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<mods:title id="0A4A4CD4F5961AAF3559B342E855133E">Fourteen new species of scuttle flies (Diptera, Phoridae) from Denmark</mods:title>
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<treatment id="D27187E87524A07D1EAFFD34FCD4682F" ID-DOI="http://doi.org/10.5281/zenodo.5929324" ID-GBIF-Taxon="154529721" ID-Zenodo-Dep="5929324" LSID="urn:lsid:plazi:treatment:D27187E87524A07D1EAFFD34FCD4682F" httpUri="http://treatment.plazi.org/id/D27187E87524A07D1EAFFD34FCD4682F" lastPageNumber="77" pageId="14" pageNumber="77">
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<heading id="012F81927524A07D1EAFFD34FE3C6CAE" bold="true" box="[151,500,660,686]" fontSize="11" level="1" pageId="14" pageNumber="77" reason="1">
<emphasis id="68ACEAEC7524A07D1EAFFD34FE3C6CAE" bold="true" box="[151,500,660,686]" pageId="14" pageNumber="77">
<taxonomicName id="9DD84D7D7524A07D1EAFFD34FE636CAE" ID-CoL="8CS7R" authority="Henry &amp; Bøggild, 2019" authorityName="Henry &amp; Bøggild" authorityYear="2019" box="[151,427,660,686]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="parhirticrus" status="sp. nov.">
<emphasis id="68ACEAEC7524A07D1EAFFD34FE636CAE" bold="true" box="[151,427,660,686]" italics="true" pageId="14" pageNumber="77">Megaselia parhirticrus</emphasis>
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<subSubSection id="12C265757524A07D1EAFFD16FEFD6CCF" box="[151,309,694,719]" pageId="14" pageNumber="77" type="description">
<paragraph id="5A6736FE7524A07D1EAFFD16FEFD6CCF" blockId="14.[151,500,660,719]" box="[151,309,694,719]" pageId="14" pageNumber="77">
(
<figureCitation id="C2E32A7B7524A07D1EA7FD16FEE46CCF" box="[159,300,694,719]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Figs 92102</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12C265757524A07D1EAFFD5EFEF76D3B" pageId="14" pageNumber="77" type="materials_examined">
<paragraph id="5A6736FE7524A07D1EAFFD5EFEF76D3B" blockId="14.[151,1437,766,1583]" pageId="14" pageNumber="77">
<emphasis id="68ACEAEC7524A07D1EAFFD5EFE4E6D17" bold="true" box="[151,390,766,791]" pageId="14" pageNumber="77">Material examined.</emphasis>
<materialsCitation id="EAB03CA37524A07D1FABFD5EFEF76D3B" ID-GBIF-Occurrence="2147524709" collectingDate="2014-06-21" collectingDateMax="2014-07-13" collectingDateMin="2014-06-21" collectorName="E. Boggild" country="Denmark" location="Draved Skov" municipality="Hojmose" pageId="14" pageNumber="77" specimenCount="1" typeStatus="holotype">
<typeStatus id="8563885C7524A07D1FABFD5EFE336D17" box="[403,507,766,791]" pageId="14" pageNumber="77" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="22CF766E7524A07D1C6BFD5EFD156D17" box="[595,733,766,791]" name="Denmark" pageId="14" pageNumber="77">DENMARk</collectingCountry>
, SJ,
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,
<collectingMunicipality id="BA03AC847524A07D1DF1FD5EFBF96D17" box="[969,1073,766,791]" pageId="14" pageNumber="77">Højmose</collectingMunicipality>
,
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,
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(CUMZ-8-59).
</materialsCitation>
</paragraph>
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<subSubSection id="12C265757524A07D1EFFFCE6FB35680A" pageId="14" pageNumber="77" type="description">
<paragraph id="5A6736FE7524A07D1EFFFCE6FA5F6AA3" blockId="14.[151,1437,766,1583]" pageId="14" pageNumber="77">
<emphasis id="68ACEAEC7524A07D1EFFFCE6FE9D6D5F" bold="true" box="[199,341,838,863]" pageId="14" pageNumber="77">Description.</emphasis>
Male. Head as
<figureCitation id="C2E32A7B7524A07D1C38FCE6FD856D5F" box="[512,589,838,863]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 92</figureCitation>
and the proboscis from below as
<figureCitation id="C2E32A7B7524A07D1D85FCE6FBC56D5F" box="[957,1037,838,863]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 93</figureCitation>
. Cheek with 5 bristles and jowl with 2 that are much longer and more robust. Postpedicels (
<figureCitation id="C2E32A7B7524A07D1CCBFCCAFC8B6D83" box="[755,835,874,899]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 92</figureCitation>
) light brown and without SPS vesicles. Thorax brown with notopleuron and mesopleuron as
<figureCitation id="C2E32A7B7524A07D1C7AFC2EFD5C6DA7" box="[578,660,910,935]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 94</figureCitation>
. Scutellum with an anterior pair of small hairs and a posterior pair of bristles (
<figureCitation id="C2E32A7B7524A07D1EC1FC12FE876DCB" box="[249,335,946,971]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 95</figureCitation>
). Abdominal tergites brown with many hairs which are moderately longer at rear of T6. Venter brownish gray with hairs on segments 36. Hypopygium as
<figureCitation id="C2E32A7B7524A07D1D0BFC76FC7A6DEF" box="[819,946,982,1007]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Figs 9698</figureCitation>
. Middle and hind legs light brown and front legs brownish yellow. Fore tarsus (
<figureCitation id="C2E32A7B7524A07D1C2AFC5AFDAA6A13" box="[530,610,1018,1043]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 99</figureCitation>
) with posterodorsal hair palisade on segments 14 and 5 longer than 4. Mid tibia and basitarsus as
<figureCitation id="C2E32A7B7524A07D1FAAFBBEFE3A6A37" box="[402,498,1054,1079]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 100</figureCitation>
. Hind femur as
<figureCitation id="C2E32A7B7524A07D1C9EFBBEFCCD6A37" box="[678,773,1054,1079]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 101</figureCitation>
. Hind tibia with about 20 differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (
<figureCitation id="C2E32A7B7524A07D1D46FBE2FC126A5B" box="[894,986,1090,1115]" captionStart="FIGURES 92102" captionStartId="15.[151,264,1953,1976]" captionTargetBox="[265,1320,198,1913]" captionTargetId="figure@15.[250,1337,193,1932]" captionTargetPageId="15" captionText="FIGURES 92102. Megaselia parhirticruss sp. n. male. 92. head. 93 proboscis from below. 94. left face of thorax. 95. scutellum. 9698. hypopygium. 99. front tarsus. 100. mid tibia and basitarsus. 101. hind femur. 102. basal half of wing." httpUri="https://zenodo.org/record/2617713/files/figure.png" pageId="14" pageNumber="77">Fig. 102</figureCitation>
) with exceptionally pale membranes,
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long. Costal index at least 0.3. Costal ratios 7.6: 4.0: 1. Costal cilia (of section 3)
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long. Hair at base of vein 3 small. With 5 axillary bristles, the outer being
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long. Sc not reaching R1. Haltere knob grayish brown.
</paragraph>
<paragraph id="5A6736FE7524A07D1EFFFB0EFB35680A" blockId="14.[151,1437,766,1583]" pageId="14" pageNumber="77">
<emphasis id="68ACEAEC7524A07D1EFFFB0EFE956AC7" bold="true" box="[199,349,1198,1223]" pageId="14" pageNumber="77">Recognition.</emphasis>
In the key of
<bibRefCitation id="3E494B0F7524A07D1FC5FB0EFD776AC7" author="Lundbeck, W." box="[509,703,1198,1223]" pageId="14" pageNumber="77" refId="ref7421" refString="Lundbeck, W. (1922) Diptera Danica. Part UI. Pipunculidae, Phoridae. Wesley, London, 455 pp." type="book" year="1922">Lundbeck (1922)</bibRefCitation>
to Group V it runs to couplet 10 lead 2 or couplet 12 lead 2, but the very short costal index and pale wing membrane immediately rules out both species. In the key to the males of the British Isles (
<bibRefCitation id="3E494B0F7524A07D1F64FB56FE3E6B0F" author="Disney, R. H. L." box="[348,502,1270,1295]" pageId="14" pageNumber="77" pagination="1 - 155" refId="ref7235" refString="Disney, R. H. L. (1989) Scuttle Flies-Diptera Phoridae Genus Megaselia. Handbooks for the Identification of British Insects, 10 (8), 1 - 155." type="journal article" year="1989">Disney, 1989</bibRefCitation>
) it runs to couplet 95, lead 2 to
<taxonomicName id="9DD84D7D7524A07D1D55FB57FBAD6B0F" authority="(Schmitz)" authorityName="Schmitz" baseAuthorityName="Schmitz" box="[877,1125,1270,1295]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="hiticrus">
<emphasis id="68ACEAEC7524A07D1D55FB57FC216B0F" box="[877,1001,1270,1295]" italics="true" pageId="14" pageNumber="77">M. hiticrus</emphasis>
(Schmitz)
</taxonomicName>
. The latter has darker legs, with the crowded hairs at the base of the hind femur only gradually increasing in size towards the base (
<figureCitation id="C2E32A7B7524A07D1B0BFABAFA5C6B33" box="[1331,1428,1306,1331]" captionStart="FIGURES 103104" captionStartId="16.[151,264,704,727]" captionTargetBox="[183,1396,219,656]" captionTargetId="figure@16.[151,1435,193,684]" captionTargetPageId="16" captionText="FIGURES 103104. Megaselia hirticrus male. 103. left face of hypopygium. 104. hind femur." httpUri="https://zenodo.org/record/2617715/files/figure.png" pageId="14" pageNumber="77">Fig. 104</figureCitation>
) and the darker hypopygium has the left hypandrial lobe forked (
<figureCitation id="C2E32A7B7524A07D1D49FA9EFC1D6B57" box="[881,981,1342,1367]" captionStart="FIGURES 103104" captionStartId="16.[151,264,704,727]" captionTargetBox="[183,1396,219,656]" captionTargetId="figure@16.[151,1435,193,684]" captionTargetPageId="16" captionText="FIGURES 103104. Megaselia hirticrus male. 103. left face of hypopygium. 104. hind femur." httpUri="https://zenodo.org/record/2617715/files/figure.png" pageId="14" pageNumber="77">Fig. 103</figureCitation>
). In
<bibRefCitation id="3E494B0F7524A07D1A31FA9EFA8A6B57" author="Schmitz, H. &amp; Delage, A." box="[1033,1346,1342,1367]" pageId="14" pageNumber="77" pagination="638 - 664" refId="ref7643" refString="Schmitz, H. &amp; Delage, A. (1974) Phoridae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. 4 (33). Lieferung 301. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 638 - 664." type="book chapter" year="1974">Schmitz &amp; Delages (1974)</bibRefCitation>
key for Abteilung V it runs to couplets 27 and 36. With the first option it runs to couplets 69 and 97 where the hairs at the base of the hind femur immediately distinguish it. With the second option it runs to couplets 27 and 36. At couplet 27 the hypopygiuym and very pale wings distinguish it. As with
<taxonomicName id="9DD84D7D7524A07D1D5EFA0CFC386BC3" authorityName="Henry &amp; Bøggild" authorityYear="2019" box="[870,1008,1450,1475]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="thomseni">
<emphasis id="68ACEAEC7524A07D1D5EFA0CFC386BC3" box="[870,1008,1450,1475]" italics="true" pageId="14" pageNumber="77">M. thomseni</emphasis>
</taxonomicName>
(see below) at couplet 36 it runs to
<taxonomicName id="9DD84D7D7524A07D1B45FA0CFE4A6BE7" authority="(Lundbeck)" authorityName="Lundbeck" baseAuthorityName="Lundbeck" baseAuthorityYear="1920" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="paupera">
<emphasis id="68ACEAEC7524A07D1B45FA0CFF3C6BE6" italics="true" pageId="14" pageNumber="77">M. paupera</emphasis>
(Lundbeck)
</taxonomicName>
, which is only known in the female. The hind femur distinguishes it from
<emphasis id="68ACEAEC7524A07D1AEDFA6FFAA16BE7" box="[1237,1385,1486,1511]" italics="true" pageId="14" pageNumber="77">
<taxonomicName id="9DD84D7D7524A07D1AEDFA6FFAAD6BE7" authorityName="Henry &amp; Bøggild" authorityYear="2019" box="[1237,1381,1486,1511]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="thomseni">M. thomseni</taxonomicName>
.
</emphasis>
The costal ratios and more numerous posterodorsal hairs of the hind tibia distinguish it from
<taxonomicName id="9DD84D7D7524A07D1A4CFA54FB3E680B" baseAuthorityName="Lundbeck" baseAuthorityYear="1920" box="[1140,1270,1524,1547]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="paupera">
<emphasis id="68ACEAEC7524A07D1A4CFA54FB3E680B" box="[1140,1270,1524,1547]" italics="true" pageId="14" pageNumber="77">M. paupera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="12C265757524A07D1EFFF9B6FCD4682F" box="[199,796,1558,1583]" pageId="14" pageNumber="77" type="etymology">
<paragraph id="5A6736FE7524A07D1EFFF9B6FCD4682F" blockId="14.[151,1437,766,1583]" box="[199,796,1558,1583]" pageId="14" pageNumber="77">
<emphasis id="68ACEAEC7524A07D1EFFF9B6FE86682F" bold="true" box="[199,334,1558,1583]" pageId="14" pageNumber="77">Etymology.</emphasis>
Named after its similarity to
<taxonomicName id="9DD84D7D7524A07D1CAFF9B7FCD4682F" baseAuthorityName="Schmitz" baseAuthorityYear="1918" box="[663,796,1558,1583]" class="Insecta" family="Phoridae" genus="Megaselia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="hirticrus">
<emphasis id="68ACEAEC7524A07D1CAFF9B7FCD4682F" box="[663,796,1558,1583]" italics="true" pageId="14" pageNumber="77">M. hirticrus</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>