272 lines
36 KiB
XML
272 lines
36 KiB
XML
<document id="5C4683E7B4BD1509DEA686B2470DCF48" ID-CLB-Dataset="53418" ID-DOI="10.11646/zootaxa.5238.1.1" ID-GBIF-Dataset="751096f2-4b5b-43c3-9748-4d07afe044c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7621793" ID-ZooBank="768E9932-2D18-4115-8359-3FF800328BCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1675871988450" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2023" docId="03C79010FFD5D73BFF707C2D20A4F89A" docLanguage="en" docName="zootaxa.5238.1.1.pdf" docOrigin="Zootaxa 5238 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5238.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chloeia boucheti Salazar-Vallejo 2023, sp. n." docType="treatment" docVersion="7" lastPageNumber="37" masterDocId="FFFEE868FFF6D71FFFE77B0B2306FFEB" masterDocTitle="Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="36" updateTime="1699474919205" updateUser="ExternalLinkService">
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<mods:title id="2CD30CC9309248BF194AEE31681433B4">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<mods:name id="35B812E235F1D67A145A0E196702C855" type="personal">
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<mods:namePart id="B65B9D49E1D91DFAC2332C752CFBFAE2">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="B6FA07F8D8CAE90BA1BBA93FCF782086">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F54E8210B9645280E61C084599306171">
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<mods:date id="5C77A9823AC07499F4C3EB763C3DD2EF">2023</mods:date>
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<mods:number id="864DB0DDA71AFEEEC9516E3566EF72C2">2023-02-07</mods:number>
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<mods:number id="EF4BD1196A1E7BE1BA7BAAD8B588DC38">5238</mods:number>
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<mods:url id="1A492581EC2E31CC51680328A098719B">http://dx.doi.org/10.11646/zootaxa.5238.1.1</mods:url>
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<mods:classification id="9A57FFCF8CE0A71C16E65AB2F4D42015">journal article</mods:classification>
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<mods:identifier id="270F2F39E7952916459002EA285CBCB7" type="CLB-Dataset">53418</mods:identifier>
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<mods:identifier id="0441C5136C05FC8B55E030DB68981BF6" type="DOI">10.11646/zootaxa.5238.1.1</mods:identifier>
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<treatment id="03C79010FFD5D73BFF707C2D20A4F89A" ID-DOI="http://doi.org/10.5281/zenodo.7621861" ID-GBIF-Taxon="209295599" ID-Zenodo-Dep="7621861" LSID="urn:lsid:plazi:treatment:03C79010FFD5D73BFF707C2D20A4F89A" httpUri="http://treatment.plazi.org/id/03C79010FFD5D73BFF707C2D20A4F89A" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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<subSubSection id="C374728DFFD5D73CFF707C2D2074F8B4" pageId="35" pageNumber="36" type="nomenclature">
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<paragraph id="8BD12106FFD5D73CFF707C2D22A9F8AA" blockId="35.[151,882,1830,1924]" box="[151,431,1830,1857]" pageId="35" pageNumber="36">
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<heading id="D099966AFFD5D73CFF707C2D22A9F8AA" bold="true" box="[151,431,1830,1857]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
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<emphasis id="B91AFD14FFD5D73CFF707C2D22A9F8AA" bold="true" box="[151,431,1830,1857]" pageId="35" pageNumber="36">
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<taxonomicName id="4C6E5A85FFD5D73CFF707C2D2263F8AA" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[151,357,1830,1857]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="35" pageNumber="36" phylum="Annelida" rank="species" species="boucheti" status="sp. nov.">
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<emphasis id="B91AFD14FFD5D73CFF707C2D2263F8AA" bold="true" box="[151,357,1830,1857]" italics="true" pageId="35" pageNumber="36">Chloeia boucheti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229406FFFD5D73CFE8A7C2C22A9F8AA" box="[365,431,1831,1857]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BD12106FFD5D73CFF707C4C2074F8B4" blockId="35.[151,882,1830,1924]" box="[151,882,1863,1887]" pageId="35" pageNumber="36">
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<uri id="FFFF2D04FFD5D73CFF707C4C2074F8B4" box="[151,882,1863,1887]" pageId="35" pageNumber="36">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC81BD3FFD5D73CFE727C4C2074F8B4" box="[405,882,1863,1887]" pageId="35" pageNumber="36">379696D1-12B7-4E11-A099-82C2C2A6D01F</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374728DFFD5D73CFF707C6223E1F86F" box="[151,231,1897,1924]" pageId="35" pageNumber="36" type="description">
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<paragraph id="8BD12106FFD5D73CFF707C6223E1F86F" blockId="35.[151,882,1830,1924]" box="[151,231,1897,1924]" pageId="35" pageNumber="36">
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<figureCitation id="13553D83FFD5D73CFF707C6223E1F86F" box="[151,231,1897,1924]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="35" pageNumber="36">Fig. 15</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C374728DFFD5D73BFF707CBA2652FEF6" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="materials_examined">
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<paragraph id="8BD12106FFD5D73BFF707CBA2652FEF6" blockId="35.[151,1436,1969,2032]" lastBlockId="36.[151,1437,151,1905]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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||
<emphasis id="B91AFD14FFD5D73CFF707CBA2238F820" bold="true" box="[151,318,1969,1995]" pageId="35" pageNumber="36">Type material</emphasis>
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||
.
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<materialsCitation id="3B062B5BFFD5D73CFEAB7CBA208EF820" ID-GBIF-Occurrence="4029882388" box="[332,904,1969,1996]" collectingDate="1991-11-04" collectionCode="MNHN" country="Indonesia" elevation="240" latitude="-9.383333" location="Indonesia" longLatPrecision="1300" longitude="131.15" pageId="35" pageNumber="36" specimenCode="2048" specimenCount="1" typeStatus="holotype">
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<emphasis id="B91AFD14FFD5D73CFEAB7CBA22C7F820" bold="true" box="[332,449,1969,1995]" pageId="35" pageNumber="36">
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||
<collectingCountry id="F3796196FFD5D73CFEAB7CBA22C7F820" box="[332,449,1969,1995]" name="Indonesia" pageId="35" pageNumber="36">Indonesia</collectingCountry>
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||
</emphasis>
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.
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<typeStatus id="54D59FA4FFD5D73CFE287CBA2131F820" box="[463,567,1969,1995]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="ED7FB9C3FFD5D73CFDA27CBA219AF820" box="[581,668,1969,1995]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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IA-TYPE
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<specimenCode id="DBC8897DFFD5D73CFCFF7CBA2054F820" box="[792,850,1969,1995]" pageId="35" pageNumber="36">2048</specimenCode>
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) and
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||
</materialsCitation>
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||
<materialsCitation id="3B062B5BFFD5D73BFC777CB92743FF3E" ID-GBIF-Occurrence="4029882428" collectingDate="1991-11-04" collectionCode="MNHN" country="Indonesia" elevation="240" lastPageId="36" lastPageNumber="37" latitude="-9.383333" location="RV Baruna Jaya I" longLatPrecision="1300" longitude="131.15" pageId="35" pageNumber="36" specimenCode="2050" specimenCount="1" stateProvince="RV Baruna Jaya I" typeStatus="paratype">
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||
<typeStatus id="54D59FA4FFD5D73CFC777CB920F6F827" box="[912,1008,1970,1996]" pageId="35" pageNumber="36" type="paratype">paratype</typeStatus>
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||
(
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||
<collectionCode id="ED7FB9C3FFD5D73CFC197CBA2753F820" box="[1022,1109,1969,1995]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
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||
IA-TYPE
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<specimenCode id="DBC8897DFFD5D73CFB367CBA260BF827" box="[1233,1293,1969,1996]" pageId="35" pageNumber="36">2050</specimenCode>
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||
),
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||
<collectingRegion id="49AAEFE4FFD5D73CFAC77CB923DDF81B" pageId="35" pageNumber="36">RV Baruna Jaya I</collectingRegion>
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||
, Sta. CP85 (
|
||
<geoCoordinate id="EE5A47C1FFD5D73CFE8F7CDE22C8F804" box="[360,462,2005,2031]" degrees="09" direction="south" minutes="22" orientation="latitude" pageId="35" pageNumber="36" precision="925" value="-9.366667">09°22´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFD5D73CFE3C7CDE2157F804" box="[475,593,2005,2031]" degrees="131" direction="east" minutes="14" orientation="longitude" pageId="35" pageNumber="36" precision="925" value="131.23334">131°14´E</geoCoordinate>
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||
), 245–
|
||
<quantity id="4C968CE3FFD5D73CFD7A7CDE21E1F81B" box="[669,743,2005,2032]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="240.0">
|
||
<elevation id="0043C635FFD5D73CFD7A7CDE21E1F81B" box="[669,743,2005,2032]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="35" pageNumber="36" unit="m" value="240.0">240 m</elevation>
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||
</quantity>
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||
, beam trawl,
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||
<date id="FFD007C6FFD5D73CFC987CDE270FF804" box="[895,1033,2005,2032]" pageId="35" pageNumber="36" value="1991-11-04">
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||
<collectingDate id="EF94FE2EFFD5D73CFC987CDE270FF804" box="[895,1033,2005,2032]" pageId="35" pageNumber="36" value="1991-11-04">4 Nov. 1991</collectingDate>
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||
</date>
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||
(paratype darker; anterior eyes reddish, minute, posterior ones not seen; caruncle grayish, median ridge blackish; branchial bases orange; anal cirri pale orange; body markedly bent ventrally;
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<quantity id="4C968CE3FFD2D73BFD9E7BB021C1FF3E" box="[633,711,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="36" pageNumber="37" unit="mm" value="46.0">46 mm</quantity>
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||
long,
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||
<quantity id="4C968CE3FFD2D73BFCEC7BB0205EFF3E" box="[779,856,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="36" pageNumber="37" unit="mm" value="12.0">12 mm</quantity>
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||
wide, 33 chaetigers).
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||
</materialsCitation>
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||
<materialsCitation id="3B062B5BFFD2D73BFBAD7BB02652FEF6" ID-GBIF-Occurrence="4029882492" collectingDate="1991-11-04" collectionCode="ECOSUR" country="Indonesia" elevation="246" latitude="-9.383333" location="RV Baruna Jaya I" longLatPrecision="1300" longitude="131.15" pageId="36" pageNumber="37" specimenCode="307" specimenCount="1" stateProvince="RV Baruna Jaya I" typeStatus="paratype">
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||
<specimenCount id="9D68EA8FFFD2D73BFBAD7BB0277FFF3E" box="[1098,1145,187,213]" pageId="36" pageNumber="37">One</specimenCount>
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||
<typeStatus id="54D59FA4FFD2D73BFB997BB027D8FF3E" box="[1150,1246,187,213]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
(
|
||
<collectionCode id="ED7FB9C3FFD2D73BFB0B7BB0265AFF3E" box="[1260,1372,187,213]" pageId="36" pageNumber="37">ECOSUR</collectionCode>
|
||
<specimenCode id="DBC8897DFFD2D73BFA857BB02696FF3E" box="[1378,1424,187,213]" pageId="36" pageNumber="37">307</specimenCode>
|
||
),
|
||
<collectingRegion id="49AAEFE4FFD2D73BFF707BD42258FF12" box="[151,350,223,249]" pageId="36" pageNumber="37">RV Baruna Jaya I</collectingRegion>
|
||
, Sta. CP84 (
|
||
<geoCoordinate id="EE5A47C1FFD2D73BFE0E7BD42148FF12" box="[489,590,223,249]" degrees="09" direction="south" minutes="23" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="-9.383333">09°23´S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5A47C1FFD2D73BFDBC7BD421D6FF12" box="[603,720,223,249]" degrees="131" direction="east" minutes="09" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="131.15">131°09´E</geoCoordinate>
|
||
), 275–
|
||
<quantity id="4C968CE3FFD2D73BFCFB7BD52063FF12" box="[796,869,222,249]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="36" pageNumber="37" unit="m" value="246.0">
|
||
<elevation id="0043C635FFD2D73BFCFB7BD52063FF12" box="[796,869,222,249]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="36" pageNumber="37" unit="m" value="246.0">246 m</elevation>
|
||
</quantity>
|
||
, beam trawl,
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||
<date id="FFD007C6FFD2D73BFC1B7BD42783FF12" box="[1020,1157,223,249]" pageId="36" pageNumber="37" value="1991-11-04">
|
||
<collectingDate id="EF94FE2EFFD2D73BFC1B7BD42783FF12" box="[1020,1157,223,249]" pageId="36" pageNumber="37" value="1991-11-04">4 Nov. 1991</collectingDate>
|
||
</date>
|
||
(paratype bent ventrally, many chaetae broken; bipinnate branchiae with orange bases; body
|
||
<quantity id="4C968CE3FFD2D73BFC6A7A0920DAFEF6" box="[909,988,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="37" unit="mm" value="25.0">25 mm</quantity>
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||
long,
|
||
<quantity id="4C968CE3FFD2D73BFBC47A082762FEF6" box="[1059,1124,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="36" pageNumber="37" unit="mm" value="8.0">8 mm</quantity>
|
||
wide, 31 chaetigers).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207A2D2070FE62" pageId="36" pageNumber="37" type="diagnosis">
|
||
<paragraph id="8BD12106FFD2D73BFF207A2D2070FE62" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207A2D223CFEAB" bold="true" box="[199,314,294,320]" pageId="36" pageNumber="37">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFD2D73BFEAB7A2C22A3FEAA" authorityName="Savigny" authorityYear="1818" box="[332,421,295,321]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="36" pageNumber="37" phylum="Annelida" rank="genus">
|
||
<emphasis id="B91AFD14FFD2D73BFEAB7A2C22A3FEAA" box="[332,421,295,321]" italics="true" pageId="36" pageNumber="37">Chloeia</emphasis>
|
||
</taxonomicName>
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||
with bippinate branchiae from chaetiger 4, progressively smaller posteriorly, branchial bases orange, with 11–12 tapered, lateral branches; anterior prostomial area pale; dorsum without middorsal spots; notochaetae furcates and harpoon-chaetae; neurochaetae furcates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207A9927AEFADA" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BD12106FFD2D73BFF207A99227EFE3A" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207A992257FE47" bold="true" box="[199,337,402,428]" pageId="36" pageNumber="37">Description</emphasis>
|
||
.
|
||
<typeStatus id="54D59FA4FFD2D73BFE877A9822CEFE46" box="[352,456,403,429]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
(MNHN IA-TYPE 2048) complete, bent ventrally; body tapered,
|
||
<quantity id="4C968CE3FFD2D73BFB527A982603FE46" box="[1205,1285,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="36" pageNumber="37" unit="mm" value="44.0">44 mm</quantity>
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||
long,
|
||
<quantity id="4C968CE3FFD2D73BFAAA7A98269BFE46" box="[1357,1437,403,429]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="36" pageNumber="37" unit="mm" value="13.0">13 mm</quantity>
|
||
wide, 34 chaetigers.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF207AD02755FDF2" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<typeStatus id="54D59FA4FFD2D73BFF207AD02229FE1E" box="[199,303,475,501]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
pale; dorsum without pigmentation (
|
||
<figureCitation id="13553D83FFD2D73BFD2B7AD02032FE1E" box="[716,820,474,501]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15A</figureCitation>
|
||
). Dorsal cirrophores orange, cirrostyles pale. Bipinnate branchiae pale, bases orange (
|
||
<figureCitation id="13553D83FFD2D73BFE007AF42149FDF2" box="[487,591,510,537]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15H</figureCitation>
|
||
). Venter pale, with a midventral whitish band.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF2079282115FD42" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">Prostomium anteriorly entire, anterior prostomial area pale. Eyes reddish, anterior ones minute, posterior ones not seen. Median antenna inserted at anterior caruncular margin, pale, ¼ as long as caruncle, about 2× longer than lateral antennae. Lateral antennae bases close to each other. Palps 2/3 as long as lateral antennae. Mouth ventral on chaetiger 2. Pharynx not exposed.</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF2079B82773FD1A" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
Caruncle grayish, trilobed, straight, tapered, reaching chaetiger 4 (
|
||
<figureCitation id="13553D83FFD2D73BFC2679B8272FFD26" box="[961,1065,690,717]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15B</figureCitation>
|
||
); middorsal ridge blackish, with about 46 vertical folds. Lateral lobes narrow, barely visible, with about 40 vertical folds.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF2079F02038FCB6" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
Bippinate branchiae from chaetiger 4, continued throughout body, parallel along most body segments, with bases orange (
|
||
<figureCitation id="13553D83FFD2D73BFED1781422B9FCD2" box="[310,447,798,825]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15C, H</figureCitation>
|
||
); progressively larger to chaetigers 10–16, diminishing in size in posterior chaetigers. In median segments each branchia with 11–12 lateral branches.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF20786C205CFBFA" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">Parapodia biramous, notopodia with cirriform branchiae along chaetigers 1–3, progressively smaller, ¼–1/5 as long as dorsal cirri. Dorsal cirri with cirrophores orange distally, cirrostyles pale, cirri slightly longer than branchiae along anterior chaetigers, about 3× longer in posterior chaetigers. Second ventral cirri with cirrophores 2× longer, and cirrostyles almost 3× longer than adjacent ones, directed dorsally. Other ventral cirri directed ventrolaterally, 2× as long as following segments in median and posterior regions.</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF207F10228BFB2E" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
Chaetae very abundant, many with broken tips. Complete chaetae with distal fragile hoods, rarely eroded. Notochaetae in anterior chaetigers furcates, major tines brownish, 4–8× longer than minor ones, (
|
||
<figureCitation id="13553D83FFD2D73BFB2F7F342637FBB2" box="[1224,1329,1086,1113]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15D</figureCitation>
|
||
). Median chaetigers with furcates and abundant harpoon-chaetae, both with short spurs (
|
||
<figureCitation id="13553D83FFD2D73BFBE47F682760FB96" box="[1027,1126,1122,1149]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15F</figureCitation>
|
||
). Neurochaetae all furcates, tips brownish, major tines 2–3× longer than minor ones in anterior chaetigers (
|
||
<figureCitation id="13553D83FFD2D73BFBF57F8C277EFB4A" box="[1042,1144,1158,1185]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15E</figureCitation>
|
||
), 2—5× longer in median chaetigers (
|
||
<figureCitation id="13553D83FFD2D73BFEFF7FA02286FB2E" box="[280,384,1194,1221]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF207FC4220AFAE6" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
Posterior region tapered; pygidium with anus terminal; anal cirri pale orange (
|
||
<figureCitation id="13553D83FFD2D73BFBAD7FC427B4FB02" box="[1098,1202,1230,1257]" captionStart="FIGURE 15" captionStartId="35.[152,255,1580,1604]" captionTargetBox="[184,1405,634,1557]" captionTargetId="figure-247@35.[182,1405,633,1557]" captionTargetPageId="35" captionText="FIGURE 15. Chloeia boucheti sp. n., holotype (MNHN IA-TYPE 2048). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Posterior region, dorsal view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, notochaetae furcate and harpoon-chaetae. G. Same, neurochaetal tips. H. Paratype (MNHN IA-TYPE 2050), chaetigers 13-16, dorsal view (most chaetae trimmed). Scale bars: A, 1.7 mm; B, 0.6 mm; C, 1.6 mm; D, 140 μm, E, 30 μm; F, 200 μm; G, 40 μm, H, 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7621852" httpUri="https://zenodo.org/record/7621852/files/figure.png" pageId="36" pageNumber="37">Fig. 15C</figureCitation>
|
||
), digitate, 4× longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BD12106FFD2D73BFF207E1D27AEFADA" blockId="36.[151,1437,151,1905]" box="[199,1192,1302,1329]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207E1D22AFFADB" bold="true" box="[199,425,1302,1328]" pageId="36" pageNumber="37">Live pigmentation.</emphasis>
|
||
Unknown. Bipinnate branchial bases and dorsal cirrophores orange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207E302767FA2A" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="8BD12106FFD2D73BFF207E302767FA2A" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207E30224EFABE" bold="true" box="[199,328,1339,1365]" pageId="36" pageNumber="37">Etymology</emphasis>
|
||
. The specific epithet is derived after Dr. Philippe Bouchet, a taxonomist of marine molluscs from the Muséum National d’Histoire Naturelle, Paris, in recognition of his many publications on molluscs, and especially because he participated in the Karubar Expedition (
|
||
<bibRefCitation id="EFFF5CF7FFD2D73BFCF57E8820F2FA76" author="Crosnier, A. & Richer de Forges, B. & Bouchet, P." box="[786,1012,1411,1437]" pageId="36" pageNumber="37" pagination="9 - 26" refId="ref104603" refString="Crosnier, A., Richer de Forges, B. & Bouchet, P. (1997) La campagne KARUBAR en Indonesie au large des iles Kai et Tanimbar. Resultats des campagnes Musorstom 16. Memoires du Museum National d'Histoire Naturelle, Paris, 172, 9 - 26." type="journal article" year="1997">
|
||
Crosnier
|
||
<emphasis id="B91AFD14FFD2D73BFC9C7E8820B4FA76" box="[891,946,1411,1437]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
), which collected the
|
||
<typeStatus id="54D59FA4FFD2D73BFB037E882612FA76" box="[1252,1300,1411,1437]" pageId="36" pageNumber="37">type</typeStatus>
|
||
material for this species. The derived name is a noun in the genitive case (
|
||
<bibRefCitation id="EFFF5CF7FFD2D73BFCAF7EAC20CBFA2A" author="ICZN & International Commission of Zoological Nomenclature" box="[840,973,1447,1473]" pageId="36" pageNumber="37" refId="ref108505" refString="ICZN (International Commission of Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London. Available from: https: // code. iczn. org (accessed 22 June 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
, Art. 31.1.2).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207EC1265AF9C6" pageId="36" pageNumber="37" type="description">
|
||
<paragraph id="8BD12106FFD2D73BFF207EC1265AF9C6" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207EC1223EFA0F" bold="true" box="[199,312,1482,1508]" pageId="36" pageNumber="37">Variation</emphasis>
|
||
. The
|
||
<typeStatus id="54D59FA4FFD2D73BFE927EC022D3FA0E" box="[373,469,1483,1509]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
(MNHN IA-TYPE 2050) is more colorful than the
|
||
<typeStatus id="54D59FA4FFD2D73BFBEC7EC02776FA0E" box="[1035,1136,1483,1509]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
|
||
, especially regarding bases of bipinnate branchiae, and tips of dorsal cirrophores. The other
|
||
<typeStatus id="54D59FA4FFD2D73BFCB97EE420B8F9E2" box="[862,958,1519,1545]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
|
||
(
|
||
<specimenCode id="DBC8897DFFD2D73BFC2B7EE42768F9E2" box="[972,1134,1519,1545]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="36" pageNumber="37" type="Museum">ECOSUR 307</specimenCode>
|
||
) is paler, orange pigmentation barely seen along bipinnate branchiae bases; eyes indistinct, and its anal cirri are 6–7× longer than wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207D3C22D9F8A6" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="8BD12106FFD2D73BFF207D3C22D9F8A6" blockId="36.[151,1437,151,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207D3C2232F9BA" bold="true" box="[199,308,1591,1617]" pageId="36" pageNumber="37">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C6E5A85FFD2D73BFEA57D3C2104F9BA" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[322,514,1591,1617]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="boucheti" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFD2D73BFEA57D3C2104F9BA" box="[322,514,1591,1617]" italics="true" pageId="36" pageNumber="37">Chloeia boucheti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFD2D73BFDED7D3C2140F9BA" box="[522,582,1591,1617]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
|
||
is being described with specimens from
|
||
<collectingCountry id="F3796196FFD2D73BFBF57D3C2779F9BA" box="[1042,1151,1591,1617]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
|
||
and belongs in the group tumida because its dorsum has no pigmentation pattern, and its bipinnate branchiae start in chaetiger 4 and are progressively smaller posteriorly. By having a tapered body, and bipinnate branchiae with 11–12 tapered lateral branches, it resembles
|
||
<taxonomicName id="4C6E5A85FFD2D73BFE477DA82109F956" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[416,527,1699,1725]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="richeri" status="sp. nov.">
|
||
<emphasis id="B91AFD14FFD2D73BFE477DA82109F956" box="[416,527,1699,1725]" italics="true" pageId="36" pageNumber="37">C. richeri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229406FFFD2D73BFDFD7DA8215FF956" box="[538,601,1699,1725]" pageId="36" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
|
||
from
|
||
<collectingCountry id="F3796196FFD2D73BFD447DA82051F956" box="[675,855,1699,1725]" name="New Caledonia" pageId="36" pageNumber="37">New Caledonia</collectingCountry>
|
||
. The main differences are the pigmentation of the anterior prostomial area, the coloration of bipinnate branchiae bases, and the
|
||
<typeStatus id="54D59FA4FFD2D73BFBE17DCC2730F90A" box="[1030,1078,1735,1761]" pageId="36" pageNumber="37">type</typeStatus>
|
||
of notochaetae. In
|
||
<taxonomicName id="4C6E5A85FFD2D73BFAF47DCC2693F90A" authorityName="Salazar-Vallejo" authorityYear="2023" box="[1299,1429,1735,1761]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="boucheti">
|
||
<emphasis id="B91AFD14FFD2D73BFAF47DCC2693F90A" box="[1299,1429,1735,1761]" italics="true" pageId="36" pageNumber="37">C. boucheti</emphasis>
|
||
</taxonomicName>
|
||
, the anterior prostomial area is pale, its bipinnate branchiae have orange bases, and its notochaetae are furcates and harpoon-chaetae, whereas in
|
||
<taxonomicName id="4C6E5A85FFD2D73BFE037C042154F8C2" authorityName="Salazar-Vallejo" authorityYear="2023" box="[484,594,1807,1833]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="richeri">
|
||
<emphasis id="B91AFD14FFD2D73BFE037C042154F8C2" box="[484,594,1807,1833]" italics="true" pageId="36" pageNumber="37">C. richeri</emphasis>
|
||
</taxonomicName>
|
||
, the anterior prostomial area is blackish, its branchial bases are pale, and only has furcate notochaetae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C374728DFFD2D73BFF207C5D20A4F89A" box="[199,930,1878,1905]" pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph id="8BD12106FFD2D73BFF207C5D20A4F89A" blockId="36.[151,1437,151,1905]" box="[199,930,1878,1905]" pageId="36" pageNumber="37">
|
||
<emphasis id="B91AFD14FFD2D73BFF207C5D225FF89B" bold="true" box="[199,345,1878,1904]" pageId="36" pageNumber="37">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3796196FFD2D73BFE807C5C22D1F89A" box="[359,471,1879,1905]" name="Indonesia" pageId="36" pageNumber="37">Indonesia</collectingCountry>
|
||
, in sediments at
|
||
<quantity id="4C968CE3FFD2D73BFD767C5D2016F89A" box="[657,784,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="2.575" metricValueMax="2.75" metricValueMin="2.4" pageId="36" pageNumber="37" unit="m" value="257.5" valueMax="275.0" valueMin="240.0">240–275 m</quantity>
|
||
water depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |