treatments-xml/data/03/C7/90/03C79010FFBAD751FF707EB4265FFB4A.xml
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<document id="72B160227944D62DD23F6793599BD0ED" ID-CLB-Dataset="53418" ID-DOI="10.11646/zootaxa.5238.1.1" ID-GBIF-Dataset="751096f2-4b5b-43c3-9748-4d07afe044c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7621793" ID-ZooBank="768E9932-2D18-4115-8359-3FF800328BCD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1675871988450" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2023" docId="03C79010FFBAD751FF707EB4265FFB4A" docLanguage="en" docName="zootaxa.5238.1.1.pdf" docOrigin="Zootaxa 5238 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5238.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chloeia keablei Salazar-Vallejo 2023, sp. n." docType="treatment" docVersion="7" lastPageNumber="79" masterDocId="FFFEE868FFF6D71FFFE77B0B2306FFEB" masterDocTitle="Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="77" updateTime="1699474919205" updateUser="ExternalLinkService">
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<mods:title id="2B188A780AE7259A7613FBFC3B90BE5E">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<mods:namePart id="67648120CDFF3A33F23647EC8DB9E2EE">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:date id="C1B0695E8A2C9D9D81E9E4B7B202A6B4">2023</mods:date>
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<treatment id="03C79010FFBAD751FF707EB4265FFB4A" ID-DOI="http://doi.org/10.5281/zenodo.7621953" ID-GBIF-Taxon="209295552" ID-Zenodo-Dep="7621953" LSID="urn:lsid:plazi:treatment:03C79010FFBAD751FF707EB4265FFB4A" httpUri="http://treatment.plazi.org/id/03C79010FFBAD751FF707EB4265FFB4A" lastPageId="78" lastPageNumber="79" pageId="76" pageNumber="77">
<subSubSection id="C374728DFFBAD753FF707EB42080FA13" pageId="76" pageNumber="77" type="nomenclature">
<paragraph id="8BD12106FFBAD753FF707EB4229BFA31" blockId="76.[151,902,1471,1565]" box="[151,413,1471,1498]" pageId="76" pageNumber="77">
<heading id="D099966AFFBAD753FF707EB4229BFA31" bold="true" box="[151,413,1471,1498]" fontSize="11" level="2" pageId="76" pageNumber="77" reason="4">
<emphasis id="B91AFD14FFBAD753FF707EB4229BFA31" bold="true" box="[151,413,1471,1498]" pageId="76" pageNumber="77">
<taxonomicName id="4C6E5A85FFBAD753FF707EB42255FA31" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[151,339,1471,1498]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="76" pageNumber="77" phylum="Annelida" rank="species" species="keablei" status="sp. nov.">
<emphasis id="B91AFD14FFBAD753FF707EB42255FA31" bold="true" box="[151,339,1471,1498]" italics="true" pageId="76" pageNumber="77">Chloeia keablei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFFBAD753FEBC7ECB229BFA31" box="[347,413,1472,1498]" pageId="76" pageNumber="77" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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urn:lsid:zoobank.org:act:
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<subSubSection id="C374728DFFBAD753FF707D0823E1F9F6" box="[151,231,1538,1565]" pageId="76" pageNumber="77" type="description">
<paragraph id="8BD12106FFBAD753FF707D0823E1F9F6" blockId="76.[151,902,1471,1565]" box="[151,231,1538,1565]" pageId="76" pageNumber="77">
<figureCitation id="13553D83FFBAD753FF707D0823E1F9F6" box="[151,231,1538,1565]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="76" pageNumber="77">Fig. 35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFBAD753FF707D402655F81A" pageId="76" pageNumber="77" type="materials_examined">
<paragraph id="8BD12106FFBAD753FF707D4023D2F93A" blockId="76.[151,1436,1610,2033]" pageId="76" pageNumber="77">
<emphasis id="B91AFD14FFBAD753FF707D402238F98F" bold="true" box="[151,318,1610,1637]" pageId="76" pageNumber="77">Type material</emphasis>
.
<materialsCitation id="3B062B5BFFBAD753FEAB7D40205BF98E" ID-GBIF-Occurrence="4029882474" accessionNumber="W24926, W53392" box="[332,861,1610,1637]" collectingDate="1991-03-17" collectingDateMax="1991-03-18" collectingDateMin="1991-03-17" collectionCode="AM" collectorName="J. K. Lowry &amp; S. J. Keable" country="Papua New Guinea" county="Lagoon" elevation="35" latitude="-5.16" location="Papua New Guinea" longLatPrecision="13" longitude="145.8295" municipality="Padoz Natun" pageId="76" pageNumber="77" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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<collectingCountry id="F3796196FFBAD753FEAB7D402134F98F" box="[332,562,1610,1637]" name="Papua New Guinea" pageId="76" pageNumber="77">Papua New Guinea</collectingCountry>
</emphasis>
.
<typeStatus id="54D59FA4FFBAD753FDA77D4021AEF98E" box="[576,680,1611,1637]" pageId="76" pageNumber="77" type="holotype">Holotype</typeStatus>
(
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<accessionNumber id="943DBCE5FFBAD753FD0B7D402057F98E" box="[748,849,1611,1637]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W24926" pageId="76" pageNumber="77">W24926</accessionNumber>
),
</materialsCitation>
<materialsCitation id="3B062B5BFFBAD753FC837D4023D2F93A" ID-GBIF-Occurrence="4029882418" accessionNumber="W24926, W53392" collectingDate="1991-03-17" collectingDateMax="1991-03-18" collectingDateMin="1991-03-17" collectingMethod="baited trap" collectionCode="AM" collectorName="J. K. Lowry &amp; S. J. Keable" country="Papua New Guinea" county="Lagoon" elevation="35" latitude="-5.16" location="Paeowai Island" longLatPrecision="13" longitude="145.8295" municipality="Padoz Natun" pageId="76" pageNumber="77" specimenCount="10" stateProvince="Madang" typeStatus="holotype">
and
<specimenCount id="9D68EA8FFFBAD753FC747D402727F98E" box="[915,1057,1611,1637]" pageId="76" pageNumber="77" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
(
<collectionCode id="ED7FB9C3FFBAD753FBD77D402759F98E" box="[1072,1119,1611,1637]" country="Australia" name="Australian Museum" pageId="76" pageNumber="77" type="Museum">AM</collectionCode>
<accessionNumber id="943DBCE5FFBAD753FB827D4127CCF98E" box="[1125,1226,1610,1637]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W53392" pageId="76" pageNumber="77">W53392</accessionNumber>
),
<collectingRegion id="49AAEFE4FFBAD753FB3A7D40263FF98E" box="[1245,1337,1611,1637]" country="Papua New Guinea" name="Madang" pageId="76" pageNumber="77">Madang</collectingRegion>
<collectingCounty id="62B0598AFFBAD753FAA77D40269EF98E" box="[1344,1432,1611,1637]" pageId="76" pageNumber="77">Lagoon</collectingCounty>
, between
<collectingMunicipality id="6BB5BB7CFFBAD753FF197D642296F962" box="[254,400,1647,1673]" pageId="76" pageNumber="77">Padoz Natun</collectingMunicipality>
reef and north end of
<location id="8EB177DDFFBAD753FD7D7D642040F962" LSID="urn:lsid:plazi:treatment:03C79010FFBAD751FF707EB4265FFB4A:8EB177DDFFBAD753FD7D7D642040F962" box="[666,838,1647,1673]" country="Papua New Guinea" county="Lagoon" latitude="-5.16" longLatPrecision="13" longitude="145.8295" municipality="Padoz Natun" name="Paeowai Island" pageId="76" pageNumber="77" stateProvince="Madang">Paeowai Island</location>
(
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,
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),
<collectingMethod id="522F5911FFBAD753FB427D642624F962" box="[1189,1314,1647,1673]" pageId="76" pageNumber="77">baited trap</collectingMethod>
, soft mud, burrows, and Ampelisca-like tubes,
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,
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<collectingDate id="EF94FE2EFFBAD753FD8B7D98202AF946" box="[620,812,1683,1709]" pageId="76" pageNumber="77" value="1991-03-17" valueMax="1991-03-18" valueMin="1991-03-17">1718 Mar. 1991</collectingDate>
</date>
,
<collectorName id="269B44D0FFBAD753FCDF7D9820B0F946" box="[824,950,1683,1709]" pageId="76" pageNumber="77">J.K. Lowry</collectorName>
&amp;
<collectorName id="269B44D0FFBAD753FC3F7D98275FF946" box="[984,1113,1683,1709]" pageId="76" pageNumber="77">S.J. Keable</collectorName>
, coll. (paratype data in variation).
</materialsCitation>
</paragraph>
<paragraph id="8BD12106FFBAD753FF207DD12655F81A" blockId="76.[151,1436,1610,2033]" pageId="76" pageNumber="77">
<emphasis id="B91AFD14FFBAD753FF207DD122BCF91F" bold="true" box="[199,442,1754,1780]" pageId="76" pageNumber="77">Additional material.</emphasis>
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<emphasis id="B91AFD14FFBAD753FE277DD12136F91F" bold="true" box="[448,560,1754,1780]" pageId="76" pageNumber="77">
<collectingCountry id="F3796196FFBAD753FE277DD12136F91F" box="[448,560,1754,1780]" name="Australia" pageId="76" pageNumber="77">Australia</collectingCountry>
</emphasis>
.
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(
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<accessionNumber id="943DBCE5FFBAD753FCC47DD1208EF91E" box="[803,904,1754,1781]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W28125" pageId="76" pageNumber="77">W28125</accessionNumber>
),
<location id="8EB177DDFFBAD753FC7C7DD02757F91E" LSID="urn:lsid:plazi:treatment:03C79010FFBAD751FF707EB4265FFB4A:8EB177DDFFBAD753FC7C7DD02757F91E" box="[923,1105,1755,1781]" country="Australia" latitude="-30.2625" longLatPrecision="12" longitude="153.36633" name="MV Cherry Lee" pageId="76" pageNumber="77">MV Cherry Lee</location>
, Sta. NSW 887 (
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,
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),
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</quantity>
,
<date id="FFD007C6FFBAD753FE7E7DF42135F8F2" box="[409,563,1791,1817]" pageId="76" pageNumber="77" value="1993-08-12">
<collectingDate id="EF94FE2EFFBAD753FE7E7DF42135F8F2" box="[409,563,1791,1817]" pageId="76" pageNumber="77" value="1993-08-12">12 Aug. 1993</collectingDate>
</date>
,
<collectorName id="269B44D0FFBAD753FDD97DF421B6F8F2" box="[574,688,1791,1817]" pageId="76" pageNumber="77">P. Berents</collectorName>
, coll. (pinkish, bent ventrally, body wall broken laterally along left size of chaetigers 1112; dorsal cirri mostly lost, colorless; branchiae colorless; anterior eyes reddish, 45× larger than posterior ones; median antenna 1/3 as long as caruncle, slightly longer than lateral antennae; body
<quantity id="4C968CE3FFBAD753FAF77C4C2658F88A" box="[1296,1374,1863,1889]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="76" pageNumber="77" unit="mm" value="30.0">30 mm</quantity>
long,
<quantity id="4C968CE3FFBAD753FF707C6023D1F86E" box="[151,215,1899,1925]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="76" pageNumber="77" unit="mm" value="8.0">8 mm</quantity>
wide, 31 chaetigers)
</materialsCitation>
.
<materialsCitation id="3B062B5BFFBAD753FE2B7C612649F81A" ID-GBIF-Occurrence="4029882344" collectingDate="2011-06-02" collectionCode="CAS" collectorName="R. Mooi" country="Philippines" county="Lubang" elevation="120" latitude="13.93" location="Lubang Islands" longLatPrecision="774" longitude="120.31" municipality="Lubang Islands" pageId="76" pageNumber="77" specimenCode="186937" specimenCount="1">
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<collectingCountry id="F3796196FFBAD753FE2B7C612154F86F" box="[460,594,1898,1924]" name="Philippines" pageId="76" pageNumber="77">Philippines</collectingCountry>
</emphasis>
.
<specimenCount id="9D68EA8FFFBAD753FDB87C6021F8F86E" box="[607,766,1899,1925]" pageId="76" pageNumber="77">One specimen</specimenCount>
(
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),
<collectingMunicipality id="6BB5BB7CFFBAD753FC537C602764F86E" box="[948,1122,1899,1925]" pageId="76" pageNumber="77">Lubang Islands</collectingMunicipality>
,
<collectingCounty id="62B0598AFFBAD753FB8A7C6027C5F86E" box="[1133,1219,1899,1925]" pageId="76" pageNumber="77">Lubang</collectingCounty>
(
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,
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), mixed bottom, rocks and sand,
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</quantity>
,
<date id="FFD007C6FFBAD753FD487C842033F842" box="[687,821,1935,1961]" pageId="76" pageNumber="77" value="2011-06-02">
<collectingDate id="EF94FE2EFFBAD753FD487C842033F842" box="[687,821,1935,1961]" pageId="76" pageNumber="77" value="2011-06-02">2 Jun. 2011</collectingDate>
</date>
,
<collectorName id="269B44D0FFBAD753FCA77C842099F842" box="[832,927,1935,1961]" pageId="76" pageNumber="77">R. Mooi</collectorName>
<emphasis id="B91AFD14FFBAD753FC417C8420D9F842" box="[934,991,1935,1961]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
, coll. (colorless, dorsal cirri purple, caruncular median ridge blackish, many notochaetae broken; anterior eyes 6× larger than posterior ones; bipinnate branchiae in median chaetigers with 1416 lateral branches; body
<quantity id="4C968CE3FFBAD753FC997CDC20CBF81A" box="[894,973,2007,2033]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="76" pageNumber="77" unit="mm" value="76.0">76 mm</quantity>
long,
<quantity id="4C968CE3FFBAD753FBF47CDD2764F81A" box="[1043,1122,2006,2033]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="76" pageNumber="77" unit="mm" value="15.0">15 mm</quantity>
wide, 37 chaetigers)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF11718EFFBBD752FF7F7FCD204FFAA1" ID-DOI="http://doi.org/10.5281/zenodo.7621925" ID-Zenodo-Dep="7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="77" pageNumber="78" startId="77.[152,255,1222,1246]" targetBox="[154,1434,183,1195]" targetPageId="77">
<paragraph id="8BD12106FFBBD752FF7F7FCD204FFAA1" blockId="77.[151,1437,1222,1355]" pageId="77" pageNumber="78">
<emphasis id="B91AFD14FFBBD752FF7F7FCD2226FB35" bold="true" box="[152,288,1222,1246]" pageId="77" pageNumber="78">FIGURE 35</emphasis>
.
<taxonomicName id="4C6E5A85FFBBD752FEC97FCD22C9FB35" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[302,463,1222,1246]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="77" pageNumber="78" phylum="Annelida" rank="species" species="keablei" status="sp. nov.">
<emphasis id="B91AFD14FFBBD752FEC97FCD22C9FB35" box="[302,463,1222,1246]" italics="true" pageId="77" pageNumber="78">Chloeia keablei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFFBBD752FE307FCC2117FB35" box="[471,529,1223,1247]" pageId="77" pageNumber="78" rank="species">
<emphasis id="B91AFD14FFBBD752FE307FCC210DFB34" bold="true" box="[471,523,1223,1247]" pageId="77" pageNumber="78">sp. n</emphasis>
.
</taxonomicNameLabel>
, holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm.
</paragraph>
</caption>
<subSubSection id="C374728DFFBBD752FF207E7320FFFA30" pageId="77" pageNumber="78" type="diagnosis">
<paragraph id="8BD12106FFBBD752FF207E7320FFFA30" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">
<emphasis id="B91AFD14FFBBD752FF207E73223CFA79" bold="true" box="[199,314,1400,1426]" pageId="77" pageNumber="78">Diagnosis</emphasis>
.
<taxonomicName id="4C6E5A85FFBBD752FEAB7E7222A3FA78" authorityName="Savigny" authorityYear="1818" box="[332,421,1401,1427]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="77" pageNumber="78" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FFBBD752FEAB7E7222A3FA78" box="[332,421,1401,1427]" italics="true" pageId="77" pageNumber="78">Chloeia</emphasis>
</taxonomicName>
with bipinnate branchiae from chaetiger 4, progressively smaller posteriorly, colorless; dorsum without pigmentation pattern; anterior eyes 4× larger than posterior ones; caruncle tapered, with about 40 folds; harpoon notochaetae without spurs; neurochaetae spurred and furcates.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFBBD751FF207EEF20CEFDF2" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78" type="description">
<paragraph id="8BD12106FFBBD752FF207EEF225FFA14" blockId="77.[151,1437,1400,2041]" box="[199,345,1508,1535]" pageId="77" pageNumber="78">
<emphasis id="B91AFD14FFBBD752FF207EEF2257FA15" bold="true" box="[199,337,1508,1534]" pageId="77" pageNumber="78">Description</emphasis>
.
</paragraph>
<paragraph id="8BD12106FFBBD752FF207D0227FFF9AC" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">
<materialsCitation id="3B062B5BFFBBD752FF207D0227FFF9AC" ID-GBIF-Occurrence="4029882491" accessionNumber="W24926" collectionCode="AM" pageId="77" pageNumber="78" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D59FA4FFBBD752FF207D022229F9C8" box="[199,303,1545,1571]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED7FB9C3FFBBD752FEDA7D02226AF9C8" box="[317,364,1545,1571]" country="Australia" name="Australian Museum" pageId="77" pageNumber="78" type="Museum">AM</collectionCode>
<accessionNumber id="943DBCE5FFBBD752FE957D0222D1F9C8" box="[370,471,1545,1571]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W24926" pageId="77" pageNumber="78">W24926</accessionNumber>
), with body fusiform, posterior end in regeneration; branchiae of chaetiger 12 removed for observing segmental pigmentation; body bent laterally,
<quantity id="4C968CE3FFBBD752FCCA7D26207AF9AC" box="[813,892,1581,1607]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="77" pageNumber="78" unit="mm" value="46.0">46 mm</quantity>
long,
<quantity id="4C968CE3FFBBD752FC247D262714F9AC" box="[963,1042,1581,1607]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="77" pageNumber="78" unit="mm" value="10.0">10 mm</quantity>
wide, 30 chaetigers.
</materialsCitation>
</paragraph>
<paragraph id="8BD12106FFBBD752FF207D5A21F6F958" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">
<typeStatus id="54D59FA4FFBBD752FF207D5A2229F980" box="[199,303,1617,1643]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
with dorsum reddish (
<figureCitation id="13553D83FFBBD752FDC97D5A219EF980" box="[558,664,1616,1643]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="77" pageNumber="78">Fig. 35A</figureCitation>
), each segment with anterior third or fourth darker than posterior region, better defined along anterior (
<figureCitation id="13553D83FFBBD752FDCF7D7E218AF964" box="[552,652,1652,1679]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="77" pageNumber="78">Fig. 35C</figureCitation>
) and median chaetigers; dorsal cirri purplish; branchiae pale; chaetae golden. Venter cream, midventral band barely visible.
</paragraph>
<paragraph id="8BD12106FFBBD752FF207DB621DBF8A8" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">
Prostomium anteriorly entire, anterior prostomial area reddish (
<figureCitation id="13553D83FFBBD752FC457DB6270AF93C" box="[930,1036,1724,1751]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="77" pageNumber="78">Fig. 35B</figureCitation>
). Eyes reddish, anterior eyes 56× larger than posterior ones. Median antenna pale, inserted at anterior caruncular margin, without tip, half as long as caruncle, slightly longer than lateral antennae. Lateral antennae bases close to each other, 2× longer than palps. Mouth ventral on chaetiger 2. Pharynx not exposed.
</paragraph>
<paragraph id="8BD12106FFBBD752FF207C4627E9F860" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">Caruncle pale, sigmoid, trilobed, tapered, reaching chaetiger 5. Median ridge plicate, colorless, with about 46 vertical folds, partially concealing lateral lobes?. Lateral lobes narrow, with about 36 vertical folds.</paragraph>
<paragraph id="8BD12106FFBBD752FF207C9E209DF838" blockId="77.[151,1437,1400,2041]" pageId="77" pageNumber="78">Bipinnate branchiae from chaetiger 4, divergent throughout body, progressively larger to chaetiger 1516, smaller posteriorly; in median segments with 1112 lateral branches.</paragraph>
<paragraph id="8BD12106FFBBD751FF207CD62044FF12" blockId="77.[151,1437,1400,2041]" lastBlockId="78.[151,1437,151,1185]" lastPageId="78" lastPageNumber="79" pageId="77" pageNumber="78">Parapodia biramous, notopodia with cirriform branchiae along chaetigers 13, ½1/3 as long as dorsal cirri, progressively smaller. Dorsal cirri slightly longer than bipinnate branchiae along body. Second ventral cirri with cirrophores 2× longer and wider, and cirrostyle 3× longer than adjacent ones, directed dorsally. Other ventral cirri directed ventrolaterally, as long as two subsequent segments.</paragraph>
<paragraph id="8BD12106FFB8D751FF207A08229FFE46" blockId="78.[151,1437,151,1185]" pageId="78" pageNumber="79">
Chaetae most complete, hoods rarely eroded. Notochaetae in anterior chaetigers furcates (
<figureCitation id="13553D83FFB8D751FB407A082608FEF6" box="[1191,1294,258,285]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="78" pageNumber="79">Fig. 35D</figureCitation>
), major tines 46× longer than minor ones. Median chaetigers with
<specimenCount id="9D68EA8FFFB8D751FD197A2C206DFEAA" box="[766,875,295,321]" pageId="78" pageNumber="79" type="generic" typeStatus="types">two types</specimenCount>
of notochaetae: a few smaller acicular or spurred chaetae, and mostly harpoon-chaetae without tines or spurs (
<figureCitation id="13553D83FFB8D751FCA07A4020AAFE8E" box="[839,940,330,357]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="78" pageNumber="79">Fig. 35F</figureCitation>
). Neurochaetae furcates; anterior chaetigers with major tines 512×longer than minor ones (
<figureCitation id="13553D83FFB8D751FD5E7A642019FE62" box="[697,799,366,393]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="78" pageNumber="79">Fig. 35E</figureCitation>
), median chaetigers with major tines 510× longer than minor ones (
<figureCitation id="13553D83FFB8D751FEC37A98228BFE46" box="[292,397,402,429]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="78" pageNumber="79">Fig. 35G</figureCitation>
).
</paragraph>
<paragraph id="8BD12106FFB8D751FF207ABC23D3FE1E" blockId="78.[151,1437,151,1185]" pageId="78" pageNumber="79">
Posterior region tapered (
<figureCitation id="13553D83FFB8D751FE0E7ABC2155FE3A" box="[489,595,438,465]" captionStart="FIGURE 35" captionStartId="77.[152,255,1222,1246]" captionTargetBox="[154,1434,183,1195]" captionTargetId="figure-18@77.[151,1436,181,1198]" captionTargetPageId="77" captionText="FIGURE 35. Chloeia keablei sp. n., holotype (AM W24926). A. Anterior region, dorsal view. B. Anterior end, dorsal view, after Methyl green staining. C. Chaetigers 9-11, dorsal view. D. Chaetiger 3, notochaetal tips. E. Same, neurochaetal tips. F. Chaetiger 15, harpoon notochaetal tips. G. Same, neurochaetal tips. H. Posterior region, dorsal view. Scale bars: A, 2.3 mm; B, C, 0.9 mm; D, 100 μm; E, 30 μm; F, 120 μm; G, 40 μm; H, 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7621925" httpUri="https://zenodo.org/record/7621925/files/figure.png" pageId="78" pageNumber="79">Fig. 35H</figureCitation>
); pygidium with anus terminal; anal cirri whitish, digitate, 4× longer than wide.
</paragraph>
<paragraph id="8BD12106FFB8D751FF207AF520CEFDF2" blockId="78.[151,1437,151,1185]" box="[199,968,510,537]" pageId="78" pageNumber="79">
<emphasis id="B91AFD14FFB8D751FF207AF522A4FDF3" bold="true" box="[199,418,510,536]" pageId="78" pageNumber="79">Live pigmentation</emphasis>
. Unknown; likely more intense than in
<typeStatus id="54D59FA4FFB8D751FCB87AF420C3FDF2" box="[863,965,511,537]" pageId="78" pageNumber="79" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFB8D751FF20792823E8FD42" pageId="78" pageNumber="79" type="etymology">
<paragraph id="8BD12106FFB8D751FF20792823E8FD42" blockId="78.[151,1437,151,1185]" pageId="78" pageNumber="79">
<emphasis id="B91AFD14FFB8D751FF207928224EFDD6" bold="true" box="[199,328,547,573]" pageId="78" pageNumber="79">Etymology</emphasis>
. The species name is derived after Dr. Stephen Keable, Collection Manager in the Australian Museum, Sydney, in recognition of his long-term support of my research activities, and especially because he collected the
<typeStatus id="54D59FA4FFB8D751FEEB7960223AFD6E" box="[268,316,619,645]" pageId="78" pageNumber="79">type</typeStatus>
specimens for this description. The specific epithet is a noun in the genitive case (
<bibRefCitation id="EFFF5CF7FFB8D751FB0779602660FD6E" author="ICZN &amp; International Commission of Zoological Nomenclature" box="[1248,1382,619,645]" pageId="78" pageNumber="79" refId="ref108505" refString="ICZN (International Commission of Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London. Available from: https: // code. iczn. org (accessed 22 June 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
, Art. 31.1.2).
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFB8D751FF2079B92013FCB6" pageId="78" pageNumber="79" type="description">
<paragraph id="8BD12106FFB8D751FF2079B92013FCB6" blockId="78.[151,1437,151,1185]" pageId="78" pageNumber="79">
<emphasis id="B91AFD14FFB8D751FF2079B9223EFD27" bold="true" box="[199,312,690,716]" pageId="78" pageNumber="79">Variation</emphasis>
.
<typeStatus id="54D59FA4FFB8D751FEA179B822B5FD26" box="[326,435,691,717]" pageId="78" pageNumber="79" type="paratype">Paratypes</typeStatus>
(AM
<accessionNumber id="943DBCE5FFB8D751FE1079B8215CFD26" box="[503,602,691,717]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W24926" pageId="78" pageNumber="79">W24926</accessionNumber>
b) slightly distorted by compression. Body pink, dorsal cirri and branchiae pale; chaetae golden; venter cream with a diffuse reddish area along a few anterior chaetigers; smaller
<specimenCount id="9D68EA8FFFB8D751FACB79DC23D4FCFE" pageId="78" pageNumber="79" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
with some dorsal cirri and branchiae purplish, three with blackish eyes, all other
<typeStatus id="54D59FA4FFB8D751FBB879F027CCFCFE" box="[1119,1226,763,789]" pageId="78" pageNumber="79" type="paratype">paratypes</typeStatus>
with eyes reddish, 23× larger than posterior ones in smaller
<typeStatus id="54D59FA4FFB8D751FD97781421D8FCD2" box="[624,734,799,825]" pageId="78" pageNumber="79" type="paratype">paratypes</typeStatus>
, larger ones with anterior eyes 46× larger than posterior ones. Body
<quantity id="4C968CE3FFB8D751FF3C78482255FCB6" box="[219,339,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="4.6" metricValueMin="1.1" pageId="78" pageNumber="79" unit="mm" value="28.5" valueMax="46.0" valueMin="11.0">1146 mm</quantity>
long,
<quantity id="4C968CE3FFB8D751FE7E78482102FCB6" box="[409,516,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="78" pageNumber="79" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">310 mm</quantity>
wide, 2435 chaetigers.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFB8D751FF20786C272EFB96" pageId="78" pageNumber="79" type="discussion">
<paragraph id="8BD12106FFB8D751FF20786C272EFB96" blockId="78.[151,1437,151,1185]" pageId="78" pageNumber="79">
<emphasis id="B91AFD14FFB8D751FF20786C2232FC6A" bold="true" box="[199,308,871,897]" pageId="78" pageNumber="79">Remarks</emphasis>
.
<taxonomicName id="4C6E5A85FFB8D751FEA1786C22FCFC6A" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[326,506,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="78" pageNumber="79" phylum="Annelida" rank="species" species="keablei" status="sp. nov.">
<emphasis id="B91AFD14FFB8D751FEA1786C22FCFC6A" box="[326,506,871,897]" italics="true" pageId="78" pageNumber="79">Chloeia keablei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFFB8D751FDE3786C2142FC6A" box="[516,580,871,897]" pageId="78" pageNumber="79" rank="species">sp. n.</taxonomicNameLabel>
is described with specimens from The
<collectingCountry id="F3796196FFB8D751FBF4786C2797FC6A" box="[1043,1169,871,897]" name="Philippines" pageId="78" pageNumber="79">Philippines</collectingCountry>
to
<collectingCountry id="F3796196FFB8D751FB5D786C2622FC6A" box="[1210,1316,871,897]" name="Australia" pageId="78" pageNumber="79">Australia</collectingCountry>
, including
<collectingCountry id="F3796196FFB8D751FF7078802275FC4E" box="[151,371,907,933]" name="Papua New Guinea" pageId="78" pageNumber="79">Papua New Guinea</collectingCountry>
, its
<typeStatus id="54D59FA4FFB8D751FE78788022C9FC4E" box="[415,463,907,933]" pageId="78" pageNumber="79">type</typeStatus>
locality. Because it has no pigmentation pattern, bipinnate branchiae from chaetiger 4, progressively smaller posteriorly, it belongs in the group tumida. Further, because it has a tapered caruncle, and colorless bipinnate branchiae, it resembles
<taxonomicName id="4C6E5A85FFB8D751FDB078D82060FC06" authority="Moore, 1911" authorityName="Moore" authorityYear="1911" box="[599,870,979,1005]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="78" pageNumber="79" phylum="Annelida" rank="species" species="pinnata">
<emphasis id="B91AFD14FFB8D751FDB078D821C8FC07" box="[599,718,979,1005]" italics="true" pageId="78" pageNumber="79">C. pinnata</emphasis>
<bibRefCitation id="EFFF5CF7FFB8D751FD3278D82060FC06" author="Moore, J. P." box="[725,870,979,1005]" pageId="78" pageNumber="79" pagination="234 - 318" refId="ref111003" refString="Moore, J. P. (1911) The polychaetous annelids dredged by the U. S. S. '' Albatross' ' off the coast of Southern California in 1904, 3. Euphrosynidae to Goniadidae. Proceedings of the Academy of Natural Sciences of Philadelphia, 63, 234 - 318, pls. 15 - 21." type="journal article" year="1911">Moore, 1911</bibRefCitation>
</taxonomicName>
, from Southern California. As indicated in the key above, these two species differ in the relative size of eyes, in the
<typeStatus id="54D59FA4FFB8D751FC8D78FC209CFBFA" box="[874,922,1015,1041]" pageId="78" pageNumber="79">type</typeStatus>
of harpoon notochaetae, and of neurochaetae. Thus, in
<taxonomicName id="4C6E5A85FFB8D751FF1F7F10226FFBDE" authorityName="Salazar-Vallejo" authorityYear="2023" box="[248,361,1051,1077]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="78" pageNumber="79" phylum="Annelida" rank="species" species="keablei">
<emphasis id="B91AFD14FFB8D751FF1F7F10226FFBDE" box="[248,361,1051,1077]" italics="true" pageId="78" pageNumber="79">C. keablei</emphasis>
</taxonomicName>
the anterior eyes are 4× larger than posterior ones; harpoon notochaetae have no spurs or smooth tines, and its neurochaetae are furcates with small minor tines, whereas in
<taxonomicName id="4C6E5A85FFB8D751FC2D7F342746FBB3" authorityName="Moore" authorityYear="1911" box="[970,1088,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="78" pageNumber="79" phylum="Annelida" rank="species" species="pinnata">
<emphasis id="B91AFD14FFB8D751FC2D7F342746FBB3" box="[970,1088,1087,1113]" italics="true" pageId="78" pageNumber="79">C. pinnata</emphasis>
</taxonomicName>
anterior eyes are 2× larger than posterior ones, its harpoon notochaetae have spurs, and neurochaetae are spurred.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFFB8D751FF207F8D265FFB4A" box="[199,1369,1158,1185]" pageId="78" pageNumber="79" type="distribution">
<paragraph id="8BD12106FFB8D751FF207F8D265FFB4A" blockId="78.[151,1437,151,1185]" box="[199,1369,1158,1185]" pageId="78" pageNumber="79">
<emphasis id="B91AFD14FFB8D751FF207F8D225FFB4B" bold="true" box="[199,345,1158,1184]" pageId="78" pageNumber="79">Distribution</emphasis>
. The
<collectingCountry id="F3796196FFB8D751FE7E7F8C211CFB4A" box="[409,538,1159,1185]" name="Philippines" pageId="78" pageNumber="79">Philippines</collectingCountry>
,
<collectingCountry id="F3796196FFB8D751FDC27F8C21F8FB4A" box="[549,766,1159,1185]" name="Papua New Guinea" pageId="78" pageNumber="79">Papua New Guinea</collectingCountry>
and
<collectingCountry id="F3796196FFB8D751FCD47F8C209BFB4A" box="[819,925,1159,1185]" name="Australia" pageId="78" pageNumber="79">Australia</collectingCountry>
, in sediments at
<quantity id="4C968CE3FFB8D751FBB17F8D27C1FB4A" box="[1110,1223,1158,1185]" metricMagnitude="1" metricUnit="m" metricValue="7.95" metricValueMax="12.4" metricValueMin="3.5" pageId="78" pageNumber="79" unit="m" value="79.5" valueMax="124.0" valueMin="35.0">35124 m</quantity>
water depth.
</paragraph>
</subSubSection>
</treatment>
</document>