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<document id="BB8D5BD4DB2F0B9642D9F06EA896D354" ID-CLB-Dataset="58036" ID-DOI="10.5281/zenodo.7910107" ID-GBIF-Dataset="c3b725f7-5bd4-440f-8808-bc172f9067e8" ID-ISSN="2305-2562" ID-Zenodo-Dep="7910107" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683577956358" checkinUser="felipe" docAuthor="Krzemiński, Wieslaw" docDate="2007" docId="03C7C359FFABD211FEA1FA40561E01AB" docLanguage="en" docName="AfrInvertebr.48.1.153–162.pdf" docOrigin="African Invertebrates 48 (1)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Hylobittacus fossilis Krzemiński 2007, comb. n." docType="treatment" docVersion="7" lastPageNumber="158" masterDocId="FFFEBB21FFAFD214FFFCFF805578072D" masterDocTitle="A revision of Eocene Bittacidae (Mecoptera) from Baltic amber, with the description of a new species" masterLastPageNumber="162" masterPageNumber="153" pageNumber="157" updateTime="1699515406999" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BDA4F98EA6C9A67D28E78C88F5CB574E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4394B930529F597BAAA91E4C96B842D0">
<mods:title id="9D446E141646EE3F3C7782B08A5EE051">A revision of Eocene Bittacidae (Mecoptera) from Baltic amber, with the description of a new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="B5233DB19B61DC5998E16CB0F1029F5F">Krzemiński, Wieslaw</mods:namePart>
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<mods:title id="CE1F2AAE3CC815A0AB51D9CD2AA1FB08">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="1E6A1E8CD84AF6B8D266460D2E4922AD">
<mods:date id="72724178BB7F66A0ECD7045605B0C305">2007</mods:date>
<mods:detail id="0E6CAF9258BA52B01843C51568808119" type="pubDate">
<mods:number id="5B242D91E0E850D06D6F0247F47B7FA1">2007-04-30</mods:number>
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<mods:number id="87CC2D412AFB55F1D7EDF7778E806F2B">48</mods:number>
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<mods:identifier id="E4C613CE03611862A83F985A3B1DD32C" type="DOI">10.5281/zenodo.7910107</mods:identifier>
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<treatment id="03C7C359FFABD211FEA1FA40561E01AB" ID-DOI="http://doi.org/10.5281/zenodo.7910806" ID-GBIF-Taxon="209589601" ID-Zenodo-Dep="7910806" LSID="urn:lsid:plazi:treatment:03C7C359FFABD211FEA1FA40561E01AB" httpUri="http://treatment.plazi.org/id/03C7C359FFABD211FEA1FA40561E01AB" lastPageId="5" lastPageNumber="158" pageId="4" pageNumber="157">
<subSubSection id="C37421C4FFABD210FEA1FA4056FA02F5" box="[349,898,1471,1496]" pageId="4" pageNumber="157" type="nomenclature">
<paragraph id="8BD1724FFFABD210FEA1FA4056FA02F5" blockId="4.[144,1104,1471,1672]" box="[349,898,1471,1496]" pageId="4" pageNumber="157">
<heading id="D099C523FFABD210FEA1FA4056FA02F5" box="[349,898,1471,1496]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="157" reason="3">
<taxonomicName id="4C6E09CCFFABD210FEA1FA40566802F5" authority="(Carpenter, 1954)" authorityName="Krzemiński" authorityYear="2007" baseAuthorityName="Carpenter" baseAuthorityYear="1954" box="[349,784,1471,1496]" class="Insecta" family="Bittacidae" genus="Hylobittacus" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="fossilis" status="comb. nov.">
<emphasis id="B91AAE5DFFABD210FEA1FA40573902F5" box="[349,577,1472,1496]" italics="true" pageId="4" pageNumber="157">Hylobittacus fossilis</emphasis>
(Carpenter, 1954)
</taxonomicName>
,
<emphasis id="B91AAE5DFFABD210FCE0FA3F56FA02F5" bold="true" box="[796,898,1471,1496]" pageId="4" pageNumber="157">
<taxonomicNameLabel id="A2291326FFABD210FCE0FA3F56FA02F5" box="[796,898,1471,1496]" pageId="4" pageNumber="157" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37421C4FFABD210FDCAFA6757D2012D" box="[566,682,1511,1536]" pageId="4" pageNumber="157" type="description">
<paragraph id="8BD1724FFFABD210FDCAFA6757D2012D" blockId="4.[144,1104,1471,1672]" box="[566,682,1511,1536]" pageId="4" pageNumber="157">
<figureCitation id="13556ECAFFABD210FDCAFA675705012D" box="[566,637,1511,1536]" captionStart="Fig" captionStartId="5.[182,215,911,931]" captionTargetBox="[192,1133,203,890]" captionTargetId="figure-265@5.[192,1133,203,900]" captionTargetPageId="5" captionText="Fig. 4. Hylobittacus fossilis (Carpenter): (A, B) male DEIM 1150/1, general appearance of wing and details of venation; (C, D) female MP/1/1/551/05/A, general appearance of wing (reversed left-right) and details of venation; (E, F) male genitalia, DEIM 1150/1 general appearance and details. Abbreviation: e – epiandrium. Scale bars: Figs 4A–D = 5 mm, Figs 4E, 4F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7910589" httpUri="https://zenodo.org/record/7910589/files/figure.png" pageId="4" pageNumber="157">Figs 4</figureCitation>
,
<figureCitation id="13556ECAFFABD210FD75FA6757D2012D" box="[649,682,1511,1536]" captionStart="Fig" captionStartId="8.[143,176,661,681]" captionTargetBox="[202,1030,203,649]" captionTargetId="figure-323@8.[200,1036,191,655]" captionTargetPageId="8" captionText="Fig. 7. Epiandrial lobes of Baltic amber Bittacidae in lateral aspect: (A) Bittacus succinus Carpenter; (B) Hylobittacus fossilis (Carpenter); (C) Hylobittacus minimus (Carpenter); (D) Hylobittacus picteti sp. n. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7910597" httpUri="https://zenodo.org/record/7910597/files/figure.png" pageId="4" pageNumber="157">7B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37421C4FFABD210FF6CF98E570F010F" box="[144,631,1550,1570]" pageId="4" pageNumber="157" type="reference_group">
<paragraph id="8BD1724FFFABD210FF6CF98E570F010F" blockId="4.[144,1104,1471,1672]" box="[144,631,1550,1570]" pageId="4" pageNumber="157">
<taxonomicName id="4C6E09CCFFABD210FF6CF98E54A4010F" authority=": Carpenter 1954: 37" authorityName=": Carpenter" authorityPageNumber="37" authorityYear="1954" box="[144,476,1550,1570]" class="Insecta" family="Bittacidae" genus="Bittacus" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B91AAE5DFFABD210FF6CF98E5467010F" box="[144,287,1550,1570]" italics="true" pageId="4" pageNumber="157">Bittacus fossilis</emphasis>
: Carpenter 1954: 37
</taxonomicName>
, 38, figs 3A, 4B.
</paragraph>
</subSubSection>
<subSubSection id="C37421C4FFABD211FF6CF9AF561E01AB" lastPageId="5" lastPageNumber="158" pageId="4" pageNumber="157" type="diagnosis">
<paragraph id="8BD1724FFFABD210FF6CF9AF558801A5" blockId="4.[144,1104,1471,1672]" pageId="4" pageNumber="157">
Diagnosis: Separated from all species of
<taxonomicName id="4C6E09CCFFABD210FDB3F9B057A60165" authorityName="Byers" authorityYear="1979" box="[591,734,1584,1608]" class="Insecta" family="Bittacidae" genus="Hylobittacus" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="157" phylum="Arthropoda" rank="genus">
<emphasis id="B91AAE5DFFABD210FDB3F9B057A60165" box="[591,734,1584,1608]" italics="true" pageId="4" pageNumber="157">Hylobittacus</emphasis>
</taxonomicName>
in Baltic amber by a combination of: Sc short, ending well before fork of Rs; epiandrial lobe with deeply incised margin dorsally.
</paragraph>
<caption id="DF1122C7FFAAD211FF4AFC0F517504C6" ID-DOI="http://doi.org/10.5281/zenodo.7910589" ID-Zenodo-Dep="7910589" httpUri="https://zenodo.org/record/7910589/files/figure.png" pageId="5" pageNumber="158" startId="5.[182,215,911,931]" targetBox="[192,1133,203,890]" targetPageId="5" targetType="figure">
<paragraph id="8BD1724FFFAAD211FF4AFC0F517504C6" blockId="5.[182,1143,911,1003]" pageId="5" pageNumber="158">
Fig. 4.
<taxonomicName id="4C6E09CCFFAAD211FF08FC0F5764048E" authority="(Carpenter)" authorityName="Krzemiński" authorityYear="2007" baseAuthorityName="Carpenter" baseAuthorityYear="1954" box="[244,540,911,931]" class="Insecta" family="Bittacidae" genus="Hylobittacus" kingdom="Animalia" order="Mecoptera" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B91AAE5DFFAAD211FF08FC0F54D4048E" box="[244,428,911,931]" italics="true" pageId="5" pageNumber="158">Hylobittacus fossilis</emphasis>
(Carpenter)
</taxonomicName>
: (A, B) male DEIM 1150/1, general appearance of wing and details of venation; (C, D) female MP/1/1/551/05/A, general appearance of wing (reversed left-right) and details of venation; (E, F) male genitalia, DEIM 1150/1 general appearance and details. Abbreviation: e – epiandrium. Scale bars: Figs 4A–D = 5 mm, Figs 4E, 4F = 1 mm.
</paragraph>
</caption>
<paragraph id="8BD1724FFFAAD211FF49FB8F54220305" blockId="5.[181,346,1039,1064]" box="[181,346,1039,1064]" pageId="5" pageNumber="158">Redescription:</paragraph>
<paragraph id="8BD1724FFFAAD211FF49FBBA558E037E" blockId="5.[181,246,1082,1107]" box="[181,246,1082,1107]" pageId="5" pageNumber="158">
<emphasis id="B91AAE5DFFAAD211FF49FBBA5597037F" box="[181,239,1082,1106]" italics="true" pageId="5" pageNumber="158">Male</emphasis>
.
</paragraph>
<paragraph id="8BD1724FFFAAD211FF49FBE551180393" blockId="5.[181,1142,1125,1310]" pageId="5" pageNumber="158">
Forewing (
<figureCitation id="13556ECAFFAAD211FED0FBE554D00353" box="[300,424,1125,1150]" captionStart="Fig" captionStartId="5.[182,215,911,931]" captionTargetBox="[192,1133,203,890]" captionTargetId="figure-265@5.[192,1133,203,900]" captionTargetPageId="5" captionText="Fig. 4. Hylobittacus fossilis (Carpenter): (A, B) male DEIM 1150/1, general appearance of wing and details of venation; (C, D) female MP/1/1/551/05/A, general appearance of wing (reversed left-right) and details of venation; (E, F) male genitalia, DEIM 1150/1 general appearance and details. Abbreviation: e – epiandrium. Scale bars: Figs 4A–D = 5 mm, Figs 4E, 4F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7910589" httpUri="https://zenodo.org/record/7910589/files/figure.png" pageId="5" pageNumber="158">Figs 4A–D</figureCitation>
)
<quantity id="4C96DFAAFFAAD211FE49FBE557520353" box="[437,554,1125,1150]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="5" pageNumber="158" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13–16 mm</quantity>
long. Single pterostigmal cross-vein; Sc short, ending well before fork of Rs; sc–r at two to four times its length before end of Sc; cubital cross-vein below fork of Mb; no additional cross-veins in radial, medial distal cells.
</paragraph>
<paragraph id="8BD1724FFFAAD211FF49FB45558C0233" blockId="5.[181,1142,1125,1310]" pageId="5" pageNumber="158">
Genitalia (
<figureCitation id="13556ECAFFAAD211FEDAFB4554DC03F3" box="[294,420,1221,1246]" captionStart="Fig" captionStartId="5.[182,215,911,931]" captionTargetBox="[192,1133,203,890]" captionTargetId="figure-265@5.[192,1133,203,900]" captionTargetPageId="5" captionText="Fig. 4. Hylobittacus fossilis (Carpenter): (A, B) male DEIM 1150/1, general appearance of wing and details of venation; (C, D) female MP/1/1/551/05/A, general appearance of wing (reversed left-right) and details of venation; (E, F) male genitalia, DEIM 1150/1 general appearance and details. Abbreviation: e – epiandrium. Scale bars: Figs 4A–D = 5 mm, Figs 4E, 4F = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7910589" httpUri="https://zenodo.org/record/7910589/files/figure.png" pageId="5" pageNumber="158">Figs 4E, 4F</figureCitation>
,
<figureCitation id="13556ECAFFAAD211FE50FB4554A903F3" box="[428,465,1221,1246]" captionStart="Fig" captionStartId="8.[143,176,661,681]" captionTargetBox="[202,1030,203,649]" captionTargetId="figure-323@8.[200,1036,191,655]" captionTargetPageId="8" captionText="Fig. 7. Epiandrial lobes of Baltic amber Bittacidae in lateral aspect: (A) Bittacus succinus Carpenter; (B) Hylobittacus fossilis (Carpenter); (C) Hylobittacus minimus (Carpenter); (D) Hylobittacus picteti sp. n. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7910597" httpUri="https://zenodo.org/record/7910597/files/figure.png" pageId="5" pageNumber="158">7B</figureCitation>
): epiandrial lobe with deeply incised margins dorsally, forming posterior lobe; upper, inner margin of epiandrial lobe with numerous spines; basistyle large.
</paragraph>
<paragraph id="8BD1724FFFAAD211FF49FAAE54FC025F" blockId="5.[181,1142,1326,1596]" pageId="5" pageNumber="158">
<materialsCitation id="3B067812FFAAD211FF49FAAE54FC025F" collectionCode="MCZ" pageId="5" pageNumber="158" specimenCode="MCZ 5209" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D5CCEDFFAAD211FF49FAAE5471026F" box="[181,265,1326,1346]" pageId="5" pageNumber="158" type="holotype">Holotype</typeStatus>
(examined):
<specimenCode id="DBC8DA34FFAAD211FE7DFAAE549C026F" box="[385,484,1326,1346]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="158" type="Museum">MCZ 5209</specimenCode>
(old no. 5117), male, mostly clearly preserved on one side, milky coating on portions of the other, some legs, half of right forewing and apical portion of right hind wing missing as a result of preparation.
</materialsCitation>
</paragraph>
<paragraph id="8BD1724FFFAAD211FF49FA0057D60111" blockId="5.[181,1142,1326,1596]" pageId="5" pageNumber="158">
Other material examined:
<materialsCitation id="3B067812FFAAD211FE5FFA00574C0281" collectionCode="MCZ" pageId="5" pageNumber="158" specimenCode="MCZ 4898" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FE5FFA00577202B9" box="[419,522,1408,1428]" collectionCode="MCZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="5" pageNumber="158" type="Museum">MCZ 4898</specimenCode>
, specimen complete except for left wings missing, clearly preserved except for some milky covering in places
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FDC2FA18544602E9" collectionCode="DEIM" collectorName="Hoffeins" pageId="5" pageNumber="158" specimenCode="DEIM 1150" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FDC2FA1857C80281" box="[574,688,1432,1452]" collectionCode="DEIM" pageId="5" pageNumber="158">DEIM 1150</specimenCode>
/1, a complete male specimen in perfect condition (
<collectorName id="269B1799FFAAD211FF47FA30547202E9" box="[187,266,1456,1476]" pageId="5" pageNumber="158">Hoffeins</collectorName>
coll.)
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FEBAFA30510A02E9" box="[326,1138,1456,1476]" collectionCode="MP" pageId="5" pageNumber="158" specimenCode="MP 1" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FEBAFA30540802E9" box="[326,368,1456,1476]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="158">MP 1</specimenCode>
/1/551/05/A, female, an entire specimen in good condition, only head partially obscured
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FF49FA4854590121" collectionCode="MP" pageId="5" pageNumber="158" specimenCode="MP 1" specimenCount="1" specimenCount-male="1">
<specimenCode id="DBC8DA34FFAAD211FF49FA48559A02F1" box="[181,226,1480,1500]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="158">MP 1</specimenCode>
/2/552/04,
<specimenCount id="9D68B9C6FFAAD211FEBBFA4854D602F1" box="[327,430,1480,1500]" pageId="5" pageNumber="158" type="male">three males</specimenCount>
in one piece of amber: two entire specimens in perfect condition; third one only partially preserved, without head, and only a portion of thorax and one wing present, abdomen obscured by milky layer
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FED6FA7851330121" box="[298,1099,1528,1548]" collectionCode="MP" pageId="5" pageNumber="158" specimenCode="MP 1" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FED6FA7854200121" box="[298,344,1528,1548]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="158">MP 1</specimenCode>
/1/552/04, female with only two wings, fragments of legs and abdomen preserved
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FBA9FA7856320109" collectionCode="MP" pageId="5" pageNumber="158" specimenCode="MP 1" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FBA9FA7855C40109" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="5" pageNumber="158">MP 1</specimenCode>
/5/552/04, sex unknown, only fragments of wings and one leg preserved
</materialsCitation>
;
<materialsCitation id="3B067812FFAAD211FCAEF99057D20111" collectionCode="MZW" pageId="5" pageNumber="158" specimenCode="MZW 19996" specimenCount="1">
<specimenCode id="DBC8DA34FFAAD211FCAEF99056B10109" box="[850,969,1552,1572]" collectionCode="MZW" pageId="5" pageNumber="158">MZW 19996</specimenCode>
, sex unknown, two wings and small fragment of thorax and legs preserved
</materialsCitation>
.
</paragraph>
<paragraph id="8BD1724FFFAAD211FF49F9CD561E01AB" blockId="5.[181,1142,1613,1670]" pageId="5" pageNumber="158">
Remarks:
<taxonomicName id="4C6E09CCFFAAD211FED7F9CD54E70148" authorityName="Krzemiński" authorityYear="2007" baseAuthorityName="Carpenter" baseAuthorityYear="1954" box="[299,415,1613,1637]" class="Insecta" family="Bittacidae" genus="Hylobittacus" kingdom="Animalia" order="Mecoptera" pageId="5" pageNumber="158" phylum="Arthropoda" rank="species" species="fossilis">
<emphasis id="B91AAE5DFFAAD211FED7F9CD54E70148" box="[299,415,1613,1637]" italics="true" pageId="5" pageNumber="158">H. fossilis</emphasis>
</taxonomicName>
seems to be the most frequent bittaciid found in Baltic amber (pers. observ.). Its venation shows remarkably little variation.
</paragraph>
</subSubSection>
</treatment>
</document>
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