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<mods:title id="49146DE469D1745106FCB3A494D9A9E2">Redescription Cyrtodactylus lateralis (Werner) (Squamata: Gekkonidae) and Phylogeny of the Prehensile-tailed Cyrtodactylus</mods:title>
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<paragraph id="3934CB59FFD6FC715BD4FA32C70E15E2" blockId="10.[151,1110,1484,1510]" box="[151,1110,1484,1510]" pageId="10" pageNumber="527">
<heading id="627C7C35FFD6FC715BD4FA32C70E15E2" bold="true" box="[151,1110,1484,1510]" fontSize="11" level="1" pageId="10" pageNumber="527" reason="1">
<emphasis id="0BFF174BFFD6FC715BD4FA32C70E15E2" bold="true" box="[151,1110,1484,1510]" pageId="10" pageNumber="527">
Phylogeny of
<taxonomicName id="FE8BB0DAFFD6FC715A03FA32C11715E2" box="[320,591,1484,1510]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD6FC715A03FA32C11715E2" bold="true" box="[320,591,1484,1510]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus lateralis</emphasis>
</taxonomicName>
and other Prehensile-tailed
<taxonomicName id="FE8BB0DAFFD6FC7158F3FA32C70E15E2" box="[944,1110,1484,1510]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD6FC7158F3FA32C70E15E2" bold="true" box="[944,1110,1484,1510]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus</emphasis>
</taxonomicName>
</emphasis>
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<paragraph id="3934CB59FFD6FC715BD4F9ECC0A9174F" blockId="10.[151,1436,1554,2011]" pageId="10" pageNumber="527">
Currently, the
<emphasis id="0BFF174BFFD6FC715A07F9EDC155162F" box="[324,525,1555,1579]" italics="true" pageId="10" pageNumber="527">Reptile Database</emphasis>
lists 206 species of
<taxonomicName id="FE8BB0DAFFD6FC71584CF9EDC79B162F" authority="Uetz &amp; Hosek 2015" authorityName="Uetz &amp; Hosek" authorityYear="2015" box="[783,1219,1554,1579]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD6FC71584CF9EDC0F4162F" box="[783,940,1555,1579]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus</emphasis>
(
<bibRefCitation id="5D1AB6A8FFD6FC715881F9EDC7E3162F" author="Uetz" box="[962,1211,1554,1579]" pageId="10" pageNumber="527" refString="Uetz, P. &amp; Hosek, J. (Eds.) (2015) The Reptile Database. Available from: http: // www. reptile-database. org (accessed 1 October 2015)" type="journal article" year="2015">Uetz &amp; Hošek 2015</bibRefCitation>
)
</taxonomicName>
. Systematists face considerable challenges when trying to identify derived morphological characters in this diverse gekkonid radiation. On the basis of morphology,
<bibRefCitation id="5D1AB6A8FFD6FC715911F9A4C0191677" box="[594,833,1626,1651]" pageId="10" pageNumber="527" refString="Grismer, L. L., Anuar, S., Quah, E., Muin, M. A., Onn, C. K., Grismer, J. L. &amp; Ahmad, N. (2010) A new spiny, prehensile-tailed species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia with a preliminary hypothesis of relationships based on morphology. Zootaxa, 2625, 40 - 52." type="journal article">
Grismer
<emphasis id="0BFF174BFFD6FC7159F5F9A5C1B61677" box="[694,750,1627,1651]" italics="true" pageId="10" pageNumber="527">et al.</emphasis>
(2010)
</bibRefCitation>
proposed a phylogeny of
<taxonomicName id="FE8BB0DAFFD6FC715F2AF9A5C65E1677" box="[1129,1286,1627,1651]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD6FC715F2AF9A5C65E1677" box="[1129,1286,1627,1651]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus</emphasis>
</taxonomicName>
with spinose prehensile tails. These authors constructed their tree using parsimony and polarized a small matrix of characters by treating all other congeners as outgroups. In their phylogeny,
<taxonomicName id="FE8BB0DAFFD6FC715800F95DC7DE16BF" box="[835,1158,1699,1723]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="0BFF174BFFD6FC715800F95DC7DE16BF" box="[835,1158,1699,1723]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus brevipalmatus</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD6FC715FF8F95DC65316BF" box="[1211,1291,1699,1723]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD6FC715FF8F95DC65316BF" box="[1211,1291,1699,1723]" italics="true" pageId="10" pageNumber="527">C. elok</emphasis>
</taxonomicName>
form a clade and share digital webbing, a square tail, and a caudal fringe.
<emphasis id="0BFF174BFFD6FC715807F938C65E16DA" box="[836,1286,1734,1758]" italics="true" pageId="10" pageNumber="527">
<taxonomicName id="FE8BB0DAFFD6FC715807F938C71416DA" box="[836,1100,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="spinosus">Cyrtodactylus spinosus</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD6FC715F1BF939C65A16DA" box="[1112,1282,1735,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="stresemanni">C. stresemanni</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FE8BB0DAFFD6FC715E7DF939C6C316DA" box="[1342,1435,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD6FC715E7DF939C6C316DA" box="[1342,1435,1734,1758]" italics="true" pageId="10" pageNumber="527">C. durio</emphasis>
</taxonomicName>
form a second clade and share spinose tubercles in the ventrolateral fold on the body. Within the second clade,
<taxonomicName id="FE8BB0DAFFD6FC715BD4F8F0C2211722" box="[151,377,1806,1830]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD6FC715BD4F8F0C2211722" box="[151,377,1806,1830]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD6FC715AF7F8F1C1041722" box="[436,604,1807,1830]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD6FC715AF7F8F1C1041722" box="[436,604,1807,1830]" italics="true" pageId="10" pageNumber="527">C. stresemanni</emphasis>
</taxonomicName>
are sister species and share enlarged, ventrolateral caudal tubercles. The authors argued that a prehensile tail is a synapomorphy of these five species.
</paragraph>
<paragraph id="3934CB59FFD6FC775B84F8A8C25A11D4" blockId="10.[151,1436,1554,2011]" lastBlockId="12.[151,1437,151,464]" lastPageId="12" lastPageNumber="529" pageId="10" pageNumber="527">
Using their methods (
<tableCitation id="7409FEE2FFD6FC715A80F8A8C143176B" box="[451,539,1878,1903]" captionStart="TABLE 1" captionStartId="12.[151,236,512,535]" captionTargetBox="[159,1347,690,1115]" captionTargetPageId="12" captionText="TABLE 1. Morphological matrix used to determine phylogeny of prehensile-tailed Cyrtodactylus. Characters are 1. Tail prehensile (1) or not (0); 2. Digits webbed (1) or not (0); 3. Ventrolateral caudal fringe present (1) or not (0); 4. Tail square (1) or round (0) in cross section; 5. Ventrolateral body fold with (1) or without (0) spinose tubercles; 6. Tail with ventrolateral caudal tubercles (1) or not (0); 7. Dorsum banded (B) or with pale vertebral stripe and 2 3 broken or complete chevrons (S), character unordered." httpUri="http://table.plazi.org/id/6DF49BD1FFD0FC775BD4FDFEC1281293" pageId="10" pageNumber="527" tableUuid="6DF49BD1FFD0FC775BD4FDFEC1281293">Table 1</tableCitation>
), we add
<taxonomicName id="FE8BB0DAFFD6FC7159D6F8A8C754176B" authority="Werner" authorityName="Werner" box="[661,1036,1878,1903]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD6FC7159D6F8A8C0C2176A" box="[661,922,1878,1902]" italics="true" pageId="10" pageNumber="527">Cyrtodactylus lateralis</emphasis>
(Werner)
</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD6FC715F58F8A9C642176B" authority="Ulber" authorityName="Ulber" box="[1051,1306,1878,1903]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD6FC715F58F8A9C797176A" box="[1051,1231,1878,1902]" italics="true" pageId="10" pageNumber="527">C. interdigitalis</emphasis>
Ulber
</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD6FC715E69F8A9C2741797" authority="Oliver et al." authorityName="Oliver et al." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD6FC715E69F8A9C6C4176A" box="[1322,1436,1878,1902]" italics="true" pageId="10" pageNumber="527">C. nuaulu</emphasis>
Oliver
<emphasis id="0BFF174BFFD6FC715BAEF885C2741797" box="[237,300,1915,1939]" italics="true" pageId="10" pageNumber="527">et al.</emphasis>
</taxonomicName>
, and
<taxonomicName id="FE8BB0DAFFD6FC715A3BF885C1161796" authority="Kraus" authorityName="Kraus" box="[376,590,1915,1938]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD6FC715A3BF885C2A51796" box="[376,509,1915,1938]" italics="true" pageId="10" pageNumber="527">C. serratus</emphasis>
Kraus
</taxonomicName>
to this morphological phylogeny (
<figureCitation id="A1B0D7DCFFD6FC7158B5F884C71A1797" box="[1014,1090,1914,1939]" captionStart="FIGURE 6" captionStartId="11.[151,249,1913,1935]" captionTargetBox="[221,1338,212,1853]" captionTargetId="figure@11.[206,1355,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Phylogenetic relationships of Cyrtodactylus with spinose, prehensile tails. (A) Phylogeny based on morphology with three species added to original hypothesis of Grismer et al. (2010). (B) Clade pruned from our larger phylogeny based on sequences of the NADH Dehydrogenase 2 gene (ND 2). Tree constructed using Bayesian inference. Nodal values are posterior probabilities, and diamonds indicate prehensile-tailed Cyrtodactylus." httpUri="https://zenodo.org/record/256584/files/figure.png" pageId="10" pageNumber="527">Fig. 6</figureCitation>
).
<bibRefCitation id="5D1AB6A8FFD6FC715F1DF884C7AF1797" author="Ulber" box="[1118,1271,1914,1939]" pageId="10" pageNumber="527" refString="Ulber, T. (1993) Bemerkungen uber cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, 69, 187 - 200. http: // dx. doi. org / 10.1002 / mmnz. 19930690202" type="journal article" year="1993">Ulber (1993)</bibRefCitation>
compared
<taxonomicName id="FE8BB0DAFFD6FC715EC1F885C27F17B2" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD6FC715EC1F885C27F17B2" italics="true" pageId="10" pageNumber="527">C. interdigitalis</emphasis>
</taxonomicName>
to
<taxonomicName id="FE8BB0DAFFD6FC715A08F861C15217B2" box="[331,522,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="0BFF174BFFD6FC715A08F861C15217B2" box="[331,522,1950,1974]" italics="true" pageId="10" pageNumber="527">C. brevipalmatus</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD6FC715902F861C1C917B2" box="[577,657,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD6FC715902F861C1C917B2" box="[577,657,1950,1974]" italics="true" pageId="10" pageNumber="527">C. elok</emphasis>
</taxonomicName>
and noted that, unlike these two species,
<taxonomicName id="FE8BB0DAFFD6FC715F20F861C64B17B2" box="[1123,1299,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="527" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD6FC715F20F861C64B17B2" box="[1123,1299,1950,1974]" italics="true" pageId="10" pageNumber="527">C. interdigitalis</emphasis>
</taxonomicName>
has a round rather than square tail. All three species share digital webbing and a ventrolateral fringe. The ventrolateral fringe consists of a distinctive row of enlarged conical tubercles separated by smaller lanceolate tubercles. The enlarged tubercles are the ventrolateral components of caudal whorls, whereas the smaller conical tubercles are the only enlarged scales separating the whorls dorsally. All three species lack spinose tubercles in the ventrolateral fold on the body.
<bibRefCitation id="5D1AB6A8FFD0FC775A46FEFAC2CF1118" author="Ulber" box="[261,407,260,285]" pageId="12" pageNumber="529" refString="Ulber, T. (1993) Bemerkungen uber cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, 69, 187 - 200. http: // dx. doi. org / 10.1002 / mmnz. 19930690202" type="journal article" year="1993">Ulber (1993)</bibRefCitation>
did not mention whether
<taxonomicName id="FE8BB0DAFFD0FC7759FAFEFAC0311118" box="[697,873,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD0FC7759FAFEFAC0311118" box="[697,873,260,284]" italics="true" pageId="12" pageNumber="529">C. interdigitalis</emphasis>
</taxonomicName>
coils its tail at rest, however we have seen photos of specimens from
<collectingCountry id="419C8BC9FFD0FC775A2EFED6C2FB113B" box="[365,419,296,319]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
exhibiting this behavior. Based on this combination of traits, we add
<taxonomicName id="FE8BB0DAFFD0FC775FE4FED6C60E1144" box="[1191,1366,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD0FC775FE4FED6C60E1144" box="[1191,1366,296,320]" italics="true" pageId="12" pageNumber="529">C. interdigitalis</emphasis>
</taxonomicName>
to the morphological phylogeny of
<bibRefCitation id="5D1AB6A8FFD0FC775AAFFEB2C1B01160" box="[492,744,332,357]" pageId="12" pageNumber="529" refString="Grismer, L. L., Anuar, S., Quah, E., Muin, M. A., Onn, C. K., Grismer, J. L. &amp; Ahmad, N. (2010) A new spiny, prehensile-tailed species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia with a preliminary hypothesis of relationships based on morphology. Zootaxa, 2625, 40 - 52." type="journal article">
Grismer
<emphasis id="0BFF174BFFD0FC775917FEB3C1C91160" box="[596,657,332,356]" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
(2010)
</bibRefCitation>
and place
<taxonomicName id="FE8BB0DAFFD0FC77582CFEB2C77D1160" box="[879,1061,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD0FC77582CFEB2C77D1160" box="[879,1061,332,356]" italics="true" pageId="12" pageNumber="529">C. interdigitalis</emphasis>
</taxonomicName>
as the sister species to a clade containing
<taxonomicName id="FE8BB0DAFFD0FC775A57FE8EC23B118C" box="[276,355,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD0FC775A57FE8EC23B118C" box="[276,355,368,392]" italics="true" pageId="12" pageNumber="529">C. elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD0FC775ADAFE8EC100118C" box="[409,600,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="0BFF174BFFD0FC775ADAFE8EC100118C" box="[409,600,368,392]" italics="true" pageId="12" pageNumber="529">C. brevipalmatus</emphasis>
</taxonomicName>
(
<figureCitation id="A1B0D7DCFFD0FC77592BFE91C1F4118C" box="[616,684,367,392]" captionStart="FIGURE 6" captionStartId="11.[151,249,1913,1935]" captionTargetBox="[221,1338,212,1853]" captionTargetId="figure@11.[206,1355,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Phylogenetic relationships of Cyrtodactylus with spinose, prehensile tails. (A) Phylogeny based on morphology with three species added to original hypothesis of Grismer et al. (2010). (B) Clade pruned from our larger phylogeny based on sequences of the NADH Dehydrogenase 2 gene (ND 2). Tree constructed using Bayesian inference. Nodal values are posterior probabilities, and diamonds indicate prehensile-tailed Cyrtodactylus." httpUri="https://zenodo.org/record/256584/files/figure.png" pageId="12" pageNumber="529">Fig. 6</figureCitation>
). We refer to this group of three species as the “mainland SE Asian clade” of prehensile-tailed
<taxonomicName id="FE8BB0DAFFD0FC775A87FE6AC13911A8" box="[452,609,404,428]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD0FC775A87FE6AC13911A8" box="[452,609,404,428]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus</emphasis>
</taxonomicName>
, because these species occur in
<collectingCountry id="419C8BC9FFD0FC775885FE6BC0A711A8" box="[966,1023,405,428]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingCountry id="419C8BC9FFD0FC775F49FE6BC72111A9" box="[1034,1145,405,429]" name="Myanmar" pageId="12" pageNumber="529">Myanmar</collectingCountry>
, Peninsular
<collectingCountry id="419C8BC9FFD0FC775E43FE6AC63211A9" box="[1280,1386,404,429]" name="Malaysia" pageId="12" pageNumber="529">Malaysia</collectingCountry>
, and
<collectingCountry id="419C8BC9FFD0FC775BD4FE49C3A611D4" box="[151,254,439,464]" name="Thailand" pageId="12" pageNumber="529">Thailand</collectingCountry>
.
</paragraph>
<caption id="6DF49BD1FFD7FC705BD4F887C01717F7" httpUri="https://zenodo.org/record/256584/files/figure.png" pageId="11" pageNumber="528" targetBox="[221,1338,212,1853]" targetPageId="11">
<paragraph id="3934CB59FFD7FC705BD4F887C01717F7" blockId="11.[151,1436,1913,2035]" pageId="11" pageNumber="528">
<emphasis id="0BFF174BFFD7FC705BD4F887C24B178A" bold="true" box="[151,275,1913,1935]" pageId="11" pageNumber="528">FIGURE 6.</emphasis>
Phylogenetic relationships of
<taxonomicName id="FE8BB0DAFFD7FC70590CF887C185178B" box="[591,733,1913,1935]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="528" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD7FC70590CF887C185178B" box="[591,733,1913,1935]" italics="true" pageId="11" pageNumber="528">Cyrtodactylus</emphasis>
</taxonomicName>
with spinose, prehensile tails. (A) Phylogeny based on morphology with three species added to original hypothesis of Grismer
<emphasis id="0BFF174BFFD7FC7059ADF862C07A17B5" box="[750,802,1947,1969]" italics="true" pageId="11" pageNumber="528">et al.</emphasis>
(2010). (B) Clade pruned from our larger phylogeny based on sequences of the NADH Dehydrogenase 2 gene (ND2). Tree constructed using Bayesian inference. Nodal values are posterior probabilities, and diamonds indicate prehensile-tailed
<taxonomicName id="FE8BB0DAFFD7FC7059F8F823C01117F7" box="[699,841,2013,2035]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="528" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD7FC7059F8F823C01117F7" box="[699,841,2013,2035]" italics="true" pageId="11" pageNumber="528">Cyrtodactylus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="6DF49BD1FFD0FC775BD4FDFEC1281293" ID-Table-UUID="6DF49BD1FFD0FC775BD4FDFEC1281293" httpUri="http://table.plazi.org/id/6DF49BD1FFD0FC775BD4FDFEC1281293" pageId="12" pageNumber="529" targetBox="[159,1347,690,1115]" targetIsTable="true" targetPageId="12">
<paragraph id="3934CB59FFD0FC775BD4FDFEC1281293" blockId="12.[151,1436,511,663]" pageId="12" pageNumber="529">
<emphasis id="0BFF174BFFD0FC775BD4FDFEC2591213" bold="true" box="[151,257,512,535]" pageId="12" pageNumber="529">TABLE 1</emphasis>
. Morphological matrix used to determine phylogeny of prehensile-tailed
<taxonomicName id="FE8BB0DAFFD0FC775F57FE01C7F11213" box="[1044,1193,511,535]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD0FC775F57FE01C7F11213" box="[1044,1193,511,535]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus</emphasis>
</taxonomicName>
. Characters are 1. Tail prehensile (1) or not (0); 2. Digits webbed (1) or not (0); 3. Ventrolateral caudal fringe present (1) or not (0); 4. Tail square (1) or round (0) in cross section; 5. Ventrolateral body fold with (1) or without (0) spinose tubercles; 6. Tail with ventrolateral caudal tubercles (1) or not (0); 7. Dorsum banded (B) or with pale vertebral stripe and 23 broken or complete chevrons (S), character unordered.
</paragraph>
</caption>
<paragraph id="3934CB59FFD0FC775986FD4CC61B145F" pageId="12" pageNumber="529">
<table id="4B8B39F9FFD003845BDCFD4CC61B145F" box="[159,1347,690,1115]" gridcols="8" gridrows="10" pageId="12" pageNumber="529">
<tr id="87BBC91BFFD003845BDCFD4CC61B12CC" box="[159,1347,690,712]" gridrow="0" pageId="12" pageNumber="529" rowspan-0="1">
<th id="C46AA067FFD003845986FD4CC18A12CC" box="[709,722,690,712]" gridcol="1" gridrow="0" pageId="12" pageNumber="529">1</th>
<th id="C46AA067FFD00384586FFD4CC06112CC" box="[812,825,690,712]" gridcol="2" gridrow="0" pageId="12" pageNumber="529">2</th>
<th id="C46AA067FFD0038458D7FD4CC0F912CC" box="[916,929,690,712]" gridcol="3" gridrow="0" pageId="12" pageNumber="529">3</th>
<th id="C46AA067FFD0038458B8FD4CC75012CC" box="[1019,1032,690,712]" gridcol="4" gridrow="0" pageId="12" pageNumber="529">4</th>
<th id="C46AA067FFD003845F20FD4CC72812CC" box="[1123,1136,690,712]" gridcol="5" gridrow="0" pageId="12" pageNumber="529">5</th>
<th id="C46AA067FFD003845F89FD4CC78F12CC" box="[1226,1239,690,712]" gridcol="6" gridrow="0" pageId="12" pageNumber="529">6</th>
<th id="C46AA067FFD003845E71FD4CC61B12CC" box="[1330,1347,690,712]" gridcol="7" gridrow="0" pageId="12" pageNumber="529">7</th>
</tr>
<tr id="87BBC91BFFD003845BDCFD04C61B1314" box="[159,1347,762,784]" gridrow="1" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFD04C2141314" box="[159,332,762,784]" gridcol="0" gridrow="1" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFD05C2661314" box="[159,318,762,784]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD0FC775BDCFD05C2661314" box="[159,318,762,784]" italics="true" pageId="12" pageNumber="529">C. interdigitalis</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FD04C18A1314" box="[709,722,762,784]" gridcol="1" gridrow="1" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFD04C0611314" box="[812,825,762,784]" gridcol="2" gridrow="1" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458D7FD04C0F91314" box="[916,929,762,784]" gridcol="3" gridrow="1" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458B8FD04C7501314" box="[1019,1032,762,784]" gridcol="4" gridrow="1" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FD04C7281314" box="[1123,1136,762,784]" gridcol="5" gridrow="1" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F89FD04C78F1314" box="[1226,1239,762,784]" gridcol="6" gridrow="1" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845E71FD04C61B1314" box="[1330,1347,762,784]" gridcol="7" gridrow="1" pageId="12" pageNumber="529">B</td>
</tr>
<tr id="87BBC91BFFD003845BDCFCDAC61B133E" box="[159,1347,804,826]" gridrow="2" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFCDAC214133E" box="[159,332,804,826]" gridcol="0" gridrow="2" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFCDAC3BE133E" box="[159,230,804,826]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD0FC775BDCFCDAC3BE133E" box="[159,230,804,826]" italics="true" pageId="12" pageNumber="529">C. elok</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FCDAC18A133E" box="[709,722,804,826]" gridcol="1" gridrow="2" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFCDAC061133E" box="[812,825,804,826]" gridcol="2" gridrow="2" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458D7FCDAC0F9133E" box="[916,929,804,826]" gridcol="3" gridrow="2" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458B8FCDAC750133E" box="[1019,1032,804,826]" gridcol="4" gridrow="2" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F20FCDAC728133E" box="[1123,1136,804,826]" gridcol="5" gridrow="2" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F89FCDAC78F133E" box="[1226,1239,804,826]" gridcol="6" gridrow="2" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845E71FCDAC61B133E" box="[1330,1347,804,826]" gridcol="7" gridrow="2" pageId="12" pageNumber="529">B</td>
</tr>
<tr id="87BBC91BFFD003845BDCFCB3C61B1367" box="[159,1347,845,867]" gridrow="3" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFCB3C2141367" box="[159,332,845,867]" gridcol="0" gridrow="3" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFCB0C2141367" box="[159,332,845,867]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="brevipalmatus">
<emphasis id="0BFF174BFFD0FC775BDCFCB0C2141367" box="[159,332,845,867]" italics="true" pageId="12" pageNumber="529">C. brevipalmatus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FCB3C18A1367" box="[709,722,845,867]" gridcol="1" gridrow="3" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFCB3C0611367" box="[812,825,845,867]" gridcol="2" gridrow="3" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458D7FCB3C0F91367" box="[916,929,845,867]" gridcol="3" gridrow="3" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD0038458B8FCB3C7501367" box="[1019,1032,845,867]" gridcol="4" gridrow="3" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F20FCB3C7281367" box="[1123,1136,845,867]" gridcol="5" gridrow="3" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F89FCB3C78F1367" box="[1226,1239,845,867]" gridcol="6" gridrow="3" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845E71FCB3C61B1367" box="[1330,1347,845,867]" gridcol="7" gridrow="3" pageId="12" pageNumber="529">B</td>
</tr>
<tr id="87BBC91BFFD003845BDCFC88C61B1389" box="[159,1347,886,909]" gridrow="4" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFC88C2141389" box="[159,332,886,909]" gridcol="0" gridrow="4" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFC89C24B1389" box="[159,275,887,909]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC775BDCFC89C24B1389" box="[159,275,887,909]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FC88C18A1389" box="[709,722,886,909]" gridcol="1" gridrow="4" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFC88C0611389" box="[812,825,886,909]" gridcol="2" gridrow="4" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FC88C0F91389" box="[916,929,886,909]" gridcol="3" gridrow="4" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FC88C7501389" box="[1019,1032,886,909]" gridcol="4" gridrow="4" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FC88C7281389" box="[1123,1136,886,909]" gridcol="5" gridrow="4" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FC88C78F1389" box="[1226,1239,886,909]" gridcol="6" gridrow="4" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845E71FC88C61B1389" box="[1330,1347,886,909]" gridcol="7" gridrow="4" pageId="12" pageNumber="529">S</td>
</tr>
<tr id="87BBC91BFFD003845BDCFC5EC61B13B2" box="[159,1347,928,950]" gridrow="5" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFC5EC21413B2" box="[159,332,928,950]" gridcol="0" gridrow="5" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFC5FC26D13B2" box="[159,309,929,950]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC775BDCFC5FC26D13B2" box="[159,309,929,950]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FC5EC18A13B2" box="[709,722,928,950]" gridcol="1" gridrow="5" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFC5EC06113B2" box="[812,825,928,950]" gridcol="2" gridrow="5" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FC5EC0F913B2" box="[916,929,928,950]" gridcol="3" gridrow="5" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FC5EC75013B2" box="[1019,1032,928,950]" gridcol="4" gridrow="5" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FC5EC72813B2" box="[1123,1136,928,950]" gridcol="5" gridrow="5" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FC5EC78F13B2" box="[1226,1239,928,950]" gridcol="6" gridrow="5" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845E71FC5EC61B13B2" box="[1330,1347,928,950]" gridcol="7" gridrow="5" pageId="12" pageNumber="529">S</td>
</tr>
<tr id="87BBC91BFFD003845BDCFC37C61B13DB" box="[159,1347,969,991]" gridrow="6" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFC37C21413DB" box="[159,332,969,991]" gridcol="0" gridrow="6" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFC34C25A13DB" box="[159,258,969,991]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC775BDCFC34C25A13DB" box="[159,258,969,991]" italics="true" pageId="12" pageNumber="529">C. nuaulu</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FC37C18A13DB" box="[709,722,969,991]" gridcol="1" gridrow="6" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFC37C06113DB" box="[812,825,969,991]" gridcol="2" gridrow="6" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FC37C0F913DB" box="[916,929,969,991]" gridcol="3" gridrow="6" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FC37C75013DB" box="[1019,1032,969,991]" gridcol="4" gridrow="6" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FC37C72813DB" box="[1123,1136,969,991]" gridcol="5" gridrow="6" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FC37C78F13DB" box="[1226,1239,969,991]" gridcol="6" gridrow="6" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845E71FC37C61B13DB" box="[1330,1347,969,991]" gridcol="7" gridrow="6" pageId="12" pageNumber="529">S</td>
</tr>
<tr id="87BBC91BFFD003845BDCFC0CC61B140C" box="[159,1347,1010,1032]" gridrow="7" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFC0CC214140C" box="[159,332,1010,1032]" gridcol="0" gridrow="7" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFC0DC257140C" box="[159,271,1011,1032]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD0FC775BDCFC0DC257140C" box="[159,271,1011,1032]" italics="true" pageId="12" pageNumber="529">C. serratus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FC0CC18A140C" box="[709,722,1010,1032]" gridcol="1" gridrow="7" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFC0CC061140C" box="[812,825,1010,1032]" gridcol="2" gridrow="7" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FC0CC0F9140C" box="[916,929,1010,1032]" gridcol="3" gridrow="7" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FC0CC750140C" box="[1019,1032,1010,1032]" gridcol="4" gridrow="7" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FC0CC728140C" box="[1123,1136,1010,1032]" gridcol="5" gridrow="7" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FC0CC78F140C" box="[1226,1239,1010,1032]" gridcol="6" gridrow="7" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845E71FC0CC61B140C" box="[1330,1347,1010,1032]" gridcol="7" gridrow="7" pageId="12" pageNumber="529">S</td>
</tr>
<tr id="87BBC91BFFD003845BDCFBE2C61B1436" box="[159,1347,1052,1074]" gridrow="8" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFBE2C2141436" box="[159,332,1052,1074]" gridcol="0" gridrow="8" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFBE2C2481436" box="[159,272,1052,1074]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD0FC775BDCFBE2C2481436" box="[159,272,1052,1074]" italics="true" pageId="12" pageNumber="529">C. lateralis</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FBE2C18A1436" box="[709,722,1052,1074]" gridcol="1" gridrow="8" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFBE2C0611436" box="[812,825,1052,1074]" gridcol="2" gridrow="8" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FBE2C0F91436" box="[916,929,1052,1074]" gridcol="3" gridrow="8" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FBE2C7501436" box="[1019,1032,1052,1074]" gridcol="4" gridrow="8" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FBE2C7281436" box="[1123,1136,1052,1074]" gridcol="5" gridrow="8" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FBE2C78F1436" box="[1226,1239,1052,1074]" gridcol="6" gridrow="8" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845E71FBE2C61B1436" box="[1330,1347,1052,1074]" gridcol="7" gridrow="8" pageId="12" pageNumber="529">B</td>
</tr>
<tr id="87BBC91BFFD003845BDCFBBBC61B145F" box="[159,1347,1093,1115]" gridrow="9" pageId="12" pageNumber="529">
<th id="C46AA067FFD003845BDCFBBBC214145F" box="[159,332,1093,1115]" gridcol="0" gridrow="9" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775BDCFBB8C3AB145F" box="[159,243,1093,1115]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC775BDCFBB8C3AB145F" box="[159,243,1093,1115]" italics="true" pageId="12" pageNumber="529">C. durio</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD003845986FBBBC18A145F" box="[709,722,1093,1115]" gridcol="1" gridrow="9" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD00384586FFBBBC061145F" box="[812,825,1093,1115]" gridcol="2" gridrow="9" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458D7FBBBC0F9145F" box="[916,929,1093,1115]" gridcol="3" gridrow="9" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD0038458B8FBBBC750145F" box="[1019,1032,1093,1115]" gridcol="4" gridrow="9" pageId="12" pageNumber="529">0</td>
<td id="C46AA067FFD003845F20FBBBC728145F" box="[1123,1136,1093,1115]" gridcol="5" gridrow="9" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845F89FBBBC78F145F" box="[1226,1239,1093,1115]" gridcol="6" gridrow="9" pageId="12" pageNumber="529">1</td>
<td id="C46AA067FFD003845E71FBBBC61B145F" box="[1330,1347,1093,1115]" gridcol="7" gridrow="9" pageId="12" pageNumber="529">B</td>
</tr>
</table>
</paragraph>
<paragraph id="3934CB59FFD0FC775B84FB5EC18516FC" blockId="12.[151,1437,1183,2037]" pageId="12" pageNumber="529">
In their phylogeny,
<bibRefCitation id="5D1AB6A8FFD0FC775AE9FB61C1C614BC" box="[426,670,1183,1208]" pageId="12" pageNumber="529" refString="Grismer, L. L., Anuar, S., Quah, E., Muin, M. A., Onn, C. K., Grismer, J. L. &amp; Ahmad, N. (2010) A new spiny, prehensile-tailed species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia with a preliminary hypothesis of relationships based on morphology. Zootaxa, 2625, 40 - 52." type="journal article">
Grismer
<emphasis id="0BFF174BFFD0FC77594CFB5FC11214BC" box="[527,586,1184,1208]" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
(2010)
</bibRefCitation>
placed
<taxonomicName id="FE8BB0DAFFD0FC7759B9FB5EC08514BC" box="[762,989,1184,1208]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC7759B9FB5EC08514BC" box="[762,989,1184,1208]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus durio</emphasis>
</taxonomicName>
as the sister species to
<taxonomicName id="FE8BB0DAFFD0FC775FB1FB5EC6C414B3" box="[1266,1436,1184,1207]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC775FB1FB5EC6C414B3" box="[1266,1436,1184,1207]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
with
<taxonomicName id="FE8BB0DAFFD0FC775B8CFB3AC20814D8" box="[207,336,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC775B8CFB3AC20814D8" box="[207,336,1220,1244]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
sister to the
<taxonomicName id="FE8BB0DAFFD0FC775A9DFB3AC16414D8" box="[478,572,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC775A9DFB3AC16414D8" box="[478,572,1220,1244]" italics="true" pageId="12" pageNumber="529">C. durio</emphasis>
</taxonomicName>
+
<taxonomicName id="FE8BB0DAFFD0FC77591AFB3AC1A714D8" box="[601,767,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC77591AFB3AC1A714D8" box="[601,767,1220,1244]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
clade. They note that
<taxonomicName id="FE8BB0DAFFD0FC7758B4FB3AC72014D8" box="[1015,1144,1220,1244]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC7758B4FB3AC72014D8" box="[1015,1144,1220,1244]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
lacks ventrolateral caudal tubercles, but did not explain how they scored this character. We count tubercles in the middle of the tail:
<emphasis id="0BFF174BFFD0FC775EC1FB16C23F1520" italics="true" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775EC1FB16C3AF1520" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="lateralis">C. lateralis</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD0FC775A40FAF2C23B1520" box="[259,355,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">C. durio</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FE8BB0DAFFD0FC775ADDFAF2C11D1520" box="[414,581,1292,1316]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC775ADDFAF2C11D1520" box="[414,581,1292,1316]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
have six spinose tubercles per caudal whorl throughout most of the tail (the count will be higher close to the body and lower distally), whereas
<taxonomicName id="FE8BB0DAFFD0FC7758C7FACEC75D1543" box="[900,1029,1328,1351]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC7758C7FACEC75D1543" box="[900,1029,1328,1351]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
has only 4 (also noted by
<bibRefCitation id="5D1AB6A8FFD0FC775E69FAD1C3AF1568" author="Linkem" pageId="12" pageNumber="529" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Mohammed, I. S., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toe gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, eastern Indonesia. Herpetologica, 64, 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem
<emphasis id="0BFF174BFFD0FC775ECBFACFC3EC1568" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
2008
</bibRefCitation>
). We add
<taxonomicName id="FE8BB0DAFFD0FC775A21FAAAC2871568" box="[354,479,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD0FC775A21FAAAC2871568" box="[354,479,1364,1388]" italics="true" pageId="12" pageNumber="529">C. lateralis</emphasis>
</taxonomicName>
to the
<taxonomicName id="FE8BB0DAFFD0FC775969FAAAC1DE1568" box="[554,646,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC775969FAAAC1DE1568" box="[554,646,1364,1388]" italics="true" pageId="12" pageNumber="529">C. durio</emphasis>
</taxonomicName>
+
<taxonomicName id="FE8BB0DAFFD0FC7759E7FAAAC0121568" box="[676,842,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC7759E7FAAAC0121568" box="[676,842,1364,1388]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
Clade, because
<taxonomicName id="FE8BB0DAFFD0FC7758BEFAAAC7011568" box="[1021,1113,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC7758BEFAAAC7011568" box="[1021,1113,1364,1388]" italics="true" pageId="12" pageNumber="529">C. durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD0FC775FCCFAAAC6541568" box="[1167,1292,1364,1388]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD0FC775FCCFAAAC6541568" box="[1167,1292,1364,1388]" italics="true" pageId="12" pageNumber="529">C. lateralis</emphasis>
</taxonomicName>
are identical for the characters used to construct the morphological phylogeny (
<tableCitation id="7409FEE2FFD0FC7758C2FA89C08D1594" box="[897,981,1399,1424]" captionStart="TABLE 1" captionStartId="12.[151,236,512,535]" captionTargetBox="[159,1347,690,1115]" captionTargetPageId="12" captionText="TABLE 1. Morphological matrix used to determine phylogeny of prehensile-tailed Cyrtodactylus. Characters are 1. Tail prehensile (1) or not (0); 2. Digits webbed (1) or not (0); 3. Ventrolateral caudal fringe present (1) or not (0); 4. Tail square (1) or round (0) in cross section; 5. Ventrolateral body fold with (1) or without (0) spinose tubercles; 6. Tail with ventrolateral caudal tubercles (1) or not (0); 7. Dorsum banded (B) or with pale vertebral stripe and 2 3 broken or complete chevrons (S), character unordered." httpUri="http://table.plazi.org/id/6DF49BD1FFD0FC775BD4FDFEC1281293" pageId="12" pageNumber="529" tableUuid="6DF49BD1FFD0FC775BD4FDFEC1281293">Table 1</tableCitation>
).
<taxonomicName id="FE8BB0DAFFD0FC7758ADFA86C7B91594" box="[1006,1249,1400,1424]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC7758ADFA86C7B91594" box="[1006,1249,1400,1424]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus nuaulu</emphasis>
</taxonomicName>
and
<emphasis id="0BFF174BFFD0FC775E5BFA86C6C41594" box="[1304,1436,1400,1424]" italics="true" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC775E5BFA86C6C01594" box="[1304,1432,1400,1424]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">C. serratus</taxonomicName>
,
</emphasis>
both from east of Wallaces Line, have conical tubercles in the ventrolateral fold, ventrolateral caudal tubercles, and tails with whorls of six spinose tubercles extending for most of the length of the tail. On the basis of morphology, we infer that the tails of these two species are prehensile, even though neither description mentions whether the species coils its tail in a manner similar to
<taxonomicName id="FE8BB0DAFFD0FC77586FF9F6C0EA161B" box="[812,946,1544,1567]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC77586FF9F6C0EA161B" box="[812,946,1544,1567]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
and the Sunda Shelf species. The unique
<typeStatus id="E63075FBFFD0FC775BD4F9D2C3A11641" box="[151,249,1580,1605]" pageId="12" pageNumber="529" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FE8BB0DAFFD0FC775A5DF9D2C2C31640" box="[286,411,1580,1604]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD0FC775A5DF9D2C2C31640" box="[286,411,1580,1604]" italics="true" pageId="12" pageNumber="529">C. serratus</emphasis>
</taxonomicName>
was collected in the Star Mountains of New
<collectingCountry id="419C8BC9FFD0FC7758D6F9D2C0BE1641" box="[917,998,1580,1605]" name="Guinea-Bissau" pageId="12" pageNumber="529">Guinea</collectingCountry>
in 1987 and
<bibRefCitation id="5D1AB6A8FFD0FC775F3AF9D3C6561640" author="Kraus" box="[1145,1294,1580,1604]" pageId="12" pageNumber="529" refString="Kraus, F. (2007) A new species of Cyrtodactylus (Squamata: Gekkonidae) from western Papua New Guinea. Zootaxa, 1425 (1), 63 - 68. http: // dx. doi. org / 10.11646 / zootaxa. 1425.1.8" type="journal article" year="2007">Kraus (2007)</bibRefCitation>
provided no information about its behavior or natural history. All adults of the
<typeStatus id="E63075FBFFD0FC7758C0F9AEC0EB166C" box="[899,947,1616,1640]" pageId="12" pageNumber="529">type</typeStatus>
series of
<taxonomicName id="FE8BB0DAFFD0FC775F62F9AEC7C9166C" box="[1057,1169,1616,1640]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC775F62F9AEC7C9166C" box="[1057,1169,1616,1640]" italics="true" pageId="12" pageNumber="529">C. nuaulu</emphasis>
</taxonomicName>
were collected on trees and vines,
<quantity id="FE7366BCFFD0FC775A53F98AC2D61688" box="[272,398,1652,1677]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="12" pageNumber="529" unit="cm" value="125.0" valueMax="200.0" valueMin="50.0">50200 cm</quantity>
aboveground on Seram (
<bibRefCitation id="5D1AB6A8FFD0FC7759EAF98AC02E1688" author="Oliver" box="[681,886,1652,1677]" pageId="12" pageNumber="529" refString="Oliver, P., Edgar, P., Mumpuni, Iskandar, D. T. &amp; Lilley, R. (2009) A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa, 2115, 47 - 55." type="journal article" year="2009">
Oliver
<emphasis id="0BFF174BFFD0FC7759BAF98BC06A1688" box="[761,818,1652,1676]" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
2009
</bibRefCitation>
). Based on these observations, we add
<taxonomicName id="FE8BB0DAFFD0FC775E6EF98AC6C41688" box="[1325,1436,1652,1676]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC775E6EF98AC6C41688" box="[1325,1436,1652,1676]" italics="true" pageId="12" pageNumber="529">C. nuaulu</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD0FC775B8AF966C21016B4" box="[201,328,1688,1712]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD0FC775B8AF966C21016B4" box="[201,328,1688,1712]" italics="true" pageId="12" pageNumber="529">C. serratus</emphasis>
</taxonomicName>
to the clade containing
<taxonomicName id="FE8BB0DAFFD0FC775927F966C05516AB" box="[612,781,1688,1711]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC775927F966C05516AB" box="[612,781,1688,1711]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
,
<emphasis id="0BFF174BFFD0FC77585DF966C0DD16B4" box="[798,901,1688,1712]" italics="true" pageId="12" pageNumber="529">
<taxonomicName id="FE8BB0DAFFD0FC77585DF966C0DA16B4" box="[798,898,1688,1712]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">C. durio</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FE8BB0DAFFD0FC775881F966C71B16B4" box="[962,1091,1688,1712]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD0FC775881F966C71B16B4" box="[962,1091,1688,1712]" italics="true" pageId="12" pageNumber="529">C. lateralis</emphasis>
</taxonomicName>
. We refer to this clade as the “insular clade” of prehensile-tailed geckos, because most of the species occur on islands. Of the five species of the insular clade, only
<taxonomicName id="FE8BB0DAFFD0FC775A2FF91EC29116FC" box="[364,457,1760,1784]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC775A2FF91EC29116FC" box="[364,457,1760,1784]" italics="true" pageId="12" pageNumber="529">C. durio</emphasis>
</taxonomicName>
occurs on the mainland.
</paragraph>
<paragraph id="3934CB59FFD0FC765B84F8FBC07F11A9" blockId="12.[151,1437,1183,2037]" lastBlockId="13.[151,1437,151,1149]" lastPageId="13" lastPageNumber="530" pageId="12" pageNumber="529">
We have shown that some distinctive characters originally suggesting a close relationship between
<taxonomicName id="FE8BB0DAFFD0FC775BD4F8D6C2211744" box="[151,377,1832,1856]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD0FC775BD4F8D6C2211744" box="[151,377,1832,1856]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD0FC775AF0F8D6C104173B" box="[435,604,1832,1855]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC775AF0F8D6C104173B" box="[435,604,1832,1855]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
are also found in
<taxonomicName id="FE8BB0DAFFD0FC77586DF8D6C0F51744" box="[814,941,1832,1856]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD0FC77586DF8D6C0F51744" box="[814,941,1832,1856]" italics="true" pageId="12" pageNumber="529">C. serratus</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD0FC7758A5F8D6C70F1744" box="[998,1111,1832,1856]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC7758A5F8D6C70F1744" box="[998,1111,1832,1856]" italics="true" pageId="12" pageNumber="529">C. nuaulu</emphasis>
</taxonomicName>
. Although the specimen has faded with age, the dorsal pattern of the
<typeStatus id="E63075FBFFD0FC77592AF8B2C1931761" box="[617,715,1868,1893]" pageId="12" pageNumber="529" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FE8BB0DAFFD0FC7759B7F8B2C0C51760" box="[756,925,1868,1892]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD0FC7759B7F8B2C0C51760" box="[756,925,1868,1892]" italics="true" pageId="12" pageNumber="529">C. stresemanni</emphasis>
</taxonomicName>
remains well-defined (
<bibRefCitation id="5D1AB6A8FFD0FC775FE6F8B2C275178C" author="Rosler" pageId="12" pageNumber="529" refString="Rosler, H. &amp; Glaw, F. (2008) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa, 1729, 8 - 22." type="journal article" year="2008">Rösler &amp; Glaw 2008, their figure 7</bibRefCitation>
). The specimen has two pairs of elongate blotches on the flanks separated by a wide, pale vertebral stripe. This dorsal pattern is unlike that of any other Sunda Shelf
<taxonomicName id="FE8BB0DAFFD0FC7758C3F86AC74517A8" box="[896,1053,1940,1964]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD0FC7758C3F86AC74517A8" box="[896,1053,1940,1964]" italics="true" pageId="12" pageNumber="529">Cyrtodactylus</emphasis>
</taxonomicName>
, however it resembles patterns in the
<typeStatus id="E63075FBFFD0FC775BFCF847C3B717D5" box="[191,239,1977,2001]" pageId="12" pageNumber="529">type</typeStatus>
series of
<taxonomicName id="FE8BB0DAFFD0FC775A19F847C29017D4" box="[346,456,1976,2000]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD0FC775A19F847C29017D4" box="[346,456,1976,2000]" italics="true" pageId="12" pageNumber="529">C. nuaulu</emphasis>
</taxonomicName>
(compare especially to MZB
<quantity id="FE7366BCFFD0FC775857F846C03117D5" box="[788,873,1976,2001]" metricMagnitude="1" metricUnit="m" metricValue="5.905499999999999" pageId="12" pageNumber="529" unit="in" value="2325.0">2325 in</quantity>
<bibRefCitation id="5D1AB6A8FFD0FC77582CF846C76E17D4" author="Oliver" box="[879,1078,1976,2001]" pageId="12" pageNumber="529" refString="Oliver, P., Edgar, P., Mumpuni, Iskandar, D. T. &amp; Lilley, R. (2009) A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa, 2115, 47 - 55." type="journal article" year="2009">
Oliver
<emphasis id="0BFF174BFFD0FC7758FEF847C0AC17D4" box="[957,1012,1976,2000]" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
2009
</bibRefCitation>
, their figure 2),
<taxonomicName id="FE8BB0DAFFD0FC775FABF847C63017D4" box="[1256,1384,1977,2000]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD0FC775FABF847C63017D4" box="[1256,1384,1977,2000]" italics="true" pageId="12" pageNumber="529">C. spinosus</emphasis>
</taxonomicName>
(see especially MZB 7029 with “less complete” chevrons,
<bibRefCitation id="5D1AB6A8FFD0FC7759BEF822C08217F0" author="Linkem" box="[765,986,2012,2037]" pageId="12" pageNumber="529" refString="Linkem, C. W., McGuire, J. A., Hayden, C. J., Mohammed, I. S., Bickford, D. P. &amp; Brown, R. M. (2008) A new species of bent-toe gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, eastern Indonesia. Herpetologica, 64, 224 - 234. http: // dx. doi. org / 10.1655 / 07 - 040.1" type="journal article" year="2008">
Linkem
<emphasis id="0BFF174BFFD0FC77581FF823C0CE17F0" box="[860,918,2012,2036]" italics="true" pageId="12" pageNumber="529">et al.</emphasis>
2008
</bibRefCitation>
: 228), and
<taxonomicName id="FE8BB0DAFFD0FC775F1AF822C78F17F0" box="[1113,1239,2012,2036]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="529" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD0FC775F1AF822C78F17F0" box="[1113,1239,2012,2036]" italics="true" pageId="12" pageNumber="529">C. serratus</emphasis>
</taxonomicName>
(
<bibRefCitation id="5D1AB6A8FFD0FC765FA4F822C25110B4" author="Kraus" lastPageId="13" lastPageNumber="530" pageId="12" pageNumber="529" refString="Kraus, F. (2007) A new species of Cyrtodactylus (Squamata: Gekkonidae) from western Papua New Guinea. Zootaxa, 1425 (1), 63 - 68. http: // dx. doi. org / 10.11646 / zootaxa. 1425.1.8" type="journal article" year="2007">Kraus 2007, his figure 1A</bibRefCitation>
). All four species share a wide pale vertebral stripe and few (23 between limbs) large, dark-brown chevron-shaped blotches. The vertebral stripe may be interrupted by the chevrons medially or divide the chevrons as it does in the
<typeStatus id="E63075FBFFD1FC765A0AFF21C2EE10FC" box="[329,438,223,248]" pageId="13" pageNumber="530" type="holotype">holotypes</typeStatus>
of
<taxonomicName id="FE8BB0DAFFD1FC765A99FF1EC10E10FC" box="[474,598,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="serratus">
<emphasis id="0BFF174BFFD1FC765A99FF1EC10E10FC" box="[474,598,224,248]" italics="true" pageId="13" pageNumber="530">C. serratus</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC7659CFFF1EC06A10F3" box="[652,818,224,247]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD1FC7659CFFF1EC06A10F3" box="[652,818,224,247]" italics="true" pageId="13" pageNumber="530">C. stresemanni</emphasis>
</taxonomicName>
and some
<typeStatus id="E63075FBFFD1FC7658EAFF1EC74C10FC" box="[937,1044,224,248]" pageId="13" pageNumber="530" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="FE8BB0DAFFD1FC765F7BFF1EC7E010F3" box="[1080,1208,224,247]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD1FC765F7BFF1EC7E010F3" box="[1080,1208,224,247]" italics="true" pageId="13" pageNumber="530">C. spinosus</emphasis>
</taxonomicName>
(MZB 7029) and
<taxonomicName id="FE8BB0DAFFD1FC765EC1FF1EC3BD1118" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="nuaulu">
<emphasis id="0BFF174BFFD1FC765EC1FF1EC3BD1118" italics="true" pageId="13" pageNumber="530">C. nuaulu</emphasis>
</taxonomicName>
(MZB 2329). Although the dark triangular blotch on the nape of
<taxonomicName id="FE8BB0DAFFD1FC7658FCFEFAC73E1118" box="[959,1126,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD1FC7658FCFEFAC73E1118" box="[959,1126,260,284]" italics="true" pageId="13" pageNumber="530">C. stresemanni</emphasis>
</taxonomicName>
might at first appear to ally it with
<taxonomicName id="FE8BB0DAFFD1FC765BAFFED6C2671144" box="[236,319,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765BAFFED6C2671144" box="[236,319,296,320]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
and
<emphasis id="0BFF174BFFD1FC765A3FFED6C0011144" box="[380,857,296,320]" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765A3FFED6C11A1144" box="[380,578,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="brevipalmatus">C. brevipalmatus</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC765913FED6C19D1144" box="[592,709,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="nuaulu">C. nuaulu</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC765990FED6C00E1144" box="[723,854,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="serratus">C. serratus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FE8BB0DAFFD1FC7658D5FED6C742113B" box="[918,1050,296,319]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="0BFF174BFFD1FC7658D5FED6C742113B" box="[918,1050,296,319]" italics="true" pageId="13" pageNumber="530">C. spinosus</emphasis>
</taxonomicName>
have triangular blotches on their napes as well.
<taxonomicName id="FE8BB0DAFFD1FC765A11FEB2C1631160" box="[338,571,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765A11FEB2C1631160" box="[338,571,332,356]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC7659C2FEB2C0501160" box="[641,776,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC7659C2FEB2C0501160" box="[641,776,332,356]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
unequivocally lack the distinctive color pattern of
<taxonomicName id="FE8BB0DAFFD1FC765EC1FEB2C2451183" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="0BFF174BFFD1FC765EC1FEB2C2451183" italics="true" pageId="13" pageNumber="530">C. stresemanni</emphasis>
</taxonomicName>
and the three other species. Like
<emphasis id="0BFF174BFFD1FC7659F3FE8EC087118C" box="[688,991,368,392]" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC7659F3FE8EC02E118C" box="[688,886,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="brevipalmatus">C. brevipalmatus</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC7658C6FE8EC083118C" box="[901,987,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">C. elok</taxonomicName>
,
</emphasis>
and
<taxonomicName id="FE8BB0DAFFD1FC765F5EFE8EC78A118C" box="[1053,1234,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD1FC765F5EFE8EC78A118C" box="[1053,1234,368,392]" italics="true" pageId="13" pageNumber="530">C. interdigitalis</emphasis>
</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC765FA0FE8EC61C118C" box="[1251,1348,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765FA0FE8EC61C118C" box="[1251,1348,368,392]" italics="true" pageId="13" pageNumber="530">C. durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765EC1FE8EC3AD11A8" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765EC1FE8EC3AD11A8" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
are banded species lacking a pale vertebral stripe.
</paragraph>
<paragraph id="3934CB59FFD1FC765B84FE46C0B812C8" blockId="13.[151,1437,151,1149]" pageId="13" pageNumber="530">
To determine polarity of the distinctive color pattern of the Wallacean and Papuan species, we reviewed recent descriptions and revisions of
<taxonomicName id="FE8BB0DAFFD1FC765A9CFE22C12411F0" box="[479,636,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD1FC765A9CFE22C12411F0" box="[479,636,476,500]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
from across the distribution of the genus. We were unable to find other species with a pale vertebral stripe and 23 dark brown chevrons.
<taxonomicName id="FE8BB0DAFFD1FC76583BFDFEC74E121C" authority="Oliver" authorityName="Oliver" box="[888,1046,511,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="zugi">
<emphasis id="0BFF174BFFD1FC76583BFDFEC09F1213" box="[888,967,512,535]" italics="true" pageId="13" pageNumber="530">C. zugi</emphasis>
Oliver
</taxonomicName>
has two large brown chevrons, but lacks a vertebral stripe. The pattern of the Wallacean and Papuan prehensile-tailed
<taxonomicName id="FE8BB0DAFFD1FC765F1BFDDAC7AD1238" box="[1112,1269,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD1FC765F1BFDDAC7AD1238" box="[1112,1269,548,572]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
is most likely derived, however, its distribution is not congruent with the morphological phylogeny regardless of polarity. If derived, the pattern is a synapomorphy of the insular clade, with a reversal to the banded pattern in
<taxonomicName id="FE8BB0DAFFD1FC765FACFD92C6131280" box="[1263,1355,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765FACFD92C6131280" box="[1263,1355,620,644]" italics="true" pageId="13" pageNumber="530">C. durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765EC1FD92C3AC12AC" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765EC1FD92C3AC12AC" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
. If the pattern is primitive, a banded pattern would have to evolve twice: once leading to the mainland SE Asia clade and once leading to a clade containing
<taxonomicName id="FE8BB0DAFFD1FC765984FD4AC07D12C8" box="[711,805,692,716]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765984FD4AC07D12C8" box="[711,805,692,716]" italics="true" pageId="13" pageNumber="530">C. durio</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765818FD4AC08112C8" box="[859,985,692,716]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765818FD4AC08112C8" box="[859,985,692,716]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3934CB59FFD1FC765B84FD29C25513A1" blockId="13.[151,1437,151,1149]" pageId="13" pageNumber="530">
Prior to this study, molecular data were available for only two of the eight species of prehensile-tailed
<taxonomicName id="FE8BB0DAFFD1FC765BD4FD02C26C1310" box="[151,308,764,788]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD1FC765BD4FD02C26C1310" box="[151,308,764,788]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
.
<bibRefCitation id="5D1AB6A8FFD1FC765A07FD02C1461310" box="[324,542,764,789]" pageId="13" pageNumber="530" refString="Wood, P. L. Jr., Heinicke, M. P., Jackman, T. R. &amp; Bauer, A. M. (2012) Phylogeny of bent-toed geckos (Cyrtodactylus) reveals a west to east pattern of diversification. Molecular Phylogenetics and Evolution, 65, 992 - 1003. http: // dx. doi. org / 10.1016 / j. ympev. 2012.08.025" type="journal article">
Wood
<emphasis id="0BFF174BFFD1FC765AD3FD03C2921310" box="[400,458,764,788]" italics="true" pageId="13" pageNumber="530">et al.</emphasis>
(2012)
</bibRefCitation>
confirmed the suspicions of
<bibRefCitation id="5D1AB6A8FFD1FC76582DFD02C7591310" author="Ulber" box="[878,1025,764,789]" pageId="13" pageNumber="530" refString="Ulber, T. (1993) Bemerkungen uber cyrtodactyline Geckos aus Thailand nebst Beschreibungen von zwei neuen Arten (Reptilia: Gekkonidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin. Zoologisches Museum und Institut fur Spezielle Zoologie, 69, 187 - 200. http: // dx. doi. org / 10.1002 / mmnz. 19930690202" type="journal article" year="1993">Ulber (1993)</bibRefCitation>
that
<taxonomicName id="FE8BB0DAFFD1FC765F7DFD02C7C81310" box="[1086,1168,764,788]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765F7DFD02C7C81310" box="[1086,1168,764,788]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
is closely related to
<emphasis id="0BFF174BFFD1FC765EC1FD02C275133C" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765EC1FD02C271133C" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">C. interdigitalis</taxonomicName>
,
</emphasis>
and these two species are sister taxa in Clade F of their phylogeny. We obtained ND2 and COI sequences for four of our new specimens of
<taxonomicName id="FE8BB0DAFFD1FC7659DAFCBAC0421358" box="[665,794,836,860]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC7659DAFCBAC0421358" box="[665,794,836,860]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
and an ND2 sequence of the
<typeStatus id="E63075FBFFD1FC765F37FCBAC78E1359" box="[1140,1238,836,861]" pageId="13" pageNumber="530" type="holotype">holotype</typeStatus>
of
<emphasis id="0BFF174BFFD1FC765E43FCBAC6301358" box="[1280,1384,836,860]" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765E43FCBAC63B1358" box="[1280,1379,836,860]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">C. durio</taxonomicName>
.
</emphasis>
Our maximum likelihood and Bayesian analyses recovered similar topologies with minor differences in support for each gene.
</paragraph>
<paragraph id="3934CB59FFD1FC765B84FC51C79A1478" blockId="13.[151,1437,151,1149]" pageId="13" pageNumber="530">
Relatively few sequences of the COI gene were available on GenBank (Appendix 1), however we were able to place
<taxonomicName id="FE8BB0DAFFD1FC765B9AFC2AC2F513E8" box="[217,429,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765B9AFC2AC2F513E8" box="[217,429,980,1004]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765AA7FC2AC13A13E8" box="[484,610,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765AA7FC2AC13A13E8" box="[484,610,980,1004]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
within a phylogeny containing 34 additional congeners and
<taxonomicName id="FE8BB0DAFFD1FC765E4FFC2AC6C413E8" box="[1292,1436,980,1004]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="gekko">
<emphasis id="0BFF174BFFD1FC765E4FFC2AC6C413E8" box="[1292,1436,980,1004]" italics="true" pageId="13" pageNumber="530">Gekko gekko</emphasis>
</taxonomicName>
as an outgroup. Our Maximum Likelihood tree contained few mainland species and had low support for most nodes. Nonetheless, we found no evidence of a close relationship between
<taxonomicName id="FE8BB0DAFFD1FC7658A8FBE2C7631430" box="[1003,1083,1052,1076]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC7658A8FBE2C7631430" box="[1003,1083,1052,1076]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765F36FBE2C7AC1430" box="[1141,1268,1052,1076]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765F36FBE2C7AC1430" box="[1141,1268,1052,1076]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
using the COI sequences. In this small matrix of species,
<taxonomicName id="FE8BB0DAFFD1FC76593BFBBEC190145C" box="[632,712,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC76593BFBBEC190145C" box="[632,712,1088,1112]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC7659BCFBBEC025145C" box="[767,893,1088,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC7659BCFBBEC025145C" box="[767,893,1088,1112]" italics="true" pageId="13" pageNumber="530">C. lateralis</emphasis>
</taxonomicName>
were placed near the base of the phylogeny and had uncorrected genetic distances greater than 19% to other species in the phylogeny (
<tableCitation id="7409FEE2FFD1FC765F1EFB9AC7ED1478" box="[1117,1205,1124,1149]" captionStart="TABLE 2" captionStartId="13.[151,236,1196,1219]" captionTargetBox="[159,1383,1310,1664]" captionTargetPageId="13" captionText="TABLE 2. Uncorrected pairwise distances (percent difference) of mitochondrial loci for representative individuals of Malaysian and Indonesian Cyrtodactylus. The upper diagonal shows distances for Cytochrome c oxidase subunit I; the lower diagonal shows distances for NADH dehydrogenase 2 gene sequences." httpUri="http://table.plazi.org/id/6DF49BD1FFD1FC765BD4FB52C0961507" pageId="13" pageNumber="530" tableUuid="6DF49BD1FFD1FC765BD4FB52C0961507">Table 2</tableCitation>
).
</paragraph>
<caption id="6DF49BD1FFD1FC765BD4FB52C0961507" ID-Table-UUID="6DF49BD1FFD1FC765BD4FB52C0961507" httpUri="http://table.plazi.org/id/6DF49BD1FFD1FC765BD4FB52C0961507" pageId="13" pageNumber="530" targetBox="[159,1383,1310,1664]" targetIsTable="true" targetPageId="13">
<paragraph id="3934CB59FFD1FC765BD4FB52C0961507" blockId="13.[151,1436,1196,1283]" pageId="13" pageNumber="530">
<emphasis id="0BFF174BFFD1FC765BD4FB52C25114C7" bold="true" box="[151,265,1196,1219]" pageId="13" pageNumber="530">TABLE 2.</emphasis>
Uncorrected pairwise distances (percent difference) of mitochondrial loci for representative individuals of Malaysian and Indonesian
<taxonomicName id="FE8BB0DAFFD1FC765AFAFB35C11614E7" box="[441,590,1227,1251]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD1FC765AFAFB35C11614E7" box="[441,590,1227,1251]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
. The upper diagonal shows distances for Cytochrome c oxidase subunit I; the lower diagonal shows distances for NADH dehydrogenase 2 gene sequences.
</paragraph>
</caption>
<paragraph id="3934CB59FFD1FC765A34FAE0C6491684" pageId="13" pageNumber="530">
<table id="4B8B39F9FFD103845BDCFAE0C63F1684" box="[159,1383,1310,1664]" gridcols="8" gridrows="9" pageId="13" pageNumber="530">
<tr id="87BBC91BFFD103845BDCFAE0C63F1530" box="[159,1383,1310,1332]" gridrow="0" pageId="13" pageNumber="530" rowspan-0="1">
<th id="C46AA067FFD103845A34FAE0C2931530" box="[375,459,1310,1332]" gridcol="1" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765A34FAE0C2931530" box="[375,459,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765A34FAE0C2931530" box="[375,459,1310,1332]" italics="true" pageId="13" pageNumber="530">lateralis</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD103845ABFFAE0C16C1530" box="[508,564,1310,1332]" gridcol="2" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765ABEFAE0C16C1530" box="[509,564,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765ABEFAE0C16C1530" box="[509,564,1310,1332]" italics="true" pageId="13" pageNumber="530">durio</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD10384591DFAE0C1821530" box="[606,730,1310,1332]" gridcol="3" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC76591DFAE0C1821530" box="[606,730,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="0BFF174BFFD1FC76591DFAE0C1821530" box="[606,730,1310,1332]" italics="true" pageId="13" pageNumber="530">consobrinus</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD103845856FAE0C0D81530" box="[789,896,1310,1332]" gridcol="4" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765856FAE0C0D81530" box="[789,896,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="paradoxus">
<emphasis id="0BFF174BFFD1FC765856FAE0C0D81530" box="[789,896,1310,1332]" italics="true" pageId="13" pageNumber="530">paradoxus</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD1038458FAFAE0C7171530" box="[953,1103,1310,1332]" gridcol="5" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC7658FAFAE0C7171530" box="[953,1103,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="0BFF174BFFD1FC7658FAFAE0C7171530" box="[953,1103,1310,1332]" italics="true" pageId="13" pageNumber="530">quadrivirgatus</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD103845FD2FAE0C7E51530" box="[1169,1213,1310,1332]" gridcol="6" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765FD2FAE0C7E31530" box="[1169,1211,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765FD2FAE0C7E31530" box="[1169,1211,1310,1332]" italics="true" pageId="13" pageNumber="530">elok</emphasis>
</taxonomicName>
</th>
<th id="C46AA067FFD103845FA6FAE0C63F1530" box="[1253,1383,1310,1332]" gridcol="7" gridrow="0" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765FA6FAE0C63F1530" box="[1253,1383,1310,1332]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD1FC765FA6FAE0C63F1530" box="[1253,1383,1310,1332]" italics="true" pageId="13" pageNumber="530">interdigitalis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="87BBC91BFFD103845BDCFAB7C63F155B" box="[159,1383,1353,1375]" gridrow="1" pageId="13" pageNumber="530" rowspan-1="1">
<th id="C46AA067FFD103845BDCFAB7C26D155B" box="[159,309,1353,1375]" gridcol="0" gridrow="1" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCFAB7C3AC155B" box="[159,244,1353,1375]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765BDCFAB7C3AC155B" box="[159,244,1353,1375]" italics="true" pageId="13" pageNumber="530">lateralis</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845ABFFAB7C16C155B" box="[508,564,1353,1375]" gridcol="2" gridrow="1" pageId="13" pageNumber="530">-</td>
<td id="C46AA067FFD10384591DFAB7C182155B" box="[606,730,1353,1375]" gridcol="3" gridrow="1" pageId="13" pageNumber="530">20.1</td>
<td id="C46AA067FFD103845856FAB7C0D8155B" box="[789,896,1353,1375]" gridcol="4" gridrow="1" pageId="13" pageNumber="530">19.1</td>
<td id="C46AA067FFD1038458FAFAB7C717155B" box="[953,1103,1353,1375]" gridcol="5" gridrow="1" pageId="13" pageNumber="530">18.8</td>
<td id="C46AA067FFD103845FD2FAB7C7E5155B" box="[1169,1213,1353,1375]" gridcol="6" gridrow="1" pageId="13" pageNumber="530">21.1</td>
<td id="C46AA067FFD103845FA6FAB7C63F155B" box="[1253,1383,1353,1375]" gridcol="7" gridrow="1" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCFA8CC63F158C" box="[159,1383,1394,1416]" gridrow="2" pageId="13" pageNumber="530" rowspan-2="1">
<th id="C46AA067FFD103845BDCFA8CC26D158C" box="[159,309,1394,1416]" gridcol="0" gridrow="2" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCFA8CC38E158C" box="[159,214,1394,1416]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD1FC765BDCFA8CC38E158C" box="[159,214,1394,1416]" italics="true" pageId="13" pageNumber="530">durio</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34FA8CC293158C" box="[375,459,1394,1416]" gridcol="1" gridrow="2" pageId="13" pageNumber="530">11.7</td>
<td id="C46AA067FFD10384591DFA8CC182158C" box="[606,730,1394,1416]" gridcol="3" gridrow="2" pageId="13" pageNumber="530">-</td>
<td id="C46AA067FFD103845856FA8CC0D8158C" box="[789,896,1394,1416]" gridcol="4" gridrow="2" pageId="13" pageNumber="530">-</td>
<td id="C46AA067FFD1038458FAFA8CC717158C" box="[953,1103,1394,1416]" gridcol="5" gridrow="2" pageId="13" pageNumber="530">-</td>
<td id="C46AA067FFD103845FD2FA8CC7E5158C" box="[1169,1213,1394,1416]" gridcol="6" gridrow="2" pageId="13" pageNumber="530">-</td>
<td id="C46AA067FFD103845FA6FA8CC63F158C" box="[1253,1383,1394,1416]" gridcol="7" gridrow="2" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCFA62C63F15B6" box="[159,1383,1436,1458]" gridrow="3" pageId="13" pageNumber="530" rowspan-3="1">
<th id="C46AA067FFD103845BDCFA62C26D15B6" box="[159,309,1436,1458]" gridcol="0" gridrow="3" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCFA62C24315B6" box="[159,283,1436,1458]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="consobrinus">
<emphasis id="0BFF174BFFD1FC765BDCFA62C24315B6" box="[159,283,1436,1458]" italics="true" pageId="13" pageNumber="530">consobrinus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34FA62C29315B6" box="[375,459,1436,1458]" gridcol="1" gridrow="3" pageId="13" pageNumber="530">23.3</td>
<td id="C46AA067FFD103845ABFFA62C16C15B6" box="[508,564,1436,1458]" gridcol="2" gridrow="3" pageId="13" pageNumber="530">22.2</td>
<td id="C46AA067FFD103845856FA62C0D815B6" box="[789,896,1436,1458]" gridcol="4" gridrow="3" pageId="13" pageNumber="530">20.2</td>
<td id="C46AA067FFD1038458FAFA62C71715B6" box="[953,1103,1436,1458]" gridcol="5" gridrow="3" pageId="13" pageNumber="530">16.9</td>
<td id="C46AA067FFD103845FD2FA62C7E515B6" box="[1169,1213,1436,1458]" gridcol="6" gridrow="3" pageId="13" pageNumber="530">19.3</td>
<td id="C46AA067FFD103845FA6FA62C63F15B6" box="[1253,1383,1436,1458]" gridcol="7" gridrow="3" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCFA3BC63F15DF" box="[159,1383,1477,1499]" gridrow="4" pageId="13" pageNumber="530" rowspan-4="1">
<th id="C46AA067FFD103845BDCFA3BC26D15DF" box="[159,309,1477,1499]" gridcol="0" gridrow="4" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCFA3BC25215DF" box="[159,266,1477,1499]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="paradoxus">
<emphasis id="0BFF174BFFD1FC765BDCFA3BC25215DF" box="[159,266,1477,1499]" italics="true" pageId="13" pageNumber="530">paradoxus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34FA3BC29315DF" box="[375,459,1477,1499]" gridcol="1" gridrow="4" pageId="13" pageNumber="530">19.1</td>
<td id="C46AA067FFD103845ABFFA3BC16C15DF" box="[508,564,1477,1499]" gridcol="2" gridrow="4" pageId="13" pageNumber="530">21.1</td>
<td id="C46AA067FFD10384591DFA3BC18215DF" box="[606,730,1477,1499]" gridcol="3" gridrow="4" pageId="13" pageNumber="530">21.5</td>
<td id="C46AA067FFD1038458FAFA3BC71715DF" box="[953,1103,1477,1499]" gridcol="5" gridrow="4" pageId="13" pageNumber="530">18.9</td>
<td id="C46AA067FFD103845FD2FA3BC7E515DF" box="[1169,1213,1477,1499]" gridcol="6" gridrow="4" pageId="13" pageNumber="530">20.4</td>
<td id="C46AA067FFD103845FA6FA3BC63F15DF" box="[1253,1383,1477,1499]" gridcol="7" gridrow="4" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCFA10C63F1600" box="[159,1383,1518,1540]" gridrow="5" pageId="13" pageNumber="530" rowspan-5="1">
<th id="C46AA067FFD103845BDCFA10C26D1600" box="[159,309,1518,1540]" gridcol="0" gridrow="5" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCFA10C26D1600" box="[159,309,1518,1540]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="0BFF174BFFD1FC765BDCFA10C26D1600" box="[159,309,1518,1540]" italics="true" pageId="13" pageNumber="530">quadrivirgatus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34FA10C2931600" box="[375,459,1518,1540]" gridcol="1" gridrow="5" pageId="13" pageNumber="530">18.7</td>
<td id="C46AA067FFD103845ABFFA10C16C1600" box="[508,564,1518,1540]" gridcol="2" gridrow="5" pageId="13" pageNumber="530">21.8</td>
<td id="C46AA067FFD10384591DFA10C1821600" box="[606,730,1518,1540]" gridcol="3" gridrow="5" pageId="13" pageNumber="530">28.7</td>
<td id="C46AA067FFD103845856FA10C0D81600" box="[789,896,1518,1540]" gridcol="4" gridrow="5" pageId="13" pageNumber="530">21.4</td>
<td id="C46AA067FFD103845FD2FA10C7E51600" box="[1169,1213,1518,1540]" gridcol="6" gridrow="5" pageId="13" pageNumber="530">19.3</td>
<td id="C46AA067FFD103845FA6FA10C63F1600" box="[1253,1383,1518,1540]" gridcol="7" gridrow="5" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCF9E6C63F162A" box="[159,1383,1560,1582]" gridrow="6" pageId="13" pageNumber="530" rowspan-6="1">
<th id="C46AA067FFD103845BDCF9E6C26D162A" box="[159,309,1560,1582]" gridcol="0" gridrow="6" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCF9E6C391162A" box="[159,201,1560,1582]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765BDCF9E6C391162A" box="[159,201,1560,1582]" italics="true" pageId="13" pageNumber="530">elok</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34F9E6C293162A" box="[375,459,1560,1582]" gridcol="1" gridrow="6" pageId="13" pageNumber="530">27.6</td>
<td id="C46AA067FFD103845ABFF9E6C16C162A" box="[508,564,1560,1582]" gridcol="2" gridrow="6" pageId="13" pageNumber="530">28.4</td>
<td id="C46AA067FFD10384591DF9E6C182162A" box="[606,730,1560,1582]" gridcol="3" gridrow="6" pageId="13" pageNumber="530">28.7</td>
<td id="C46AA067FFD103845856F9E6C0D8162A" box="[789,896,1560,1582]" gridcol="4" gridrow="6" pageId="13" pageNumber="530">28.7</td>
<td id="C46AA067FFD1038458FAF9E6C717162A" box="[953,1103,1560,1582]" gridcol="5" gridrow="6" pageId="13" pageNumber="530">30.2</td>
<td id="C46AA067FFD103845FA6F9E6C63F162A" box="[1253,1383,1560,1582]" gridcol="7" gridrow="6" pageId="13" pageNumber="530">-</td>
</tr>
<tr id="87BBC91BFFD103845BDCF9BFC63F1653" box="[159,1383,1601,1623]" gridrow="7" pageId="13" pageNumber="530" rowspan-7="1">
<th id="C46AA067FFD103845BDCF9BFC26D1653" box="[159,309,1601,1623]" gridcol="0" gridrow="7" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCF9BFC2791653" box="[159,289,1601,1623]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD1FC765BDCF9BFC2791653" box="[159,289,1601,1623]" italics="true" pageId="13" pageNumber="530">interdigitalis</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34F9BFC2931653" box="[375,459,1601,1623]" gridcol="1" gridrow="7" pageId="13" pageNumber="530">26.7</td>
<td id="C46AA067FFD103845ABFF9BFC16C1653" box="[508,564,1601,1623]" gridcol="2" gridrow="7" pageId="13" pageNumber="530">27.4</td>
<td id="C46AA067FFD10384591DF9BFC1821653" box="[606,730,1601,1623]" gridcol="3" gridrow="7" pageId="13" pageNumber="530">26.7</td>
<td id="C46AA067FFD103845856F9BFC0D81653" box="[789,896,1601,1623]" gridcol="4" gridrow="7" pageId="13" pageNumber="530">26.5</td>
<td id="C46AA067FFD1038458FAF9BFC7171653" box="[953,1103,1601,1623]" gridcol="5" gridrow="7" pageId="13" pageNumber="530">27.5</td>
<td id="C46AA067FFD103845FD2F9BFC7E51653" box="[1169,1213,1601,1623]" gridcol="6" gridrow="7" pageId="13" pageNumber="530">22.9</td>
</tr>
<tr id="87BBC91BFFD103845BDCF994C63F1684" box="[159,1383,1642,1664]" gridrow="8" pageId="13" pageNumber="530">
<th id="C46AA067FFD103845BDCF994C26D1684" box="[159,309,1642,1664]" gridcol="0" gridrow="8" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765BDCF995C24D1684" box="[159,277,1643,1664]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="semicinctus">
<emphasis id="0BFF174BFFD1FC765BDCF995C24D1684" box="[159,277,1643,1664]" italics="true" pageId="13" pageNumber="530">semicinctus</emphasis>
</taxonomicName>
</th>
<td id="C46AA067FFD103845A34F994C2931684" box="[375,459,1642,1664]" gridcol="1" gridrow="8" pageId="13" pageNumber="530">22.1</td>
<td id="C46AA067FFD103845ABFF994C16C1684" box="[508,564,1642,1664]" gridcol="2" gridrow="8" pageId="13" pageNumber="530">21.6</td>
<td id="C46AA067FFD10384591DF994C1821684" box="[606,730,1642,1664]" gridcol="3" gridrow="8" pageId="13" pageNumber="530">22.9</td>
<td id="C46AA067FFD103845856F994C0D81684" box="[789,896,1642,1664]" gridcol="4" gridrow="8" pageId="13" pageNumber="530">76.2</td>
<td id="C46AA067FFD1038458FAF994C7171684" box="[953,1103,1642,1664]" gridcol="5" gridrow="8" pageId="13" pageNumber="530">22.9</td>
<td id="C46AA067FFD103845FD2F994C7E51684" box="[1169,1213,1642,1664]" gridcol="6" gridrow="8" pageId="13" pageNumber="530">23.7</td>
<td id="C46AA067FFD103845FA6F994C63F1684" box="[1253,1383,1642,1664]" gridcol="7" gridrow="8" pageId="13" pageNumber="530">25.6</td>
</tr>
</table>
</paragraph>
<paragraph id="3934CB59FFD1FC765B84F939C08A1797" blockId="13.[151,1436,1734,2011]" pageId="13" pageNumber="530">
Our ND2 matrix included
<emphasis id="0BFF174BFFD1FC765AB3F938C7D916DA" box="[496,1153,1734,1758]" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC765AB3F938C18B16DA" box="[496,723,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="durio">Cyrtodactylus durio</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC76599CF939C06916DA" box="[735,817,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">C. elok</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC76587EF939C0A816DA" box="[829,1008,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">C. interdigitalis</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC7658BFF939C72516DA" box="[1020,1149,1734,1758]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">C. lateralis</taxonomicName>
,
</emphasis>
83 additional congeners, 26 gecko species as near outgroups, and
<taxonomicName id="FE8BB0DAFFD1FC765919F915C0681707" box="[602,816,1771,1795]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="carolinensis">
<emphasis id="0BFF174BFFD1FC765919F915C0681707" box="[602,816,1771,1795]" italics="true" pageId="13" pageNumber="530">Anolis carolinensis</emphasis>
</taxonomicName>
as a distant outgroup. We only discuss the region of the tree containing prehensile-tailed
<taxonomicName id="FE8BB0DAFFD1FC765947F8F0C1F91722" box="[516,673,1806,1830]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="genus">
<emphasis id="0BFF174BFFD1FC765947F8F0C1F91722" box="[516,673,1806,1830]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus</emphasis>
</taxonomicName>
and have collapsed several clades in figure 6 to make the phylogeny easier to read. The base of the tree, including
<emphasis id="0BFF174BFFD1FC7659E7F8CDC08A174F" box="[676,978,1843,1867]" italics="true" pageId="13" pageNumber="530">
<taxonomicName id="FE8BB0DAFFD1FC7659E7F8CDC06D174F" box="[676,821,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="angularis">C. angularis</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD1FC765801F8CDC095174F" box="[834,973,1843,1867]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="tibetanus">C. tibetanus</taxonomicName>
,
</emphasis>
outgroups, and a clade of the
<collectingCountry id="419C8BC9FFD1FC765E6CF8CDC6C4174F" box="[1327,1436,1843,1867]" name="Myanmar" pageId="13" pageNumber="530">Myanmar</collectingCountry>
species, is not shown. Overall, our ND2 phylogenies had high bootstrap support and posterior probabilities; it showed substantial congruence with the phylogeny of
<bibRefCitation id="5D1AB6A8FFD1FC7659BBF884C0961797" box="[760,974,1914,1939]" pageId="13" pageNumber="530" refString="Wood, P. L. Jr., Heinicke, M. P., Jackman, T. R. &amp; Bauer, A. M. (2012) Phylogeny of bent-toed geckos (Cyrtodactylus) reveals a west to east pattern of diversification. Molecular Phylogenetics and Evolution, 65, 992 - 1003. http: // dx. doi. org / 10.1016 / j. ympev. 2012.08.025" type="journal article">
Wood
<emphasis id="0BFF174BFFD1FC765802F885C0211797" box="[833,889,1915,1939]" italics="true" pageId="13" pageNumber="530">et al.</emphasis>
(2012)
</bibRefCitation>
.
</paragraph>
<paragraph id="3934CB59FFD1FC755B84F861C7221144" blockId="13.[151,1436,1734,2011]" lastBlockId="14.[151,1437,151,536]" lastPageId="14" lastPageNumber="531" pageId="13" pageNumber="530">
<collectingCountry id="419C8BC9FFD1FC765B84F861C3BE17B3" box="[199,230,1951,1975]" name="American Samoa" pageId="13" pageNumber="530">As</collectingCountry>
for the COI gene, we found no evidence of a close relationship between
<taxonomicName id="FE8BB0DAFFD1FC765F56F860C64E17B2" box="[1045,1302,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD1FC765F56F860C64E17B2" box="[1045,1302,1950,1974]" italics="true" pageId="13" pageNumber="530">Cyrtodactylus lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC765E0FF861C6C417B2" box="[1356,1436,1950,1974]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC765E0FF861C6C417B2" box="[1356,1436,1950,1974]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
using the ND2 gene (
<figureCitation id="A1B0D7DCFFD1FC765AC9F83CC28817DF" box="[394,464,1986,2011]" captionStart="FIGURE 6" captionStartId="11.[151,249,1913,1935]" captionTargetBox="[221,1338,212,1853]" captionTargetId="figure@11.[206,1355,193,1868]" captionTargetPageId="11" captionText="FIGURE 6. Phylogenetic relationships of Cyrtodactylus with spinose, prehensile tails. (A) Phylogeny based on morphology with three species added to original hypothesis of Grismer et al. (2010). (B) Clade pruned from our larger phylogeny based on sequences of the NADH Dehydrogenase 2 gene (ND 2). Tree constructed using Bayesian inference. Nodal values are posterior probabilities, and diamonds indicate prehensile-tailed Cyrtodactylus." httpUri="https://zenodo.org/record/256584/files/figure.png" pageId="13" pageNumber="530">Fig. 6</figureCitation>
). We found that
<taxonomicName id="FE8BB0DAFFD1FC7659CCF83DC1B817DF" box="[655,736,1987,2011]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD1FC7659CCF83DC1B817DF" box="[655,736,1987,2011]" italics="true" pageId="13" pageNumber="530">C. elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD1FC76585AF83DC09317DF" box="[793,971,1987,2011]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="530" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD1FC76585AF83DC09317DF" box="[793,971,1987,2011]" italics="true" pageId="13" pageNumber="530">C. interdigitalis</emphasis>
</taxonomicName>
are closely related and belong to a clade containing the
<taxonomicName id="FE8BB0DAFFD2FC755A03FF66C28D10B4" box="[320,469,152,176]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="0BFF174BFFD2FC755A03FF66C28D10B4" box="[320,469,152,176]" italics="true" pageId="14" pageNumber="531">C. pulchellus</emphasis>
</taxonomicName>
Group, a large radiation of 18 Australo-Papuan species, and the mainland species
<emphasis id="0BFF174BFFD2FC755EC1FF66C1B310D0" italics="true" pageId="14" pageNumber="531">
<taxonomicName id="FE8BB0DAFFD2FC755EC1FF66C3AE10D0" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="oldhami">C. oldhami</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD2FC755A42FF42C2D310D0" box="[257,395,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="tygroides">C. tygroides</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD2FC755AD4FF42C16210D0" box="[407,570,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="hontreensis">C. hontreensis</taxonomicName>
,
<taxonomicName id="FE8BB0DAFFD2FC755905FF42C1B310D0" box="[582,747,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="intermedius">C. intermedius</taxonomicName>
</emphasis>
, and
<taxonomicName id="FE8BB0DAFFD2FC75586BFF42C0EF10D0" box="[808,951,188,212]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="0BFF174BFFD2FC75586BFF42C0EF10D0" box="[808,951,188,212]" italics="true" pageId="14" pageNumber="531">C. peguensis</emphasis>
</taxonomicName>
. This result agrees with that of
<bibRefCitation id="5D1AB6A8FFD2FC755E58FF42C3BC10FC" pageId="14" pageNumber="531" refString="Wood, P. L. Jr., Heinicke, M. P., Jackman, T. R. &amp; Bauer, A. M. (2012) Phylogeny of bent-toed geckos (Cyrtodactylus) reveals a west to east pattern of diversification. Molecular Phylogenetics and Evolution, 65, 992 - 1003. http: // dx. doi. org / 10.1016 / j. ympev. 2012.08.025" type="journal article">
Wood
<emphasis id="0BFF174BFFD2FC755E27FF43C6C410D0" box="[1380,1436,188,212]" italics="true" pageId="14" pageNumber="531">et al.</emphasis>
(2012)
</bibRefCitation>
, although they found the
<taxonomicName id="FE8BB0DAFFD2FC755943FF1EC1CB10FC" box="[512,659,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="pulchellus">
<emphasis id="0BFF174BFFD2FC755943FF1EC1CB10FC" box="[512,659,224,248]" italics="true" pageId="14" pageNumber="531">C. pulchellus</emphasis>
</taxonomicName>
Group to be sister to the
<taxonomicName id="FE8BB0DAFFD2FC7558EEFF1EC0A410FC" box="[941,1020,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD2FC7558EEFF1EC0A410FC" box="[941,1020,224,248]" italics="true" pageId="14" pageNumber="531">C. elok</emphasis>
</taxonomicName>
+
<taxonomicName id="FE8BB0DAFFD2FC755F5AFF1EC79210FC" box="[1049,1226,224,248]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD2FC755F5AFF1EC79210FC" box="[1049,1226,224,248]" italics="true" pageId="14" pageNumber="531">C. interdigitalis</emphasis>
</taxonomicName>
clade, whereas we found the mainland species to be sister to this pair. Nonetheless,
<taxonomicName id="FE8BB0DAFFD2FC75583BFEFAC0911118" box="[888,969,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD2FC75583BFEFAC0911118" box="[888,969,260,284]" italics="true" pageId="14" pageNumber="531">C. elok</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD2FC755F42FEFAC7EB1118" box="[1025,1203,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD2FC755F42FEFAC7EB1118" box="[1025,1203,260,284]" italics="true" pageId="14" pageNumber="531">C. interdigitalis</emphasis>
</taxonomicName>
have relatively long branch lengths and an uncorrected genetic distance of 22.9% from one another (
<tableCitation id="7409FEE2FFD2FC755F56FED9C7351144" box="[1045,1133,295,320]" captionStart="TABLE 2" captionStartId="13.[151,236,1196,1219]" captionTargetBox="[159,1383,1310,1664]" captionTargetPageId="13" captionText="TABLE 2. Uncorrected pairwise distances (percent difference) of mitochondrial loci for representative individuals of Malaysian and Indonesian Cyrtodactylus. The upper diagonal shows distances for Cytochrome c oxidase subunit I; the lower diagonal shows distances for NADH dehydrogenase 2 gene sequences." httpUri="http://table.plazi.org/id/6DF49BD1FFD1FC765BD4FB52C0961507" pageId="14" pageNumber="531" tableUuid="6DF49BD1FFD1FC765BD4FB52C0961507">Table 2</tableCitation>
).
</paragraph>
<paragraph id="3934CB59FFD2FC755B84FEB3C2C8121C" blockId="14.[151,1437,151,536]" pageId="14" pageNumber="531">
In contrast,
<taxonomicName id="FE8BB0DAFFD2FC755A10FEB2C1001160" box="[339,600,332,356]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD2FC755A10FEB2C1001160" box="[339,600,332,356]" italics="true" pageId="14" pageNumber="531">Cyrtodactylus lateralis</emphasis>
</taxonomicName>
belongs to a large clade of mostly Sunda Shelf and
<collectingCountry id="419C8BC9FFD2FC755F84FEB2C6621161" box="[1223,1338,332,357]" name="Philippines" pageId="14" pageNumber="531">Philippine</collectingCountry>
species.
<taxonomicName id="FE8BB0DAFFD2FC755BD4FE8EC2C1118C" box="[151,409,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD2FC755BD4FE8EC2C1118C" box="[151,409,368,392]" italics="true" pageId="14" pageNumber="531">Cyrtodactylus lateralis</emphasis>
</taxonomicName>
is sister to
<taxonomicName id="FE8BB0DAFFD2FC75595CFE8EC125118C" box="[543,637,368,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="durio">
<emphasis id="0BFF174BFFD2FC75595CFE8EC125118C" box="[543,637,368,392]" italics="true" pageId="14" pageNumber="531">C. durio</emphasis>
</taxonomicName>
and their ND2 sequences have diverged by 11.7%. These two species have yet to show any clear affinitiess to three clades in the crown clade of figure 6, and we show all four lineages as an unresolved polytomy. The distant relationship of
<taxonomicName id="FE8BB0DAFFD2FC7559B9FE46C02311D4" box="[762,891,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD2FC7559B9FE46C02311D4" box="[762,891,440,464]" italics="true" pageId="14" pageNumber="531">C. lateralis</emphasis>
</taxonomicName>
to the
<taxonomicName id="FE8BB0DAFFD2FC755897FE46C77F11D4" box="[980,1063,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD2FC755897FE46C77F11D4" box="[980,1063,440,464]" italics="true" pageId="14" pageNumber="531">C. elok</emphasis>
</taxonomicName>
+
<taxonomicName id="FE8BB0DAFFD2FC755F0FFE46C65911D4" box="[1100,1281,440,464]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD2FC755F0FFE46C65911D4" box="[1100,1281,440,464]" italics="true" pageId="14" pageNumber="531">C. interdigitalis</emphasis>
</taxonomicName>
clade is also reflected in high uncorrected genetic distances between
<taxonomicName id="FE8BB0DAFFD2FC755859FE22C0C211F0" box="[794,922,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD2FC755859FE22C0C211F0" box="[794,922,476,500]" italics="true" pageId="14" pageNumber="531">C. lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD2FC755895FE22C77011F0" box="[982,1064,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="elok">
<emphasis id="0BFF174BFFD2FC755895FE22C77011F0" box="[982,1064,476,500]" italics="true" pageId="14" pageNumber="531">C. elok</emphasis>
</taxonomicName>
(27.6%) and
<taxonomicName id="FE8BB0DAFFD2FC755F84FE22C61F11F0" box="[1223,1351,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="0BFF174BFFD2FC755F84FE22C61F11F0" box="[1223,1351,476,500]" italics="true" pageId="14" pageNumber="531">C. lateralis</emphasis>
</taxonomicName>
and
<taxonomicName id="FE8BB0DAFFD2FC755EC1FE22C27F121C" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="531" phylum="Chordata" rank="species" species="interdigitalis">
<emphasis id="0BFF174BFFD2FC755EC1FE22C27F121C" italics="true" pageId="14" pageNumber="531">C. interdigitalis</emphasis>
</taxonomicName>
(26.7%).
</paragraph>
</subSubSection>
</treatment>
</document>