259 lines
36 KiB
XML
259 lines
36 KiB
XML
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<mods:title id="7BC34DC627131FFC004A21DDE2A4CFC7">Manual of North American Agromyzidae (Diptera, Schizophora), with revision of the fauna of the " Delmarva " states</mods:title>
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<mods:namePart id="009AA1D4C8CC2FA3031A85DFC4E88EBF">Lonsdale, Owen</mods:namePart>
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<mods:affiliation id="D73A121B4E6D44F493F9A69C1EB2975A">Agriculture & Agri-Food Canada, 960 Carling Avenue, Ottawa, ON, K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:nameIdentifier id="BDF6E9A4BB13B6B7A9ABEC6BFFCA68A3" type="email">neoxabea@hotmail.com</mods:nameIdentifier>
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<mods:date id="877ACEF24F3F6D8CB095EC56B84E6441">2021</mods:date>
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<mods:identifier id="DB165439EE5356B5D86060FE8BFD05EF" type="DOI">http://dx.doi.org/10.3897/zookeys.1051.64603</mods:identifier>
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<treatment id="5A3D36355C6E1A29DC20BEE5C76B0268" ID-GBIF-Taxon="183833215" LSID="urn:lsid:plazi:treatment:5A3D36355C6E1A29DC20BEE5C76B0268" httpUri="http://treatment.plazi.org/id/5A3D36355C6E1A29DC20BEE5C76B0268" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Agromyzidae" scope_order="Diptera" scope_subFamily="Agromyzinae">
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<subSubSection id="0E5021C5D64FC2D9B9301A6FCA9C3669" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="F83A81F47FC4F553DEF1672A216F8CB5" pageId="0" pageNumber="1">
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<taxonomicName id="57CF1CDA5B4C4D6163E81DFBA367D3E4" ID-CoL="B8FP" authority="Spencer" authorityName="Spencer" authorityYear="1969" class="Insecta" family="Agromyzidae" genus="Agromyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agromyza bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bispinata">Agromyza bispinata Spencer</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="0AB7B809CF6FAD249681E7F5A93CB840" pageId="0" pageNumber="1" type="description">
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<paragraph id="905E72E6E4985D2FB43F66F9F13BB339" pageId="0" pageNumber="1">
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<figureCitation id="2068C0ABD61ACAB48AED6D6A8010C203" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">Figs 38</figureCitation>
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<figureCitation id="4E8310D2F540C88D1C7220BA290E5DCB" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">, 41</figureCitation>
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<figureCitation id="B838AFDFDE9912F22FA59EAEDF62225A" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">, 42</figureCitation>
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<figureCitation id="CD20D2ED8ABF154EFF00C4495D03D1C7" captionStart="Figures 167–174" captionStartId="F24" captionText="Figures 167 - 174. Agromyza bispinata Spencer, " typical " male genitalia 167 hypandrium and postgonite, ventral 168 phallus, ventral 169 hypandrial complex, left lateral 170 external genitalia, left lateral 171 external genitalia, posterior 172 external genitalia, anterior 173 external genitalia, ventral 174 A. bispinata, longer, multi-chambered " atypical " male (Maryland specimen), phallus, ventral." figureDoi="10.3897/zookeys.1051.64603.figures167-174" httpUri="https://binary.pensoft.net/fig/572123" pageId="0" pageNumber="1">, 167-174</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="5411CF9E4C74769C575E5D238E19E47E" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="E5B94BB949E72BEBD4BAAEE54CA2FF42" pageId="0" pageNumber="1">
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<taxonomicName id="D79D7A89F273DB7D4802887D6FD9AD4C" authorityName="Spencer" authorityYear="1969" class="Insecta" family="Agromyzidae" genus="Agromyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agromyza bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bispinata">Agromyza bispinata</taxonomicName>
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Spencer, 1969: 39.
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<bibRefCitation id="A4124CD7469CAE55F4BDDA14FD715891" author="Spencer, KA" journalOrPublisher="Agriculture Handbook" pageId="0" pageNumber="1" pagination="1 - 478" publicationUrl="https://www.biodiversitylibrary.org/page/51258090" refId="B261" refString="Spencer, KA, Steyskal, GC, 1986b. Manual of the Agromyzidae (Diptera) of the United States. U.S.D.A. Agriculture Handbook 638: 1 - 478, https://www.biodiversitylibrary.org/page/51258090" title="Manual of the Agromyzidae (Diptera) of the United States. U. S. D. A." url="https://www.biodiversitylibrary.org/page/51258090" volume="638" year="1986 b">Spencer and Steyskal 1986b</bibRefCitation>
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: 263;
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<bibRefCitation id="104F0D1542C3F34227A7CEEF64FE2E8C" DOI="https://doi.org/10.11646/zootaxa.4479.1.1" author="Eiseman, CS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 156" refId="B49" refString="Eiseman, CS, Lonsdale, O, 2018. New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species. Zootaxa 4479 (1): 1 - 156, DOI: https://doi.org/10.11646/zootaxa.4479.1.1" title="New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species." url="https://doi.org/10.11646/zootaxa.4479.1.1" volume="4479" year="2018">Eiseman and Lonsdale 2018</bibRefCitation>
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: 8.
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</paragraph>
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</subSubSection>
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<subSubSection id="BCF69CC87A3B7188B419B075A5DF217F" pageId="0" pageNumber="1" type="description">
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<paragraph id="9FD46246E0EBFFB883F13FD61E9F5508" pageId="0" pageNumber="1">Description</paragraph>
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<paragraph id="71068114F2E9356D7C331B47E10D9A12" pageId="0" pageNumber="1">
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<emphasis id="4DD8AE8983F1C501E4867EA02E5D69EA" bold="true" pageId="0" pageNumber="1">
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(Figs
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<figureCitation id="E6FCF94C0C927AFD8FD373715A7E722F" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">38</figureCitation>
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,
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<figureCitation id="99399117CE072E58B5389B81CCA3634F" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">41</figureCitation>
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,
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<figureCitation id="D1F5117891ED83C14EEE4A34C3617E67" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">42</figureCitation>
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).
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</emphasis>
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Wing length 2.2-2.4 mm (♂). Female unknown [see discussion below]. Length of ultimate section of vein M4 divided by penultimate section: 0.5-0.7. Eye height divided by gena height: 4.2-6.0. Male first flagellomere large (usually at least 30% longer than pedicel, but not much longer than pedicel in a minority of specimens) and varying in shape from elongate and subovate to (more commonly) circular; at least distal 1/2 (usually distal 2/3) covered with long hairs. Ocellar triangle relatively small with corners slightly rounded. Fronto-orbital plate slightly projecting.
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</paragraph>
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<paragraph id="2B37AF917F45A58C42C812553AE152CC" pageId="0" pageNumber="1">
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<emphasis id="BD0AF0137A74F9078E5A4736D40F368A" bold="true" italics="true" pageId="0" pageNumber="1">Chaetotaxy</emphasis>
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: Two ori; two ors. Three dorsocentrals, decreasing in length anteriorly with anterior seta not much longer than surrounding setulae. Mid tibia with two posteromedial setae.
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</paragraph>
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<paragraph id="A1C39F2247D3165584A3DEE5E14336A5" pageId="0" pageNumber="1">
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<emphasis id="3CC2EBB93491189C7AC98DD043B193A2" bold="true" italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
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: Body primarily dark brown with halter white. Base of first flagellomere and distal margin of pedicel orange; orange region sometimes either extending to basal 1/3 of first flagellomere or strongly reduced (particularly if spines on surstylus reduced). Gena sometimes paler, excluding ventral margin. Calypter white with margin sometimes yellowish or slightly brown, and hairs light brown. Base of fore tibia paler or segment paler overall. Tarsi yellow to orange-brown.
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</paragraph>
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<paragraph id="60C442A30C974A54D600E6CD329A83AC" pageId="0" pageNumber="1">
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<emphasis id="4180844560418F30C1A1084FE404B37B" bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
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: (Figs
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<figureCitation id="4E6FD0BD5812CB90ED9844A818F3ED07" captionStart="Figures 167–174" captionStartId="F24" captionText="Figures 167 - 174. Agromyza bispinata Spencer, " typical " male genitalia 167 hypandrium and postgonite, ventral 168 phallus, ventral 169 hypandrial complex, left lateral 170 external genitalia, left lateral 171 external genitalia, posterior 172 external genitalia, anterior 173 external genitalia, ventral 174 A. bispinata, longer, multi-chambered " atypical " male (Maryland specimen), phallus, ventral." figureDoi="10.3897/zookeys.1051.64603.figures167-174" httpUri="https://binary.pensoft.net/fig/572123" pageId="0" pageNumber="1">167-173</figureCitation>
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) Surstylus small, lobate, margin darkly pigmented; with two or three medial setae and two large posterior spines; spines sometimes minute and rounded and distiphallus more elongate with relatively pronounced internally haired medial section; surstylus barely visible when viewed laterally and with basal suture obliterated. Hypandrium relatively narrow and tapered apically with membranous window in lobe. Postgonite relatively small and upcurved. Halves of basiphallus converging to, and overlapping at base; with small mediolateral membranous lobe on each sclerite; twisted medially and broad apically, with converging ventrodistal points. Distiphallus of "
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<taxonomicName id="C2061A21B5CB2CE4189C17EE0E05CB61" authorityName="Meigen" authorityYear="1830" class="Insecta" family="Boraginaceae" genus="Agromyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agromyza nigripes" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nigripes">
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<emphasis id="8D05577587221E2680ACB6E1521E17D5" italics="true" pageId="0" pageNumber="1">Agromyza nigripes</emphasis>
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</taxonomicName>
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-type" (capsule-shaped with subbasal opening for entry of ejaculatory duct, pronounced dorsobasal collar, and medial convolution); relatively narrow and pale with sides parallel, being only slightly wider apically.
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</paragraph>
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<paragraph id="AF1E643E5B1148D994531FFEEAE1BCCE" pageId="0" pageNumber="1">
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<emphasis id="869F83FDBA68CF14F56CC25F1847BDBD" bold="true" italics="true" pageId="0" pageNumber="1">Variation</emphasis>
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: Phallus sometimes elongate with pronounced medial section with densely spinulose inner surface and elongate medial membranous projection on right sclerite of mesophallus (Fig.
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<figureCitation id="F7E5A1B2D914E348BFB5AE737846FA2C" captionStart="Figures 167–174" captionStartId="F24" captionText="Figures 167 - 174. Agromyza bispinata Spencer, " typical " male genitalia 167 hypandrium and postgonite, ventral 168 phallus, ventral 169 hypandrial complex, left lateral 170 external genitalia, left lateral 171 external genitalia, posterior 172 external genitalia, anterior 173 external genitalia, ventral 174 A. bispinata, longer, multi-chambered " atypical " male (Maryland specimen), phallus, ventral." figureDoi="10.3897/zookeys.1051.64603.figures167-174" httpUri="https://binary.pensoft.net/fig/572123" pageId="0" pageNumber="1">174</figureCitation>
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); if so, spines on epandrium reduced to absent. Possibly a new species, but
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<normalizedToken id="1C2184C805C95F67D968BD980AFBB5B9" originalValue="“typical”">"typical"</normalizedToken>
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males show intermediate degrees of spine reduction on epandrium that resemble the state seen in these other males.
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</paragraph>
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</subSubSection>
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<subSubSection id="ADDF91704DF718A00A50490E0B6F4BA6" pageId="0" pageNumber="1" type="host">
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<paragraph id="E0A37F1FA0DF080790CFB4F540FE9311" pageId="0" pageNumber="1">Host.</paragraph>
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<paragraph id="97F3B7AB7296213FEB76D39AD033FFD9" pageId="0" pageNumber="1">
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<taxonomicName id="4339550C34181DA62FEE6D0AB714D812" class="Insecta" family="Poaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Poaceae</taxonomicName>
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-
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<taxonomicName id="F84D80FD7169869EE7ACB72618938779" class="Monocotyledoneae" family="Poaceae" genus="Elymus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Elymus hystrix" order="Graminales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="hystrix">
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<emphasis id="241BB7C9273ABB2A6D29005C0B20DA22" italics="true" pageId="0" pageNumber="1">Elymus hystrix</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="B5662334823DC89E5C4AD82C2EFD4223" DOI="https://doi.org/10.11646/zootaxa.4479.1.1" author="Eiseman, CS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 156" refId="B49" refString="Eiseman, CS, Lonsdale, O, 2018. New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species. Zootaxa 4479 (1): 1 - 156, DOI: https://doi.org/10.11646/zootaxa.4479.1.1" title="New state and host records for Agromyzidae (Diptera) in the United States, with the description of thirty new species." url="https://doi.org/10.11646/zootaxa.4479.1.1" volume="4479" year="2018">Eiseman and Lonsdale 2018</bibRefCitation>
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||
).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="F28889B4FFD89F7616B84BCC068F8E2E" pageId="0" pageNumber="1" type="distribution">
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<paragraph id="BEE412DA0BA2DE0F440FAB0A5991E3B8" pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph id="1755D34AF2B856B515446DDB176F7D67" pageId="0" pageNumber="1">
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<emphasis id="8FD5DE84E9D145C18E2C27833BCC91E1" bold="true" pageId="0" pageNumber="1">Canada</emphasis>
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: ON, MB*.
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<emphasis id="351CBFCE9F1881BEFAAEA82E1DC49321" bold="true" pageId="0" pageNumber="1">USA</emphasis>
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: CT*, GA, IA, MD, NC, NH*, NY, PA*, UT, VA, WV*.
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</paragraph>
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</subSubSection>
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<subSubSection id="03CAF6995159B6388BDE01191E53CA18" pageId="0" pageNumber="1" type="type material">
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<paragraph id="5561F0CD2E55A9371D40D3596506053C" pageId="0" pageNumber="1">Type material.</paragraph>
|
||
<paragraph id="C55512C8467AC2B628B97AA46A3E7BE9" pageId="0" pageNumber="1">
|
||
<emphasis id="3AE9D9FA3879ECA07DE6B6BCE6979BCB" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="F6A70C67D35C691A67E86F3E5F5230AD" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
|
||
: Canada. ON
|
||
</emphasis>
|
||
: Simcoe, 5.vi.1939, G.S. Steyskal (1♂, CNC).
|
||
</paragraph>
|
||
<paragraph id="BCA9347B52166C288ADC0F0AEFC13A64" pageId="0" pageNumber="1">
|
||
<emphasis id="770A5DF17FCA1CF4E88ABD8E2011CFB5" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
|
||
:
|
||
<emphasis id="C648B494737B04D7823752600C7A03E8" bold="true" pageId="0" pageNumber="1">Canada. ON</emphasis>
|
||
: Midland, 20.vii.1955, J.G. Chillcott, CNC352821 (1♂, CNC), Pt. Ryerse, 1.vi.1939, G.E. Shewell, CNC352822, (1♂, CNC), Simcoe, 5.vi.1939, G.E. Shewell, CNC352819 (1♂, CNC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DE32744D2DC81E73BCECD63F6A80532" pageId="0" pageNumber="1" type="males examined">
|
||
<paragraph id="C6F340CFEBD05ECC4AA84F94A4A187D6" pageId="0" pageNumber="1">Males examined.</paragraph>
|
||
<paragraph id="9402FB05C8B66E8F07ECFB4EEE7B7556" pageId="0" pageNumber="1">
|
||
<emphasis id="E08AB32BEE21411095860C666BFDDD42" bold="true" pageId="0" pageNumber="1">Canada. MB</emphasis>
|
||
: Ninette, 28.vii.1958, "sq. fringe darker spines on surs. larger aed. larger", J.G. Chillcott, CNC352820 (1♂, CNC).
|
||
<emphasis id="383D3BBA1A4062DD44762DA756F815B4" bold="true" pageId="0" pageNumber="1">USA. CT</emphasis>
|
||
: Redding, 11.vi.1929, A.L. Melander (1♂, USNM), Rabin Co., 13.vii.1957, W.R. Richards, CNC352824 (1♂, CNC),
|
||
<emphasis id="428E12055F8A2BF2BF54EBFC867D351C" bold="true" pageId="0" pageNumber="1">IA</emphasis>
|
||
: Winneshiek Co.,
|
||
<geoCoordinate id="FBB929E5027D2B3FD5D6E332B0D98939" degrees="43" direction="north" minutes="25" orientation="latitude" precision="1" seconds="55.97" value="43.432217">43°25'55.97"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3347065E3A32BB1F94AF94D325D51887" degrees="92" direction="west" minutes="0" orientation="longitude" precision="1" seconds="34.78" value="-92.00966">92°0'34.78"W</geoCoordinate>
|
||
, 16.vii.2015, C.S. Eiseman,
|
||
<taxonomicName id="F2CAE8CB03CA57C83D648E446A75389F" class="Monocotyledoneae" family="Poaceae" genus="Elymus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Elymus hystrix" order="Graminales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="hystrix">
|
||
<emphasis id="4974D56AEB3E21F2D487475708678CEE" italics="true" pageId="0" pageNumber="1">Elymus hystrix</emphasis>
|
||
</taxonomicName>
|
||
em. 7.viii.2015, #CSE1973, CNC564662 (1♂, CNC),
|
||
<emphasis id="2B62DC3E7041E83C1A23CD700FB0C07A" bold="true" pageId="0" pageNumber="1">MD</emphasis>
|
||
: Bethseda, 30.v.1980, G.C. Steyskal, swept ex.
|
||
<taxonomicName id="DB145C1FF6775C36D0DA434A38A7AF97" class="Insecta" family="Hesperiidae" genus="Viola" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Viola papilionaceae" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="papilionaceae">
|
||
<emphasis id="2EDBA976DB24A10479D68E3176BA693C" italics="true" pageId="0" pageNumber="1">Viola papilionaceae</emphasis>
|
||
</taxonomicName>
|
||
(2♂, USNM), Bethseda, 14.v.1981, G.C. Steyskal (1♂, USNM), 11.viii.1981 (1♂, USNM), 14.viii.1981 (2♂, USNM), 16.viii.1981 (1♂, USNM), 1.viii.1981 (3♂, USNM), 17.v.1969 (1♂, USNM), 7.viii.1981 (2♂, USNM), 13.ix.1981 (1♂, USNM), 27.v.1972 (1♂, USNM), Plummers Isl., 19.vi.1913, R.C. Shannon (1♂, USNM), Montgomery Co., Dickerson, 14.vii1974, G.A. Foster (13♂, USNM), 4mi SW of Ashton, G.F. and J.F. Hevel, 16.viii.1986 (1♂, USNM), 24.vii.1982 (1♂, USNM), 27.viii.1981 (1♂, USNM), Cabin John, 20.vi.1931, A.L. Melander (1♂, USNM),
|
||
<emphasis id="C363618B5B460AD6A73CC3412D8B204F" bold="true" pageId="0" pageNumber="1">NC</emphasis>
|
||
: Macon Co., Wayah Gap, 3800', 29.vii.1957, J.G. Chillcott (1♂, USNM), Jackson Co., Cherokee, 609 m, 25.vii.1957, CNC352823 (1♂, CNC),
|
||
<emphasis id="8A1F8049A05C8C91550813D4CA256829" bold="true" pageId="0" pageNumber="1">NH</emphasis>
|
||
: Bretton Wda, 1.vii.1936, A.L. Melander (1♂, USNM),
|
||
<emphasis id="961C43D9F81806F3FFA7298F1244B0D0" bold="true" pageId="0" pageNumber="1">PA</emphasis>
|
||
: Allegeny Co., Little Orleans, Little Orleans campground, 6.vi.1998, sweeping, C.R. Bartlett (1♂, UDCC),
|
||
<emphasis id="B30CBF934DC212270E68A8C099416F49" bold="true" pageId="0" pageNumber="1">VA</emphasis>
|
||
: Falls Church, Holmes Run, 26.vi.1961, light trap, W.W. Wirth (1♂, USNM), Shenandoah, Big. Meadows, A.L. Melander, 1.vii.1939 (1♂, USNM), 2.vii.1939 (2♂, USNM), 5.vii.1939 (1♂, USNM), Great Falls, 9.vii.1926, A.L. Melander (1♂, USNM), Glencarlyn, 2.vi.1925, J.R. Malloch (1♂, USNM), Fairfax Co., Dead Run, 22.vi.1916, R.C. Shannon (1♂, USNM), Fairfax Co., Turkey Run Park, nr. mouth of Turkey Run,
|
||
<geoCoordinate id="9E676A4EC8D367AB89013192DD81D753" degrees="38" direction="north" minutes="57.9" orientation="latitude" precision="92" value="38.965">38°57.9'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="30D830A1A2D3949146A6B6ADFBD130B7" degrees="7" direction="west" minutes="09.4" orientation="longitude" precision="92" value="-7.1566668">7°09.4'W</geoCoordinate>
|
||
, Malaise trap, 18-30.v.2007, D.R. Smith (1♂, USNM),
|
||
<emphasis id="AD65724DE73D96CA5EEBBE38DD6E4DEB" bold="true" pageId="0" pageNumber="1">WV</emphasis>
|
||
: Parkersburg, 21.vi.1970, G. Steyskal (1♂, USNM), Morgan Co., nr. Great Cacapon, 1.ix.1984, G.F. and J.F. Hevel (1♂, USNM), Hardy Co., Lost River St. Pk., 1-14.viii.1960, K.V. Krombein (1♂, USNM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3E88069E99998E93321F38E751067BB6" pageId="0" pageNumber="1" type="females examined">
|
||
<paragraph id="9C43E49CD404CC856C46F26E2E340A82" pageId="0" pageNumber="1">Females examined.</paragraph>
|
||
<paragraph id="E12A72917091A195E9A32F99F7AAFCCB" pageId="0" pageNumber="1">
|
||
<emphasis id="B65421A71479C0A5532589728B92DF67" bold="true" pageId="0" pageNumber="1">USA. CT</emphasis>
|
||
: Colebrook, 28.viii.1941, A.L. Melander (1♀, USNM),
|
||
<emphasis id="DBC2A0BB799663C06B743F746CFBB5C5" bold="true" pageId="0" pageNumber="1">MD</emphasis>
|
||
: Coleville, W.W. Wirth, 21.v.1977 (1♀, USNM), 11.v.1977 (1♀, USNM),
|
||
<emphasis id="34B2373CC89B14D90EAC3367288081D9" bold="true" pageId="0" pageNumber="1">NC</emphasis>
|
||
: Macon Co., Wayah Bald, 1066 m, 13.vii.1957, W.R. Richards, CNC352828 (1♀, CNC), Macon Co., Wayah Gap, 16.vii.1957, J.G. Chillcott, CNC352827 (1♀, CNC),
|
||
<emphasis id="38F3AFB1AF074F2F760DB411FC35A360" bold="true" pageId="0" pageNumber="1">NH</emphasis>
|
||
: Franconia Notch, 8.vii.1931, J.M. Aldrich (1♀, USNM),
|
||
<emphasis id="EEA6EE89880B0588771505081B395560" bold="true" pageId="0" pageNumber="1">NY</emphasis>
|
||
: Ithaca, 31.v.1914, A.L. Melander (3♀, USNM),
|
||
<emphasis id="DA0E6F3BE3C298E423D34350DD70C415" bold="true" pageId="0" pageNumber="1">TN</emphasis>
|
||
: Smokies, Chimneys, 25.vi.1941, A.L. Melander (1♀, USNM),
|
||
<emphasis id="A34666EAF489F25D6873DF83EC4390B1" bold="true" pageId="0" pageNumber="1">VA</emphasis>
|
||
: Falls Church, 18.vii.1960, W.W. Wirth (1♀, USNM), Shenandoah, Big Meadows, A.L. Melander, 3.vii.1939 (1♀, USNM), 2.vii.1929 (1♀, USNM), Shenandoah, Lewis falls, 3.vii.1939, A.L. Melander (1♀, USNM), Fairfax Co., Dead Run, 28.vii.1915, R.C. Shannon (1♀, USNM), Shenandoah Co., Mt. Jackson, 25.v.1962, J.G. Chillcott, J.R. Vockeroth, CNC352825, CNC352826 (2♀, CNC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8C0E390EEE47424E003F67C3056F1B96" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="32013FCE2FFB5E03629CC4A83D3D9900" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="83EBA71923E1D47E16C25DE5C1AEC737" pageId="0" pageNumber="1">
|
||
<taxonomicName id="E35CB4642CD19A17AC3D2127B9BC4852" authorityName="Spencer" authorityYear="1969" class="Insecta" family="Boraginaceae" genus="Agromyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agromyza bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bispinata">
|
||
<emphasis id="43073EA81EF99A1A3335D46A007318C6" italics="true" pageId="0" pageNumber="1">Agromyza bispinata</emphasis>
|
||
</taxonomicName>
|
||
belongs to a complex of species defined by an enlarged and long-haired male first flagellomere, whose species appear to be separable only on the basis of male genitalic morphology. The females of these species, which are currently indistinguishable from one another, differ from the males in having an elongate oval first flagellomere with the long pale hairs restricted to the distal margin; the examined females are listed in the material examined section for this species. The sparse, long hairs on the first flagellomere of these species (Figs
|
||
<figureCitation id="C2397279263079F41E9444B484F293DA" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">41</figureCitation>
|
||
,
|
||
<figureCitation id="9EB1D59AE8A46C54524ACCB19299646F" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">42</figureCitation>
|
||
) should not be confused with the discrete apical tuft of hairs characteristic of many other
|
||
<taxonomicName id="51CE8506529531357A3E4DB2CE7F2B05" class="Insecta" family="Boraginaceae" genus="Agromyza" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agromyza" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4ACCD3334F112614F60F2C0ED1116FD6" italics="true" pageId="0" pageNumber="1">Agromyza</emphasis>
|
||
</taxonomicName>
|
||
species, including
|
||
<taxonomicName id="D5D290386992B27BE8194E58869E04DA" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. canadensis" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="canadensis">
|
||
<emphasis id="EF2D867C256D54BCFD66162AD3E8FCAB" italics="true" pageId="0" pageNumber="1">A. canadensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="1953165CD50D7630D87D73E326236273" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. kincaidi" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="kincaidi">
|
||
<emphasis id="2AA6737273BD713FD2D5C70D9447268F" italics="true" pageId="0" pageNumber="1">A. kincaidi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="B60A6682C88153AF6105E220B2F0873F" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. pudica" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pudica">
|
||
<emphasis id="78087D3F79374B51E29C59DB5660745B" italics="true" pageId="0" pageNumber="1">A. pudica</emphasis>
|
||
</taxonomicName>
|
||
(see Figs
|
||
<figureCitation id="54B58FB779C42E306107427BB33925ED" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">39</figureCitation>
|
||
,
|
||
<figureCitation id="A56E75178AA5E8DE558741420AEE0840" captionStart="Figures 37–44" captionStartId="F5" captionText="Figures 37 - 44. 37 Agromyza aristata Malloch, female lateral 38 Agromyza bispinata Spencer, female lateral with detail of stridulatory file 39, 40 Agromyza kincaidi Malloch, male head (arrow indicates discrete apical tuft of hairs) 39 lateral 40 same, anterior 41, 42 Agromyza bispinata, male head 41 lateral 42 same, anterior 43 Agromyza sp., female (S. A. Marshall - Ontario, Hornings Mills) 44 Euhexomyza schineri (Giraud) (T. Murray - Massachusetts, Groton)." figureDoi="10.3897/zookeys.1051.64603.figures37-44" httpUri="https://binary.pensoft.net/fig/572104" pageId="0" pageNumber="1">40</figureCitation>
|
||
). This complex currently includes
|
||
<taxonomicName id="0FF10F5DFF0054D122249A962CEE0E6B" authority="hockingi" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. echinalis subsp. A." order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="echinalis" subSpecies="A.">A. echinalis in the east Nearctic, A. hockingi</taxonomicName>
|
||
Spencer in Canada and the western United States (but expected in the eastern United States), and the widespread
|
||
<taxonomicName id="4A72BBC53C343715A1A80D6C7F9746CE" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. tacita" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tacita">
|
||
<emphasis id="711480B1D14A050D685F68D9860FE878" italics="true" pageId="0" pageNumber="1">A. tacita</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D944B82B7EB54CC2A70D650D423BF9C1" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bispinata">
|
||
<emphasis id="EA5180C0A8335722D2CFDE7F88D2EB87" italics="true" pageId="0" pageNumber="1">A. bispinata</emphasis>
|
||
</taxonomicName>
|
||
. There is a relatively large amount of genitalic variation within the latter two species, including distiphallus morphology in
|
||
<taxonomicName id="5B6B869097F9597B827C48FF3828470C" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bispinata">
|
||
<emphasis id="5FB03D0B937715C6C0F09D7C0B88435C" italics="true" pageId="0" pageNumber="1">A. bispinata</emphasis>
|
||
</taxonomicName>
|
||
and epandrial spine number in
|
||
<taxonomicName id="7B7097B1BA0AAE3D39704E437DDC8912" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. tacita" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="tacita">
|
||
<emphasis id="7AA43CA71B4B4A8CD604F4EF64DCC8AA" italics="true" pageId="0" pageNumber="1">A. tacita</emphasis>
|
||
</taxonomicName>
|
||
, which suggests the presence of cryptic taxa. The presence of only two epandrial spines, which is likely the derived state, currently defines
|
||
<taxonomicName id="0761F0B14C57E9A736FCBF3D5ED8AA91" class="Magnoliopsida" family="Amaranthaceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="A. bispinata" order="Diptera" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bispinata">
|
||
<emphasis id="28646318B254B791CFA3B87464C32840" italics="true" pageId="0" pageNumber="1">A. bispinata</emphasis>
|
||
</taxonomicName>
|
||
within this group, but more rigorous analysis of the clade as a whole is required to better delineate species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |