
<emphasis id="65C59475FFD30B3AFC29FE40687CBDF7" box="[1013,1064,439,464]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. 2018
</bibRefCitation>
). These glands have been shown to have a repellent effect on ants or play a role in chemical mimicry and are identified by 
<bibRefCitation id="33203596FFD30B3AFB16FE2569C8BDD3" author="Seago, A. E. &amp; Giorgi, J. A. &amp; Li, J. &amp; Slipinski, A." box="[1226,1436,476,501]" pageId="17" pageNumber="272" pagination="137 - 151" refId="ref22430" refString="Seago, A. E., Giorgi, J. A., Li, J. &amp; Slipinski, A. (2011) Phylogeny, classification and evolution of ladybird beetles (Coleoptera: Coccinellidae) based on simultaneous analysis of molecular and morphological data. Molecular Phylogenetics and Evolution, 60 (1), 137 - 151. https: // doi. org / 10.1016 / j. ympev. 2011.03.015" type="journal article" year="2011">
Seago 
<emphasis id="65C59475FFD30B3AFAC8FE246910BDD2" box="[1300,1348,476,501]" italics="true" pageId="17" pageNumber="272">et al</emphasis>
. (2011)
</bibRefCitation>
as an important step in the transition to feeding on ant-protected prey. Presumably they have been rendered obsolete by the enclosed environment provided by the woven larval tent of 
<taxonomicName id="90B133E4FFD30B3AFC3FFDDD682EBE1A" authorityName="Vandenberg &amp; Hanson" authorityYear="2019" box="[995,1146,548,573]" class="Insecta" family="Coccinellidae" genus="Moiradiomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="65C59475FFD30B3AFC3FFDDD682EBE1A" box="[995,1146,548,573]" italics="true" pageId="17" pageNumber="272">Moiradiomus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>