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<document id="858AC819B733BD280B853BB8A2E64929" ID-DOI="10.5281/zenodo.5355546" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355546" ID-ZooBank="43B1BFEE-71F6-4D05-813F-AA5357310547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630215601964" checkinUser="valdenar" docAuthor="Hughes, Lauren E. &amp; Kaji, Tomonari" docDate="2016" docId="283C0A34FF98274F57A3FEC21792F9C0" docLanguage="en" docName="64rbz351-359.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Quadrivisio meufong Hughes &amp; Kaji 2016, new species" docType="treatment" docUuid="CA01B47B-5DA7-4F66-9F5E-B6CF663A6B95" docUuidSource="ZooBank" docVersion="4" lastPageNumber="358" masterDocId="D405724CFF9A27485465FFDF154DFFA1" masterDocTitle="Description of a new species of Quadrivisio Stebbing, 1907, from Songkhla Lake, Thailand (Crustacea: Peracarida: Amphipoda: Maeridae)" masterLastPageNumber="359" masterPageNumber="351" pageNumber="353" updateTime="1700278603914" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3446962E2BB4D25DA2FC0E94487F2E9E">Description of a new species of Quadrivisio Stebbing, 1907, from Songkhla Lake, Thailand (Crustacea: Peracarida: Amphipoda: Maeridae)</mods:title>
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<mods:namePart id="0D2489BE7C4614AF96FB5914FECDA906">Hughes, Lauren E.</mods:namePart>
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<mods:namePart id="67B661C4125027FCC3881BBABB0A32F1">Kaji, Tomonari</mods:namePart>
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<mods:title id="24E9DD1D18ED087281911847A072CE36">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="05A837F8F76FBE21C87FD1E81980FD57">
<mods:date id="F06A69E592CB25018766C37D3F0CEF81">2016</mods:date>
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<mods:number id="C53BF82F500D9F4CFBD76CDF85EDA228">2016-11-21</mods:number>
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<mods:number id="6B375ED3B781755E68783C36261C58BD">64</mods:number>
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<treatment id="283C0A34FF98274F57A3FEC21792F9C0" LSID="urn:lsid:zoobank.org:act:CA01B47B-5DA7-4F66-9F5E-B6CF663A6B95" httpUri="http://treatment.plazi.org/id/283C0A34FF98274F57A3FEC21792F9C0" lastPageId="7" lastPageNumber="358" pageId="2" pageNumber="353">
<subSubSection id="E88FE8A9FF98274A57A3FEC2100DFE94" box="[966,1344,285,309]" pageId="2" pageNumber="353" type="nomenclature">
<paragraph id="A02ABB22FF98274A57A3FEC2100DFE94" blockId="2.[966,1344,285,341]" box="[966,1344,285,309]" pageId="2" pageNumber="353">
<heading id="FB620C4EFF98274A57A3FEC2100DFE94" bold="true" box="[966,1344,285,309]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="353" reason="2">
<emphasis id="92E16730FF98274A57A3FEC2100DFE94" bold="true" box="[966,1344,285,309]" pageId="2" pageNumber="353">
<taxonomicName id="6795C0A1FF98274A57A3FEC211FFFE94" ID-CoL="4QZDH" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[966,1202,285,309]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF98274A57A3FEC211FFFE94" bold="true" box="[966,1202,285,309]" italics="true" pageId="2" pageNumber="353">Quadrivisio meufong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89D2DA4BFF98274A50DFFEC2100DFE94" box="[1210,1344,285,309]" pageId="2" pageNumber="353" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="E88FE8A9FF98274A5022FEE211F3FEF4" box="[1095,1214,317,341]" pageId="2" pageNumber="353" type="description">
<paragraph id="A02ABB22FF98274A5022FEE211F3FEF4" blockId="2.[966,1344,285,341]" box="[1095,1214,317,341]" pageId="2" pageNumber="353">
(
<figureCitation id="38AEA7A7FF98274A502AFEE211FBFEF4" box="[1103,1206,317,341]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[136,171,1985,2007]" captionStartId-1="5.[113,148,1180,1202]" captionStartId-2="6.[136,171,1551,1573]" captionStartId-3="7.[113,148,1058,1080]" captionTargetBox-0="[137,1477,297,1968]" captionTargetBox-1="[114,1451,190,1163]" captionTargetBox-2="[130,1487,179,1542]" captionTargetBox-3="[113,1451,189,1043]" captionTargetId-0="figure-8@4.[131,1486,288,1968]" captionTargetId-1="figure-9@5.[110,1454,186,1163]" captionTargetId-2="figure-8@6.[130,1487,179,1542]" captionTargetId-3="figure-489@7.[113,1451,189,1043]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 3. Quadrivisio meufong, new species, holotype, male, 9.8 mm, PSUZC-CR 0301, Songkhla Lake, Thailand. Scale bars = 0.1 mm." captionText-1="Fig. 4. Quadrivisio meufong, new species, holotype, male, 9.8 mm, PSUZC-CR 0301, Songkhla Lake, Thailand. Scale bars = 0.1 mm." captionText-2="Fig. 5. Quadrivisio meufong, new species, paratype, female, 10.5 mm, PSUZC-CR 0304, Songkhla Lake, Thailand. Scale bars = 0.1 mm." captionText-3="Fig. 6. 3D reconstructed male gnathopod 2 of Quadrivisio meufong, new species, male 13.5 mm, PSUZC-CR 0302. A, overall image of carpus, propodus, and dactylus of the male gnathopod 2; B, magnified, proximal view of the expanded palm cuticle. Arrow heads indicate expanded palm structure; oc, outer layer of cuticle; ic, inner layer of cuticle. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5355552" figureDoi-1="http://doi.org/10.5281/zenodo.5355554" figureDoi-2="http://doi.org/10.5281/zenodo.5355556" figureDoi-3="http://doi.org/10.5281/zenodo.5355558" httpUri-0="https://zenodo.org/record/5355552/files/figure.png" httpUri-1="https://zenodo.org/record/5355554/files/figure.png" httpUri-2="https://zenodo.org/record/5355556/files/figure.png" httpUri-3="https://zenodo.org/record/5355558/files/figure.png" pageId="2" pageNumber="353">Figs. 36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF98274A5726FEA3110DFE0E" pageId="2" pageNumber="353" type="reference_group">
<paragraph id="A02ABB22FF98274A5726FEA3108FFE30" blockId="2.[835,1474,380,431]" box="[835,1474,380,402]" pageId="2" pageNumber="353">
<treatmentCitationGroup id="80859C0CFF98274A5726FEA3108FFE30" box="[835,1474,380,402]" pageId="2" pageNumber="353">
?
<treatmentCitation id="21349D33FF98274A572BFEA31054FE30" author="Bussarawich S" box="[846,1305,380,402]" page="4" pageId="2" pageNumber="353" year="1985">
<taxonomicName id="6795C0A1FF98274A572BFEA31054FE30" ID-CoL="4QZDC" authority=", Bussarawich, 1985: 4" authorityName="Bussarawich" authorityPageNumber="4" authorityYear="1985" box="[846,1305,380,402]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="92E16730FF98274A572BFEA31175FE30" box="[846,1080,380,401]" italics="true" pageId="2" pageNumber="353">Quadrivisio bengalensis</emphasis>
,
<bibRefCitation id="C404C6D3FF98274A5023FEA31054FE30" author="Bussarawich S" box="[1094,1305,380,402]" pageId="2" pageNumber="353" refId="ref3768" refString="Bussarawich S (1985) Gammaridean Amphipoda from mangroves in southern Thailand. Fifth Seminar on Mangrove Ecosystems Phuket, Thailand: 1 - 7, 11 figs." type="journal volume" year="1985">Bussarawich, 1985: 4</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 5, fig. 3;
<bibRefCitation id="C404C6D3FF98274A5119FEA3108FFE30" author="Lowry JK" box="[1404,1474,380,401]" pageId="2" pageNumber="353" pagination="309 - 342" refId="ref4003" refString="Lowry JK (2000) Taxonomic status of amphipod Crustaceans in the South China Sea with a checklist of known species. The Raffles Bulletin of Zoology, Supplement 8: 309 - 342." type="journal article" year="2000">Lowry,</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="A02ABB22FF98274A570CFE46110DFE0E" blockId="2.[835,1474,380,431]" box="[873,1088,409,431]" pageId="2" pageNumber="353">2000: 319 (checklist).</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF98274A5726FE081040FBCE" pageId="2" pageNumber="353" type="materials_examined">
<paragraph id="A02ABB22FF98274A5726FE0816E5FCAE" blockId="2.[835,1476,471,783]" pageId="2" pageNumber="353">
<emphasis id="92E16730FF98274A5726FE0816A7FE4E" bold="true" box="[835,1002,471,495]" pageId="2" pageNumber="353">Type material.</emphasis>
<typeStatus id="7F2E0580FF98274A5795FE08111BFE4E" box="[1008,1110,471,495]" pageId="2" pageNumber="353" type="holotype">Holotype</typeStatus>
, male,
<quantity id="676D16C7FF98274A50C5FE0811BEFE4F" box="[1184,1267,471,495]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="2" pageNumber="353" unit="mm" value="9.8">9.8 mm</quantity>
, dissected, 4 slides,
<collectionCode id="C68423E7FF98274A5726FE28168AFDAE" box="[835,967,503,527]" pageId="2" pageNumber="353">PSUZC-CR</collectionCode>
0301, Khu Khut Bird Sanctuary,
<collectingRegion id="625175C0FF98274A513DFE28108CFDAE" box="[1368,1473,503,527]" country="Thailand" name="Songkhla" pageId="2" pageNumber="353">Songkhla</collectingRegion>
Lake,
<collectingCountry id="D882FBB2FF98274A57EFFDC816A1FD8E" box="[906,1004,535,559]" name="Thailand" pageId="2" pageNumber="353">Thailand</collectingCountry>
,
<geoCoordinate id="C5A1DDE5FF98274A579FFDC811D9FD8E" box="[1018,1172,535,559]" degrees="7" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="21.14" value="7.455872">7°2721.14”N</geoCoordinate>
,
<geoCoordinate id="C5A1DDE5FF98274A50C7FDC81019FD8E" box="[1186,1364,535,559]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="20.21" value="100.40562">100°2420.21”E</geoCoordinate>
, in shore, on
<taxonomicName id="6795C0A1FF98274A570FFDE8105BFDEE" authority="(Vahl) Lye" authorityName="Lye" authorityYear="1971" baseAuthorityName="Vahl" box="[874,1302,567,591]" class="Liliopsida" family="Cyperaceae" genus="Schoenoplectus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="353" phylum="Tracheophyta" rank="species" species="subulatus">
<emphasis id="92E16730FF98274A570FFDE811C0FDEE" box="[874,1165,567,591]" italics="true" pageId="2" pageNumber="353">Schoenoplectus subulatus</emphasis>
(Vahl) Lye
</taxonomicName>
(original label
<taxonomicName id="6795C0A1FF98274A5726FD881103FDCE" authority="Schrad." authorityName="Schrad." box="[835,1102,599,623]" class="Liliopsida" family="Cyperaceae" genus="Scirpus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="353" phylum="Tracheophyta" rank="species" species="litoralis">
<emphasis id="92E16730FF98274A5726FD8816BDFDCE" box="[835,1008,599,623]" italics="true" pageId="2" pageNumber="353">Scirpus litoralis</emphasis>
Schrad.
</taxonomicName>
), coll. J.K. Lowry &amp;
<collectionCode id="C68423E7FF98274A515CFD881002FDCF" box="[1337,1359,599,622]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="353" type="Museum">V</collectionCode>
. Lheknim,
<date id="D42B9DE2FF98274A5720FDA81152FD2E" box="[837,1055,631,655]" pageId="2" pageNumber="353" value="1999-12-13">13 December 1999</date>
.
<typeStatus id="7F2E0580FF98274A5048FDA811D1FD2E" box="[1069,1180,631,655]" pageId="2" pageNumber="353" type="paratype">Paratypes</typeStatus>
: male,
<quantity id="676D16C7FF98274A5094FDA81015FD2F" box="[1265,1368,631,655]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="353" unit="mm" value="13.5">13.5 mm</quantity>
, PSUZC- CR 0302; male,
<quantity id="676D16C7FF98274A579FFD481113FD0F" box="[1018,1118,663,687]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="2" pageNumber="353" unit="mm" value="12.3">12.3 mm</quantity>
,
<collectionCode id="C68423E7FF98274A5009FD4811A6FD0E" box="[1132,1259,663,687]" pageId="2" pageNumber="353">PSUZC-CR</collectionCode>
0303; female,
<quantity id="676D16C7FF98274A51F6FD48163DFD6F" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="353" unit="mm" value="10.5">10.5 mm</quantity>
, dissected, 1 slide,
<collectionCode id="C68423E7FF98274A5031FD68119BFD6E" box="[1108,1238,695,719]" pageId="2" pageNumber="353">PSUZC-CR</collectionCode>
0304; 40+ juveniles and females,
<collectionCode id="C68423E7FF98274A57BDFD081115FD4E" box="[984,1112,727,751]" pageId="2" pageNumber="353">PSUZC-CR</collectionCode>
0305; all from same location as
<typeStatus id="7F2E0580FF98274A5726FD2816E9FCAE" box="[835,932,759,783]" pageId="2" pageNumber="353" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="A02ABB22FF98274A5726FCE81174FBAE" blockId="2.[834,1474,823,1039]" pageId="2" pageNumber="353">
<emphasis id="92E16730FF98274A5726FCE811E9FCEE" bold="true" box="[835,1188,823,847]" pageId="2" pageNumber="353">Additional material examined.</emphasis>
<materialsCitation id="10FDB17FFF98274A50C9FCE816EBFC6E" collectingDate="1999-12-13" collectionCode="V" collectorName="Royle &amp; J. K. Lowry &amp; Lheknim" country="Thailand" latitude="7.45" location="Khu Khut Bird Sanctuary" longLatPrecision="1303" longitude="100.4" municipality="Lake Basin" pageId="2" pageNumber="353" specimenCode="PSUZC-CR 0306" specimenCount="10" specimenCount-juvenile="10" stateProvince="Songkhla">
<specimenCount id="B69370ABFF98274A50C9FCE8107DFCEE" box="[1196,1328,823,847]" pageId="2" pageNumber="353" type="juvenile">10 juveniles</specimenCount>
and females,
<specimenCode id="F0331359FF98274A5726FC88114DFCCE" box="[835,1024,855,879]" collectionCode="PSUZC-CR" pageId="2" pageNumber="353">PSUZC-CR 0306</specimenCode>
,
<location id="A54AEDF9FF98274A506CFC881054FCCE" LSID="urn:lsid:plazi:treatment:283C0A34FF98274F57A3FEC21792F9C0:A54AEDF9FF98274A506CFC881054FCCE" box="[1033,1305,855,879]" country="Thailand" latitude="7.45" longLatPrecision="1303" longitude="100.4" municipality="Lake Basin" name="Khu Khut Bird Sanctuary" pageId="2" pageNumber="353" stateProvince="Songkhla">Khu Khut Bird Sanctuary</location>
,
<collectingRegion id="625175C0FF98274A5146FC8810C5FCCE" box="[1315,1416,855,879]" country="Thailand" name="Songkhla" pageId="2" pageNumber="353">Songkhla</collectingRegion>
<collectingMunicipality id="404E2158FF98274A51E8FC8816C9FC2E" pageId="2" pageNumber="353">Lake Basin</collectingMunicipality>
,
<collectingCountry id="D882FBB2FF98274A57F6FCA816B5FC2E" box="[915,1016,887,911]" name="Thailand" pageId="2" pageNumber="353">Thailand</collectingCountry>
,
<geoCoordinate id="C5A1DDE5FF98274A5062FCA81117FC2E" box="[1031,1114,887,911]" degrees="7" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="353" precision="925" value="7.45">7°27N</geoCoordinate>
,
<geoCoordinate id="C5A1DDE5FF98274A503BFCA81180FC2E" box="[1118,1229,887,911]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="2" pageNumber="353" precision="925" value="100.4">100°24E</geoCoordinate>
,
<taxonomicName id="6795C0A1FF98274A50BBFCA8108CFC2E" box="[1246,1473,887,911]" class="Liliopsida" family="Hydrocharitaceae" genus="Hydrilla" kingdom="Plantae" order="Alismatales" pageId="2" pageNumber="353" phylum="Tracheophyta" rank="species" species="verticillata">
<emphasis id="92E16730FF98274A50BBFCA8108CFC2E" box="[1246,1473,887,911]" italics="true" pageId="2" pageNumber="353">Hydrilla verticillata</emphasis>
</taxonomicName>
(L. f.)
<collectorName id="0D60DEF4FF98274A57EBFC48169DFC0E" box="[910,976,919,943]" pageId="2" pageNumber="353">Royle</collectorName>
,
<date id="D42B9DE2FF98274A57B8FC4811FDFC0E" box="[989,1200,919,943]" pageId="2" pageNumber="353" value="1999-12-13">
<collectingDate id="C46F640AFF98274A57B8FC4811FDFC0E" box="[989,1200,919,943]" pageId="2" pageNumber="353" value="1999-12-13">13 December 1999</collectingDate>
</date>
, coll.
<collectorName id="0D60DEF4FF98274A5091FC481022FC0E" box="[1268,1391,919,943]" pageId="2" pageNumber="353">J.K. Lowry</collectorName>
and
<collectionCode id="C68423E7FF98274A51CDFC4810F3FC0F" box="[1448,1470,919,942]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="353" type="Museum">V</collectionCode>
.
<collectorName id="0D60DEF4FF98274A5726FC6816EBFC6E" box="[835,934,951,975]" pageId="2" pageNumber="353">Lheknim</collectorName>
</materialsCitation>
;
<materialsCitation id="10FDB17FFF98274A57D5FC681178FBAE" collectionCode="AM" country="Thailand" location="Lake" pageId="2" pageNumber="353" specimenCount="32" specimenCount-juvenile="29" specimenCount-male="3" stateProvince="Songkhla">
<specimenCount id="B69370ABFF98274A57D5FC68114EFC6E" box="[944,1027,951,975]" pageId="2" pageNumber="353" type="male">3 males</specimenCount>
,
<specimenCount id="B69370ABFF98274A5069FC6811C1FC6E" box="[1036,1164,951,975]" pageId="2" pageNumber="353" type="juvenile">29 juveniles</specimenCount>
and females,
<collectionCode id="C68423E7FF98274A517EFC68100BFC6E" box="[1307,1350,951,975]" country="Australia" name="Australian Museum" pageId="2" pageNumber="353" type="Museum">AM</collectionCode>
P. PSUZC- CR 0307,
<collectingRegion id="625175C0FF98274A57DEFC08116EFC4E" box="[955,1059,983,1007]" country="Thailand" name="Songkhla" pageId="2" pageNumber="353">Songkhla</collectingRegion>
<location id="A54AEDF9FF98274A5048FC08112AFC4E" LSID="urn:lsid:plazi:treatment:283C0A34FF98274F57A3FEC21792F9C0:A54AEDF9FF98274A5048FC08112AFC4E" box="[1069,1127,983,1007]" country="Thailand" name="Lake" pageId="2" pageNumber="353" stateProvince="Songkhla">Lake</location>
,
<collectingCountry id="D882FBB2FF98274A5013FC081196FC4E" box="[1142,1243,983,1007]" name="Thailand" pageId="2" pageNumber="353">Thailand</collectingCountry>
, 1987-1989 (several collections combined)
</materialsCitation>
.
</paragraph>
<paragraph id="A02ABB22FF98274A5726FBE81040FBCE" blockId="2.[835,1474,1079,1135]" pageId="2" pageNumber="353">
<materialsCitation id="10FDB17FFF98274A5726FBE81044FBCE" country="Thailand" latitude="7.455872" location="Khu Khut Bird Sanctuary" longLatPrecision="1" longitude="100.40562" municipality="Lake" pageId="2" pageNumber="353" specimenCount="1" stateProvince="Songkhla" typeStatus="holotype">
<emphasis id="92E16730FF98274A5726FBE816ACFBEE" bold="true" box="[835,993,1079,1103]" pageId="2" pageNumber="353">
<typeStatus id="7F2E0580FF98274A5726FBE81630FBEE" box="[835,893,1079,1103]" pageId="2" pageNumber="353">Type</typeStatus>
locality.
</emphasis>
<location id="A54AEDF9FF98274A578EFBE81044FBEE" LSID="urn:lsid:plazi:treatment:283C0A34FF98274F57A3FEC21792F9C0:A54AEDF9FF98274A578EFBE81044FBEE" box="[1003,1289,1079,1103]" country="Thailand" latitude="7.455872" longLatPrecision="1" longitude="100.40562" municipality="Lake" name="Khu Khut Bird Sanctuary" pageId="2" pageNumber="353" stateProvince="Songkhla">Khu Khut Bird Sanctuary</location>
,
<collectingRegion id="625175C0FF98274A5172FBE81031FBEE" box="[1303,1404,1079,1103]" country="Thailand" name="Songkhla" pageId="2" pageNumber="353">Songkhla</collectingRegion>
<collectingMunicipality id="404E2158FF98274A51E3FBE810F3FBEE" box="[1414,1470,1079,1103]" pageId="2" pageNumber="353">Lake</collectingMunicipality>
,
<collectingCountry id="D882FBB2FF98274A5726FB8816E8FBCE" box="[835,933,1111,1135]" name="Thailand" pageId="2" pageNumber="353">Thailand</collectingCountry>
,
<geoCoordinate id="C5A1DDE5FF98274A57D4FB881107FBCE" box="[945,1098,1111,1135]" degrees="7" direction="north" minutes="27" orientation="latitude" pageId="2" pageNumber="353" precision="1" seconds="21.14" value="7.455872">7°2721.14”N</geoCoordinate>
,
<geoCoordinate id="C5A1DDE5FF98274A5032FB881044FBCE" box="[1111,1289,1111,1135]" degrees="100" direction="east" minutes="24" orientation="longitude" pageId="2" pageNumber="353" precision="1" seconds="20.21" value="100.40562">100°2420.21”E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF98274A5726FB48114CFB4E" pageId="2" pageNumber="353" type="etymology">
<paragraph id="A02ABB22FF98274A5726FB48114CFB4E" blockId="2.[834,1474,1175,1263]" pageId="2" pageNumber="353">
<emphasis id="92E16730FF98274A5726FB481689FB0E" bold="true" box="[835,964,1175,1199]" pageId="2" pageNumber="353">Etymology.</emphasis>
From the Thai meu - hand and fong bubble, in reference to the adult male gnathopod 2 propodus palm expanded cuticle.
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF98274A5726FAC81131F90E" pageId="2" pageNumber="353" type="diagnosis">
<paragraph id="A02ABB22FF98274A5726FAC81131F90E" blockId="2.[834,1475,1303,1711]" pageId="2" pageNumber="353">
<emphasis id="92E16730FF98274A5726FAC816F5FA8E" bold="true" box="[835,952,1303,1327]" pageId="2" pageNumber="353">Diagnosis.</emphasis>
Antenna 1 accessory flagellum long, with 9-10 articles. Mandibular palp with 2 apical setae. Gnathopod 2 palm subacute, two thirds length of propodus posterior margin, subrectangular distomedial shelf with robust setae, midpalm dominated by expanded cuticle covered in fine setae, palmar margin defined by rounded corner with 2 robust setae; dactylus without posteroproximal shelf, closing along and equal to length of palm. Pereonite 7 without carina. Pleonites 13 bicarinate. Epimeron 12 posterodistal corner with 3 serrate teeth Epimeron 3 ventral margin serrate, posterodistal corner with 2 teeth. Urosomites 13 bicarinate. Telson each lobe with 2 long and 12 short apical robust setae, inner margins smooth.
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF98274E5726F9081645F83A" lastPageId="6" lastPageNumber="357" pageId="2" pageNumber="353" type="description">
<paragraph id="A02ABB22FF98274D5726F9081445F9C1" blockId="2.[835,1476,1751,2095]" lastBlockId="5.[112,754,1257,1633]" lastPageId="5" lastPageNumber="356" pageId="2" pageNumber="353">
<emphasis id="92E16730FF98274A5726F908169CF94E" bold="true" box="[835,977,1751,1775]" pageId="2" pageNumber="353">Description.</emphasis>
Based on
<typeStatus id="7F2E0580FF98274A5028F90811E1F94E" box="[1101,1196,1751,1775]" pageId="2" pageNumber="353" type="holotype">holotype</typeStatus>
male,
<quantity id="676D16C7FF98274A5098F908101AF94F" box="[1277,1367,1751,1775]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="2" pageNumber="353" unit="mm" value="9.8">9.8 mm</quantity>
, PSUZC- CR 0301. Head. Eyes ommatidia split into two pairs on each side of head; lateral cephalic lobe broad, truncated, anteroventral margin with notch/slit. Antenna 1 longer than antenna 2; peduncular article 1 shorter than article 2, without robust setae along posterior margin; article 2 longer than article 3; flagellum articles longer than broad, with 38 articles; accessory flagellum long, with 9-10 articles. Antenna 2 peduncular article 2 cone gland not reaching to end of peduncular article 3; article 4 shorter than article 5; flagellum with 34 articles. Maxilla 1 inner plate unknown; palp 2-articulate with 6 apical robust setae. Lower lip inner and outer lobes apically setose; mandibular lobes apically subacute. Mandible accessory setal row well developed with 8 setae; molar well developed, columnar, triturative; mandibular palp well developed, 3-articulate; article 1 length 1.8 times as long as broad, 0.8 times article 2; article 2 length subequal to article 3, without setae; article 3 rectilinear, short (3.5 times as long as broad), 1.4 times article 1, with 2 apical setae. Maxilliped basal endite with 4 short apical robust setae; ischial endite medial margin lined with robust setae; palp 4-articulate, article 4 lined with slender setae, unguis preset.
</paragraph>
<caption id="F4EAEBAAFF99274B5414F7FD1026F799" ID-DOI="http://doi.org/10.5281/zenodo.5355550" ID-Zenodo-Dep="5355550" box="[113,1387,2082,2104]" httpUri="https://zenodo.org/record/5355550/files/figure.png" pageId="3" pageNumber="354" startId="3.[113,148,2082,2104]" targetBox="[117,1450,200,2066]" targetPageId="3">
<paragraph id="A02ABB22FF99274B5414F7FD1026F799" blockId="3.[113,1387,2082,2104]" box="[113,1387,2082,2104]" pageId="3" pageNumber="354">
Fig. 2.
<taxonomicName id="6795C0A1FF99274B54DFF7FD170CF796" authority="Stebbing, 1907" authorityName="Stebbing" authorityYear="1907" box="[186,577,2082,2104]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="92E16730FF99274B54DFF7FD14EBF796" box="[186,422,2082,2103]" italics="true" pageId="3" pageNumber="354">Quadrivisio bengalensis</emphasis>
<bibRefCitation id="C404C6D3FF99274B55C9F7FD170CF796" author="Stebbing TRR" box="[428,577,2082,2104]" pageId="3" pageNumber="354" pagination="159 - 162" refId="ref4226" refString="Stebbing TRR (1907) The fauna of brackish ponds at Port Canning, Lower Bengal, Part V. Definition of a new genus of Amphipoda, and description of the typical species. Records of the Indian Museum, 1: 159 - 162." type="journal article" year="1907">Stebbing, 1907</bibRefCitation>
</taxonomicName>
, male, 11 mm, PSUZC-CR 0308, Songkhla Lake, Thailand. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="F4EAEBAAFF9E274C54EDF81E10FEF877" ID-DOI="http://doi.org/10.5281/zenodo.5355552" ID-Zenodo-Dep="5355552" box="[136,1459,1985,2007]" httpUri="https://zenodo.org/record/5355552/files/figure.png" pageId="4" pageNumber="355" startId="4.[136,171,1985,2007]" targetBox="[137,1477,297,1968]" targetPageId="4">
<paragraph id="A02ABB22FF9E274C54EDF81E10FEF877" blockId="4.[136,1459,1985,2007]" box="[136,1459,1985,2007]" pageId="4" pageNumber="355">
Fig. 3.
<taxonomicName id="6795C0A1FF9E274C54B4F81E14D0F877" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[209,413,1985,2006]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9E274C54B4F81E14D0F877" box="[209,413,1985,2006]" italics="true" pageId="4" pageNumber="355">Quadrivisio meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9E274C55CFF81E176FF876" box="[426,546,1985,2007]" pageId="4" pageNumber="355" rank="species">new species</taxonomicNameLabel>
, holotype, male, 9.8 mm, PSUZC-CR 0301, Songkhla Lake, Thailand. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="F4EAEBAAFF9F274D5414FB4310D1FB10" ID-DOI="http://doi.org/10.5281/zenodo.5355554" ID-Zenodo-Dep="5355554" box="[113,1436,1180,1202]" httpUri="https://zenodo.org/record/5355554/files/figure.png" pageId="5" pageNumber="356" startId="5.[113,148,1180,1202]" targetBox="[114,1451,190,1163]" targetPageId="5">
<paragraph id="A02ABB22FF9F274D5414FB4310D1FB10" blockId="5.[113,1436,1180,1202]" box="[113,1436,1180,1202]" pageId="5" pageNumber="356">
Fig. 4.
<taxonomicName id="6795C0A1FF9F274D54DFFB4314CBFB10" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[186,390,1180,1201]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9F274D54DFFB4314CBFB10" box="[186,390,1180,1201]" italics="true" pageId="5" pageNumber="356">Quadrivisio meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9F274D55F1FB431746FB13" box="[404,523,1180,1202]" pageId="5" pageNumber="356" rank="species">new species</taxonomicNameLabel>
, holotype, male, 9.8 mm, PSUZC-CR 0301, Songkhla Lake, Thailand. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A02ABB22FF9F274D5414F9561034F9A0" blockId="5.[113,754,1673,2113]" lastBlockId="5.[811,1453,1257,1537]" pageId="5" pageNumber="356">Pereon. Gnathopod 1 coxa anterior margin straight concave, anteroventral corner not produced, rounded, posteroventral margin with a few serration and robust seta; carpus 2.8 times as long as broad, longer than propodus; propodus medial surface with comb, palm near transverse, convex, entire, defined by posterodistal corner, with 4 robust setae. Gnathopod 2 subchelate; coxa posteroventral margin with robust seta; basis slender, anterodistal corner without lobes; ischium anterior margin without lobes; merus distoventrally truncate; carpus compressed, lobate, projecting between merus and propodus with rows of dense setae; propodus massive, palm subacute, two thirds length of propodus posterior margin, subrectangular distomedial shelf with robust setae, midpalm dominated by expanded cuticle covered in fine setae, palmar margin defined by rounded corner with 2 robust setae, posterior proximal margin lined with long slender setae; dactylus without posteroproximal shelf, closing along and equal to length of palm. Pereopod 4 coxa posteroventral lobe well developed, with subrectangular posteromedial corner. Pereopods 56 basis weakly expanded, posterior margin straight; merus and carpus not broadened. Pereopod 7 basis expanded, posterior margin convex; merus and carpus not broadened. Pereonite 7 without carina.</paragraph>
<paragraph id="A02ABB22FF9F274D574AF9F611B4F840" blockId="5.[812,1453,1577,2017]" pageId="5" pageNumber="356">Pleon. Pleonites 13 bicarinate. Epimeron 12 posterodistal corner with 3 serrate teeth Epimeron 3 ventral margin serrate, posterodistal corner with 2 teeth. Urosomites 13 bicarinate. Uropod 1 peduncle with 1 basofacial seta, longer than rami; outer ramus subequal to inner ramus with apical and marginal robust setae. Uropod 2 peduncle longer than rami, outer ramus shorter than inner ramus with apical and marginal robust setae. Uropod 3 peduncle shorter than rami; rami foliaceous, subequal in length, lined with short robust and slender setae. Telson deeply cleft (90%), longer as broad, tapering distally, with short inner and long outer apical cusps, apical conical extension reaching less than one third along longest seta, each lobe with 2 long and 12 short apical robust setae, inner margins smooth.</paragraph>
<paragraph id="A02ABB22FF9F274D5748F7D610E1F7E1" blockId="5.[813,1452,2057,2113]" pageId="5" pageNumber="356">
Female (sexually dimorphic characters), based on
<typeStatus id="7F2E0580FF9F274D5135F7D610E1F780" box="[1360,1452,2057,2081]" pageId="5" pageNumber="356" type="paratype">paratype</typeStatus>
female,
<quantity id="676D16C7FF9F274D57EBF7F616A6F7E1" box="[910,1003,2089,2113]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="356" unit="mm" value="7.3">7.3 mm</quantity>
, PSUZC-CR 0304. Gnathopod 2 coxa
</paragraph>
<paragraph id="A02ABB22FF9C274E5653FFAB115EFF2D" blockId="6.[566,1043,116,140]" box="[566,1043,116,140]" pageId="6" pageNumber="357">
<emphasis id="92E16730FF9C274E5653FFAB1698FF2A" box="[566,981,116,140]" italics="true" pageId="6" pageNumber="357">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2016
</paragraph>
<caption id="F4EAEBAAFF9C274E54EDF9D0108EF985" ID-DOI="http://doi.org/10.5281/zenodo.5355556" ID-Zenodo-Dep="5355556" box="[136,1475,1551,1573]" httpUri="https://zenodo.org/record/5355556/files/figure.png" pageId="6" pageNumber="357" startId="6.[136,171,1551,1573]" targetBox="[130,1487,179,1542]" targetPageId="6">
<paragraph id="A02ABB22FF9C274E54EDF9D0108EF985" blockId="6.[136,1475,1551,1573]" box="[136,1475,1551,1573]" pageId="6" pageNumber="357">
Fig. 5.
<taxonomicName id="6795C0A1FF9C274E54AAF9D014D6F985" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[207,411,1551,1572]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9C274E54AAF9D014D6F985" box="[207,411,1551,1572]" italics="true" pageId="6" pageNumber="357">Quadrivisio meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9C274E55CDF9D01752F984" box="[424,543,1551,1573]" pageId="6" pageNumber="357" rank="species">new species</taxonomicNameLabel>
, paratype, female, 10.5 mm, PSUZC-CR 0304, Songkhla Lake, Thailand. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A02ABB22FF9C274E54EDF9BC1767F89B" blockId="6.[134,776,1635,1851]" pageId="6" pageNumber="357">posterodistal corner with robust seta; carpus subtriangular, 0.9 times propodus length; propodus rectilinear, palm subacute, without distomedial shelf, palmar margin irregular, entire, lined with short robust setae, defined by posteroventral corner, with 3 spaced long robust setae; dactylus weakly recurved, closing short of palm, margins smooth. Telson lobes with 1 long and 2 short apical robust setae.</paragraph>
<paragraph id="A02ABB22FF9C274E54EDF8BC1645F83A" blockId="6.[136,776,1891,1947]" pageId="6" pageNumber="357">
<emphasis id="92E16730FF9C274E54EDF8BC144DF8DA" bold="true" box="[136,256,1891,1915]" pageId="6" pageNumber="357">Variation.</emphasis>
Uropod 3 increases with growth stage (often regenerating or missing, broken off during sample processing).
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF9C274F54EDF81C143CF9A0" lastPageId="7" lastPageNumber="358" pageId="6" pageNumber="357" type="discussion">
<paragraph id="A02ABB22FF9C274E54EDF81C104AF83A" blockId="6.[136,775,1987,2107]" lastBlockId="6.[834,1474,1635,1947]" pageId="6" pageNumber="357">
<emphasis id="92E16730FF9C274E54EDF81C15B6F87A" bold="true" box="[136,251,1987,2011]" pageId="6" pageNumber="357">Remarks.</emphasis>
The male gnathopod 2 propodus palm of
<taxonomicName id="6795C0A1FF9C274E5689F81C15ABF85A" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9C274E5689F81C15ABF85A" italics="true" pageId="6" pageNumber="357">Q. meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9C274E549DF83C14CCF85A" box="[248,385,2019,2043]" pageId="6" pageNumber="357" rank="species">new species</taxonomicNameLabel>
, has a large bubble-like expanded cuticle covered in fine setae. In the few male specimens on hand, the expanded palm cuticle may be collapsed or inflated, and it is unclear if this is related to preservation technique (with the tegument collapsing during fixation/ preservation) or growth stage (larger males have a more inflated structure) (
<figureCitation id="38AEA7A7FF9C274E5062F91C1104F97A" box="[1031,1097,1731,1755]" captionStart="Fig" captionStartId="5.[113,148,1180,1202]" captionTargetBox="[114,1451,190,1163]" captionTargetId="figure-9@5.[110,1454,186,1163]" captionTargetPageId="5" captionText="Fig. 4. Quadrivisio meufong, new species, holotype, male, 9.8 mm, PSUZC-CR 0301, Songkhla Lake, Thailand. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5355554" httpUri="https://zenodo.org/record/5355554/files/figure.png" pageId="6" pageNumber="357">Figs 4</figureCitation>
,
<figureCitation id="38AEA7A7FF9C274E5036F91C112FF97A" box="[1107,1122,1731,1755]" captionStart="Fig" captionStartId="7.[113,148,1058,1080]" captionTargetBox="[113,1451,189,1043]" captionTargetId="figure-489@7.[113,1451,189,1043]" captionTargetPageId="7" captionText="Fig. 6. 3D reconstructed male gnathopod 2 of Quadrivisio meufong, new species, male 13.5 mm, PSUZC-CR 0302. A, overall image of carpus, propodus, and dactylus of the male gnathopod 2; B, magnified, proximal view of the expanded palm cuticle. Arrow heads indicate expanded palm structure; oc, outer layer of cuticle; ic, inner layer of cuticle. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5355558" httpUri="https://zenodo.org/record/5355558/files/figure.png" pageId="6" pageNumber="357">6</figureCitation>
). The palmar structure is enclosed by an outer and inner layer of cuticle (
<figureCitation id="38AEA7A7FF9C274E5095F93C100BF95A" box="[1264,1350,1763,1787]" captionStart="Fig" captionStartId="7.[113,148,1058,1080]" captionTargetBox="[113,1451,189,1043]" captionTargetId="figure-489@7.[113,1451,189,1043]" captionTargetPageId="7" captionText="Fig. 6. 3D reconstructed male gnathopod 2 of Quadrivisio meufong, new species, male 13.5 mm, PSUZC-CR 0302. A, overall image of carpus, propodus, and dactylus of the male gnathopod 2; B, magnified, proximal view of the expanded palm cuticle. Arrow heads indicate expanded palm structure; oc, outer layer of cuticle; ic, inner layer of cuticle. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5355558" httpUri="https://zenodo.org/record/5355558/files/figure.png" pageId="6" pageNumber="357">Fig. 6B</figureCitation>
), and there are no hemolymphal connection to propodus observed. The cuticular layers are seemingly peeled and inflated structure derived from ordinary cuticular composition, that is the inner layer might be comparable to endo and exocuticle, while the outer layer is comparable with epicuticle.
</paragraph>
<paragraph id="A02ABB22FF9C274F5726F81C14DAFAC0" blockId="6.[835,1475,1987,2107]" lastBlockId="7.[112,753,1193,1377]" lastPageId="7" lastPageNumber="358" pageId="6" pageNumber="357">
<taxonomicName id="6795C0A1FF9C274E5726F81C1169F87A" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[835,1060,1987,2011]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9C274E5726F81C1169F87A" box="[835,1060,1987,2011]" italics="true" pageId="6" pageNumber="357">Quadrivisio meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9C274E5055F81C11FFF87A" box="[1072,1202,1987,2011]" pageId="6" pageNumber="357" rank="species">new species</taxonomicNameLabel>
, is strikingly similar to
<taxonomicName id="6795C0A1FF9C274E51C2F81C1107F85A" authority="(Shoemaker, 1933)" baseAuthorityName="Shoemaker" baseAuthorityYear="1933" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="92E16730FF9C274E51C2F81C163CF85A" italics="true" pageId="6" pageNumber="357">Q. lutzi</emphasis>
(
<bibRefCitation id="C404C6D3FF9C274E57E7F83C110FF85A" author="Shoemaker CR" box="[898,1090,2019,2043]" pageId="6" pageNumber="357" pagination="1 - 24" refId="ref4203" refString="Shoemaker CR (1933) Amphipoda from Florida and the West Indies. American Museum Novitates, 598: 1 - 24." type="journal article" year="1933">Shoemaker, 1933</bibRefCitation>
)
</taxonomicName>
from Georgetown,
<collectingCountry id="D882FBB2FF9C274E5146F83C108FF85A" box="[1315,1474,2019,2043]" name="Guyana" pageId="6" pageNumber="357">British Guiana</collectingCountry>
on the northern coast of South America. Both species have a similarly shaped palm of the male gnathopod 2. The presence of two carinae on pleonites 13 and urosomite 1 separate
<taxonomicName id="6795C0A1FF9D274F54B5FB161400FB40" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[208,333,1225,1249]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="meufong">
<emphasis id="92E16730FF9D274F54B5FB1615AEFB41" box="[208,227,1225,1248]" italics="true" pageId="7" pageNumber="358">Q</emphasis>
.
<emphasis id="92E16730FF9D274F5494FB161400FB40" box="[241,333,1225,1249]" italics="true" pageId="7" pageNumber="358">meufong</emphasis>
</taxonomicName>
from
<taxonomicName id="6795C0A1FF9D274F55F5FB161493FB40" baseAuthorityName="Shoemaker" baseAuthorityYear="1933" box="[400,478,1225,1249]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="92E16730FF9D274F55F5FB161493FB40" box="[400,478,1225,1249]" italics="true" pageId="7" pageNumber="358">Q. lutzi</emphasis>
</taxonomicName>
, which has no carinae on the pleonites, two carinae on urosomite 1, and four carinae in urosomite
<quantity id="676D16C7FF9D274F5487FAD61441FA80" box="[226,268,1289,1313]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="358" unit="in" value="2.0">2 in</quantity>
female and juveniles and, notably, a smooth dorsum in males (i.e., carina formula 2-2-2-2-0 vs 0-0-0-2-4 or 0-0-0-0-0, respectively).
</paragraph>
<caption id="F4EAEBAAFF9D274F5414FBFD1176FBD0" ID-DOI="http://doi.org/10.5281/zenodo.5355558" ID-Zenodo-Dep="5355558" httpUri="https://zenodo.org/record/5355558/files/figure.png" pageId="7" pageNumber="358" startId="7.[113,148,1058,1080]" targetBox="[113,1451,189,1043]" targetPageId="7">
<paragraph id="A02ABB22FF9D274F5414FBFD1176FBD0" blockId="7.[113,1453,1058,1138]" pageId="7" pageNumber="358">
Fig. 6. 3D reconstructed male gnathopod 2 of
<taxonomicName id="6795C0A1FF9D274F5659FBFD164AFB96" authority="Hughes &amp; Kaji, 2016" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[572,775,1058,1079]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="meufong" status="sp. nov.">
<emphasis id="92E16730FF9D274F5659FBFD164AFB96" box="[572,775,1058,1079]" italics="true" pageId="7" pageNumber="358">Quadrivisio meufong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="89D2DA4BFF9D274F5770FBFC16C0FB99" box="[789,909,1058,1080]" pageId="7" pageNumber="358" rank="species">new species</taxonomicNameLabel>
, male 13.5 mm, PSUZC-CR 0302. A, overall image of carpus, propodus, and dactylus of the male gnathopod 2; B, magnified, proximal view of the expanded palm cuticle. Arrow heads indicate expanded palm structure; oc, outer layer of cuticle; ic, inner layer of cuticle. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A02ABB22FF9D274F5416FA56143CF9A0" blockId="7.[112,752,1417,1537]" pageId="7" pageNumber="358">
Material identified as
<taxonomicName id="6795C0A1FF9D274F553DFA5614B8FA00" authorityName="Stebbing" authorityYear="1907" box="[344,501,1417,1441]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="92E16730FF9D274F553DFA5614B8FA00" box="[344,501,1417,1441]" italics="true" pageId="7" pageNumber="358">Q. bengalensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="C404C6D3FF9D274F567EFA5617BDFA00" author="Bussarawich S" box="[539,752,1417,1441]" pageId="7" pageNumber="358" refId="ref3768" refString="Bussarawich S (1985) Gammaridean Amphipoda from mangroves in southern Thailand. Fifth Seminar on Mangrove Ecosystems Phuket, Thailand: 1 - 7, 11 figs." type="journal volume" year="1985">Bussarawich (1985)</bibRefCitation>
from
<collectingCountry id="D882FBB2FF9D274F54D5FA76145FFA60" box="[176,274,1449,1473]" name="Thailand" pageId="7" pageNumber="358">Thailand</collectingCountry>
do not have robust setae lining the inner margin of the telson and, therefore, may represent juveniles of
<taxonomicName id="6795C0A1FF9D274F54F5FA361442F9A0" authorityName="Hughes &amp; Kaji" authorityYear="2016" box="[144,271,1513,1537]" class="Malacostraca" family="Maeridae" genus="Quadrivisio" kingdom="Animalia" order="Amphipoda" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="meufong">
<emphasis id="92E16730FF9D274F54F5FA361442F9A0" box="[144,271,1513,1537]" italics="true" pageId="7" pageNumber="358">Q. meufong</emphasis>
</taxonomicName>
, instead.
</paragraph>
</subSubSection>
<subSubSection id="E88FE8A9FF9D274F5414F9F61792F9C0" pageId="7" pageNumber="358" type="distribution">
<paragraph id="A02ABB22FF9D274F5414F9F61792F9C0" blockId="7.[113,753,1577,1633]" pageId="7" pageNumber="358">
<emphasis id="92E16730FF9D274F5414F9F61449F9E0" bold="true" box="[113,260,1577,1601]" pageId="7" pageNumber="358">Distribution.</emphasis>
<collectingCountry id="D882FBB2FF9D274F556EF9F61420F9E0" box="[267,365,1577,1601]" name="Thailand" pageId="7" pageNumber="358">Thailand</collectingCountry>
.
<collectingRegion id="625175C0FF9D274F551DF9F61490F9E0" box="[376,477,1577,1601]" country="Thailand" name="Songkhla" pageId="7" pageNumber="358">Songkhla</collectingRegion>
Lake (current study). Ao Yon, Ao Nam Bor, Klong Pak Pan (
<bibRefCitation id="C404C6D3FF9D274F5664F996179EF9C0" author="Bussarawich S" box="[513,723,1609,1633]" pageId="7" pageNumber="358" refId="ref3768" refString="Bussarawich S (1985) Gammaridean Amphipoda from mangroves in southern Thailand. Fifth Seminar on Mangrove Ecosystems Phuket, Thailand: 1 - 7, 11 figs." type="journal volume" year="1985">Bussarawich, 1985</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>