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<document id="15543D989E8D7EE0D3552130C6EF5DAB" ID-CLB-Dataset="21809" ID-DOI="10.5852/ejt.2020.652" ID-GBIF-Dataset="64fd24bb-ee1d-44d7-ac6a-25e8d5f96777" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860236" ID-ZooBank="DC65C142-53F1-4416-A916-8F78C27DCF93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590592466678" checkinUser="valdenar" docAuthor="Garbin, Rafaella C., Bandyopadhyay, Saswati &amp; Joyce, Walter G." docDate="2020" docId="03C7CF0BC0751D36FF6CA84AFAD09122" docLanguage="en" docName="ejt-652_garbin_bandyopadhyay_joyce.pdf.imf" docOrigin="European Journal of Taxonomy 652" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Geoemydidae Theobald 1868" docType="key" docVersion="8" lastPageNumber="67" masterDocId="FFFEB773C0341D74FFD1A908FFA79736" masterDocTitle="A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan" masterLastPageNumber="67" masterPageNumber="1" pageNumber="66" updateTime="1698841626966" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3E11D4295997DCDC053349312BC8B619">A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan</mods:title>
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<mods:namePart id="883FCB204F285F16EFEA39477BD3727A">Garbin, Rafaella C.</mods:namePart>
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<mods:affiliation id="2173D31C11B35C4E9D7F48FF65397E65">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. &amp; urn: lsid: zoobank. org: author: A 8 FCEF 2 B- 7524 - 46 C 3 - 808 C-D 9 CBFBC 08017 &amp; Corresponding author: garbin. rafaella @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="CC130DD6E5B23F50423DAE8E736F5CAF" type="email">garbin.rafaella@gmail.com</mods:nameIdentifier>
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<mods:namePart id="48C8EC3F42629ABDE9DDCC763C4F8FB7">Bandyopadhyay, Saswati</mods:namePart>
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<mods:affiliation id="1D9D28BFAD33435142641151A658205C">Geological Studies Unit, Indian Statistical Institute, Kolkata; Barrackpore Trunk Road 233, Kolkata, India. &amp; Email: saswati @ isical. ac. in &amp; urn: lsid: zoobank. org: author: 02005 D 35 - 6 DA 9 - 4 A 7 E-B 838 - 16 F 4 FEF 108 DA</mods:affiliation>
<mods:nameIdentifier id="F85883A6D51DF068E2FC2881734D0856" type="email">saswati@isical.ac.in</mods:nameIdentifier>
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<mods:namePart id="EB332AE5BCF703F0E7DE514034802FD5">Joyce, Walter G.</mods:namePart>
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<mods:affiliation id="F4DEF543AAF1825B4C13F9AA1B7F438E">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. &amp; Email: walter. joyce @ unifr. ch &amp; urn: lsid: zoobank. org: author: C 30 D 317 A- 0029 - 439 C- 8446 - 31281771 ADA 5</mods:affiliation>
<mods:nameIdentifier id="56514950BF529504869A2A4D796F3941" type="email">walter.joyce@unifr.ch</mods:nameIdentifier>
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<mods:title id="719E041F7F089523A11B77D2E91E7A19">European Journal of Taxonomy</mods:title>
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<treatment id="03C7CF0BC0751D36FF6CA84AFAD09122" ID-DOI="http://doi.org/10.5281/zenodo.3861093" ID-GBIF-Taxon="164362565" ID-Zenodo-Dep="3861093" LSID="urn:lsid:plazi:treatment:03C7CF0BC0751D36FF6CA84AFAD09122" httpUri="http://treatment.plazi.org/id/03C7CF0BC0751D36FF6CA84AFAD09122" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<subSubSection id="C3742D96C0751D35FF6CA84AFB79966A" box="[189,1246,321,348]" pageId="65" pageNumber="66" type="nomenclature">
<paragraph id="8BD17E1DC0751D35FF6CA84AFB79966A" blockId="65.[189,1246,321,348]" box="[189,1246,321,348]" pageId="65" pageNumber="66">
<heading id="D099C971C0751D35FF6CA84AFB79966A" bold="true" box="[189,1246,321,348]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="B91AA20FC0751D35FF6CA84AFB79966A" bold="true" box="[189,1246,321,348]" pageId="65" pageNumber="66">
Key for the identification of adult Indian
<taxonomicName id="4C6E059EC0751D35FD11A849FCE0966D" authority="Theobald" authorityName="Theobald" authorityYear="1868" box="[704,839,321,347]" class="Reptilia" family="Geoemydidae" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="family">geoemydid</taxonomicName>
turtles with regard of their shell
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0751D36FF6CA879FAD09122" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66" type="key">
<paragraph id="8BD17E1DC0751D35FF6CA879FDFA9509" blockId="65.[189,1399,369,575]" pageId="65" pageNumber="66">
We created this key to simplify the identification of geoemydid shell specimens that occur since Miocene and as of today at the Indian subcontinent, belonging to the following species:
<taxonomicName id="4C6E059EC0751D35FB4EA89EFAC89699" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[1183,1391,405,431]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="kachuga">
<emphasis id="B91AA20FC0751D35FB4EA89EFAC89699" box="[1183,1391,405,431]" italics="true" pageId="65" pageNumber="66">Batagur kachuga</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FF6CA8B2FE3D96E5" baseAuthorityName="Murray" baseAuthorityYear="1884" box="[189,410,441,467]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="dhongoka">
<emphasis id="B91AA20FC0751D35FF6CA8B2FE3D96E5" box="[189,410,441,467]" italics="true" pageId="65" pageNumber="66">Batagur dhongoka</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FE7AA8B2FDFF96E5" box="[427,600,441,467]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="baska">
<emphasis id="B91AA20FC0751D35FE7AA8B2FDFF96E5" box="[427,600,441,467]" italics="true" pageId="65" pageNumber="66">Batagur baska</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FDB8A8B1FCF796E5" baseAuthorityName="Riche in Daudin" baseAuthorityYear="1801" box="[617,848,441,467]" class="Reptilia" family="Geoemydidae" genus="Cuora" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="amboinensis">
<emphasis id="B91AA20FC0751D35FDB8A8B1FCF796E5" box="[617,848,441,467]" italics="true" pageId="65" pageNumber="66">Cuora amboinensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FCB0A8B1FBBA96E5" baseAuthorityName="Gray" baseAuthorityYear="1862" box="[865,1053,441,467]" class="Reptilia" family="Geoemydidae" genus="Cuora" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="mouhotti">
<emphasis id="B91AA20FC0751D35FCB0A8B1FBBA96E5" box="[865,1053,441,467]" italics="true" pageId="65" pageNumber="66">Cuora mouhotti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FBFFA8B1FB5296E5" box="[1070,1269,441,467]" class="Reptilia" family="Geoemydidae" genus="Cyclemys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="gemeli">
<emphasis id="B91AA20FC0751D35FBFFA8B1FB5296E5" box="[1070,1269,441,467]" italics="true" pageId="65" pageNumber="66">Cyclemys gemeli</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FAD7A8B1FF5C96C1" class="Reptilia" family="Geoemydidae" genus="Cyclemys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B91AA20FC0751D35FAD7A8B1FF5C96C1" italics="true" pageId="65" pageNumber="66">Cyclemys fusca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FED9A8D5FDA096C1" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[264,519,477,503]" class="Reptilia" family="Geoemydidae" genus="Geoclemys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="hamiltonii">
<emphasis id="B91AA20FC0751D35FED9A8D5FDA096C1" box="[264,519,477,503]" italics="true" pageId="65" pageNumber="66">Geoclemys hamiltonii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FDC5A8D5FD6C96C1" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[532,715,477,503]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0751D35FDC5A8D5FD6C96C1" box="[532,715,477,503]" italics="true" pageId="65" pageNumber="66">Hardella thurjii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FD08A8D5FC7696C0" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[729,977,477,503]" class="Reptilia" family="Geoemydidae" genus="Melanochelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="trijuga">
<emphasis id="B91AA20FC0751D35FD08A8D5FC7696C0" box="[729,977,477,503]" italics="true" pageId="65" pageNumber="66">Melanochelys trijuga</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FC0FA8D5FAA296C0" baseAuthorityName="Blyth" baseAuthorityYear="1856" box="[990,1285,477,503]" class="Reptilia" family="Geoemydidae" genus="Melanochelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="tricarinata">
<emphasis id="B91AA20FC0751D35FC0FA8D5FAA296C0" box="[990,1285,477,503]" italics="true" pageId="65" pageNumber="66">Melanochelys tricarinata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FAC3A8D5FEAA952C" authorityName="Anderson" authorityYear="1879" class="Reptilia" family="Geoemydidae" genus="Morenia" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91AA20FC0751D35FAC3A8D5FEAA952C" italics="true" pageId="65" pageNumber="66">Morenia petersi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FECEAB09FDA5952C" authorityName="Joyce &amp; Lyson" authorityYear="2010" box="[287,514,513,539]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="tatrotia">
<emphasis id="B91AA20FC0751D35FECEAB09FDA5952C" box="[287,514,513,539]" italics="true" pageId="65" pageNumber="66">Pangshura tatrotia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FDC5AB09FCB8952D" authorityName="Jerdon" authorityYear="1870" box="[532,799,513,539]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="sylhetensis">
<emphasis id="B91AA20FC0751D35FDC5AB09FCB8952D" box="[532,799,513,539]" italics="true" pageId="65" pageNumber="66">Pangshura sylhetensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FCE0AB09FC51952C" baseAuthorityName="Joyce &amp; Lyson" baseAuthorityYear="2010" box="[817,1014,513,539]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="tecta">
<emphasis id="B91AA20FC0751D35FCE0AB09FC51952C" box="[817,1014,513,539]" italics="true" pageId="65" pageNumber="66">Pangshura tecta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FBD9AB09FB43952D" baseAuthorityName="Gray" baseAuthorityYear="1863" box="[1032,1252,513,539]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="smithii">
<emphasis id="B91AA20FC0751D35FBD9AB09FB43952D" box="[1032,1252,513,539]" italics="true" pageId="65" pageNumber="66">Pangshura smithii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E059EC0751D35FB27AB09FEBD9508" baseAuthorityName="Gray" baseAuthorityYear="1834" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="tentoria">
<emphasis id="B91AA20FC0751D35FB27AB09FEBD9508" italics="true" pageId="65" pageNumber="66">Pangshura tentoria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E059EC0751D35FE82AB2CFDF19508" baseAuthorityName="Henderson" baseAuthorityYear="1912" box="[339,598,548,574]" class="Reptilia" family="Geoemydidae" genus="Vijayachelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="sylvatica">
<emphasis id="B91AA20FC0751D35FE82AB2CFDF19508" box="[339,598,548,574]" italics="true" pageId="65" pageNumber="66">Vijayachelys sylvatica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD17E1DC0751D35FF6CAB6FFEAB95DB" blockId="65.[189,1399,615,749]" pageId="65" pageNumber="66">This key was created based on the scoring of characters from our matrix and should be used together with descriptions and illustrations from the literature (Boulenger 1889; Ernst &amp; Barbour 1989; Das 1991, 1994) for correct identification. This key can only be used for adult shells deprived of horny scutes.</paragraph>
<key id="FFFF292CC0751D36FF6CAA1DFAD09122" lastPageId="66" lastPageNumber="67" pageId="65" pageNumber="66">
<keyStep id="309A66B8C0751D35FF6CAA1DFAD09441" pageId="65" pageNumber="66">
<paragraph id="8BD17E1DC0751D35FF6CAA1DFAD09419" blockId="65.[189,1399,788,887]" box="[189,1399,788,815]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAA1DFAD09419" box="[189,1399,788,815]" pageId="65" pageNumber="66">1. Carapace with three longitudinal keels.............................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF10AA31FAD09441" blockId="65.[189,1399,788,887]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AA31FAD09441" pageId="65" pageNumber="66">
Carapace without longitudinal keels or with a median keel only; carapace median length more than
<quantity id="4C96D3F8C0751D35FF3BAA54FEEB9441" box="[234,332,860,887]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="mm" value="150.0">150 mm</quantity>
.............................................................................................................................................. 7
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BD17E1DC0751D35FF6CAA97FAD094EB" blockId="65.[189,1399,926,989]" pageId="65" pageNumber="66">
<keyStep id="309A66B8C0751D35FF6CAA97FAD094EB" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAA97FAD094EB" pageId="65" pageNumber="66">
2. Neural bones III-VI hexagonal, with anterior short sides...........
<taxonomicName id="4C6E059EC0751D35FC03AA96FAD1948F" authority="(Gray, 1830)" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[978,1398,926,953]" class="Reptilia" family="Geoemydidae" genus="Geoclemys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="hamiltonii">
<emphasis id="B91AA20FC0751D35FC03AA96FB7D948F" bold="true" box="[978,1242,926,953]" italics="true" pageId="65" pageNumber="66">Geoclemys hamiltonii</emphasis>
(Gray, 1830)
</taxonomicName>
Neural bones IIIVI hexagonal, with posterior short sides ............................................................... 3
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF6CAD0DFAD09351" blockId="65.[189,1399,1029,1127]" pageId="65" pageNumber="66">
<keyStep id="309A66B8C0751D35FF6CAD0DFAD09351" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAD0DFAD09351" pageId="65" pageNumber="66">3. Presence of a plastral hinge ............................................................................................................... 6 Absence of a plastral hinge; pectoroabdominal sulcus and hyo-hypoplastron suture do not contact. ........................................................................................................................................................... 4</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF6CAD87FAD093C7" blockId="65.[189,1399,1167,1265]" pageId="65" pageNumber="66">
<keyStep id="309A66B8C0751D35FF6CAD87FAD093C7" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAD87FAD093C7" pageId="65" pageNumber="66">
4. Neural bone II hexagonal; pygal bone overlapped by fifth vertebral scute......................................... ...........................................................................................
<taxonomicName id="4C6E059EC0751D35FC5DADBAFAD193FB" authority="(Henderson, 1912)" baseAuthorityName="Henderson" baseAuthorityYear="1912" box="[908,1398,1202,1229]" class="Reptilia" family="Geoemydidae" genus="Vijayachelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="sylvatica">
<emphasis id="B91AA20FC0751D35FC5DADBAFB3493FB" bold="true" box="[908,1171,1202,1229]" italics="true" pageId="65" pageNumber="66">Vijayachelys sylvatica</emphasis>
(Henderson, 1912)
</taxonomicName>
Neural bone II octagonal; pygal bone not overlapped by fifth vertebral scute.................................. 5
</keyLead>
</keyStep>
</paragraph>
<keyStep id="309A66B8C0751D35FF6CAC11FAD092F5" pageId="65" pageNumber="66">
<paragraph id="8BD17E1DC0751D35FF6CAC11FAD1924D" blockId="65.[189,1399,1305,1475]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAC11FAD1924D" pageId="65" pageNumber="66">
5. Second to fourth vertebral scutes longer than wide or as long as wide; cervical scute very small; longitudinal gutter on the lateral of the carapace formed by the uplift of peripherals ........................ ..........................................................................................
<taxonomicName id="4C6E059EC0751D35FC59AC68FAD1924D" authority="(Schweigger, 1812)" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[904,1398,1376,1403]" class="Reptilia" family="Geoemydidae" genus="Melanochelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="trijuga">
<emphasis id="B91AA20FC0751D35FC59AC68FB2E924D" bold="true" box="[904,1161,1376,1403]" italics="true" pageId="65" pageNumber="66">Melanochelys trijuga</emphasis>
(Schweigger, 1812)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF10AC8DFAD092F5" blockId="65.[189,1399,1305,1475]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AC8DFAD092F5" pageId="65" pageNumber="66">
Second to fourth vertebral scutes broader than long; first vertebral scute anteriorly wider; cervical scute large, carapace with straight lateral sides ......................
<taxonomicName id="4C6E059EC0751D35FC64ACA0FAD092F5" authority="Blyth, 1856" authorityName="Blyth" authorityYear="1856" box="[949,1399,1448,1475]" class="Reptilia" family="Geoemydidae" genus="Melanochelys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="tricarinata">
<emphasis id="B91AA20FC0751D35FC64ACA0FB4292F5" bold="true" box="[949,1253,1448,1475]" italics="true" pageId="65" pageNumber="66">Melanochelys tricarinata</emphasis>
Blyth, 1856
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A66B8C0751D35FF6CACE3FAD19147" pageId="65" pageNumber="66">
<paragraph id="8BD17E1DC0751D35FF6CACE3FAD0911F" blockId="65.[189,1399,1515,1649]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CACE3FAD0911F" pageId="65" pageNumber="66">
6. Carapace ovoid rounded on top; median vertebral keel weakly developed; posterior margin of the carapace not serrated ...................................................
<taxonomicName id="4C6E059EC0751D35FCB7AF07FAD0911F" authority="(Riche in Daudin, 1801)" baseAuthorityName="Riche in Daudin" baseAuthorityYear="1801" box="[870,1399,1550,1577]" class="Reptilia" family="Geoemydidae" genus="Cuora" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="amboinensis">
<emphasis id="B91AA20FC0751D35FCB7AF07FBF5911F" bold="true" box="[870,1106,1550,1577]" italics="true" pageId="65" pageNumber="66">Cuora amboinensis</emphasis>
(Riche
<emphasis id="B91AA20FC0751D35FB7EAF18FB61911F" box="[1199,1222,1552,1577]" italics="true" pageId="65" pageNumber="66">in</emphasis>
Daudin, 1801)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF10AF3BFAD19147" blockId="65.[189,1399,1515,1649]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AF3BFAD19147" pageId="65" pageNumber="66">
Carapace flattened between second and fourth vertebral scutes; anterior margin of the carapace slightly serrated; posterior margin of the carapace slightly serrated ....
<taxonomicName id="4C6E059EC0751D35FBC9AF5EFAD19147" authority="(Gray, 1862)" baseAuthorityName="Gray" baseAuthorityYear="1862" box="[1048,1398,1622,1649]" class="Reptilia" family="Geoemydidae" genus="Cuora" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="mouhotii">
<emphasis id="B91AA20FC0751D35FBC9AF5EFB7E9147" bold="true" box="[1048,1241,1622,1649]" italics="true" pageId="65" pageNumber="66">Cuora mouhotii</emphasis>
(Gray, 1862)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A66B8C0751D35FF10AF91FAD0900B" pageId="65" pageNumber="66">
<paragraph id="8BD17E1DC0751D35FF10AF91FAD091E1" blockId="65.[193,1399,1689,1751]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AF91FAD091E1" pageId="65" pageNumber="66">
7. Entoplastron intersected by humeropectoral sulcus; plastral buttresses absent or very reduced................................................................................................................
<taxonomicName id="4C6E059EC0751D35FB52AFB4FAD091E1" authority="Bell, 1834" authorityName="Bell" authorityYear="1834" box="[1155,1399,1724,1751]" class="Reptilia" family="Geoemydidae" genus="Cyclemys" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC0751D35FB52AFB4FB5291E1" bold="true" box="[1155,1269,1724,1751]" italics="true" pageId="65" pageNumber="66">Cyclemys</emphasis>
Bell, 1834
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF10AFF7FAD0900B" blockId="65.[193,1399,1791,1853]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AFF7FAD0900B" pageId="65" pageNumber="66"> Entoplastron not intersected by humeropectoral sulcus; presence of well developed plastral buttresses............................................................................................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A66B8C0751D35FF6CAE6DFAD090DD" pageId="65" pageNumber="66">
<paragraph id="8BD17E1DC0751D35FF6CAE6DFAD09095" blockId="65.[189,1399,1893,2027]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF6CAE6DFAD09095" pageId="65" pageNumber="66">8. Presence of intercostal fontanelle in adults (sexually dimorphic); fourth to sixth marginal scutes can overlap adjacent costal bones ............................................................................................................ 9</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0751D35FF10AEA5FAD090DD" blockId="65.[189,1399,1893,2027]" pageId="65" pageNumber="66">
<keyLead id="309FDD28C0751D35FF10AEA5FAD090DD" pageId="65" pageNumber="66">
Absence of intercostal fontanelle in adults; fourth to sixth marginal scutes never overlap costal bones ................................................................................................
<taxonomicName id="4C6E059EC0751D35FC27AED9FAD090DD" authority="Anderson, 1879" authorityName="Anderson" authorityYear="1879" box="[1014,1399,2001,2027]" class="Reptilia" family="Geoemydidae" genus="Morenia" kingdom="Animalia" order="Testudines" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="petersi">
<emphasis id="B91AA20FC0751D35FC27AED9FB1290DD" bold="true" box="[1014,1205,2001,2027]" italics="true" pageId="65" pageNumber="66">Morenia petersi</emphasis>
Anderson, 1879
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BD17E1DC0761D36FF6CA803FAD1967F" blockId="66.[189,1398,267,329]" pageId="66" pageNumber="67">
<keyStep id="309A66B8C0761D36FF6CA803FAD1967F" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CA803FAD1967F" pageId="66" pageNumber="67">9. Long fourth vertebral scute, with its anterior margin over neural IV.............................................. 10 Short fourth vertebral scute, anterior margin crossing neural V...................................................... 12</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF6CA878FAD2969B" blockId="66.[189,1399,368,430]" pageId="66" pageNumber="67">
<keyStep id="309A66B8C0761D36FF6CA878FAD2969B" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CA878FAD2969B" pageId="66" pageNumber="67">
10. Fourth vertebral bottleneck anteriorly; neural IV octagonal ..........................
<taxonomicName id="4C6E059EC0761D36FBB6A878FAD096BD" authority="Gray, 1856" authorityName="Gray" authorityYear="1856" box="[1127,1399,368,395]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="genus">
<emphasis id="B91AA20FC0761D36FBB6A878FB4A96BD" bold="true" box="[1127,1261,368,395]" italics="true" pageId="66" pageNumber="67">Pangshura</emphasis>
Gray, 1856
</taxonomicName>
Fourth vertebral straight laterally; neural IV hexagonal with anterior short-sides...........................11
</keyLead>
</keyStep>
</paragraph>
<keyStep id="309A66B8C0761D36FF6CA8DDFAD0954A" pageId="66" pageNumber="67">
<paragraph id="8BD17E1DC0761D36FF6CA8DDFAD19503" blockId="66.[189,1399,469,636]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CA8DDFAD19503" pageId="66" pageNumber="67">
11. Second vertebral scute with a strong process on posterior margin with invagination into third vertebral; gulohumeral sulcus forming a right angle; humeropectoral sulcus straight........................ .........................................................................................................
<taxonomicName id="4C6E059EC0761D36FC27AB13FAD19503" authority="(Gray, 1832)" baseAuthorityName="Gray" baseAuthorityYear="1832" box="[1014,1398,539,566]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="dhongoka">
<emphasis id="B91AA20FC0761D36FC27AB13FB7D9500" bold="true" box="[1014,1242,539,566]" italics="true" pageId="66" pageNumber="67">Batagur dhongoka</emphasis>
(Gray, 1832)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF10AB37FAD0954A" blockId="66.[189,1399,469,636]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF10AB37FAD0954A" pageId="66" pageNumber="67">
Second vertebral scute large, as long as wide, with straight lateral margins; third vertebral scute with lateral margin with posterior shorter side ..........................................
<taxonomicName id="4C6E059EC0761D36FBD7AB6AFAD0954A" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[1030,1399,609,636]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="kachuga">
<emphasis id="B91AA20FC0761D36FBD7AB6AFB7E954A" bold="true" box="[1030,1241,609,636]" italics="true" pageId="66" pageNumber="67">Batagur kachuga</emphasis>
(Gray, 1831)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A66B8C0761D36FF6CABABFAD0947C" pageId="66" pageNumber="67">
<paragraph id="8BD17E1DC0761D36FF6CABABFAD195D6" blockId="66.[189,1399,675,842]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CABABFAD195D6" pageId="66" pageNumber="67">
12. Fourth vertebral scute shorter than second and third vertebrals; anterior plastral margin straight; anal notch absent, or if present, in a wide angle.............................................
<taxonomicName id="4C6E059EC0761D36FBF8ABCEFAD195D6" authority="(Gray, 1830)" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[1065,1398,710,737]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="baska">
<emphasis id="B91AA20FC0761D36FBF8ABCEFB7E95D7" bold="true" box="[1065,1241,710,737]" italics="true" pageId="66" pageNumber="67">Batagur baska</emphasis>
(Gray, 1830)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF10ABE2FAD0947C" blockId="66.[189,1399,675,842]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF10ABE2FAD0947C" pageId="66" pageNumber="67">
Carapace median keel present; fourth vertebral scute about the same length and width as second and third vertebrals; first vertebral wider posteriorly; strong gular inflection; anal notch rounded .................................................................................................
<taxonomicName id="4C6E059EC0761D36FBCBAA27FAD0947C" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[1050,1399,815,842]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0761D36FBCBAA27FB7D947C" bold="true" box="[1050,1242,815,842]" italics="true" pageId="66" pageNumber="67">Hardella thurjii</emphasis>
(Gray, 1831)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A66B8C0761D36FF6CAA79FAD194C3" pageId="66" pageNumber="67">
<paragraph id="8BD17E1DC0761D36FF6CAA79FAD09498" blockId="66.[189,1399,881,1013]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CAA79FAD09498" pageId="66" pageNumber="67">
13. Second vertebral scute much shorter than third vertebral, embracing two neural bones; posterior margin of carapace strongly serrated .........................................
<taxonomicName id="4C6E059EC0761D36FC19AA9CFAD09498" authority="Jerdon, 1870" authorityName="Jerdon" authorityYear="1870" box="[968,1399,916,943]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="sylhetensis">
<emphasis id="B91AA20FC0761D36FC19AA9CFB709499" bold="true" box="[968,1239,916,943]" italics="true" pageId="66" pageNumber="67">Pangshura sylhetensis</emphasis>
Jerdon, 1870
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF10AAB0FAD194C3" blockId="66.[189,1399,881,1013]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF10AAB0FAD194C3" pageId="66" pageNumber="67"> Second vertebral scute as long as broad, embracing three neural bones; posterior margin of fourth vertebral over neural VIII or posterior............................................................................................. 14</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BD17E1DC0761D36FF6CAD14FAD1934B" blockId="66.[189,1398,1052,1149]" pageId="66" pageNumber="67">
<keyStep id="309A66B8C0761D36FF6CAD14FAD1934B" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CAD14FAD1934B" pageId="66" pageNumber="67">
14. Third vertebral scute much longer than wide, with almost parallel lateral margins............................ .........................................................................................................
<taxonomicName id="4C6E059EC0761D36FC28AD37FAD1936C" authority="(Gray, 1863)" baseAuthorityName="Gray" baseAuthorityYear="1863" box="[1017,1398,1087,1114]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="smithii">
<emphasis id="B91AA20FC0761D36FC28AD37FB7E936C" bold="true" box="[1017,1241,1087,1114]" italics="true" pageId="66" pageNumber="67">Pangshura smithii</emphasis>
(Gray, 1863)
</taxonomicName>
Third vertebral scute pentagonal, pointed behind............................................................................ 15
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF6CADACFAD193D7" blockId="66.[189,1398,1188,1250]" pageId="66" pageNumber="67">
<keyStep id="309A66B8C0761D36FF6CADACFAD193D7" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CADACFAD193D7" pageId="66" pageNumber="67">
15. Second vertebral scute hexagonal to square/rectangular ............
<taxonomicName id="4C6E059EC0761D36FC00ADACFAD19388" authority="(Murray, 1884)" baseAuthorityName="Murray" baseAuthorityYear="1884" box="[977,1398,1188,1215]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="tentoria">
<emphasis id="B91AA20FC0761D36FC00ADACFB1A9388" bold="true" box="[977,1213,1188,1215]" italics="true" pageId="66" pageNumber="67">Pangshura tentoria</emphasis>
(Murray, 1884)
</taxonomicName>
Second vertebral scute hexagonal, greatly constricted posteriorly.................................................. 16
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF6CAC00FAD0925F" blockId="66.[189,1399,1288,1386]" pageId="66" pageNumber="67">
<keyStep id="309A66B8C0761D36FF6CAC00FAD0925F" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CAC00FAD0925F" pageId="66" pageNumber="67">
16. First vertebral scute significantly constricted posteriorly....................
<taxonomicName id="4C6E059EC0761D36FBC2AC00FAD19215" authority="(Gray, 1830)" baseAuthorityName="Gray" baseAuthorityYear="1830" box="[1043,1398,1288,1315]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="tecta">
<emphasis id="B91AA20FC0761D36FBC2AC00FB7E9215" bold="true" box="[1043,1241,1288,1315]" italics="true" pageId="66" pageNumber="67">Pangshura tecta</emphasis>
(Gray, 1830)
</taxonomicName>
First vertebral scute rectangular or anteriorly constricted ................................................................... ...........................................................................................
<taxonomicName id="4C6E059EC0761D36FC40AC47FAD0925F" authority="Joyce &amp; Lyson, 2010" authorityName="Joyce &amp; Lyson" authorityYear="2010" box="[913,1399,1359,1386]" class="Reptilia" family="Geoemydidae" genus="Pangshura" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="tatrotia">
<emphasis id="B91AA20FC0761D36FC40AC47FBD1925F" bold="true" box="[913,1142,1359,1386]" italics="true" pageId="66" pageNumber="67">Pangshura tatrotia</emphasis>
Joyce &amp; Lyson, 2010
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<keyStep id="309A66B8C0761D36FF6CAC99FAD09122" pageId="66" pageNumber="67">
<paragraph id="8BD17E1DC0761D36FF6CAC99FAD092F8" blockId="66.[189,1399,1424,1557]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CAC99FAD092F8" pageId="66" pageNumber="67">
17. Carapace elongated to rectangular; crown of the head brown, but not lighter than temporal region.. ..................................
<taxonomicName id="4C6E059EC0761D36FE3AACBBFAD092F8" authority="Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer, 2008" authorityName="Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer" authorityYear="2008" box="[491,1399,1459,1486]" class="Reptilia" family="Geoemydidae" genus="Cyclemys" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="gemeli">
<emphasis id="B91AA20FC0761D36FE3AACBBFD1292F8" bold="true" box="[491,693,1459,1486]" italics="true" pageId="66" pageNumber="67">Cyclemys gemeli</emphasis>
Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer, 2008
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD17E1DC0761D36FF6CACDFFAD09122" blockId="66.[189,1399,1424,1557]" pageId="66" pageNumber="67">
<keyLead id="309FDD28C0761D36FF6CACDFFAD09122" pageId="66" pageNumber="67">
Carapace ovoid when viewed from above; crown of the head lighter than temporal region .............. ....................................
<taxonomicName id="4C6E059EC0761D36FE2BACF2FAD09122" authority="Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer, 2008" authorityName="Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer" authorityYear="2008" box="[506,1399,1530,1557]" class="Reptilia" family="Geoemydidae" genus="Cyclemys" kingdom="Animalia" order="Testudines" pageId="66" pageNumber="67" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B91AA20FC0761D36FE2BACF2FD139122" bold="true" box="[506,692,1530,1557]" italics="true" pageId="66" pageNumber="67">Cyclemys fusca</emphasis>
Fritz, Guicking, Auer, Sommer, Wink &amp; Hundsdorfer, 2008
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>