177 lines
20 KiB
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177 lines
20 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5557619" ID-GBIF-Dataset="9de32fb5-ae5c-4552-afdb-9cfdcad56ce9" ID-GBIF-Taxon="188614998" ID-Zenodo-Dep="5557619" approvalRequired="2" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" checkinTime="1633713770872" checkinUser="carolina" docAuthor="Deschodt, Christian M. & Davis, Adrian L. V." docDate="2017" docId="03C7F607FFFDFFC7A58D3157EEB061AD" docLanguage="en" docName="AnnDNMNH.7.109-114.pdf.imf" docOrigin="Annals of the Ditsong National Museum of Natural History 7 (7)" docTitle="Namakwanus scholtzi Deschodt & Davis 2017, spec. nov." docType="treatment" docVersion="6" lastPageId="3" lastPageNumber="112" masterDocId="FFFE8E7FFFFCFFC4A72A3246EF2D645E" masterDocTitle="Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="114" masterPageNumber="109" pageId="1" pageNumber="110" updateTime="1634154866724" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Transfer of three species of Namakwanus Scholtz & Howden to Versicorpus Deschodt, Davis & Scholtz or to Namaphilus gen. nov., with descriptions of two new species (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Deschodt, Christian M.</mods:namePart>
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<mods:affiliation>Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Davis, Adrian L. V.</mods:namePart>
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<mods:affiliation>Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Annals of the Ditsong National Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2017-05-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>7</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>7</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>109</mods:start>
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<mods:end>114</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.5562912</mods:identifier>
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<mods:identifier type="GBIF-Dataset">9de32fb5-ae5c-4552-afdb-9cfdcad56ce9</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5562912</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5557619" ID-GBIF-Taxon="188614998" ID-Zenodo-Dep="5557619" LSID="urn:lsid:plazi:treatment:03C7F607FFFDFFC7A58D3157EEB061AD" httpUri="http://treatment.plazi.org/id/03C7F607FFFDFFC7A58D3157EEB061AD" lastPageId="3" lastPageNumber="112" pageId="1" pageNumber="110">
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<subSubSection box="[679,1154,785,807]" pageId="1" pageNumber="110" type="nomenclature">
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<paragraph blockId="1.[679,1154,785,807]" box="[679,1154,785,807]" pageId="1" pageNumber="110">
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<heading box="[679,1154,785,807]" level="2" pageId="1" pageNumber="110">
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<emphasis bold="true" box="[679,1048,785,807]" pageId="1" pageNumber="110">
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<taxonomicName authority="Deschodt & Davis, 2017" authorityName="Deschodt & Davis" authorityYear="2017" box="[679,927,785,807]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="110" phylum="Arthropoda" rank="species" species="scholtzi" status="spec. nov.">Namakwanus scholtzi</taxonomicName>
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<taxonomicNameLabel box="[934,1048,785,807]" pageId="1" pageNumber="110" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<figureCitation box="[1061,1154,785,807]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[639,669,1614,1631]" captionStartId-1="3.[639,669,1187,1204]" captionTargetBox-0="[195,1130,199,1593]" captionTargetBox-1="[218,1104,185,1174]" captionTargetId-0="figure-14@2.[188,1138,185,1602]" captionTargetId-1="figure-305@3.[230,1104,185,1174]" captionTargetPageId-0="2" captionText-0="Fig. 1 Plate showing the habitus and aedeagus of one new species and habitus of all other species treated in this paper:A: Namakwanus irishi Scholtz & Howden,1987;B: Namakwanus scholtzi spec.nov.;C: Aedeagus of Namakwanus scholtzi spec.nov.;D: Namaphilus davisi (Deschodt & Scholtz,2007) comb.nov.;E: Namaphilus endroedyi (Deschodt,Davis & Scholtz,2011) comb.nov.;F: Namaphilus ameibensis spec.nov.;G: Versicorpus streyi (Frolov,2005) comb.nov.;H: Versicorpus erongoense Deschodt,Davis & Scholtz,2011." captionText-1="Fig. 2 Map showing the collecting localities of all the known species of the Byrrhidium group.The black outline indicates the distribution of rock hyraxes (shapefile supplied by IUCN). See online version for map in colour." figureDoi-0="http://doi.org/10.5281/zenodo.5562914" figureDoi-1="http://doi.org/10.5281/zenodo.5562916" httpUri-0="https://zenodo.org/record/5562914/files/figure.png" httpUri-1="https://zenodo.org/record/5562916/files/figure.png" pageId="1" pageNumber="110">Figs 1–2</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="110" type="materials_examined">
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<paragraph blockId="1.[679,1201,841,917]" pageId="1" pageNumber="110">
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<materialsCitation ID-GBIF-Occurrence="3382697307" collectingDate="1959-05-11" collectionCode="TMSA" collectorName="H. Dick Brown" country="Namibia" location="Kuiseb River" municipality="Gobabeb" pageId="1" pageNumber="110" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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SPECIMEN EXAMINED.
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<typeStatus box="[946,1040,841,863]" pageId="1" pageNumber="110">Holotype</typeStatus>
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<specimenCount box="[1053,1063,844,864]" pageId="1" pageNumber="110" type="male">♂</specimenCount>
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: [
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<collectingCountry box="[1087,1192,841,863]" name="Namibia" pageId="1" pageNumber="110" value="Namibia">NAMIBIA</collectingCountry>
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]:
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<collectingMunicipality box="[679,781,868,890]" pageId="1" pageNumber="110">Gobabeb</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:03C7F607FFFDFFC7A58D3157EEB061AD:8EB111CAFFFDFFC5A43D3122ECB66724" box="[791,923,868,890]" country="Namibia" municipality="Gobabeb" name="Kuiseb River" pageId="1" pageNumber="110">Kuiseb River</location>
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,
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<location LSID="urn:lsid:plazi:treatment:03C7F607FFFDFFC7A58D3157EEB061AD:8EB111CAFFFDFFC5A48C3122EB6D6724" box="[934,1088,868,890]" country="Namibia" municipality="Gobabeb" name="Central Namib" pageId="1" pageNumber="110">Central Namib</location>
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, S. W. Afr.,
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<date box="[679,780,895,917]" pageId="1" pageNumber="110" value="1959-05-11">
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<collectingDate box="[679,780,895,917]" pageId="1" pageNumber="110" value="1959-05-11">11.v.1959</collectingDate>
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</date>
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,
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<collectorName box="[790,942,895,917]" pageId="1" pageNumber="110">H. Dick Brown</collectorName>
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.– (
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<collectionCode box="[968,1038,895,917]" collectionName="South Africa, Gauteng, Pretoria, Transvaal Museum" pageId="1" pageNumber="110">TMSA</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="110" type="etymology">
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<paragraph blockId="1.[679,1200,950,1026]" pageId="1" pageNumber="110">ETYMOLOGY. This species is named in honour of Professor Clarke H. Scholtz, for his extensive and lifelong contribution to entomology.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="112" pageId="1" pageNumber="110" type="description">
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<paragraph blockId="1.[679,1201,1059,1724]" box="[679,1056,1059,1081]" pageId="1" pageNumber="110">DESCRIPTION. Dark brown, convex.</paragraph>
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<paragraph blockId="1.[679,1201,1059,1724]" pageId="1" pageNumber="110">Head: two long, distinct, slightly upturned clypeal teeth and one much smaller tooth medially on lower margin of clypeus. Slight red tinge laterally.</paragraph>
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<paragraph blockId="1.[679,1201,1059,1724]" pageId="1" pageNumber="110">Pronotum: punctate with round punctures of variable size, surface shagreened. Laterally clearly flattenedwith ridge in median one-third justinside of lateral margin. Ridge parallel with outside margin anteriorly and curving inward posteriorly. Lateral sides with slight red tinge.</paragraph>
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<paragraph blockId="1.[679,1201,1059,1724]" pageId="1" pageNumber="110">Protibia: surface shagreened, outside margin anteriorly tridentate in last one-third and slightly sinuous to dentate in first two-thirds. Outside two denticles larger than median denticle. Median dorsal ridge present, angulate at posterior denticle of outside margin. Golden brown setae present, longer terminally.Ventral surface withmedian ridge.</paragraph>
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<paragraph blockId="1.[679,1201,1059,1724]" pageId="1" pageNumber="110">Elytra: convex. Surface shagreened, punctures on second to fourth intervals more or less in straight double lines, punctures on all other intervals forming more or less straight single lines. Punctures and striae not well defined.</paragraph>
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<paragraph blockId="1.[679,1201,1059,1724]" lastBlockId="3.[126,648,1287,1389]" lastPageId="3" lastPageNumber="112" pageId="1" pageNumber="110">Sterna: prosternum with clear round and adjacent punctures. Mesometasternal suture distinct and very slightly curved, punctures on mesosternum larger and further apart than on metasternum, surface shagreened.</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5562914" ID-Zenodo-Dep="5562914" httpUri="https://zenodo.org/record/5562914/files/figure.png" pageId="2" pageNumber="111" startId="2.[639,669,1614,1631]" targetBox="[195,1130,199,1593]" targetPageId="2">
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<paragraph blockId="2.[126,1200,1614,1723]" pageId="2" pageNumber="111">
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<heading bold="true" box="[639,686,1614,1631]" centered="true" fontSize="7" level="2" pageId="2" pageNumber="111" reason="4">
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<emphasis bold="true" box="[639,686,1614,1631]" pageId="2" pageNumber="111">Fig. 1</emphasis>
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</heading>
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Plate showing the habitus and aedeagus of one new species and habitus of all other species treated in this paper:
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<emphasis bold="true" box="[1062,1082,1641,1658]" pageId="2" pageNumber="111">A:</emphasis>
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<taxonomicName authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="irishi">
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Namakwanus irishi
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<bibRefCitation author="SCHOLTZ, C. H. & HOWDEN, H. F." box="[169,372,1663,1680]" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="2" pageNumber="111" pagination="75 - 119" part="50" refId="ref4071" refString="SCHOLTZ, C. H. and HOWDEN, H. F., 1987. A revision of the southern African Cathonina [sic] (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa 50: 75 - 119." title="A revision of the southern African Cathonina [sic] (Coleoptera: Scarabaeidae: Scarabaeinae)" type="journal article" year="1987">Scholtz & Howden,1987</bibRefCitation>
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</taxonomicName>
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;
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<emphasis bold="true" box="[378,397,1663,1680]" pageId="2" pageNumber="111">B:</emphasis>
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<taxonomicName authority="Deschodt & Davis, 2017" authorityName="Deschodt & Davis" authorityYear="2017" box="[401,580,1663,1680]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="scholtzi" status="spec.nov.">Namakwanus scholtzi</taxonomicName>
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<taxonomicNameLabel box="[584,670,1663,1680]" pageId="2" pageNumber="111" rank="species">spec.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[673,693,1663,1680]" pageId="2" pageNumber="111">C:</emphasis>
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Aedeagus of
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<taxonomicName authority="Deschodt & Davis, 2017" authorityName="Deschodt & Davis" authorityYear="2017" box="[805,983,1663,1680]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="scholtzi" status="spec.nov.">Namakwanus scholtzi</taxonomicName>
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<taxonomicNameLabel box="[988,1073,1663,1680]" pageId="2" pageNumber="111" rank="species">spec.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[1077,1097,1663,1680]" pageId="2" pageNumber="111">D:</emphasis>
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<taxonomicName authority="(Deschodt & Scholtz, 2007)" authorityName="Deschodt & Davis" authorityYear="2017" baseAuthorityName="Deschodt & Scholtz" baseAuthorityYear="2007" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="davisi" status="comb.nov.">
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Namaphilus davisi (
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<bibRefCitation author="DESCHODT, C. M. & KRYGER, U. & SCHOLTZ, C. H." box="[181,394,1684,1701]" journalOrPublisher="Insect Systematics & Evolution" pageId="2" pageNumber="111" pagination="361 - 376" part="38" refId="ref3667" refString="DESCHODT, C. M., KRYGER, U. and SCHOLTZ, C. H., 2007. New taxa of relictual Canthonini dung beetles (Scarabaeidae: Scarabaeinae) utilizing rock hyrax middens as refuges in south-western Africa. Insect Systematics & Evolution 38: 361 - 376." title="New taxa of relictual Canthonini dung beetles (Scarabaeidae: Scarabaeinae) utilizing rock hyrax middens as refuges in south-western Africa" type="journal article" year="2007">Deschodt & Scholtz,2007</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[402,493,1684,1701]" pageId="2" pageNumber="111" rank="species">comb.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[496,515,1684,1701]" pageId="2" pageNumber="111">E:</emphasis>
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<taxonomicName authority="(Deschodt, Davis & Scholtz, 2011)" authorityName="Deschodt & Davis" authorityYear="2017" baseAuthorityName="Deschodt, Davis & Scholtz" baseAuthorityYear="2011" box="[518,983,1684,1701]" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="endroedyi" status="comb.nov.">
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Namaphilus endroedyi (
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<bibRefCitation author="DESCHODT, C. M. & DAVIS, A. L. V. & SCHOLTZ, C. H." box="[710,978,1684,1701]" journalOrPublisher="Insect Systematics & Evolution" pageId="2" pageNumber="111" pagination="277 - 294" part="42" refId="ref3549" refString="DESCHODT, C. M., DAVIS, A. L. V. and SCHOLTZ, C. H., 2011. New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa. Insect Systematics & Evolution 42 (3): 277 - 294." title="New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa" type="journal article" year="2011">Deschodt,Davis & Scholtz,2011</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[986,1076,1684,1701]" pageId="2" pageNumber="111" rank="species">comb.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[1080,1097,1684,1701]" pageId="2" pageNumber="111">F:</emphasis>
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<taxonomicName authority="Deschodt & Davis, 2017" authorityName="Deschodt & Davis" authorityYear="2017" class="Insecta" family="Scarabaeidae" genus="Namaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="ameibensis" status="spec.nov.">Namaphilus ameibensis</taxonomicName>
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<taxonomicNameLabel box="[223,308,1706,1723]" pageId="2" pageNumber="111" rank="species">spec.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[311,332,1706,1723]" pageId="2" pageNumber="111">G:</emphasis>
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<taxonomicName authority="(Frolov, 2005)" authorityName="Deschodt & Davis" authorityYear="2017" baseAuthorityName="Frolov" baseAuthorityYear="2005" box="[335,594,1706,1723]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="streyi" status="comb.nov.">
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Versicorpus streyi (
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<bibRefCitation author="FROLOV, A. V." box="[488,590,1706,1723]" editor="KONSTANTINOV, A. & TISHECHKIN, A. & PENEV, L." journalOrPublisher="Pensoft, Sofia" pageId="2" pageNumber="111" pagination="245 - 247" refId="ref3720" refString="FROLOV, A. V., 2005. A new species of the dung beetle genus Namakwanus (Coleoptera, Scarabaeidae) from Namibia. In: KONSTANTINOV, A., TISHECHKIN, A. and PENEV, L., eds, Contributions to systematics and biology of beetles, papers celebrating the 80 th birthday of Igor Konstantinovich Lopatin, pp. 245 - 247. Pensoft, Sofia." title="A new species of the dung beetle genus Namakwanus (Coleoptera, Scarabaeidae) from Namibia" type="book chapter" volumeTitle="Contributions to systematics and biology of beetles, papers celebrating the 80 th birthday of Igor Konstantinovich Lopatin" year="2005">Frolov,2005</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel box="[598,688,1706,1723]" pageId="2" pageNumber="111" rank="species">comb.nov.</taxonomicNameLabel>
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;
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<emphasis bold="true" box="[691,710,1706,1723]" pageId="2" pageNumber="111">H:</emphasis>
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<taxonomicName authority="Deschodt, Davis & Scholtz, 2011" authorityName="Deschodt, Davis & Scholtz" authorityYear="2011" box="[713,1182,1706,1723]" class="Insecta" family="Scarabaeidae" genus="Versicorpus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="erongoense">
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Versicorpus erongoense
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<bibRefCitation author="DESCHODT, C. M. & DAVIS, A. L. V. & SCHOLTZ, C. H." box="[916,1182,1706,1723]" journalOrPublisher="Insect Systematics & Evolution" pageId="2" pageNumber="111" pagination="277 - 294" part="42" refId="ref3549" refString="DESCHODT, C. M., DAVIS, A. L. V. and SCHOLTZ, C. H., 2011. New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa. Insect Systematics & Evolution 42 (3): 277 - 294." title="New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa" type="journal article" year="2011">Deschodt,Davis & Scholtz,2011</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.5562916" ID-Zenodo-Dep="5562916" httpUri="https://zenodo.org/record/5562916/files/figure.png" pageId="3" pageNumber="112" startId="3.[639,669,1187,1204]" targetBox="[218,1104,185,1174]" targetPageId="3">
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<paragraph blockId="3.[126,1200,1187,1251]" pageId="3" pageNumber="112">
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<heading bold="true" box="[639,686,1187,1204]" centered="true" fontSize="7" level="2" pageId="3" pageNumber="112" reason="4">
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<emphasis bold="true" box="[639,686,1187,1204]" pageId="3" pageNumber="112">Fig. 2</emphasis>
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</heading>
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Map showing the collecting localities of all the known species of the
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<taxonomicName authorityName="OvaLe Harold" authorityYear="1869" box="[684,773,1213,1230]" class="Insecta" family="Scarabaeidae" genus="Byrrhidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="112" phylum="Arthropoda" rank="genus">Byrrhidium</taxonomicName>
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group.The black outline indicates the distribution of rock hyraxes (shapefile supplied by IUCN). See online version for map in colour.
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</paragraph>
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</caption>
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<paragraph blockId="3.[126,648,1287,1389]" pageId="3" pageNumber="112">Pygidium: surface shagreened and punctate, puncturessmalland further apartthantheirdiameter.</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="112" type="discussion">
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<paragraph blockId="3.[126,647,1421,1523]" pageId="3" pageNumber="112">
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DISCUSSION. This new species can be separated from
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<taxonomicName authorityName="Scholtz & Howden" authorityYear="1987" box="[180,256,1448,1470]" class="Insecta" family="Scarabaeidae" genus="Namakwanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="111" phylum="Arthropoda" rank="species" species="irishi">
|
||
<emphasis box="[180,256,1448,1470]" italics="true" pageId="3" pageNumber="112">N. irishi</emphasis>
|
||
</taxonomicName>
|
||
by the tips of the clypeal horns being slightly further apart, more upturned and slightly pointing inward at the apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |