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<document id="70800DDE5B3C0CD393ADBB77A7EDBC60" ID-DOI="10.11646/zootaxa.5399.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10493947" ID-ZooBank="EC9BEA58-70C4-4BA2-B7FA-094F49A911A1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1705011990532" checkinUser="plazi" docAuthor="Wang, Shuxia &amp; Xu, Zhixin" docDate="2024" docId="D27187D8AC3FFFDD33C09300287EF96E" docLanguage="en" docName="zootaxa.5399.2.1.pdf" docOrigin="Zootaxa 5399 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5399.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.17:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="17" docTitle="Periacma solitaridens Wang &amp; Xu 2024, sp. nov." docType="treatment" docVersion="3" lastPageNumber="118" masterDocId="2E48FFA0AC2EFFCC3357924B292BFFF7" masterDocTitle="Taxonomy of the genus Periacma Meyrick (Lepidoptera: Autostichidae) from China, with a checklist of the world species" masterLastPageNumber="129" masterPageNumber="101" pageNumber="118" updateTime="1705410171724" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="75F8010306B469D4B9096E95DBC6D243">Taxonomy of the genus Periacma Meyrick (Lepidoptera: Autostichidae) from China, with a checklist of the world species</mods:title>
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<mods:namePart id="2360EA4A5941197BC7B2285004BAB87F">Wang, Shuxia</mods:namePart>
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<mods:namePart id="5B07FF17D35592D218080CC680B331E5">Xu, Zhixin</mods:namePart>
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<mods:affiliation id="21FE3C50A583F7A91B8039DF95E0789B">College of Life Sciences, Nankai University, Tianjin 300071, China.</mods:affiliation>
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<paragraph id="5A6736CEAC3FFFDD33C093002B70FE92" blockId="17.[151,603,330,393]" box="[151,603,330,357]" pageId="17" pageNumber="118">
<heading id="012F81A2AC3FFFDD33C093002B70FE92" bold="true" box="[151,603,330,357]" fontSize="11" level="1" pageId="17" pageNumber="118" reason="1">
<emphasis id="68ACEADCAC3FFFDD33C093002B70FE92" bold="true" box="[151,603,330,357]" pageId="17" pageNumber="118">
<taxonomicName id="9DD84D4DAC3FFFDD33C0930028DBFE92" authority="Wang" authorityName="Wang &amp; Xu" authorityYear="2024" box="[151,496,330,357]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="solitaridens" status="sp. nov.">
<emphasis id="68ACEADCAC3FFFDD33C0930028B5FE92" bold="true" box="[151,414,330,357]" italics="true" pageId="17" pageNumber="118">Periacma solitaridens</emphasis>
Wang
</taxonomicName>
,
<taxonomicNameLabel id="739F57A7AC3FFFDD32AA93002B70FE92" box="[509,603,331,357]" pageId="17" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="12C26545AC3FFFDD33C09324280CFE7E" box="[151,295,367,393]" pageId="17" pageNumber="118" type="description">
<paragraph id="5A6736CEAC3FFFDD33C09324280CFE7E" blockId="17.[151,603,330,393]" box="[151,295,367,393]" pageId="17" pageNumber="118">
(
<figureCitation id="C2E32A4BAC3FFFDD33C8932429DDFE7E" box="[159,246,367,393]" captionStart="FIGURES 1018" captionStartId="4.[151,264,1816,1840]" captionTargetBox="[170,1417,194,1780]" captionTargetId="figure-22@4.[158,1430,181,1793]" captionTargetPageId="4" captionText="FIGURES 1018. Adults of Periacma spp. 10. P. platygnatha sp. nov., holotype; 11. P. ludingensis sp. nov., holotype; 12. P. mediacantha sp. nov., holotype; 13. P. nakhonnayokensis; 14. P. nepalensis; 15. P. rectignatha; 16. P. securiformis sp. nov., holotype; 17. P. solitaridens sp. nov., holotype; 18. P. subaequalis sp. nov., holotype. Scales = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10493951" httpUri="https://zenodo.org/record/10493951/files/figure.png" pageId="17" pageNumber="118">Figs 17</figureCitation>
,
<figureCitation id="C2E32A4BAC3FFFDD325593242834FE7E" box="[258,287,367,393]" captionStart="FIGURES 2932" captionStartId="16.[151,264,1132,1156]" captionTargetBox="[171,1414,194,1096]" captionTargetId="figure-22@16.[159,1427,181,1108]" captionTargetPageId="16" captionText="FIGURES 2932. Male genitalia of Periacma spp.29.P.nepalensis, slide No. XZX20769; 30. P. securiformis sp. nov., holotype, slide No. LiSR16146; 31. P. solitaridens sp. nov., holotype, slide No. XZX20615; 32. P. subaequalis sp. nov., holotype, slide No. XZX20520. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10493957" httpUri="https://zenodo.org/record/10493957/files/figure.png" pageId="17" pageNumber="118">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="12C26545AC3FFFDD33C093FC2D08FE02" pageId="17" pageNumber="118" type="materials_examined">
<paragraph id="5A6736CEAC3FFFDD33C093FC2D08FE02" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD33C093FC286DFE27" bold="true" box="[151,326,438,465]" pageId="17" pageNumber="118">Type material.</emphasis>
<materialsCitation id="EAB03C93AC3FFFDD321A93FD2B52FE02" collectingDate="2016-06-30" collectionCode="XZX" collectorName="KJ Teng &amp; XF Yang" country="China" elevation="1695" latitude="29.36" location="Hailuogou" longLatPrecision="4873" longitude="102.6" pageId="17" pageNumber="118" specimenCode="XZX20615" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="68ACEADCAC3FFFDD321A93FD2B08FE27" bold="true" box="[333,547,438,464]" pageId="17" pageNumber="118">
<collectingCountry id="22CF765EAC3FFFDD321A93FD2887FE27" box="[333,428,438,464]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
,
<collectingRegion id="981CF82CAC3FFFDD32EE93FD2B35FE27" box="[441,542,438,464]" country="China" name="Sichuan" pageId="17" pageNumber="118">Sichuan</collectingRegion>
:
</emphasis>
<typeStatus id="8563886CAC3FFFDD317C93FC2BB8FE26" box="[555,659,439,465]" pageId="17" pageNumber="118" type="holotype">Holotype</typeStatus>
♂,
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(
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,
<geoCoordinate id="3FEC5009AC3FFFDD30E793FC2D26FE26" box="[944,1037,439,465]" degrees="102.6" direction="east" orientation="longitude" pageId="17" pageNumber="118" precision="5555" value="102.6">102.6ºE</geoCoordinate>
),
<location id="5F076015AC3FFFDD377693FC2D5EFE26" LSID="urn:lsid:plazi:treatment:D27187D8AC3FFFDD33C09300287EF96E:5F076015AC3FFFDD377693FC2D5EFE26" box="[1057,1141,439,465]" country="China" latitude="29.36" longLatPrecision="4873" longitude="102.6" name="Luding" pageId="17" pageNumber="118" stateProvince="Sichuan">Luding</location>
,
<quantity id="9D209B2BAC3FFFDD37D693FD2DF3FE26" box="[1153,1240,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="17" pageNumber="118" unit="m" value="1695.0">
<elevation id="D1F5D1FDAC3FFFDD37D693FD2DF3FE26" box="[1153,1240,438,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6949999999999998" pageId="17" pageNumber="118" unit="m" value="1695.0">1695 m</elevation>
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,
<date id="2E66100EAC3FFFDD37B293FC2C4CFE26" box="[1253,1383,439,465]" pageId="17" pageNumber="118" value="2016-06-30">
<collectingDate id="3E22E9E6AC3FFFDD37B293FC2C4CFE26" box="[1253,1383,439,465]" pageId="17" pageNumber="118" value="2016-06-30">30.VI.2016</collectingDate>
</date>
, leg.
<collectorName id="F72D5318AC3FFFDD33C0939029DEFE02" box="[151,245,475,501]" pageId="17" pageNumber="118">KJ Teng</collectorName>
&amp;
<collectorName id="F72D5318AC3FFFDD324F93902854FE02" box="[280,383,475,501]" pageId="17" pageNumber="118">XF Yang</collectorName>
, slide No.
<specimenCode id="0A7E9EB5AC3FFFDD32A193912B52FE02" box="[502,633,474,501]" pageId="17" pageNumber="118">XZX20615</specimenCode>
</materialsCitation>
.
<materialsCitation id="EAB03C93AC3FFFDD31D493902D0BFE02" box="[643,1056,475,503]" collectingDate="2016-06-30" collectionCode="XZX" collectorName="KJ Teng &amp; XF Yang" country="China" elevation="1695" latitude="29.36" location="Hailuogou" longLatPrecision="4873" longitude="102.6" pageId="17" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="8563886CAC3FFFDD31D493902BC2FE02" box="[643,745,475,501]" pageId="17" pageNumber="118" type="paratype">Paratype</typeStatus>
:
<specimenCount id="4CDEFD47AC3FFFDD31A393972A3CFE00" box="[756,791,476,503]" count="1" pageId="17" pageNumber="118" type="male">1♂</specimenCount>
, sama data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="12C26545AC3FFFDD339093B52AFCFCE2" pageId="17" pageNumber="118" type="diagnosis">
<paragraph id="5A6736CEAC3FFFDD339093B52AFCFCE2" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD339093B52869FDEF" bold="true" box="[199,322,510,536]" pageId="17" pageNumber="118">Diagnosis.</emphasis>
The new species is similar to
<taxonomicName id="9DD84D4DAC3FFFDD31FE93B42D15FDEE" authority="Wang &amp; Li, 2002" authorityName="Wang &amp; Li" authorityYear="2002" box="[681,1086,511,537]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="kangdingensis">
<emphasis id="68ACEADCAC3FFFDD31FE93B42A43FDEE" box="[681,872,511,537]" italics="true" pageId="17" pageNumber="118">P. kangdingensis</emphasis>
<bibRefCitation id="3E494B3FAC3FFFDD302593B42D15FDEE" author="Wang, S. X. &amp; Li, H. H." box="[882,1086,511,537]" pageId="17" pageNumber="118" pagination="565 - 571" refId="ref17786" refString="Wang, S. X. &amp; Li, H. H. (2002) Four newspecies and two newrecords of the genus Periacma Meyrick from China (Lepidoptera: Oecophoridae). Acta Zootaxonomica Sinica, 27 (3), 565 - 571." type="journal article" year="2002">
Wang
<emphasis id="68ACEADCAC3FFFDD30EB93B42AF9FDEE" box="[956,978,511,537]" italics="true" pageId="17" pageNumber="118">&amp;</emphasis>
Li, 2002
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="9DD84D4DAC3FFFDD372C93B4280EFDCA" authority="Wang &amp; Zheng, 1995" authorityName="Wang &amp; Zheng" authorityYear="1995" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="fengxianensis">
<emphasis id="68ACEADCAC3FFFDD372C93B42C1AFDEE" box="[1147,1329,511,537]" italics="true" pageId="17" pageNumber="118">P. fengxianensis</emphasis>
<bibRefCitation id="3E494B3FAC3FFFDD366B93B4280EFDCA" author="Wang, S. X. &amp; Zheng, Z. M." pageId="17" pageNumber="118" pagination="5 - 14" refId="ref17995" refString="Wang, S. X. &amp; Zheng, Z. M. (1995) Five new species and one new record of the genus Periacma Meyrick from China (Lepidoptera: Oecophoridae). Acta Agriculturae Boreali Occidentalis Sinica, 4 (1), 5 - 14." type="journal article" year="1995">
Wang
<emphasis id="68ACEADCAC3FFFDD36D193B42CB7FDEE" box="[1414,1436,511,537]" italics="true" pageId="17" pageNumber="118">&amp;</emphasis>
Zheng,
<quantity id="9D209B2BAC3FFFDD33BA9069280EFDCA" box="[237,293,546,573]" metricMagnitude="1" metricUnit="m" metricValue="5.0673" pageId="17" pageNumber="118" unit="in" value="1995.0">1995</quantity>
</bibRefCitation>
</taxonomicName>
in male genitalia. It can be distinguished from
<taxonomicName id="9DD84D4DAC3FFFDD306F90682ADFFDCA" authorityName="Wang &amp; Li" authorityYear="2002" box="[824,1012,547,573]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="kangdingensis">
<emphasis id="68ACEADCAC3FFFDD306F90682ADFFDCA" box="[824,1012,547,573]" italics="true" pageId="17" pageNumber="118">P. kangdingensis</emphasis>
</taxonomicName>
by the uncus distinctly widened from basal 1/3 to a broadly rounded apex, the costal part of the valva elliptical distally, and the sacculus half as long as the costal part of the valva; in
<taxonomicName id="9DD84D4DAC3FFFDD329390202BABFD72" authorityName="Wang &amp; Li" authorityYear="2002" box="[452,640,619,645]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="kangdingensis">
<emphasis id="68ACEADCAC3FFFDD329390202BABFD72" box="[452,640,619,645]" italics="true" pageId="17" pageNumber="118">P. kangdingensis</emphasis>
</taxonomicName>
, the uncus is slightly widened to the obtuse apex, the costal part of the valva is subparallel distally, and the sacculus is longer than half the length of the costal part of the valva. The new species can be distinguished from
<taxonomicName id="9DD84D4DAC3FFFDD314B90F82BFBFD3A" authorityName="Wang &amp; Zheng" authorityYear="1995" box="[540,720,691,717]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="fengxianensis">
<emphasis id="68ACEADCAC3FFFDD314B90F82BFBFD3A" box="[540,720,691,717]" italics="true" pageId="17" pageNumber="118">P. fengxianensis</emphasis>
</taxonomicName>
by the basal width of the uncus less than 1/2 of its widest part, and the aedeagus with a preapical denticle on one side; in
<taxonomicName id="9DD84D4DAC3FFFDD304F909C2AE0FD06" authorityName="Wang &amp; Zheng" authorityYear="1995" box="[792,971,727,753]" class="Insecta" family="Autostichidae" genus="Periacma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="species" species="fengxianensis">
<emphasis id="68ACEADCAC3FFFDD304F909C2AE0FD06" box="[792,971,727,753]" italics="true" pageId="17" pageNumber="118">P. fengxianensis</emphasis>
</taxonomicName>
, the basal width of the uncus is more than 1/2 of its widest part, and the aedeagus has several denticles laterodistally.
</paragraph>
</subSubSection>
<subSubSection id="12C26545AC3FFFDD3390915528B9F9DA" pageId="17" pageNumber="118" type="description">
<paragraph id="5A6736CEAC3FFFDD339091552BD3FCCE" blockId="17.[151,1437,438,1689]" box="[199,760,798,825]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD339091552872FCCF" bold="true" box="[199,345,798,824]" pageId="17" pageNumber="118">Description.</emphasis>
Adult (
<figureCitation id="C2E32A4BAC3FFFDD32F891542B29FCCE" box="[431,514,799,825]" captionStart="FIGURES 1018" captionStartId="4.[151,264,1816,1840]" captionTargetBox="[170,1417,194,1780]" captionTargetId="figure-22@4.[158,1430,181,1793]" captionTargetPageId="4" captionText="FIGURES 1018. Adults of Periacma spp. 10. P. platygnatha sp. nov., holotype; 11. P. ludingensis sp. nov., holotype; 12. P. mediacantha sp. nov., holotype; 13. P. nakhonnayokensis; 14. P. nepalensis; 15. P. rectignatha; 16. P. securiformis sp. nov., holotype; 17. P. solitaridens sp. nov., holotype; 18. P. subaequalis sp. nov., holotype. Scales = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10493951" httpUri="https://zenodo.org/record/10493951/files/figure.png" pageId="17" pageNumber="118">Fig. 17</figureCitation>
). Wingspan 16.0 mm.
</paragraph>
<paragraph id="5A6736CEAC3FFFDD339091082B86FC52" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD33909108282FFCAA" box="[199,260,835,861]" italics="true" pageId="17" pageNumber="118">Head</emphasis>
. Vertex and frons yellow. Labial palpus yellow, second segment with distal 2/3 black on outer surface. Antenna with scape white on ventral surface, dark brown on dorsal surface; flagellum yellow on ventral surface, black with fine yellow annuli on dorsal surface.
</paragraph>
<paragraph id="5A6736CEAC3FFFDD33B391E42AF8FB32" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD33B391E42816FC3E" box="[228,317,943,969]" italics="true" pageId="17" pageNumber="118">Thorax.</emphasis>
Mesonotum dark brown, surrounded by yellow scales; tegula yellow, with dense dark brown scales basally. Forewing yellow, with brown scales denser distally and between fold and dorsum; costal margin dark brown in basal 1/6; dorsum with a large dark brown spot near base; dark brown speckle along above fold to middle of cell; discal spot dark brown, above preceding speckle; postmedial fascia dark brown, extending from about costal 2/3 to tornus, narrower medially; fringe yellow. Hindwing and fringe grey. Legs yellow, except three distal tarsomeres of fore- and midlegs black; on dorsal surface, foreleg with femur black, midleg with black scales on femur, tibiae of fore- and midlegs black, tarsi black at middle of first tarsomere and at base of second tarsomere; on outer surface, hindleg with tarsus black at base of each tarsomere except first tarsomere.
</paragraph>
<paragraph id="5A6736CEAC3FFFDD339096842B5CF9FE" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD339096842845FB1E" box="[199,366,1231,1257]" italics="true" pageId="17" pageNumber="118">Male genitalia</emphasis>
(
<figureCitation id="C2E32A4BAC3FFFDD3229968428FFFB1E" box="[382,468,1231,1257]" captionStart="FIGURES 2932" captionStartId="16.[151,264,1132,1156]" captionTargetBox="[171,1414,194,1096]" captionTargetId="figure-22@16.[159,1427,181,1108]" captionTargetPageId="16" captionText="FIGURES 2932. Male genitalia of Periacma spp.29.P.nepalensis, slide No. XZX20769; 30. P. securiformis sp. nov., holotype, slide No. LiSR16146; 31. P. solitaridens sp. nov., holotype, slide No. XZX20615; 32. P. subaequalis sp. nov., holotype, slide No. XZX20520. Scales = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10493957" httpUri="https://zenodo.org/record/10493957/files/figure.png" pageId="17" pageNumber="118">Fig. 31</figureCitation>
). Uncus narrow and parallel sided in basal 1/3, with long setae at basal 1/6 laterally, inflated from basal 1/3 to broadly rounded apex. Gnathos with basal arm gradually widened; ventral plate narrow and parallel sided posteriorly, widened and spinulous anteriorly, narrowed outward to obtuse end anterolaterally, slightly concave at middle on anterior margin. Valva with costal part uniformly wide from base to basal 2/5, distal 3/5 inflated elliptically, obtusely rounded at apex; costa straight basally, arched distally; dorso-proximal process dilated distally, setose; sacculus about half length of costal part of valva, wide at base, narrowed to middle length, sharply produced from middle to a slender, apex-pointed process bent inward distally. Saccus short, trapezoidal. Aedeagus about half length of costal part of valva, gradually widened from base to 1/2, then tapered to before apex; apex widened, with a denticle before apex.
</paragraph>
<paragraph id="5A6736CEAC3FFFDD3390945828B9F9DA" blockId="17.[151,1437,438,1689]" box="[199,402,1555,1581]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD339094582830F9DA" box="[199,283,1555,1581]" italics="true" pageId="17" pageNumber="118">Female</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="12C26545AC3FFFDD3390947D2B0DF9A6" box="[199,550,1590,1617]" pageId="17" pageNumber="118" type="distribution">
<paragraph id="5A6736CEAC3FFFDD3390947D2B0DF9A6" blockId="17.[151,1437,438,1689]" box="[199,550,1590,1617]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD3390947D284AF9A7" bold="true" box="[199,353,1590,1616]" pageId="17" pageNumber="118">Distribution.</emphasis>
<collectingCountry id="22CF765EAC3FFFDD3230947C2881F9A6" box="[359,426,1591,1617]" name="China" pageId="17" pageNumber="118">China</collectingCountry>
(
<collectingRegion id="981CF82CAC3FFFDD32EE947C2B31F9A6" box="[441,538,1591,1617]" country="China" name="Sichuan" pageId="17" pageNumber="118">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="12C26545AC3FFFDD33909410287EF96E" pageId="17" pageNumber="118" type="etymology">
<paragraph id="5A6736CEAC3FFFDD33909410287EF96E" blockId="17.[151,1437,438,1689]" pageId="17" pageNumber="118">
<emphasis id="68ACEADCAC3FFFDD339094102865F982" bold="true" box="[199,334,1627,1653]" pageId="17" pageNumber="118">Etymology.</emphasis>
The specific epithet is derived from the Latin
<emphasis id="68ACEADCAC3FFFDD300C94102AEEF982" box="[859,965,1627,1653]" italics="true" pageId="17" pageNumber="118">solitarius</emphasis>
and
<emphasis id="68ACEADCAC3FFFDD30AB94102D16F982" box="[1020,1085,1627,1653]" italics="true" pageId="17" pageNumber="118">dens</emphasis>
, referring to the aedeagus with a single denticle.
</paragraph>
</subSubSection>
</treatment>
</document>