373 lines
51 KiB
XML
373 lines
51 KiB
XML
<document ID-DOI="10.3390/insects13090796" ID-GBIF-Dataset="dd0a5d02-944b-4fbf-8856-edf45197ae1b" ID-Zenodo-Dep="7046409" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1662215982534" checkinUser="felipe" docAuthor="Boudinot, Brendon E., Richter, Adrian K., Hammel, Jörg U., Szwedo, Jacek, Bojarski, Błażej & Perrichot, Vincent" docDate="2022" docId="220587AFFFB9FFA614C9FE4CFB541E29" docLanguage="en" docName="Insects.13.796.1-19.pdf" docOrigin="Insects 73 (796)" docSource="http://dx.doi.org/10.3390/insects13090796" docTitle="Desyopone hereon Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="13" masterDocId="DE3CFFD7FFB1FFAA153CFFA3FFA7196B" masterDocTitle="Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="9" updateTime="1662217461897" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Genomic-Phenomic Reciprocal Illumination: Desyopone hereon gen. et sp. nov., an Exceptional Aneuretine-like Fossil Ant from Ethiopian Amber (Hymenoptera: Formicidae: Ponerinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Boudinot, Brendon E.</mods:namePart>
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<mods:affiliation>Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, Vor dem Neutor 1, 07743 Jena, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">boudinotb@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Richter, Adrian K.</mods:namePart>
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<mods:affiliation>Institut für Zoologie und Evolutionsforschung, Friedrich-Schiller-Universität Jena, Vor dem Neutor 1, 07743 Jena, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hammel, Jörg U.</mods:namePart>
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<mods:affiliation>Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Strasse 1, 21502 Geesthacht, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szwedo, Jacek</mods:namePart>
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<mods:affiliation>Laboratory of Evolutionary Entomology and Museum of Amber Inclusions, Faculty of Biology, University of Gdansk, 59 Wita Stwosza Street, 80 - 309 Gdansk, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bojarski, Błażej</mods:namePart>
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<mods:affiliation>Laboratory of Evolutionary Entomology and Museum of Amber Inclusions, Faculty of Biology, University of Gdansk, 59 Wita Stwosza Street, 80 - 309 Gdansk, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perrichot, Vincent</mods:namePart>
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<mods:affiliation>CNRS, Géosciences Rennes, University Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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<mods:nameIdentifier type="email">vincent.perrichot@univ-rennes1.fr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Insects</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>73</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>796</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>19</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3390/insects13090796</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.3390/insects13090796</mods:identifier>
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<mods:identifier type="GBIF-Dataset">dd0a5d02-944b-4fbf-8856-edf45197ae1b</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7046409</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:220587AFFFB9FFA614C9FE4CFB541E29" httpUri="http://treatment.plazi.org/id/220587AFFFB9FFA614C9FE4CFB541E29" lastPageId="12" lastPageNumber="13" pageId="8" pageNumber="9">
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<subSubSection box="[501,1120,495,520]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph blockId="8.[443,1497,261,754]" box="[501,1120,495,520]" pageId="8" pageNumber="9">
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<heading box="[501,1120,495,520]" level="2" pageId="8" pageNumber="9">
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<emphasis bold="true" box="[501,1120,495,520]" pageId="8" pageNumber="9">
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†
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<taxonomicName authority="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot, 2022" authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[514,729,495,519]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hereon" status="sp. nov.">
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<emphasis bold="true" box="[514,729,495,519]" italics="true" pageId="8" pageNumber="9">Desyopone hereon</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[735,828,496,520]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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Boudinot and Perrichot
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="description">
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<paragraph blockId="8.[443,1497,261,754]" box="[500,652,529,553]" pageId="8" pageNumber="9">
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(
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<figureCitation box="[508,643,529,553]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStartId-0="3.[444,514,1899,1921]" captionStartId-1="8.[444,514,1802,1824]" captionStartId-2="9.[444,514,1859,1881]" captionStartId-3="10.[444,514,638,660]" captionStartId-4="10.[444,514,1885,1907]" captionTargetBox-0="[190,1397,624,1866]" captionTargetBox-1="[152,1435,788,1767]" captionTargetBox-2="[140,1450,262,1821]" captionTargetBox-3="[204,1378,265,583]" captionTargetBox-4="[257,1318,815,1804]" captionTargetId-0="figure-388@3.[190,1397,624,1866]" captionTargetId-1="figure-301@8.[152,1435,788,1767]" captionTargetId-2="figure-174@9.[140,1450,262,1826]" captionTargetId-3="figure-543@10.[199,1386,260,602]" captionTargetId-4="figure-941@10.[217,1370,803,1848]" captionTargetPageId-0="3" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="10" captionText-0="Figure 1. Photograph of entire amber piece MAIG 6016, with indication of type specimens (labeled H for holotype, P1–P12 for paratypes) of †Desyopone hereon gen. et sp. nov., and with detailed views of seven of them (A–D). (A) paratype 4; (B) holotype; (C) paratypes 1–3; (D) paratypes 5–6. Scale bars: 0.5 mm." captionText-1="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." captionText-2="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." captionText-3="Figure 4. Volume renders of the segmented data from the head of †Desyopone hereon gen. et sp. nov., holotype MAIG 6016, showing the highly unusual mandibles. (A) full-face view; (B) ventral view; (C) dorsolateral anterior oblique view; (D) oral view. Scale bar approximate due to slightly unequal scaling of images for depiction." captionText-4="Figure 5. Diagrammatic representations of wing venation (top) and cell identities (bottom) based on the MAIG 6016 paratype 1 of †Desyopone hereon gen. et sp. nov. Cell names: CC = costal cell; RC1-2/SMC1 = first and second radial cells or submarginal cell 1; RC3/MC1 = third radial or first marginal cell; MC1/DC1 = first medial or first discal cell; RsC2/SMC2 = second sectorial or second submarginal cell; CuC1/SBC = first cubital or first subbasal cell; CuC2/SDC = second cubital or first subdiscal cell." figureDoi-0="http://doi.org/10.5281/zenodo.7046411" figureDoi-1="http://doi.org/10.5281/zenodo.7046413" figureDoi-2="http://doi.org/10.5281/zenodo.7046415" figureDoi-3="http://doi.org/10.5281/zenodo.7046417" figureDoi-4="http://doi.org/10.5281/zenodo.7046419" httpUri-0="https://zenodo.org/record/7046411/files/figure.png" httpUri-1="https://zenodo.org/record/7046413/files/figure.png" httpUri-2="https://zenodo.org/record/7046415/files/figure.png" httpUri-3="https://zenodo.org/record/7046417/files/figure.png" httpUri-4="https://zenodo.org/record/7046419/files/figure.png" pageId="8" pageNumber="9">Figures 1–5</figureCitation>
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)
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||
</paragraph>
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||
<paragraph blockId="8.[443,1497,261,754]" box="[500,1497,562,587]" pageId="8" pageNumber="9">
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||
<emphasis box="[500,661,563,587]" italics="true" pageId="8" pageNumber="9">
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ZooBank
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<collectionCode box="[602,661,563,586]" pageId="8" pageNumber="9">LSID</collectionCode>
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||
</emphasis>
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||
:
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||
<uri box="[674,1492,562,586]" pageId="8" pageNumber="9">
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||
urn: lsid:zoobank.org:act:
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||
<uuid box="[959,1492,562,586]" pageId="8" pageNumber="9">9E345965-6AA1-42D8-A468-187BABBB38D2</uuid>
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||
</uri>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="8" pageNumber="9" type="etymology">
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<paragraph blockId="8.[443,1497,261,754]" pageId="8" pageNumber="9">
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<emphasis box="[500,616,596,620]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
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. The specific epithet gratefully recognizes the Helmholtz-Zentrum Hereon, the research center which funds and operates the Imaging Beamline (P05) at
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<collectionCode box="[1359,1429,630,654]" pageId="8" pageNumber="9">DESY</collectionCode>
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, thus making the present work possible.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph blockId="8.[443,1497,261,754]" pageId="8" pageNumber="9">
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<materialsCitation collectionCode="MAIG" country="Ethiopia" location="Bashilo river gorge near Weldiya" pageId="8" pageNumber="9" specimenCode="MAIG 6016" specimenCount="1" specimenCount-male="1" stateProvince="Amhara Region" typeStatus="holotype">
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||
<emphasis box="[500,597,696,720]" italics="true" pageId="8" pageNumber="9">
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||
<typeStatus box="[500,597,696,720]" pageId="8" pageNumber="9">Holotype</typeStatus>
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||
</emphasis>
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||
.
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||
<specimenCount box="[612,670,696,720]" pageId="8" pageNumber="9" type="male">Male</specimenCount>
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||
(m),
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<specimenCode box="[731,865,696,721]" pageId="8" pageNumber="9">MAIG 6016</specimenCode>
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||
(
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||
<figureCitation box="[880,1005,696,721]" captionStart="Figure 1" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[190,1397,624,1866]" captionTargetId="figure-388@3.[190,1397,624,1866]" captionTargetPageId="3" captionText="Figure 1. Photograph of entire amber piece MAIG 6016, with indication of type specimens (labeled H for holotype, P1–P12 for paratypes) of †Desyopone hereon gen. et sp. nov., and with detailed views of seven of them (A–D). (A) paratype 4; (B) holotype; (C) paratypes 1–3; (D) paratypes 5–6. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7046411" httpUri="https://zenodo.org/record/7046411/files/figure.png" pageId="8" pageNumber="9">Figures 1B</figureCitation>
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||
,
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||
<figureCitation box="[1017,1076,697,721]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="8" pageNumber="9">2A,B</figureCitation>
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||
,
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<figureCitation box="[1086,1099,697,721]" captionStart="Figure 3" captionStartId="9.[444,514,1859,1881]" captionTargetBox="[140,1450,262,1821]" captionTargetId="figure-174@9.[140,1450,262,1826]" captionTargetPageId="9" captionText="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." figureDoi="http://doi.org/10.5281/zenodo.7046415" httpUri="https://zenodo.org/record/7046415/files/figure.png" pageId="8" pageNumber="9">3</figureCitation>
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and
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<figureCitation box="[1157,1170,697,721]" captionStart="Figure 4" captionStartId="10.[444,514,638,660]" captionTargetBox="[204,1378,265,583]" captionTargetId="figure-543@10.[199,1386,260,602]" captionTargetPageId="10" captionText="Figure 4. Volume renders of the segmented data from the head of †Desyopone hereon gen. et sp. nov., holotype MAIG 6016, showing the highly unusual mandibles. (A) full-face view; (B) ventral view; (C) dorsolateral anterior oblique view; (D) oral view. Scale bar approximate due to slightly unequal scaling of images for depiction." figureDoi="http://doi.org/10.5281/zenodo.7046417" httpUri="https://zenodo.org/record/7046417/files/figure.png" pageId="8" pageNumber="9">4</figureCitation>
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), deposited in the Museum of Amber Inclusions at University of Gda´nsk, Poland.
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</materialsCitation>
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7046413" ID-Zenodo-Dep="7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="8" pageNumber="9" startId="8.[444,514,1802,1824]" targetBox="[152,1435,788,1767]" targetPageId="8">
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||
<paragraph blockId="8.[443,1496,1802,1961]" pageId="8" pageNumber="9">
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<emphasis bold="true" box="[444,538,1802,1824]" pageId="8" pageNumber="9">Figure 2.</emphasis>
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Photographs of †
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<taxonomicName authority="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot, 2022" authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[722,893,1802,1824]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="hereon" status="gen. et sp. nov.">
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||
<emphasis box="[722,893,1802,1824]" italics="true" pageId="8" pageNumber="9">Desyopone hereon</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel box="[898,1057,1802,1823]" pageId="8" pageNumber="9" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, MAIG 6016. (
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||
<emphasis bold="true" box="[1205,1224,1802,1823]" pageId="8" pageNumber="9">A</emphasis>
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||
,
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<emphasis bold="true" box="[1230,1246,1802,1823]" pageId="8" pageNumber="9">B</emphasis>
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||
) holotype, anterodorsolateral views of head and metasoma; (
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||
<emphasis bold="true" box="[830,847,1836,1858]" pageId="8" pageNumber="9">C</emphasis>
|
||
) paratype 1, wing view; (
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||
<emphasis bold="true" box="[1107,1127,1836,1857]" pageId="8" pageNumber="9">D</emphasis>
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||
) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm.
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||
</paragraph>
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||
</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7046415" ID-Zenodo-Dep="7046415" httpUri="https://zenodo.org/record/7046415/files/figure.png" pageId="9" pageNumber="10" startId="9.[444,514,1859,1881]" targetBox="[140,1450,262,1821]" targetPageId="9">
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||
<paragraph blockId="9.[443,1491,1859,1984]" pageId="9" pageNumber="10">
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||
<emphasis bold="true" box="[444,538,1859,1881]" pageId="9" pageNumber="10">Figure 3.</emphasis>
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Amira volume renders of the raw data from the †
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||
<taxonomicName authority="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot, 2022" authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[1058,1229,1859,1881]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="hereon" status="gen. et sp. nov.">
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||
<emphasis box="[1058,1229,1859,1881]" italics="true" pageId="9" pageNumber="10">Desyopone hereon</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[1235,1393,1860,1881]" pageId="9" pageNumber="10" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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holotype MAIG 6016. (
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||
<emphasis bold="true" box="[601,620,1893,1914]" pageId="9" pageNumber="10">A</emphasis>
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||
) body in lateral view; (
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||
<emphasis bold="true" box="[885,901,1894,1915]" pageId="9" pageNumber="10">B</emphasis>
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||
) mesosoma in dorsal view; (
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||
<emphasis bold="true" box="[1224,1241,1893,1915]" pageId="9" pageNumber="10">C</emphasis>
|
||
) head, prothorax, and mesothorax in ventral view; (
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||
<emphasis bold="true" box="[756,776,1928,1949]" pageId="9" pageNumber="10">D</emphasis>
|
||
) petiole in dorsal view; (
|
||
<emphasis bold="true" box="[1041,1056,1928,1949]" pageId="9" pageNumber="10">E</emphasis>
|
||
) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite.
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||
</paragraph>
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</caption>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7046417" ID-Zenodo-Dep="7046417" httpUri="https://zenodo.org/record/7046417/files/figure.png" pageId="10" pageNumber="11" startId="10.[444,514,638,660]" targetBox="[204,1378,265,583]" targetPageId="10">
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||
<paragraph blockId="10.[443,1494,638,763]" pageId="10" pageNumber="11">
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||
<emphasis bold="true" box="[444,538,638,660]" pageId="10" pageNumber="11">Figure 4.</emphasis>
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Volume renders of the segmented data from the head of †
|
||
<taxonomicName authority="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot, 2022" authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[1150,1324,638,660]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hereon" status="gen. et sp. nov.">
|
||
<emphasis box="[1150,1324,638,660]" italics="true" pageId="10" pageNumber="11">Desyopone hereon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1330,1491,639,660]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, holotype MAIG 6016, showing the highly unusual mandibles. (
|
||
<emphasis bold="true" box="[1123,1142,672,693]" pageId="10" pageNumber="11">A</emphasis>
|
||
) full-face view; (
|
||
<emphasis bold="true" box="[1323,1339,673,694]" pageId="10" pageNumber="11">B</emphasis>
|
||
) ventral view; (
|
||
<emphasis bold="true" box="[451,468,706,728]" pageId="10" pageNumber="11">C</emphasis>
|
||
) dorsolateral anterior oblique view; (
|
||
<emphasis bold="true" box="[862,882,707,728]" pageId="10" pageNumber="11">D</emphasis>
|
||
) oral view. Scale bar approximate due to slightly unequal scaling of images for depiction.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7046419" ID-Zenodo-Dep="7046419" httpUri="https://zenodo.org/record/7046419/files/figure.png" pageId="10" pageNumber="11" startId="10.[444,514,1885,1907]" targetBox="[257,1318,815,1804]" targetPageId="10">
|
||
<paragraph blockId="10.[444,1494,1885,2078]" pageId="10" pageNumber="11">
|
||
<emphasis bold="true" box="[444,539,1885,1907]" pageId="10" pageNumber="11">Figure 5.</emphasis>
|
||
Diagrammatic representations of wing venation (
|
||
<emphasis bold="true" box="[1080,1116,1886,1907]" pageId="10" pageNumber="11">top</emphasis>
|
||
) and cell identities (
|
||
<emphasis bold="true" box="[1336,1414,1886,1907]" pageId="10" pageNumber="11">bottom</emphasis>
|
||
) based on the MAIG 6016 paratype 1 of †
|
||
<taxonomicName authority="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot, 2022" authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[816,995,1920,1941]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hereon" status="gen. et sp. nov.">
|
||
<emphasis box="[816,995,1920,1941]" italics="true" pageId="10" pageNumber="11">Desyopone hereon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1002,1173,1920,1941]" pageId="10" pageNumber="11" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Cell names: CC = costal cell; RC1-2/SMC1 = first and second radial cells or submarginal cell 1; RC3/MC1 = third radial or first marginal cell; MC1/DC1 = first medial or first discal cell; RsC2/SMC2 = second sectorial or second submarginal cell; CuC1/SBC = first cubital or first subbasal cell; CuC2/SDC = second cubital or first subdiscal cell.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<materialsCitation collectionCode="MAIG" pageId="11" pageNumber="12" specimenCode="MAIG 6016" specimenCount="12" specimenCount-male="12" typeStatus="paratype">
|
||
<emphasis box="[500,608,262,285]" italics="true" pageId="11" pageNumber="12">
|
||
<typeStatus box="[500,608,262,285]" pageId="11" pageNumber="12">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<specimenCount box="[634,741,261,285]" pageId="11" pageNumber="12" type="male">12 males</specimenCount>
|
||
(m). Synincluded with holotype in amber piece MAIG 6016 (
|
||
<figureCitation box="[451,634,295,319]" captionStart="Figure 1" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[190,1397,624,1866]" captionTargetId="figure-388@3.[190,1397,624,1866]" captionTargetPageId="3" captionText="Figure 1. Photograph of entire amber piece MAIG 6016, with indication of type specimens (labeled H for holotype, P1–P12 for paratypes) of †Desyopone hereon gen. et sp. nov., and with detailed views of seven of them (A–D). (A) paratype 4; (B) holotype; (C) paratypes 1–3; (D) paratypes 5–6. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7046411" httpUri="https://zenodo.org/record/7046411/files/figure.png" pageId="11" pageNumber="12">Figures 1A,C,D</figureCitation>
|
||
,
|
||
<figureCitation box="[646,705,295,319]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="11" pageNumber="12">2C,D</figureCitation>
|
||
and
|
||
<figureCitation box="[763,776,295,319]" captionStart="Figure 5" captionStartId="10.[444,514,1885,1907]" captionTargetBox="[257,1318,815,1804]" captionTargetId="figure-941@10.[217,1370,803,1848]" captionTargetPageId="10" captionText="Figure 5. Diagrammatic representations of wing venation (top) and cell identities (bottom) based on the MAIG 6016 paratype 1 of †Desyopone hereon gen. et sp. nov. Cell names: CC = costal cell; RC1-2/SMC1 = first and second radial cells or submarginal cell 1; RC3/MC1 = third radial or first marginal cell; MC1/DC1 = first medial or first discal cell; RsC2/SMC2 = second sectorial or second submarginal cell; CuC1/SBC = first cubital or first subbasal cell; CuC2/SDC = second cubital or first subdiscal cell." figureDoi="http://doi.org/10.5281/zenodo.7046419" httpUri="https://zenodo.org/record/7046419/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,637,328,352]" italics="true" pageId="11" pageNumber="12">Type locality</emphasis>
|
||
. Exact locality unknown in the
|
||
<location LSID="urn:lsid:plazi:treatment:220587AFFFB9FFA614C9FE4CFB541E29:AF736062FFBAFFA116CCFEEBFAD4180B" box="[1008,1395,328,353]" country="Ethiopia" name="Bashilo river gorge near Weldiya" pageId="11" pageNumber="12" stateProvince="Amhara Region">Bashilo river gorge near Weldiya</location>
|
||
, Semien Wollo Zone,
|
||
<collectingRegion box="[591,780,362,386]" country="Ethiopia" name="Amara" pageId="11" pageNumber="12">Amhara Region</collectingRegion>
|
||
,
|
||
<collectingCountry box="[791,894,362,386]" name="Ethiopia" pageId="11" pageNumber="12">Ethiopia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="11.[442,1497,261,1892]" box="[500,1361,395,420]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,639,395,419]" italics="true" pageId="11" pageNumber="12">Type horizon</emphasis>
|
||
. A fine siltstone/mudstone of Early Miocene age (16–23 Ma).
|
||
</paragraph>
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,591,429,453]" italics="true" pageId="11" pageNumber="12">Material</emphasis>
|
||
.
|
||
<typeStatus box="[606,715,429,453]" pageId="11" pageNumber="12">Holotype</typeStatus>
|
||
and
|
||
<typeStatus box="[770,884,429,453]" pageId="11" pageNumber="12">paratypes</typeStatus>
|
||
1–4 are nearly complete, preserved without apparent distortion but with integument entirely covered by a white, opaque (bacterial?) coat for
|
||
<typeStatus box="[443,562,496,520]" pageId="11" pageNumber="12">paratypes</typeStatus>
|
||
1, 2, 4 (
|
||
<figureCitation box="[656,815,496,520]" captionStart="Figure 1" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[190,1397,624,1866]" captionTargetId="figure-388@3.[190,1397,624,1866]" captionTargetPageId="3" captionText="Figure 1. Photograph of entire amber piece MAIG 6016, with indication of type specimens (labeled H for holotype, P1–P12 for paratypes) of †Desyopone hereon gen. et sp. nov., and with detailed views of seven of them (A–D). (A) paratype 4; (B) holotype; (C) paratypes 1–3; (D) paratypes 5–6. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7046411" httpUri="https://zenodo.org/record/7046411/files/figure.png" pageId="11" pageNumber="12">Figures 1A,C</figureCitation>
|
||
and
|
||
<figureCitation box="[877,914,496,520]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="11" pageNumber="12">2A</figureCitation>
|
||
).
|
||
<typeStatus box="[935,1054,496,520]" pageId="11" pageNumber="12">Paratypes</typeStatus>
|
||
5–12 mostly complete but variously preserved, more or less distorted by apparent dehydration (e.g.,
|
||
<figureCitation box="[1198,1317,529,553]" captionStart="Figure 1" captionStartId="3.[444,514,1899,1921]" captionTargetBox="[190,1397,624,1866]" captionTargetId="figure-388@3.[190,1397,624,1866]" captionTargetPageId="3" captionText="Figure 1. Photograph of entire amber piece MAIG 6016, with indication of type specimens (labeled H for holotype, P1–P12 for paratypes) of †Desyopone hereon gen. et sp. nov., and with detailed views of seven of them (A–D). (A) paratype 4; (B) holotype; (C) paratypes 1–3; (D) paratypes 5–6. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7046411" httpUri="https://zenodo.org/record/7046411/files/figure.png" pageId="11" pageNumber="12">Figure 1D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,610,563,586]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
. †
|
||
<taxonomicName authorityName="Boudinot & Richter & Hammel & Szwedo & Bojarski & Perrichot" authorityYear="2022" box="[649,850,563,587]" class="Insecta" family="Formicidae" genus="Desyopone" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="hereon">
|
||
<emphasis box="[649,850,563,587]" italics="true" pageId="11" pageNumber="12">Desyopone hereon</emphasis>
|
||
</taxonomicName>
|
||
is uniquely identifiable among all
|
||
<taxonomicName authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[1283,1404,563,587]" pageId="11" pageNumber="7" rank="subFamily" subFamily="Ponerinae">Ponerinae</taxonomicName>
|
||
by the distinctly elongate petiolar peduncle and the enlarged, lobate mandibles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,623,630,653]" italics="true" pageId="11" pageNumber="12">Description</emphasis>
|
||
.
|
||
<emphasis box="[639,793,630,653]" italics="true" pageId="11" pageNumber="12">Measurements</emphasis>
|
||
:
|
||
<typeStatus box="[807,916,630,654]" pageId="11" pageNumber="12">Holotype</typeStatus>
|
||
(
|
||
<typeStatus box="[931,1047,630,654]" pageId="11" pageNumber="12">paratypes</typeStatus>
|
||
) (taken from
|
||
<typeStatus box="[1202,1316,630,654]" pageId="11" pageNumber="12">paratypes</typeStatus>
|
||
1,2,4)—BL 3.11 (3.00–3.50), HW 0.57 (0.50–0.60), HWE 0.70 (0.66–0.70), SL 0.12 (0.10–0.12), EL 0.23, OLL 0.08 (0.08), OIL 0.16 (0.16), WL 1.25 (1.25),
|
||
<collectionCode box="[946,988,697,720]" country="France" name="Musee de Lectoure" pageId="11" pageNumber="12">ML</collectionCode>
|
||
0.52 (0.50–0.56),
|
||
<collectionCode box="[1188,1241,697,720]" country="Netherlands" name="Museum Wasmann" pageId="11" pageNumber="12">MW</collectionCode>
|
||
0.49, FWL (2.75),
|
||
<collectingCountry box="[1453,1492,697,720]" name="Philippines" pageId="11" pageNumber="12">PH</collectingCountry>
|
||
0.26 (0.30),
|
||
<collectionCode box="[575,607,731,754]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="11" pageNumber="12" type="Museum">PL</collectionCode>
|
||
~0.40 (0.40), GL 1.20 (1.00–1.50), GW 0.62 (0.65–0.80), HWI 81 (76–86), SI 21 (20), OCI 50 (50),
|
||
<collectionCode box="[646,671,764,787]" country="Russia" name="Paleontological Institute" pageId="11" pageNumber="12">PI</collectionCode>
|
||
154 (133).
|
||
</paragraph>
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,557,797,821]" italics="true" pageId="11" pageNumber="12">Head</emphasis>
|
||
(
|
||
<figureCitation box="[574,713,797,821]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="8.[444,514,1802,1824]" captionStartId-1="9.[444,514,1859,1881]" captionStartId-2="10.[444,514,638,660]" captionTargetBox-0="[152,1435,788,1767]" captionTargetBox-1="[140,1450,262,1821]" captionTargetBox-2="[204,1378,265,583]" captionTargetId-0="figure-301@8.[152,1435,788,1767]" captionTargetId-1="figure-174@9.[140,1450,262,1826]" captionTargetId-2="figure-543@10.[199,1386,260,602]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." captionText-1="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." captionText-2="Figure 4. Volume renders of the segmented data from the head of †Desyopone hereon gen. et sp. nov., holotype MAIG 6016, showing the highly unusual mandibles. (A) full-face view; (B) ventral view; (C) dorsolateral anterior oblique view; (D) oral view. Scale bar approximate due to slightly unequal scaling of images for depiction." figureDoi-0="http://doi.org/10.5281/zenodo.7046413" figureDoi-1="http://doi.org/10.5281/zenodo.7046415" figureDoi-2="http://doi.org/10.5281/zenodo.7046417" httpUri-0="https://zenodo.org/record/7046413/files/figure.png" httpUri-1="https://zenodo.org/record/7046415/files/figure.png" httpUri-2="https://zenodo.org/record/7046417/files/figure.png" pageId="11" pageNumber="12">Figures 2–4</figureCitation>
|
||
): Head capsule ovoid-elliptical in lateral view; in full-face view, posterior head margin broadly and evenly convex to compound eyes; oral region of head, i.e., the malar areas, clypeus, and mouthparts, narrower than distance between compound eyes; postgenal bridge short, about 2/5 the length of the head in full-face view as measured from the postocciput to the hypostoma; malar areas distinctly developed; clypeus medially bulging, laterally depressed. Compound eyes situated almost entirely in anterior half of head; eyes subspherical and relatively small, with their length being about 1/3 head length. Ocelli distant from compound eyes, with the lateral ocelli separated from the compound eyes by slightly more than one compound eye length. Antennal toruli located at about head midlength in full-face view, distinctly posterad the posterior clypeal margin. Antennal scapes short, just barely longer than wide, with their length distinctly <2 lateral ocellus diameters; scape length slightly more than one pedicel length, but not more than two. Pedicel about 1/3 the length of the first flagellomere. Flagellum narrow and long, with their length greatly exceeding mesosoma length. Mandibles flat and lobate in appearance, without distinct masticatory and basal margins; medial mandibular margin convex, curving more-or-less evenly around apex, which does not have incurvature; mandible length slightly greater than compound eye length. Labrum and paraglossae dangling at rest, both distinctly narrower than the distance between the mandibular bases. Maxillary stipes without transverse ridging.
|
||
</paragraph>
|
||
<paragraph blockId="11.[442,1497,261,1892]" pageId="11" pageNumber="12">
|
||
<emphasis box="[500,614,1434,1457]" italics="true" pageId="11" pageNumber="12">Mesosoma</emphasis>
|
||
(
|
||
<figureCitation box="[630,741,1433,1457]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="11" pageNumber="12">Figures 2</figureCitation>
|
||
and
|
||
<figureCitation box="[800,814,1433,1457]" captionStart="Figure 3" captionStartId="9.[444,514,1859,1881]" captionTargetBox="[140,1450,262,1821]" captionTargetId="figure-174@9.[140,1450,262,1826]" captionTargetPageId="9" captionText="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." figureDoi="http://doi.org/10.5281/zenodo.7046415" httpUri="https://zenodo.org/record/7046415/files/figure.png" pageId="11" pageNumber="12">3</figureCitation>
|
||
): Pronotum short and simple but with distinct muscular convexity as seen in lateral and dorsal views; posterad the anteromedian pronotal lobe (“nuchal lobe”), pronotum in the form of a simple arch, without distinct dorsal and ventral surfaces. Propleurae widely emarginate posteromedially, together forming a broad arch for the prosternum. Prosternum with basisternum apparently arcuate anteriorly; prosternal process developed. Mesoscutum somewhat narrow, with the anteroposterior length slightly greater than the lateromedial width. Notauli developed, and Y-shaped; meeting in the posterior half of the mesoscutum. Parapsides developed, although indistinct. Scutoscutellar sulcus broad, with at least five cross-costae. Mesoscutellum simple, convex, longer than broad in full-face view. Oblique mesopleural sulcus developed. Spiracular lobe (ventrad wing insertions) absent. Mesopleural area divided into dorsal and ventral regions; both regions dorsoventrally taller than anteroposteriorly long. Mesosternal and metasternal regions without processes. Propodeum convex, without armature or distinct sculpturation. Propodeal lobes developed, weak.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,548,262,285]" italics="true" pageId="12" pageNumber="13">Legs</emphasis>
|
||
: Mesotibiae with no spur, metatibiae with a single, pectinate spur. Pretarsal claws narrow and simple, without additional teeth. Arolia well-developed but not very large.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,622,329,352]" italics="true" pageId="12" pageNumber="13">Fore wings</emphasis>
|
||
(
|
||
<figureCitation box="[638,796,328,352]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="12" pageNumber="13">Figures 2C,D</figureCitation>
|
||
and
|
||
<figureCitation box="[857,871,328,352]" captionStart="Figure 5" captionStartId="10.[444,514,1885,1907]" captionTargetBox="[257,1318,815,1804]" captionTargetId="figure-941@10.[217,1370,803,1848]" captionTargetPageId="10" captionText="Figure 5. Diagrammatic representations of wing venation (top) and cell identities (bottom) based on the MAIG 6016 paratype 1 of †Desyopone hereon gen. et sp. nov. Cell names: CC = costal cell; RC1-2/SMC1 = first and second radial cells or submarginal cell 1; RC3/MC1 = third radial or first marginal cell; MC1/DC1 = first medial or first discal cell; RsC2/SMC2 = second sectorial or second submarginal cell; CuC1/SBC = first cubital or first subbasal cell; CuC2/SDC = second cubital or first subdiscal cell." figureDoi="http://doi.org/10.5281/zenodo.7046419" httpUri="https://zenodo.org/record/7046419/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
|
||
): Costal vein (C) present, complete. Rsf1 diverging from Sc+
|
||
<collectionCode box="[554,573,362,386]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
|
||
+Rs well proximad pterostigma, with Sc+
|
||
<collectionCode box="[1082,1101,362,386]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
|
||
abscissa about 1/3 pterostigma length. Pterostigma well-developed, long, and narrow, with its length> 5 × its width. Rf distad pterostigma tubular. M+Cu tubular. Mf1 diverging from M+Cu at or slightly distad crossvein cu-a; this abscissa very weakly curved and meeting Rf1 at a distinct oblique angle. Rs+M tubular. Rsf2–3 diverging from Rs+M proximad 1m-cu. Crossvein 1r-rs absent. Crossvein 2r-rs anterior juncture at 2/3 length of pterostigma; this crossvein meeting Rsf proximad crossvein 2rs-m, which is tubular (
|
||
<typeStatus box="[968,1074,563,587]" pageId="12" pageNumber="13">paratype</typeStatus>
|
||
4 has 2rs-m duplicated on the right wing, see
|
||
<figureCitation box="[555,671,596,620]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="12" pageNumber="13">Figure 2D</figureCitation>
|
||
). Rsf4+ tubular, meeting Rf distally). Mf2 (=abscissa between Rs+M and 1m-cu) short, shorter than 2r-rs and 2rs-m. Mf3 similar in length to but distinctly shorter than Rsf2–3. Mf4+ tubular proximally, becoming nebulous distally, with this occurring at a distance that is about 3 × the length of 2rs-m. Cuf3 (=abscissa of Cu after 1m-cu) joined to 1A posteriorly; 1A is tubular for its entire length. Submarginal cells 1 and 2 similar in size and shape, but with 2 distinctly smaller than 1. Marginal cell 1 long and narrow, with a length that is about 1.5 × pterostigma length. Discal cell 1 subrhomboidal, its length slightly less than 2 × its anteroposterior width. Subdiscal cell closed.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,626,864,888]" italics="true" pageId="12" pageNumber="13">Hind wings</emphasis>
|
||
(
|
||
<figureCitation box="[640,764,864,888]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="12" pageNumber="13">Figures 2C</figureCitation>
|
||
and
|
||
<figureCitation box="[822,835,864,888]" captionStart="Figure 5" captionStartId="10.[444,514,1885,1907]" captionTargetBox="[257,1318,815,1804]" captionTargetId="figure-941@10.[217,1370,803,1848]" captionTargetPageId="10" captionText="Figure 5. Diagrammatic representations of wing venation (top) and cell identities (bottom) based on the MAIG 6016 paratype 1 of †Desyopone hereon gen. et sp. nov. Cell names: CC = costal cell; RC1-2/SMC1 = first and second radial cells or submarginal cell 1; RC3/MC1 = third radial or first marginal cell; MC1/DC1 = first medial or first discal cell; RsC2/SMC2 = second sectorial or second submarginal cell; CuC1/SBC = first cubital or first subbasal cell; CuC2/SDC = second cubital or first subdiscal cell." figureDoi="http://doi.org/10.5281/zenodo.7046419" httpUri="https://zenodo.org/record/7046419/files/figure.png" pageId="12" pageNumber="13">5</figureCitation>
|
||
): Wing with eight distal hamuli. Jugal lobe absent. C not distinctly developed.
|
||
<collectionCode box="[698,716,898,921]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
|
||
splitting from Sc+
|
||
<collectionCode box="[931,950,897,921]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
|
||
+Rs distad crossvein 1rs-m; Rf incomplete, not reaching anterior wing margin. Rsf tubular for a distance that is about 2 × the length of Sc+
|
||
<collectionCode box="[486,505,964,988]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="12" pageNumber="13">R</collectionCode>
|
||
+Rs distad 1rs-m. M+Cu splitting well distad crossvein cu-a. Mf1 meeting rs-m at a broad, oblique angle; Mf developed as a stud distad this juncture. Cuf developed distad the split of M+Cu, but exact condition uncertain. Crossvein cu-a long, slightly longer than the length of Mf1; this crossvein situated proximad split of M+Cu by about twice its length. Anal vein (A) tubular past its juncture with cu-a. Anal cell relatively long; M+Cu proximad cu-a distinctly longer than M+Cu distad cu-a.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,608,1166,1189]" italics="true" pageId="12" pageNumber="13">Metasoma</emphasis>
|
||
(
|
||
<figureCitation box="[623,745,1165,1189]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="12" pageNumber="13">Figures 2B</figureCitation>
|
||
and
|
||
<figureCitation box="[802,887,1165,1189]" captionStart="Figure 3" captionStartId="9.[444,514,1859,1881]" captionTargetBox="[140,1450,262,1821]" captionTargetId="figure-174@9.[140,1450,262,1826]" captionTargetPageId="9" captionText="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." figureDoi="http://doi.org/10.5281/zenodo.7046415" httpUri="https://zenodo.org/record/7046415/files/figure.png" pageId="12" pageNumber="13">3A,D,E</figureCitation>
|
||
): Petiole nodiform and distinctly pedunculate, albeit without a marked inflection between its anterior portion and the anterior surface of the petiolar node; peduncle about 2/5 petiole length; petiolar node height about 3/5 entire petiole length; node broad and convex; posterior collar well-developed; tergosternal fusion absent, laterotergites present; sternum low and very weakly sinuate in lateral view, without an anteroventral (=subpetiolar) process, nor with a posterior process; posterior sternal margin distinctly notched. Helcium infraaxial (=below abdominal segment
|
||
<collectionCode box="[1325,1352,1367,1390]" pageId="12" pageNumber="13">III</collectionCode>
|
||
midheight); helcial tergite broad and overlapping sternite laterally. Prora not distinctly developed. Gastral segments homonomous in appearance and gradually decreasing in length from abdominal segment III to VIII. Abdominal segment IV without cinctus (=constriction). Abdominal spiracles IV–VIII obscured by preceding tergites. Abdominal tergum VIII apparently simple. Abdominal sternum IX lobate and somewhat narrow, with a length that is about twice its width. Cerci (=pygostyles) developed.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,601,1600,1624]" italics="true" pageId="12" pageNumber="13">Genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[617,746,1600,1624]" captionStart="Figure 2" captionStartId="8.[444,514,1802,1824]" captionTargetBox="[152,1435,788,1767]" captionTargetId="figure-301@8.[152,1435,788,1767]" captionTargetPageId="8" captionText="Figure 2. Photographs of †Desyopone hereon gen. et sp. nov., MAIG 6016. (A,B) holotype, anterodorsolateral views of head and metasoma; (C) paratype 1, wing view; (D) paratype 4, wing view. AtIII/MtII: abdominal tergite III/metasomal tergite II; AsIX/MsVIII: abdominal sternite IX/metasomal sternite VIII. Note that the specimen figured in D has a duplicated crossvein 2rs-m on right fore wing. Scale bars: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7046413" httpUri="https://zenodo.org/record/7046413/files/figure.png" pageId="12" pageNumber="13">Figures 2B</figureCitation>
|
||
and
|
||
<figureCitation box="[807,844,1600,1624]" captionStart="Figure 3" captionStartId="9.[444,514,1859,1881]" captionTargetBox="[140,1450,262,1821]" captionTargetId="figure-174@9.[140,1450,262,1826]" captionTargetPageId="9" captionText="Figure 3. Amira volume renders of the raw data from the †Desyopone hereon gen. et sp. nov. holotype MAIG 6016. (A) body in lateral view; (B) mesosoma in dorsal view; (C) head, prothorax, and mesothorax in ventral view; (D) petiole in dorsal view; (E) petiole in ventral view, highlighting the petiolar tergum, laterotergites, and sternum, and the helcial tergite and sternite." figureDoi="http://doi.org/10.5281/zenodo.7046415" httpUri="https://zenodo.org/record/7046415/files/figure.png" pageId="12" pageNumber="13">3A</figureCitation>
|
||
): Cupula present (only visible in µ- CT cross-sections). Gonopods longer than tall. Gonostyli broad proximally, indistinctly set off from gonocoxa, and lobate in appearance, being narrowly rounded apically. Lateropenites (=digiti) thickened apically, downcurved. Penites (=penisvalvae) apparently longer than tall, curving strongly to their narrowly lobate apices.
|
||
</paragraph>
|
||
<paragraph blockId="12.[442,1497,261,1859]" pageId="12" pageNumber="13">
|
||
<emphasis box="[500,589,1768,1791]" italics="true" pageId="12" pageNumber="13">Setation</emphasis>
|
||
: Body with two primary hair classes: (1) short pubescence, which covers the head and all segments of the antennae, mesosoma, legs, and metasoma; (2) long hairs, which are sparse on all body regions, but are denser on the metasoma.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |