treatments-xml/data/03/C7/90/03C79010FF81D764FF707E7C268CFB2E.xml

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<mods:title id="3ECE866E719E02BFE6F8C779F24BED81">Revision of Chloeia Savigny in Lamarck, 1818 (Annelida, Amphinomidae)</mods:title>
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<treatment id="03C79010FF81D764FF707E7C268CFB2E" ID-DOI="http://doi.org/10.5281/zenodo.7641443" ID-GBIF-Taxon="209295553" ID-Zenodo-Dep="7641443" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E" httpUri="http://treatment.plazi.org/id/03C79010FF81D764FF707E7C268CFB2E" lastPageId="123" lastPageNumber="124" pageId="119" pageNumber="120">
<subSubSection id="C374728DFF81D768FF707E7C207BFA5B" pageId="119" pageNumber="120" type="nomenclature">
<paragraph id="8BD12106FF81D768FF707E7C22A1FA79" blockId="119.[151,893,1399,1493]" box="[151,423,1399,1426]" pageId="119" pageNumber="120">
<heading id="D099966AFF81D768FF707E7C22A1FA79" bold="true" box="[151,423,1399,1426]" fontSize="11" level="2" pageId="119" pageNumber="120" reason="4">
<emphasis id="B91AFD14FF81D768FF707E7C22A1FA79" bold="true" box="[151,423,1399,1426]" pageId="119" pageNumber="120">
<taxonomicName id="4C6E5A85FF81D768FF707E7C225BFA79" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[151,349,1399,1426]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="119" pageNumber="120" phylum="Annelida" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B91AFD14FF81D768FF707E7C225BFA79" bold="true" box="[151,349,1399,1426]" italics="true" pageId="119" pageNumber="120">Chloeia zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFF81D768FE827E7322A1FA79" box="[357,423,1400,1426]" pageId="119" pageNumber="120" rank="species">sp. n.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act: 
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<subSubSection id="C374728DFF81D768FF707EB02212FA3E" box="[151,276,1466,1493]" pageId="119" pageNumber="120" type="description">
<paragraph id="8BD12106FF81D768FF707EB02212FA3E" blockId="119.[151,893,1399,1493]" box="[151,276,1466,1493]" pageId="119" pageNumber="120">
<figureCitation id="13553D83FF81D768FF707EB023EAFA3E" box="[151,236,1466,1493]" captionStart="FIGURE 58" captionStartId="120.[152,255,899,923]" captionTargetBox="[179,1408,348,874]" captionTargetId="figure-101@120.[177,1411,345,876]" captionTargetPageId="120" captionText="FIGURE 58. Chloeia zibrowii sp. n., holotype (MNHN IA-TYPE 2056). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 14-16, dorsal view, branchiae removed from chaetiger 14. Scale bars: A, 2.7 mm; B, 1.6 mm; C, 0.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7621998" httpUri="https://zenodo.org/record/7621998/files/figure.png" pageId="119" pageNumber="120">Figs 58</figureCitation>
, 
<figureCitation id="13553D83FF81D768FF1F7EB12212FA3E" box="[248,276,1466,1493]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="119" pageNumber="120">59</figureCitation>
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<emphasis id="B91AFD14FF81D768FF707EF62238F9FC" bold="true" box="[151,318,1533,1559]" pageId="119" pageNumber="120">Type material</emphasis>
. 
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<collectingCountry id="F3796196FF81D768FEAB7EF6211DF9FC" box="[332,539,1533,1559]" name="French Polynesia" pageId="119" pageNumber="120">French Polynesia</collectingCountry>
, 
<collectingRegion id="49AAEFE4FF81D768FDC07EF6200CF9FC" box="[551,778,1533,1559]" country="French Polynesia" name="Iles Marquises" pageId="119" pageNumber="120">Marquesas Islands</collectingRegion>
.
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<paragraph id="8BD12106FF81D768FF207D2A277DF894" blockId="119.[151,1437,1533,2028]" pageId="119" pageNumber="120">
<materialsCitation id="3B062B5BFF81D768FF207D2A2126F9B4" ID-GBIF-Occurrence="4029882359" collectingDate="1997-09-09" collectingDateMax="1997-09-09" collectingDateMin="1997-08-27" collectionCode="MNHN" collectorName="RV Alis &amp; Sta. DR" county="Nuku Hiva Island" elevation="305" latitude="-8.818334" location="Alis" longLatPrecision="129" longitude="-140.285" municipality="Nuku Hiva Island" pageId="119" pageNumber="120" specimenCode="2056" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D59FA4FF81D768FF207D2A2229F9D0" box="[199,303,1569,1595]" pageId="119" pageNumber="120" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED7FB9C3FF81D768FED87D2A2290F9D0" box="[319,406,1569,1595]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="119" pageNumber="120">MNHN</collectionCode>
IA-TYPE 
<specimenCode id="DBC8897DFF81D768FDF37D2A2156F9D7" box="[532,592,1569,1596]" pageId="119" pageNumber="120">2056</specimenCode>
), RV 
<location id="8EB177DDFF81D768FD687D2A21C6F9D0" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF81D768FD687D2A21C6F9D0" box="[655,704,1569,1595]" county="Nuku Hiva Island" latitude="-8.818334" longLatPrecision="129" longitude="-140.285" municipality="Nuku Hiva Island" name="Alis" pageId="119" pageNumber="120">Alis</location>
, Sta. DR1298 (
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, 
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), off 
<location id="8EB177DDFF81D768FB2B7D2A269EF9D0" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF81D768FB2B7D2A269EF9D0" box="[1228,1432,1569,1595]" county="Nuku Hiva Island" latitude="-8.818334" longLatPrecision="129" longitude="-140.285" municipality="Nuku Hiva Island" name="Nuku Hiva Island" pageId="119" pageNumber="120">Nuku Hiva Island</location>
, rock dredge, 
<quantity id="4C968CE3FF81D768FED67D4E227BF98B" box="[305,381,1605,1632]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="119" pageNumber="120" unit="m" value="305.0">
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, 
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<collectingDate id="EF94FE2EFF81D768FE6B7D4E211DF9B4" box="[396,539,1605,1631]" pageId="119" pageNumber="120" value="1997-09-09">9 Sep. 1997</collectingDate>
</date>
.
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<materialsCitation id="3B062B5BFF81D768FDCD7D4E22D4F8F8" ID-GBIF-Occurrence="4029882503" collectingDate="1997-08-27" collectingDateMax="1997-09-09" collectingDateMin="1997-08-27" collectionCode="MNHN, ECOSUR" collectorName="RV Alis &amp; Sta. DR" county="Nuku Hiva Island" elevation="350" latitude="-8.818334" location="Nuku Hiva Island" longLatPrecision="129" longitude="-140.285" municipality="Nuku Hiva Island" pageId="119" pageNumber="120" specimenCode="2057" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D68EA8FFF81D768FDCD7D4E215AF9B4" box="[554,604,1605,1631]" pageId="119" pageNumber="120">Five</specimenCount>
<typeStatus id="54D59FA4FF81D768FD817D4D21D7F98B" box="[614,721,1606,1632]" pageId="119" pageNumber="120" type="paratype">paratypes</typeStatus>
(3: 
<collectionCode id="ED7FB9C3FF81D768FCE27D4D205DF98B" box="[773,859,1606,1632]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="119" pageNumber="120">MNHN</collectionCode>
IA-TYPE 
<specimenCode id="DBC8897DFF81D768FC3B7D4E2711F98B" box="[988,1047,1605,1632]" pageId="119" pageNumber="120">2057</specimenCode>
; 2: 
<collectionCode id="ED7FB9C3FF81D768FBA07D4E27B3F9B4" box="[1095,1205,1605,1631]" pageId="119" pageNumber="120">ECOSUR</collectionCode>
310), 
<collectorName id="269B44D0FF81D768FAE37D4D2664F9B4" box="[1284,1378,1605,1632]" pageId="119" pageNumber="120">RV Alis</collectorName>
, 
<collectorName id="269B44D0FF81D768FA967D4E23B9F968" pageId="119" pageNumber="120">Sta. DR</collectorName>
1197 (
<geoCoordinate id="EE5A47C1FF81D768FEE17D622284F968" box="[262,386,1641,1668]" degrees="08" direction="south" minutes="57.4" orientation="latitude" pageId="119" pageNumber="120" precision="92" value="-8.956667">08°57.4´S</geoCoordinate>
, 
<geoCoordinate id="EE5A47C1FF81D768FE777D622120F968" box="[400,550,1641,1667]" degrees="140" direction="west" minutes="01.9" orientation="longitude" pageId="119" pageNumber="120" precision="92" value="-140.03166">140°01.9´W</geoCoordinate>
), off 
<collectingMunicipality id="6BB5BB7CFF81D768FD847D622036F968" box="[611,816,1641,1667]" pageId="119" pageNumber="120">Nuku Hiva Island</collectingMunicipality>
, rock dredge, 
<quantity id="4C968CE3FF81D768FC347D622718F96F" box="[979,1054,1641,1668]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="119" pageNumber="120" unit="m" value="350.0">
<elevation id="0043C635FF81D768FC347D622718F96F" box="[979,1054,1641,1668]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="119" pageNumber="120" unit="m" value="350.0">350 m</elevation>
</quantity>
, 
<date id="FFD007C6FF81D768FBCC7D6227C0F968" box="[1067,1222,1641,1668]" pageId="119" pageNumber="120" value="1997-08-27">
<collectingDate id="EF94FE2EFF81D768FBCC7D6227C0F968" box="[1067,1222,1641,1668]" pageId="119" pageNumber="120" value="1997-08-27">27 Aug. 1997</collectingDate>
</date>
(complete 
<collectionCode id="ED7FB9C3FF81D768FAA07D61269BF96F" box="[1351,1437,1642,1668]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="119" pageNumber="120">MNHN</collectionCode>
paratype used for description; 
<collectionCode id="ED7FB9C3FF81D768FE007D85213BF943" box="[487,573,1678,1704]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="119" pageNumber="120">MNHN</collectionCode>
paratypes anterior fragments, one with most chaetae broken; 
<quantity id="4C968CE3FF81D768FB017D862658F943" box="[1254,1374,1677,1704]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.2" metricValueMin="2.2" pageId="119" pageNumber="120" unit="mm" value="27.0" valueMax="32.0" valueMin="22.0">22–32 mm</quantity>
long, 
<quantity id="4C968CE3FF81D768FF707DBA2217F927" box="[151,273,1713,1740]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="119" pageNumber="120" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
wide, 12–17 chaetigers; 
<collectionCode id="ED7FB9C3FF81D768FDCA7DBA219DF920" box="[557,667,1713,1739]" pageId="119" pageNumber="120">ECOSUR</collectionCode>
paratypes one complete, markedly bent laterally, with most chaetae broken, 
<quantity id="4C968CE3FF81D768FF157DDE2246F91B" box="[242,320,1749,1776]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="119" pageNumber="120" unit="mm" value="60.0">60 mm</quantity>
long, 
<quantity id="4C968CE3FF81D768FE637DDE22D4F91B" box="[388,466,1749,1776]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="119" pageNumber="120" unit="mm" value="17.0">17 mm</quantity>
wide, 33 chaetigers; an anterior fragment 
<quantity id="4C968CE3FF81D768FC427DDE20F5F91B" box="[933,1011,1749,1776]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="119" pageNumber="120" unit="mm" value="30.0">30 mm</quantity>
long, 
<quantity id="4C968CE3FF81D768FBD07DDE2771F91B" box="[1079,1143,1749,1776]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="119" pageNumber="120" unit="mm" value="9.0">9 mm</quantity>
wide, 17 chaetigers; some features used for variation).
</materialsCitation>
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<specimenCount id="9D68EA8FFF81D768FE3E7DF2210EF8F8" box="[473,520,1785,1811]" pageId="119" pageNumber="120">One</specimenCount>
<typeStatus id="54D59FA4FF81D768FDF67DF12177F8FF" box="[529,625,1786,1812]" pageId="119" pageNumber="120" type="paratype">paratype</typeStatus>
(
<specimenCode id="DBC8897DFF81D768FD667DF22020F8F8" box="[641,806,1785,1811]" collectionCode="ECOSUR" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="119" pageNumber="120" type="Museum">ECOSUR 311</specimenCode>
), RV 
<location id="8EB177DDFF81D768FC817DF22091F8F8" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF81D768FC817DF22091F8F8" box="[870,919,1785,1811]" county="Nuku Hiva Island" latitude="-8.818334" longLatPrecision="129" longitude="-140.285" municipality="Nuku Hiva Island" name="Alis" pageId="119" pageNumber="120">Alis</location>
, Sta. DR1298 (
<geoCoordinate id="EE5A47C1FF81D768FBA07DF227C5F8F8" box="[1095,1219,1785,1811]" degrees="08" direction="south" minutes="49.1" orientation="latitude" pageId="119" pageNumber="120" precision="92" value="-8.818334">08°49.1´S</geoCoordinate>
, 
<geoCoordinate id="EE5A47C1FF81D768FB367DF22661F8F8" box="[1233,1383,1785,1811]" degrees="140" direction="west" minutes="17.1" orientation="longitude" pageId="119" pageNumber="120" precision="92" value="-140.285">140°17.1´W</geoCoordinate>
), off 
<collectingCounty id="62B0598AFF81D768FF707C162262F8DC" box="[151,356,1821,1847]" pageId="119" pageNumber="120">Nuku Hiva Island</collectingCounty>
, rock dredge, 
<quantity id="4C968CE3FF81D768FDE17C162156F8D3" box="[518,592,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="119" pageNumber="120" unit="m" value="305.0">
<elevation id="0043C635FF81D768FDE17C162156F8D3" box="[518,592,1821,1848]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="119" pageNumber="120" unit="m" value="305.0">305 m</elevation>
</quantity>
, 
<date id="FFD007C6FF81D768FDBA7C1621E3F8DC" box="[605,741,1821,1847]" pageId="119" pageNumber="120" value="1997-09-09">
<collectingDate id="EF94FE2EFF81D768FDBA7C1621E3F8DC" box="[605,741,1821,1847]" pageId="119" pageNumber="120" value="1997-09-09">9 Sep. 1997</collectingDate>
</date>
(without posterior end; caruncle black; eyes reddish; median antenna half as long as caruncle; cirriform branchiae with tips blackish; bipinnate branchiae with brownish lateral branches; middorsal bands almost faded; body 
<quantity id="4C968CE3FF81D768FD417C6E21F3F86B" box="[678,757,1893,1920]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="119" pageNumber="120" unit="mm" value="64.0">64 mm</quantity>
long, 
<quantity id="4C968CE3FF81D768FCDB7C6E208DF86B" box="[828,907,1893,1920]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="119" pageNumber="120" unit="mm" value="16.0">16 mm</quantity>
wide, 27 chaetigers).
</materialsCitation>
</paragraph>
<paragraph id="8BD12106FF81D767FF207C82221CFEAA" blockId="119.[151,1437,1533,2028]" lastBlockId="120.[151,1437,150,321]" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
<emphasis id="B91AFD14FF81D768FF207C8222B2F848" bold="true" box="[199,436,1929,1955]" pageId="119" pageNumber="120">Additional material</emphasis>
. 
<materialsCitation id="3B062B5BFF81D767FE247C82209AFF5A" ID-GBIF-Occurrence="4029882450" collectingDate="1997-08-25" collectingDateMax="1997-09-02" collectingDateMin="1997-08-25" collectionCode="MNHN, ECOSUR" collectorName="RV Alis &amp; Sta. CP" country="French Polynesia" elevation="260" lastPageId="120" lastPageNumber="121" latitude="-9.786667" location="Alis" longLatPrecision="129" longitude="-139.63667" municipality="Two" pageId="119" pageNumber="120" specimenCount="1" stateProvince="Marquesas Islands">
<emphasis id="B91AFD14FF81D768FE247C82208FF848" bold="true" box="[451,905,1929,1955]" pageId="119" pageNumber="120">
<collectingCountry id="F3796196FF81D768FE247C822195F848" box="[451,659,1929,1955]" name="French Polynesia" pageId="119" pageNumber="120">French Polynesia</collectingCountry>
, 
<collectingRegion id="49AAEFE4FF81D768FD467C822082F848" box="[673,900,1929,1955]" country="French Polynesia" name="Iles Marquises" pageId="119" pageNumber="120">Marquesas Islands</collectingRegion>
.
</emphasis>
<specimenCount id="9D68EA8FFF81D768FC767C812739F848" box="[913,1087,1929,1956]" pageId="119" pageNumber="120">Two specimens</specimenCount>
(1: 
<collectionCode id="ED7FB9C3FF81D768FB897C8127C1F84F" box="[1134,1223,1930,1956]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="119" pageNumber="120">MNHN</collectionCode>
; 1: 
<collectionCode id="ED7FB9C3FF81D768FB167C822662F848" box="[1265,1380,1929,1955]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="119" pageNumber="120" type="Museum">ECOSUR</collectionCode>
), RV 
<location id="8EB177DDFF81D768FF707CA623CEF82C" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF81D768FF707CA623CEF82C" box="[151,200,1965,1991]" country="French Polynesia" latitude="-9.786667" longLatPrecision="129" longitude="-139.63667" municipality="Two" name="Alis" pageId="119" pageNumber="120" stateProvince="Marquesas Islands">Alis</location>
, Sta. CP1176 (
<geoCoordinate id="EE5A47C1FF81D768FE967CA622EBF82C" box="[369,493,1965,1991]" degrees="08" direction="south" minutes="44.8" orientation="latitude" pageId="119" pageNumber="120" precision="92" value="-8.746667">08°44.8´S</geoCoordinate>
, 
<geoCoordinate id="EE5A47C1FF81D768FE1C7CA62196F82C" box="[507,656,1965,1992]" degrees="140" direction="west" minutes="14.5" orientation="longitude" pageId="119" pageNumber="120" precision="92" value="-140.24167">140°14.5´W</geoCoordinate>
), off 
<location id="8EB177DDFF81D768FD2A7CA6209FF82C" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF81D768FD2A7CA6209FF82C" box="[717,921,1965,1991]" country="French Polynesia" latitude="-9.786667" longLatPrecision="129" longitude="-139.63667" municipality="Two" name="Nuku Hiva Island" pageId="119" pageNumber="120" stateProvince="Marquesas Islands">Nuku Hiva Island</location>
, bean trawl, 
<quantity id="4C968CE3FF81D768FBCC7CA62773F823" box="[1067,1141,1965,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="119" pageNumber="120" unit="m" value="260.0">
<elevation id="0043C635FF81D768FBCC7CA62773F823" box="[1067,1141,1965,1992]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="119" pageNumber="120" unit="m" value="260.0">260 m</elevation>
</quantity>
, 
<date id="FFD007C6FF81D768FB657CA6261AF82C" box="[1154,1308,1965,1992]" pageId="119" pageNumber="120" value="1997-08-25">
<collectingDate id="EF94FE2EFF81D768FB657CA6261AF82C" box="[1154,1308,1965,1992]" pageId="119" pageNumber="120" value="1997-08-25">25 Aug. 1997</collectingDate>
</date>
(complete; dorsum pale; median antenna 1/3 as long as caruncle, caruncle blackish; eyes blackish; cirriform branchiae with tips blackish; body 
<quantity id="4C968CE3FF8ED767FEA37B9C22BBFF5A" box="[324,445,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" metricValueMax="7.8" metricValueMin="6.4" pageId="120" pageNumber="121" unit="mm" value="71.0" valueMax="78.0" valueMin="64.0">64–78 mm</quantity>
long, 
<quantity id="4C968CE3FF8ED767FDE27B9D2179FF5A" box="[517,639,150,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="120" pageNumber="121" unit="mm" value="16.5" valueMax="18.0" valueMin="15.0">15–18 mm</quantity>
wide, 33–34 chaetigers).
</materialsCitation>
<materialsCitation id="3B062B5BFF8ED767FC447B9C2210FEAA" ID-GBIF-Occurrence="4029882342" collectingDate="1997-09-02" collectingDateMax="1997-09-02" collectingDateMin="1997-08-25" collectionCode="MNHN, ECOSUR" collectorName="RV Alis &amp; Sta. CP" country="French Polynesia" elevation="575" latitude="-9.786667" location="Alis" longLatPrecision="129" longitude="-139.63667" municipality="Two" pageId="120" pageNumber="121" specimenCount="1" stateProvince="Iles Marquises">
<specimenCount id="9D68EA8FFF8ED767FC447B9C2757FF5A" box="[931,1105,151,177]" pageId="120" pageNumber="121">Five specimens</specimenCount>
(3 
<collectionCode id="ED7FB9C3FF8ED767FB9F7B9C27D6FF5A" box="[1144,1232,151,177]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="120" pageNumber="121">MNHN</collectionCode>
, 2 
<collectionCode id="ED7FB9C3FF8ED767FB157B9C2663FF5A" box="[1266,1381,151,177]" country="Mexico" httpUri="http://grbio.org/cool/dtqq-2t5p" name="El Colegio de la Frontera Sur (Mexico)" pageId="120" pageNumber="121" type="Museum">ECOSUR</collectionCode>
), RV 
<location id="8EB177DDFF8ED767FF707BB023CEFF3E" LSID="urn:lsid:plazi:treatment:03C79010FF81D764FF707E7C268CFB2E:8EB177DDFF8ED767FF707BB023CEFF3E" box="[151,200,187,213]" country="French Polynesia" latitude="-9.786667" longLatPrecision="129" longitude="-139.63667" municipality="Two" name="Alis" pageId="120" pageNumber="121" stateProvince="Iles Marquises">Alis</location>
, Sta. CP1251 (
<geoCoordinate id="EE5A47C1FF8ED767FE8A7BB022E1FF3E" box="[365,487,187,213]" degrees="09" direction="south" minutes="47.2" orientation="latitude" pageId="120" pageNumber="121" precision="92" value="-9.786667">09°47.2´S</geoCoordinate>
, 
<geoCoordinate id="EE5A47C1FF8ED767FE147BB02180FF3E" box="[499,646,187,213]" degrees="139" direction="west" minutes="38.2" orientation="longitude" pageId="120" pageNumber="121" precision="92" value="-139.63667">139°38.2´W</geoCoordinate>
), Dumont d’Urville Deep, 
<quantity id="4C968CE3FF8ED767FC567BB12734FF3E" box="[945,1074,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="120" pageNumber="121" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">
<elevation id="0043C635FF8ED767FC567BB12734FF3E" box="[945,1074,186,213]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="120" pageNumber="121" unit="m" value="575.0" valueMax="650.0" valueMin="500.0">500–650 m</elevation>
</quantity>
, beam trawl, 
<date id="FFD007C6FF8ED767FB207BB0264CFF3E" box="[1223,1354,187,213]" pageId="120" pageNumber="121" value="1997-09-02">
<collectingDate id="EF94FE2EFF8ED767FB207BB0264CFF3E" box="[1223,1354,187,213]" pageId="120" pageNumber="121" value="1997-09-02">2 Sep. 1997</collectingDate>
</date>
(colorless, many chaetae broken; complete, anterior fragment 
<quantity id="4C968CE3FF8ED767FCEC7BD4205CFF12" box="[779,858,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="120" pageNumber="121" unit="mm" value="30.0">30 mm</quantity>
long, 
<quantity id="4C968CE3FF8ED767FC477BD420E8FF12" box="[928,1006,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="120" pageNumber="121" unit="mm" value="11.0">11 mm</quantity>
wide, 15 chaetigers; smaller complete specimens with anal cirri 6–7× longer than wide, 5× in larger specimens; 
<quantity id="4C968CE3FF8ED767FC3C7A082753FEF6" box="[987,1109,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="6.7" metricValueMin="2.2" pageId="120" pageNumber="121" unit="mm" value="44.5" valueMax="67.0" valueMin="22.0">22–67 mm</quantity>
long, 
<quantity id="4C968CE3FF8ED767FB7A7A09260FFEF6" box="[1181,1289,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="120" pageNumber="121" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
wide, 29–32 chaetigers)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF11718EFF8ED767FF7F788823C0FC08" ID-DOI="http://doi.org/10.5281/zenodo.7621998" ID-Zenodo-Dep="7621998" httpUri="https://zenodo.org/record/7621998/files/figure.png" pageId="120" pageNumber="121" startId="120.[152,255,899,923]" targetBox="[179,1408,348,874]" targetPageId="120">
<paragraph id="8BD12106FF8ED767FF7F788823C0FC08" blockId="120.[151,1437,899,995]" pageId="120" pageNumber="121">
<emphasis id="B91AFD14FF8ED767FF7F78882227FC70" bold="true" box="[152,289,899,923]" pageId="120" pageNumber="121">FIGURE 58</emphasis>
. 
<taxonomicName id="4C6E5A85FF8ED767FED7788822DDFC70" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[304,475,899,923]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="120" pageNumber="121" phylum="Annelida" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B91AFD14FF8ED767FED7788822DDFC70" box="[304,475,899,923]" italics="true" pageId="120" pageNumber="121">Chloeia zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFF8ED767FE0378882119FC70" box="[484,543,899,923]" pageId="120" pageNumber="121" rank="species">
<emphasis id="B91AFD14FF8ED767FE037888211EFC70" bold="true" box="[484,536,899,923]" pageId="120" pageNumber="121">sp. n</emphasis>
.
</taxonomicNameLabel>
, holotype (MNHN IA-TYPE 2056). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 14-16, dorsal view, branchiae removed from chaetiger 14. Scale bars: A, 2.7 mm; B, 1.6 mm; C, 0.9 mm.
</paragraph>
</caption>
<subSubSection id="C374728DFF8ED767FF207F1A272CFB98" pageId="120" pageNumber="121" type="diagnosis">
<paragraph id="8BD12106FF8ED767FF207F1A272CFB98" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">
<emphasis id="B91AFD14FF8ED767FF207F1A223CFBC0" bold="true" box="[199,314,1041,1067]" pageId="120" pageNumber="121">Diagnosis</emphasis>
. 
<taxonomicName id="4C6E5A85FF8ED767FEAE7F1A22A4FBC0" authorityName="Savigny" authorityYear="1818" box="[329,418,1041,1067]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="120" pageNumber="121" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FF8ED767FEAE7F1A22A4FBC0" box="[329,418,1041,1067]" italics="true" pageId="120" pageNumber="121">Chloeia</emphasis>
</taxonomicName>
with bipinnate branchiae from chaetiger 4, decreasing in size posteriorly, each with 11–12 lateral branches in median segments; middorsal band wide, tapered posteriorly; caruncle blackish, with about 24 folds; notochaetae harpoon shaped with spurs; neurochaetae spurred and furcates.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFF8ED766FF207F762015F894" lastPageId="121" lastPageNumber="122" pageId="120" pageNumber="121" type="description">
<paragraph id="8BD12106FF8ED767FF207F762216FB34" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">
<emphasis id="B91AFD14FF8ED767FF207F762257FB7C" bold="true" box="[199,337,1149,1175]" pageId="120" pageNumber="121">Description</emphasis>
. 
<typeStatus id="54D59FA4FF8ED767FEB97F7622C0FB7C" box="[350,454,1149,1175]" pageId="120" pageNumber="121" type="holotype">Holotype</typeStatus>
(MNHN IA-TYPE 2056) an anterior fragment, 
<quantity id="4C968CE3FF8ED767FC3D7F76272EFB73" box="[986,1064,1149,1176]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="120" pageNumber="121" unit="mm" value="40.0">40 mm</quantity>
long, 
<quantity id="4C968CE3FF8ED767FB8B7F7627BFFB73" box="[1132,1209,1149,1176]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="120" pageNumber="121" unit="mm" value="17.0">17 mm</quantity>
wide, 21 chaetigers. Complete 
<typeStatus id="54D59FA4FF8ED767FEEB7FA9226AFB57" box="[268,364,1186,1212]" pageId="120" pageNumber="121" type="paratype">paratype</typeStatus>
(MNHN IA-TYPE 2057) complete, bent laterally; body tapered, 
<quantity id="4C968CE3FF8ED767FBB07FAA27A1FB57" box="[1111,1191,1185,1212]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="120" pageNumber="121" unit="mm" value="53.0">53 mm</quantity>
long, 
<quantity id="4C968CE3FF8ED767FB177FAA2635FB57" box="[1264,1331,1185,1212]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="120" pageNumber="121" unit="mm" value="9.0">9 mm</quantity>
wide, 32 chaetigers.
</paragraph>
<paragraph id="8BD12106FF8ED767FF207FE22608FA78" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">
<typeStatus id="54D59FA4FF8ED767FF207FE22229FAE8" box="[199,303,1257,1283]" pageId="120" pageNumber="121" type="holotype">Holotype</typeStatus>
with an ill-defined T-shaped brownish dorsal band, with paler core in a few anterior chaetigers (
<figureCitation id="13553D83FF8ED767FA887FE223CDFAC3" captionStart="FIGURE 58" captionStartId="120.[152,255,899,923]" captionTargetBox="[179,1408,348,874]" captionTargetId="figure-101@120.[177,1411,345,876]" captionTargetPageId="120" captionText="FIGURE 58. Chloeia zibrowii sp. n., holotype (MNHN IA-TYPE 2056). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 14-16, dorsal view, branchiae removed from chaetiger 14. Scale bars: A, 2.7 mm; B, 1.6 mm; C, 0.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7621998" httpUri="https://zenodo.org/record/7621998/files/figure.png" pageId="120" pageNumber="121">Fig. 58A</figureCitation>
), solid along following ones; lateral bands poorly defined, paler; caruncle black, middorsal ridge with 26 vertical folds (
<figureCitation id="13553D83FF8ED767FEE17E3A226BFAA7" box="[262,365,1329,1356]" captionStart="FIGURE 58" captionStartId="120.[152,255,899,923]" captionTargetBox="[179,1408,348,874]" captionTargetId="figure-101@120.[177,1411,345,876]" captionTargetPageId="120" captionText="FIGURE 58. Chloeia zibrowii sp. n., holotype (MNHN IA-TYPE 2056). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 14-16, dorsal view, branchiae removed from chaetiger 14. Scale bars: A, 2.7 mm; B, 1.6 mm; C, 0.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7621998" httpUri="https://zenodo.org/record/7621998/files/figure.png" pageId="120" pageNumber="121">Fig. 58B</figureCitation>
); eyes reddish, in a slightly darker area, anterior eyes 2–3× larger than posterior ones; cirriform branchiae with black tips; bipinnate branchiae brownish with darker lateral branches, each with 6–7 filaments per side, with darker tips (
<figureCitation id="13553D83FF8ED767FE767E7222FEFA7F" box="[401,504,1401,1428]" captionStart="FIGURE 58" captionStartId="120.[152,255,899,923]" captionTargetBox="[179,1408,348,874]" captionTargetId="figure-101@120.[177,1411,345,876]" captionTargetPageId="120" captionText="FIGURE 58. Chloeia zibrowii sp. n., holotype (MNHN IA-TYPE 2056). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetigers 14-16, dorsal view, branchiae removed from chaetiger 14. Scale bars: A, 2.7 mm; B, 1.6 mm; C, 0.9 mm." figureDoi="http://doi.org/10.5281/zenodo.7621998" httpUri="https://zenodo.org/record/7621998/files/figure.png" pageId="120" pageNumber="121">Fig. 58C</figureCitation>
); many chaetae broken. Venter brownish, with a pale midventral band.
</paragraph>
<paragraph id="8BD12106FF8ED767FF207E9627AEFA37" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">
Complete 
<typeStatus id="54D59FA4FF8ED767FEDC7E95229DFA53" box="[315,411,1438,1464]" pageId="120" pageNumber="121" type="paratype">paratype</typeStatus>
(MNHN IA-TYPE 2057) pale. Dorsum without pigmentation (
<figureCitation id="13553D83FF8ED767FB847E9627CAFA53" box="[1123,1228,1437,1464]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="120" pageNumber="121">Fig. 59A</figureCitation>
). Dorsal cirri pale. Interramal areas pale. Bipinnate branchiae pale. Venter paler, with a whitish band (
<figureCitation id="13553D83FF8ED767FBD37ECA279DFA37" box="[1076,1179,1473,1500]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="120" pageNumber="121">Fig. 59C</figureCitation>
).
</paragraph>
<paragraph id="8BD12106FF8ED767FF207EEE2223F964" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">
Prostomium anteriorly entire, anterior prostomial area pale (
<figureCitation id="13553D83FF8ED767FC8D7EEE20D7F9EB" box="[874,977,1509,1536]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="120" pageNumber="121">Fig. 59B</figureCitation>
). Eyes reddish, anterior eyes 2–3× larger than posterior ones. Median antenna inserted at anterior caruncular margin, pale, almost half as long as caruncle (4/5 as long in one additional specimen), apparently 2× longer than laterals (without tips). Lateral antennae bases close to each other. Palps broken, slightly narrower than lateral antennae. Mouth ventral on chaetiger 2. Pharynx not exposed.
</paragraph>
<paragraph id="8BD12106FF8ED767FF207D92206AF93C" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">Caruncle blackish, trilobed, slightly curved, reaching chaetiger 4; middorsal ridge black, with 24 vertical folds. Lateral lobes narrow, barely visible, with about 30 vertical folds.</paragraph>
<paragraph id="8BD12106FF8ED767FF207DEA229DF8A8" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">Bipinnate branchiae from chaetiger 4, continued throughout body, parallel along most body segments, progressively larger to chaetigers 13–16, diminishing in size in posterior chaetigers. In median segments each branchia with 11–12 lateral branches.</paragraph>
<paragraph id="8BD12106FF8ED767FF207C46219BF813" blockId="120.[151,1437,1041,2040]" pageId="120" pageNumber="121">Parapodia biramous, notopodia with cirriform branchiae along chaetigers 1–3, progressively smaller, 4/5–1/3 as long as dorsal cirri; one complete with black tip. Dorsal cirri about 2× longer than branchiae in median and posterior chaetigers. Second ventral cirri with cirrophores 2–3×longer and wider than adjacent ones, and cirrostyles 2× longer than adjacent ones, directed dorsally. Other ventral cirri directed ventrolaterally, as long as two segments in median region, up to three segments in posterior ones.</paragraph>
<caption id="DF11718EFF8FD766FF7F7E082676FA6D" ID-DOI="http://doi.org/10.5281/zenodo.7622002" ID-Zenodo-Dep="7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122" startId="121.[152,255,1283,1307]" targetBox="[154,1434,185,1256]" targetPageId="121">
<paragraph id="8BD12106FF8FD766FF7F7E082676FA6D" blockId="121.[151,1437,1283,1415]" pageId="121" pageNumber="122">
<emphasis id="B91AFD14FF8FD766FF7F7E082218FAF0" bold="true" box="[152,286,1283,1307]" pageId="121" pageNumber="122">FIGURE 59</emphasis>
. 
<taxonomicName id="4C6E5A85FF8FD766FECC7E0822D5FAF0" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[299,467,1283,1307]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="121" pageNumber="122" phylum="Annelida" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B91AFD14FF8FD766FECC7E0822D5FAF0" box="[299,467,1283,1307]" italics="true" pageId="121" pageNumber="122">Chloeia zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFF8FD766FE3E7E082117FAF1" box="[473,529,1283,1307]" pageId="121" pageNumber="122" rank="species">
<emphasis id="B91AFD14FF8FD766FE3E7E08210DFAF0" bold="true" box="[473,523,1283,1307]" pageId="121" pageNumber="122">sp. n</emphasis>
.
</taxonomicNameLabel>
, paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm.
</paragraph>
</caption>
<paragraph id="8BD12106FF8FD766FF207EBE2213F96F" blockId="121.[151,1437,1461,2028]" pageId="121" pageNumber="122">
Chaetae whitish to golden, neurochaetae with a pale green hue (observed in 
<specimenCount id="9D68EA8FFF8FD766FBCD7EBD27C6FA24" box="[1066,1216,1461,1488]" pageId="121" pageNumber="122" type="generic" typeStatus="paratype">one paratype</specimenCount>
). Complete chaetae with distal fragile hoods. Notochaetae in anterior chaetigers spurred and furcates (
<figureCitation id="13553D83FF8FD766FBAD7ED227B3FA1F" box="[1098,1205,1497,1524]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122">Fig. 59D</figureCitation>
), major tines 5–12× longer than minor ones. Median chaetigers with spurred, harpoon-chaetae (
<figureCitation id="13553D83FF8FD766FBE57EF6276FF9F3" box="[1026,1129,1533,1560]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122">Fig. 59F</figureCitation>
), denticulate tines 9—12× longer than smooth ones. Neurochaetae spurred and furcates with tiny minor tines, anterior chaetigers with major tines 5—8× longer than minor ones (
<figureCitation id="13553D83FF8FD766FDDE7D4E2199F98B" box="[569,671,1605,1632]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122">Fig. 59E</figureCitation>
), median chaetigers with major tines 4–10× longer than minor ones (
<figureCitation id="13553D83FF8FD766FF787D62220EF96F" box="[159,264,1641,1668]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122">Fig. 59G</figureCitation>
).
</paragraph>
<paragraph id="8BD12106FF8FD766FF207D8623D3F920" blockId="121.[151,1437,1461,2028]" pageId="121" pageNumber="122">
Posterior region tapered (
<figureCitation id="13553D83FF8FD766FE047D86214DF943" box="[483,587,1677,1704]" captionStart="FIGURE 59" captionStartId="121.[152,255,1283,1307]" captionTargetBox="[154,1434,185,1256]" captionTargetId="figure-18@121.[151,1436,181,1259]" captionTargetPageId="121" captionText="FIGURE 59. Chloeia zibrowii sp. n., paratype (MNHN IA-TYPE 2057). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior region, ventral view. D. Chaetiger 4, notochaetal tips. E. Same, neurochaetae (insets: neurochaetal tips). F. Chaetiger 15, notochaetal tips (insets: basal denticulate sections). G. Same, neurochaetae (inset: neurochaetal tip). H. Posterior region, ventral view. Scale bars: A, 1.4 mm; B, 0.6 mm; C, 1.2 mm; D, 110 μm; E, 80 μm; F, 60 μm; G, 140 μm; H, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7622002" httpUri="https://zenodo.org/record/7622002/files/figure.png" pageId="121" pageNumber="122">Fig. 59H</figureCitation>
); pygidium with anus terminal; anal cirri whitish, digitate, 5–6× longer than wide.
</paragraph>
<paragraph id="8BD12106FF8FD766FF207DDE2120F904" blockId="121.[151,1437,1461,2028]" box="[199,550,1749,1776]" pageId="121" pageNumber="122">
<emphasis id="B91AFD14FF8FD766FF207DDE22A4F904" bold="true" box="[199,418,1749,1775]" pageId="121" pageNumber="122">Live pigmentation</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BD12106FF8FD766FF207DF22015F894" blockId="121.[151,1437,1461,2028]" pageId="121" pageNumber="122">
<emphasis id="B91AFD14FF8FD766FF207DF2223EF8F8" bold="true" box="[199,312,1785,1811]" pageId="121" pageNumber="122">Variation</emphasis>
. Incomplete 
<typeStatus id="54D59FA4FF8FD766FE2E7DF12132F8FF" box="[457,564,1786,1812]" pageId="121" pageNumber="122" type="paratype">paratypes</typeStatus>
(MNHN IA-TYPE 2057) with body wall darker than complete 
<typeStatus id="54D59FA4FF8FD766FB1F7DF1265AF8FF" box="[1272,1372,1786,1812]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
. Eyes reddish, anterior eyes 2–3× larger than posterior ones. Caruncles darker than in complete 
<typeStatus id="54D59FA4FF8FD766FB657C1527E3F8D3" box="[1154,1253,1822,1848]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
, one incomplete 
<typeStatus id="54D59FA4FF8FD766FF707C4923F1F8B7" box="[151,247,1858,1884]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
with cirriform branchiae complete, all with black tips. Bipinnate branchiae with lateral branches darker than in complete 
<typeStatus id="54D59FA4FF8FD766FEBE7C6D22BAF86B" box="[345,444,1894,1920]" pageId="121" pageNumber="122" type="paratype">paratype</typeStatus>
, resembling those in 
<typeStatus id="54D59FA4FF8FD766FD4D7C6E2016F894" box="[682,784,1893,1919]" pageId="121" pageNumber="122" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFF8FD765FF207C8227ACFF3E" lastPageId="122" lastPageNumber="123" pageId="121" pageNumber="122" type="etymology">
<paragraph id="8BD12106FF8FD765FF207C8227ACFF3E" blockId="121.[151,1437,1461,2028]" lastBlockId="122.[151,1437,151,573]" lastPageId="122" lastPageNumber="123" pageId="121" pageNumber="122">
<emphasis id="B91AFD14FF8FD766FF207C82224EF848" bold="true" box="[199,328,1929,1955]" pageId="121" pageNumber="122">Etymology</emphasis>
. The specific epithet is after Dr Helmut Zibrowius, a specialist of serpulid polychaetes and deep-sea ecologist that worked in the Station Marine d’Endoume, Marseille, 
<collectingCountry id="F3796196FF8FD766FC6C7CA520DCF823" box="[907,986,1966,1992]" name="France" pageId="121" pageNumber="122">France</collectingCountry>
, in recognition of his many publications on polychaetes, and especially because he participated in the Musorstom 9 cruise to the Marquesas Archipelago (
<bibRefCitation id="EFFF5CF7FF8CD765FF787B9C22D8FF5A" author="Richer de Forges, B. &amp; Poupin, J. &amp; Laboute, P." box="[159,478,151,177]" pageId="122" pageNumber="123" pagination="9 - 29" refId="ref112376" refString="Richer de Forges, B., Poupin, J. &amp; Laboute, P. (1999) La campagne Musorstom 9 dans l'archipel des iles Marquises (Polynesie francaise). Compte rendu et liste des stations. Resultats des Campagnes Musorstom 20. Memoires du Museum National d'Histoire Naturelle, 180, 9 - 29." type="journal article" year="1999">
Richer de Forges 
<emphasis id="B91AFD14FF8CD765FE827B9C229AFF5A" box="[357,412,151,177]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
1999
</bibRefCitation>
), which included the collection of the 
<typeStatus id="54D59FA4FF8CD765FC607B9C20B1FF5A" box="[903,951,151,177]" pageId="122" pageNumber="123">type</typeStatus>
material for this species. The derived name is a noun in the genitive case (
<bibRefCitation id="EFFF5CF7FF8CD765FE0E7BB02168FF3E" author="ICZN &amp; International Commission of Zoological Nomenclature" box="[489,622,187,213]" pageId="122" pageNumber="123" refId="ref108505" refString="ICZN (International Commission of Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, The Natural History Museum, London. Available from: https: // code. iczn. org (accessed 22 June 2022)" type="url" year="1999">ICZN 1999</bibRefCitation>
, Art. 31.1.1) after regarding the last name as Latin.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFF8CD765FF207BD42680FDF2" pageId="122" pageNumber="123" type="discussion">
<paragraph id="8BD12106FF8CD765FF207BD42680FDF2" blockId="122.[151,1437,151,573]" pageId="122" pageNumber="123">
<emphasis id="B91AFD14FF8CD765FF207BD42232FF12" bold="true" box="[199,308,223,249]" pageId="122" pageNumber="123">Remarks</emphasis>
. 
<taxonomicName id="4C6E5A85FF8CD765FEA57BD422F8FF12" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[322,510,223,249]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="zibrowii" status="sp. nov.">
<emphasis id="B91AFD14FF8CD765FEA57BD422F8FF12" box="[322,510,223,249]" italics="true" pageId="122" pageNumber="123">Chloeia zibrowii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFF8CD765FDE27BD42147FF12" box="[517,577,223,249]" pageId="122" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
is described with specimens from several localities in the 
<collectingCountry id="F3796196FF8CD765FB357BD4269FFF12" box="[1234,1433,223,249]" name="French Polynesia" pageId="122" pageNumber="123">French Polynesia</collectingCountry>
; it belongs in the group viridis because it has a complex pigmentation pattern, and its bipinnate branchiae start in chaetiger 4, becoming progressively smaller posteriorly. It resembles 
<taxonomicName id="4C6E5A85FF8CD765FC497A2C275EFEAA" authority="Salazar-Vallejo, 2023" authorityName="Salazar-Vallejo" authorityYear="2023" box="[942,1112,295,321]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="hutchingsae" status="sp. nov.">
<emphasis id="B91AFD14FF8CD765FC497A2C275EFEAA" box="[942,1112,295,321]" italics="true" pageId="122" pageNumber="123">C. hutchingsae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229406FFF8CD765FB867A2C2798FEAA" box="[1121,1182,295,321]" pageId="122" pageNumber="123" rank="species">sp. n.</taxonomicNameLabel>
described above from The 
<collectingCountry id="F3796196FF8CD765FF2E7A402241FE8E" box="[201,327,331,357]" name="Philippines" pageId="122" pageNumber="123">Philippines</collectingCountry>
to 
<collectingCountry id="F3796196FF8CD765FE8E7A4022D6FE8E" box="[361,464,331,357]" name="Australia" pageId="122" pageNumber="123">Australia</collectingCountry>
and 
<collectingCountry id="F3796196FF8CD765FDE37A402165FE8E" box="[516,611,331,357]" name="Vanuatu" pageId="122" pageNumber="123">Vanuatu</collectingCountry>
, because both species have a well-defined middorsal band, without lateral bands. Additional similarities are found in the size of eyes and 
<typeStatus id="54D59FA4FF8CD765FCBF7A64208EFE62" box="[856,904,367,393]" pageId="122" pageNumber="123">type</typeStatus>
of chaetae. These species differ, however, after their caruncles (pigmentation and number of vertical folds), and in the relative development of bipinnate branchiae, including the number of lateral branches in median segments. In 
<taxonomicName id="4C6E5A85FF8CD765FC867ABC20DDFE3A" authorityName="Salazar-Vallejo" authorityYear="2023" box="[865,987,439,465]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="zibrowii">
<emphasis id="B91AFD14FF8CD765FC867ABC20DDFE3A" box="[865,987,439,465]" italics="true" pageId="122" pageNumber="123">C. zibrowii</emphasis>
</taxonomicName>
the caruncle is blackish, with 24 vertical folds, and the branchiae are massive, with 6–7 lateral branches. On the contrary, 
<taxonomicName id="4C6E5A85FF8CD765FBF17AD027BBFE1E" authorityName="Salazar-Vallejo" authorityYear="2023" box="[1046,1213,475,501]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="hutchingsae">
<emphasis id="B91AFD14FF8CD765FBF17AD027BBFE1E" box="[1046,1213,475,501]" italics="true" pageId="122" pageNumber="123">C. hutchingsae</emphasis>
</taxonomicName>
has a pale caruncle, with blackish median ridge, and 34 vertical folds, and its branchiae are delicate, each has 11–12 lateral branches.
</paragraph>
</subSubSection>
<subSubSection id="C374728DFF8CD764FF207929268CFB2E" lastPageId="123" lastPageNumber="124" pageId="122" pageNumber="123" type="distribution">
<paragraph id="8BD12106FF8CD765FF20792920F1FDD6" blockId="122.[151,1437,151,573]" box="[199,1015,546,573]" pageId="122" pageNumber="123">
<emphasis id="B91AFD14FF8CD765FF207929225FFDD7" bold="true" box="[199,345,546,572]" pageId="122" pageNumber="123">Distribution</emphasis>
. 
<collectingCountry id="F3796196FF8CD765FE807928212AFDD6" box="[359,556,547,573]" name="French Polynesia" pageId="122" pageNumber="123">French Polynesia</collectingCountry>
, in sediments at 
<quantity id="4C968CE3FF8CD765FD0179292063FDD6" box="[742,869,546,573]" metricMagnitude="2" metricUnit="m" metricValue="4.55" metricValueMax="6.5" metricValueMin="2.6" pageId="122" pageNumber="123" unit="m" value="455.0" valueMax="650.0" valueMin="260.0">260–650 m</quantity>
water depth.
</paragraph>
<paragraph id="8BD12106FF8CD765FF7079852285FD43" blockId="122.[151,387,654,680]" box="[151,387,654,680]" pageId="122" pageNumber="123">
<emphasis id="B91AFD14FF8CD765FF7079852285FD43" bold="true" box="[151,387,654,680]" pageId="122" pageNumber="123">General discussion</emphasis>
</paragraph>
<paragraph id="8BD12106FF8CD765FF7079DD22AFFD1A" blockId="122.[151,425,726,753]" box="[151,425,726,753]" pageId="122" pageNumber="123">
<emphasis id="B91AFD14FF8CD765FF7079DD22A4FD1A" bold="true" box="[151,418,726,753]" pageId="122" pageNumber="123">Pigmentation patterns</emphasis>
.
</paragraph>
<paragraph id="8BD12106FF8CD765FF7078142026FBFA" blockId="122.[151,1437,799,1833]" pageId="122" pageNumber="123">
In two species, specimens can have a pigmentation pattern mostly reddish, against being dark purple or blackish. This was shown for 
<taxonomicName id="4C6E5A85FF8CD765FE9A784822E0FCB6" authorityName="Schmarda" authorityYear="1861" box="[381,486,835,861]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="viridis">
<emphasis id="B91AFD14FF8CD765FE9A784822E0FCB6" box="[381,486,835,861]" italics="true" pageId="122" pageNumber="123">C. viridis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3796196FF8CD765FDCC7848218EFCB6" box="[555,648,835,861]" name="Jamaica" pageId="122" pageNumber="123">Jamaica</collectingCountry>
, and for 
<taxonomicName id="4C6E5A85FF8CD765FD0B7848204EFCB6" authorityName="sensu Potts" authorityYear="1909" box="[748,840,835,861]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FF8CD765FD0B7848204EFCB6" box="[748,840,835,861]" italics="true" pageId="122" pageNumber="123">C. fusca</emphasis>
</taxonomicName>
from 
<collectingCountry id="F3796196FF8CD765FC69784820F9FCB6" box="[910,1023,835,861]" name="Indonesia" pageId="122" pageNumber="123">Indonesia</collectingCountry>
; the former species was treated elsewhere (
<bibRefCitation id="EFFF5CF7FF8CD765FF0B786C21A6FC6A" author="Yanez-Rivera, B. &amp; Salazar-Vallejo, S. I." box="[236,672,871,897]" pageId="122" pageNumber="123" pagination="503 - 537" refId="ref114234" refString="Yanez-Rivera, B. &amp; Salazar-Vallejo, S. I. (2022) Revision of Chloeia Savigny in Lamarck, 1818 from tropical American seas (Annelida. Amphinomidae). Zootaxa, 5128 (4), 503 - 537. https: // doi. org / 10.11646 / zootaxa. 5128.4.3" type="journal article" year="2022">Yáñez-Rivera &amp; Salazar-Vallejo 2022</bibRefCitation>
), whereas the latter needs some explanations. Thus, 
<taxonomicName id="4C6E5A85FF8CD765FB1F786C2650FC6A" authorityName="sensu Potts" authorityYear="1909" box="[1272,1366,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="fusca">
<emphasis id="B91AFD14FF8CD765FB1F786C2650FC6A" box="[1272,1366,871,897]" italics="true" pageId="122" pageNumber="123">C. fusca</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3796196FF8CD765FEA0788022B1FC4E" box="[327,439,907,933]" name="Indonesia" pageId="122" pageNumber="123">Indonesia</collectingCountry>
, and has been confirmed for 
<collectingCountry id="F3796196FF8CD765FD1D788020D3FC4E" box="[762,981,907,933]" name="Papua New Guinea" pageId="122" pageNumber="123">Papua New Guinea</collectingCountry>
, and distant archipelagos like the 
<collectingCountry id="F3796196FF8CD765FAA978802201FC22" name="French Polynesia" pageId="122" pageNumber="123">French Polynesia</collectingCountry>
, the Loyalty Islands, and Hawai’i. Because the only difference was in the color of the pigmentation pattern (reddish against dark purple or blackish), the different populations were regarded as conspecific, pending a further analysis including molecular or genetic indicators.
</paragraph>
<paragraph id="8BD12106FF8CD765FF207F102154FA92" blockId="122.[151,1437,799,1833]" pageId="122" pageNumber="123">
A different issue regards to the loss of additional banding, such that only the middorsal spots remain. In at least three colorful species, 
<emphasis id="B91AFD14FF8CD765FE707F342183FBB2" box="[407,645,1087,1113]" italics="true" pageId="122" pageNumber="123">
<taxonomicName id="4C6E5A85FF8CD765FE707F342126FBB2" box="[407,544,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="amphora">C. amphora</taxonomicName>
, 
<taxonomicName id="4C6E5A85FF8CD765FDCB7F342183FBB2" authorityName=": de Quatrefages" authorityYear="1866" box="[556,645,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="flava">C. flava</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C6E5A85FF8CD765FD5A7F342041FBB2" authorityName="Baird" authorityYear="1868" box="[701,839,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="pulchella">
<emphasis id="B91AFD14FF8CD765FD5A7F342041FBB2" box="[701,839,1087,1113]" italics="true" pageId="122" pageNumber="123">C. pulchella</emphasis>
</taxonomicName>
, the additional dorsal bands can fade off completely. This has been regarded as an effect of the specimens being left in ethanol for a long time, some pigment damage due to light, or both. However, there are some videos available in internet showing living specimens with a pale dorsum and only the middorsal spots present (
<bibRefCitation id="EFFF5CF7FF8CD765FDA57FA0200FFB2E" author="Iromongara" box="[578,777,1195,1221]" pageId="122" pageNumber="123" refId="ref108752" refString="Iromongara. (2009) Umikemushi. YouTube. Available from: https: // www. youtube. com / watch? v = PGLLgLJJG-Y (accessed 14 July 2022)" type="url" year="2009">Iromongara 2009</bibRefCitation>
, 
<bibRefCitation id="EFFF5CF7FF8CD765FCF37FA02093FB2E" author="Senja, K." box="[788,917,1195,1221]" pageId="122" pageNumber="123" refId="ref112948" refString="Senja, K. (2021) Ulat bulu laut beracun (Chloeia flava). YouTube. Available from: https: // www. youtube. com / watch? v = m 3 I 3 Ci WB 1 Wo &amp; t = 44 s (accessed 14 July 2022)" type="url" year="2021">Senja 2021</bibRefCitation>
, 0:00 
<taxonomicName id="4C6E5A85FF8CD765FC3E7FA02734FB2E" authorityName=": de Quatrefages" authorityYear="1866" box="[985,1074,1195,1221]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B91AFD14FF8CD765FC3E7FA02734FB2E" box="[985,1074,1195,1221]" italics="true" pageId="122" pageNumber="123">C. flava</emphasis>
</taxonomicName>
, 0:28 + 1:27 
<taxonomicName id="4C6E5A85FF8CD765FB2D7FA02655FB2E" authorityName="Baird" authorityYear="1868" box="[1226,1363,1195,1221]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="pulchella">
<emphasis id="B91AFD14FF8CD765FB2D7FA02655FB2E" box="[1226,1363,1195,1221]" italics="true" pageId="122" pageNumber="123">C. pulchella</emphasis>
</taxonomicName>
). This would imply either a large, likely clinal variation, or the presence of more than a single biological species. The distribution data are as follows: 
<taxonomicName id="4C6E5A85FF8CD765FE047FF8216FFAE6" box="[483,617,1267,1293]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="amphora">
<emphasis id="B91AFD14FF8CD765FE047FF8216FFAE6" box="[483,617,1267,1293]" italics="true" pageId="122" pageNumber="123">C. amphora</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3796196FF8CD765FCB17FF820C5FAE6" box="[854,963,1267,1293]" name="Indonesia" pageId="122" pageNumber="123">Indonesia</collectingCountry>
and ranges to the 
<collectingCountry id="F3796196FF8CD765FB737FF82658FAE6" box="[1172,1374,1267,1293]" name="Philippines" pageId="122" pageNumber="123">Philippine Islands</collectingCountry>
from the intertidal to 
<quantity id="4C968CE3FF8CD765FEAC7E1D2283FADA" box="[331,389,1302,1329]" metricMagnitude="1" metricUnit="m" metricValue="4.5" pageId="122" pageNumber="123" unit="m" value="45.0">45 m</quantity>
depth; 
<taxonomicName id="4C6E5A85FF8CD765FE3D7E1C2132FADA" authorityName=": de Quatrefages" authorityYear="1866" box="[474,564,1303,1329]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="flava">
<emphasis id="B91AFD14FF8CD765FE3D7E1C2132FADA" box="[474,564,1303,1329]" italics="true" pageId="122" pageNumber="123">C. flava</emphasis>
</taxonomicName>
was described from 
<collectingCountry id="F3796196FF8CD765FCC77E1C205AFADA" box="[800,860,1303,1329]" name="India" pageId="122" pageNumber="123">India</collectingCountry>
, and extends to 
<collectingRegion id="49AAEFE4FF8CD765FBF47E1C279CFADA" box="[1043,1178,1303,1329]" country="Australia" name="Queensland" pageId="122" pageNumber="123">Queensland</collectingRegion>
, 
<collectingCountry id="F3796196FF8CD765FB437E1C2608FADA" box="[1188,1294,1303,1329]" name="Australia" pageId="122" pageNumber="123">Australia</collectingCountry>
, in intertidal to 
<quantity id="4C968CE3FF8CD765FF537E3123EBFABE" box="[180,237,1338,1365]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="122" pageNumber="123" unit="m" value="50.0">50 m</quantity>
depth; and 
<taxonomicName id="4C6E5A85FF8CD765FE967E3022FFFABE" authorityName="Baird" authorityYear="1868" box="[369,505,1339,1365]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="species" species="pulchella">
<emphasis id="B91AFD14FF8CD765FE967E3022FFFABE" box="[369,505,1339,1365]" italics="true" pageId="122" pageNumber="123">C. pulchella</emphasis>
</taxonomicName>
was described from northeastern 
<collectingCountry id="F3796196FF8CD765FC947E3020DCFABE" box="[883,986,1339,1365]" name="Australia" pageId="122" pageNumber="123">Australia</collectingCountry>
and has been found from 
<collectingCountry id="F3796196FF8CD765FB187E30263FFABE" box="[1279,1337,1339,1365]" name="India" pageId="122" pageNumber="123">India</collectingCountry>
to 
<collectingCountry id="F3796196FF8CD765FABB7E30269AFABE" box="[1372,1436,1339,1365]" name="Japan" pageId="122" pageNumber="123">Japan</collectingCountry>
from the intertidal to 
<quantity id="4C968CE3FF8CD765FE607E5422C6FA92" box="[391,448,1375,1401]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="122" pageNumber="123" unit="m" value="22.0">22 m</quantity>
water depth.
</paragraph>
<paragraph id="8BD12106FF8CD765FF207E882058F972" blockId="122.[151,1437,799,1833]" pageId="122" pageNumber="123">
Assessing the geographic or depth variation in pigmentation patterns is an interesting research focus for colleagues interested in genetics of pigmentation (
<bibRefCitation id="EFFF5CF7FF8CD765FD437EAC2049FA2A" author="Alvarado, S. G." box="[676,847,1447,1473]" pageId="122" pageNumber="123" pagination="1531 - 1543" refId="ref101933" refString="Alvarado, S. G. (2020) Molecular plasticity in animal pigmentation: emerging processes undelying color changes. Integrative and Comparative Biology, 60 (6), 1531 - 1543. https: // doi. org / 10.1093 / icb / icaa 142" type="journal article" year="2020">Alvarado 2020</bibRefCitation>
, 
<bibRefCitation id="EFFF5CF7FF8CD765FCBD7EAC2748FA2A" author="Orteu, A. &amp; Jiggins, C. D." box="[858,1102,1447,1473]" pageId="122" pageNumber="123" pagination="461 - 475" refId="ref111547" refString="Orteu, A. &amp; Jiggins, C. D. (2020) The genomics of colorations provides insights into adaptive evolution. Nature Reviews, Genetics, 21, 461 - 475. https: // doi. org / 10.1038 / s 41576 - 020 - 0234 - z" type="journal article" year="2020">Orteu &amp; Jiggins 2020</bibRefCitation>
, vonHoldt 
<emphasis id="B91AFD14FF8CD765FB2E7EAC2607FA2A" box="[1225,1281,1447,1473]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2021), a field that has made spectacular progresses in other colorful marine organisms, especially regarding fishes (
<bibRefCitation id="EFFF5CF7FF8CD765FAC67EC023F1F9E2" author="Behrens, K. A. &amp; Girasek, Q. L. &amp; Sickler, A. &amp; Hyde, J. &amp; Buonaccorsi, V. P." pageId="122" pageNumber="123" pagination="4259 - 4275" refId="ref103359" refString="Behrens, K. A., Girasek, Q. L., Sickler, A., Hyde, J., &amp; Buonaccorsi, V. P. (2021) Regions of genetic divergence in depth-sepa- rated Sebastes rockfish species pairs: Depth as a potential driver of speciation. Molecular Ecology, 30 (17), 4259 - 4275. https: // doi. org / 10.1111 / mec. 16046" type="journal article" year="2021">
Behrens 
<emphasis id="B91AFD14FF8CD765FA6F7EC023B2F9E2" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2021
</bibRefCitation>
, 
<bibRefCitation id="EFFF5CF7FF8CD765FEE47EE42290F9E2" author="Parichi, D. M." box="[259,406,1519,1545]" pageId="122" pageNumber="123" pagination="88 - 96" refId="ref111704" refString="Parichi, D. M. (2021) Evolution of pigment cells and patterns: recent insights from teleost fishes. Current Opinion in Genetics and Development, 69, 88 - 96. https: // doi. org / 10.1016 / j. gde. 2021.02.006" type="journal article" year="2021">Parichi 2021</bibRefCitation>
), or benthic invertebrates, including some molluscs (Korshunoa 
<emphasis id="B91AFD14FF8CD765FB967EE427ACF9E2" box="[1137,1194,1519,1545]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2020, 
<bibRefCitation id="EFFF5CF7FF8CD765FB1E7EE423D5F9C6" author="Neuhaus, J. &amp; Rauch, C. &amp; Bakken, T. &amp; Picton, B. &amp; Pola, M. &amp; Malaquias, M. A. E." pageId="122" pageNumber="123" pagination="1 - 31" refId="ref111315" refString="Neuhaus, J., Rauch, C., Bakken, T., Picton, B., Pola, M. &amp; Malaquias, M. A. E. (2021) The genus Jorunna (Nudibranchia: Discodorididae) in Europe: a new species and a possible cause of incipient speciation. Journal of Molluscan Studies, 87 (eyab 028), 1 - 31. https: // doi. org / 10.1093 / mollus / eyab 028" type="journal article" year="2021">
Neuhaus 
<emphasis id="B91AFD14FF8CD765FA847EE4269BF9E2" box="[1379,1437,1519,1545]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2021
</bibRefCitation>
). On the basis of morphological features, the different populations were regarded as conspecific, pending a further analysis including molecular or genetic indicators. The reason for this perspective is that most marine species are limited to certain geographic areas, and this applies for both, shallow water (Costello 
<emphasis id="B91AFD14FF8CD765FB5E7D5027F7F99E" box="[1209,1265,1627,1653]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2017, 
<bibRefCitation id="EFFF5CF7FF8CD765FAD87D502276F972" author="Costello, M. J. &amp; Chaudhary, C." pageId="122" pageNumber="123" pagination="527" refId="ref104422" refString="Costello, M. J. &amp; Chaudhary, C. (2017) Marine biodiversity, biogeography, deep-sea gradients, and conservation. Current Biology, 27, R 511 - R 527. https: // doi. org / 10.1016 / j. cub. 2017.04.060" type="journal article" year="2017">Costello &amp; Chaudhary 2017</bibRefCitation>
) and deep-sea species (
<bibRefCitation id="EFFF5CF7FF8CD765FD917D742055F972" author="Watling, L. &amp; Guinotte, J. &amp; Clark, M. R. &amp; Smith, C. R." box="[630,851,1663,1689]" pageId="122" pageNumber="123" pagination="91 - 112" refId="ref114013" refString="Watling, L., Guinotte, J., Clark, M. R. &amp; Smith, C. R. (2013) A proposed biogeography of the deep ocean floor. Progress in Oceanography, 111, 91 - 112. https: // doi. org / 10.1016 / j. pocean. 2012.11.003" type="journal article" year="2013">
Watling 
<emphasis id="B91AFD14FF8CD765FD307D742009F972" box="[727,783,1663,1689]" italics="true" pageId="122" pageNumber="123">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD12106FF8CD765FF207DA92777F8C2" blockId="122.[151,1437,799,1833]" pageId="122" pageNumber="123">
<emphasis id="B91AFD14FF8CD765FF207DA9227CF956" bold="true" box="[199,378,1698,1725]" pageId="122" pageNumber="123">Species groups</emphasis>
. The species of 
<taxonomicName id="4C6E5A85FF8CD765FDDE7DA82194F956" authorityName="Savigny" authorityYear="1818" box="[569,658,1699,1725]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="122" pageNumber="123" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FF8CD765FDDE7DA82194F956" box="[569,658,1699,1725]" italics="true" pageId="122" pageNumber="123">Chloeia</emphasis>
</taxonomicName>
can be grouped after several features such as the 
<typeStatus id="54D59FA4FF8CD765FB327DA82603F956" box="[1237,1285,1699,1725]" pageId="122" pageNumber="123">type</typeStatus>
of branchiae (pinnate or bipinnate), the start of branchiae (chaetigers 3–5), the dorsal pigmentation pattern (single middorsal bands, single middorsal spots, double longitudinal bands, complex patterns, or none), and some chaetal features. As indicated above, after the combination of these features, the groups are listed in Table 1.
</paragraph>
<paragraph id="8BD12106FF8CD765FF707C5C2148F89A" blockId="122.[151,754,1879,2013]" box="[151,590,1879,1905]" pageId="122" pageNumber="123">1. Branchiae pinnate (a single species).</paragraph>
<paragraph id="8BD12106FF8CD765FF707C702289F87E" blockId="122.[151,754,1879,2013]" box="[151,399,1915,1941]" pageId="122" pageNumber="123">2. Branchie bipinnate.</paragraph>
<paragraph id="8BD12106FF8CD765FF207C9421F4F852" blockId="122.[151,754,1879,2013]" box="[199,754,1951,1977]" pageId="122" pageNumber="123">2.1 Branchiae from chaetiger 3 (a single species).</paragraph>
<paragraph id="8BD12106FF8CD765FF207CC8212CF836" blockId="122.[151,754,1879,2013]" box="[199,554,1987,2013]" pageId="122" pageNumber="123">2.2 Branchiae from chaetiger 4.</paragraph>
<paragraph id="8BD12106FF8DD764FF107B9C2019FF5A" blockId="123.[199,868,151,537]" box="[247,799,151,177]" pageId="123" pageNumber="124">2.2.1 Branchiae progressively smaller posteriorly</paragraph>
<paragraph id="8BD12106FF8DD764FEB07BB02159FF3E" blockId="123.[199,868,151,537]" box="[343,607,187,213]" pageId="123" pageNumber="124">2.2.1.1 Middorsal spots</paragraph>
<paragraph id="8BD12106FF8DD764FEB07BD42160FF12" blockId="123.[199,868,151,537]" box="[343,614,223,249]" pageId="123" pageNumber="124">2.2.1.2 Middorsal bands</paragraph>
<paragraph id="8BD12106FF8DD764FEB07A082174FEF6" blockId="123.[199,868,151,537]" box="[343,626,259,285]" pageId="123" pageNumber="124">2.2.1.3 Complex patterns</paragraph>
<paragraph id="8BD12106FF8DD764FEB07A2C205FFEAA" blockId="123.[199,868,151,537]" box="[343,857,295,321]" pageId="123" pageNumber="124">2.2.1.4 Dorsum without pigmentation pattern.</paragraph>
<paragraph id="8BD12106FF8DD764FF107A402062FE8E" blockId="123.[199,868,151,537]" box="[247,868,331,357]" pageId="123" pageNumber="124">2.2.2 Branchiae abruptly smaller after a few chaetigers.</paragraph>
<paragraph id="8BD12106FF8DD764FF207A64212CFE62" blockId="123.[199,868,151,537]" box="[199,554,366,393]" pageId="123" pageNumber="124">2.3 Branchiae from chaetiger 5.</paragraph>
<paragraph id="8BD12106FF8DD764FF107A9822F7FE46" blockId="123.[199,868,151,537]" box="[247,497,403,429]" pageId="123" pageNumber="124">2.3.1 Middorsal bands</paragraph>
<paragraph id="8BD12106FF8DD764FEB07ABC21B4FE3A" blockId="123.[199,868,151,537]" box="[343,690,439,465]" pageId="123" pageNumber="124">2.3.1.1 Single middorsal bands</paragraph>
<paragraph id="8BD12106FF8DD764FEB07AD021BBFE1E" blockId="123.[199,868,151,537]" box="[343,701,475,501]" pageId="123" pageNumber="124">2.3.1.2 Double middorsal bands</paragraph>
<paragraph id="8BD12106FF8DD764FF107AF421E2FDF2" blockId="123.[199,868,151,537]" box="[247,740,511,537]" pageId="123" pageNumber="124">2.3.2 Dorsum without pigmentation pattern.</paragraph>
<paragraph id="8BD12106FF8DD764FF20794C22AAFD42" blockId="123.[151,1437,583,1221]" pageId="123" pageNumber="124">It must be kept in mind that for those species having pale pigmentation, or even for those having a colorful body, once the pigments are faded off, they might be included in the colorless groups, but fresh specimens would clarify their correct assignation.</paragraph>
<paragraph id="8BD12106FF8DD764FF2079B9212EFC22" blockId="123.[151,1437,583,1221]" pageId="123" pageNumber="124">
<emphasis id="B91AFD14FF8DD764FF2079B922BAFD27" bold="true" box="[199,444,690,716]" pageId="123" pageNumber="124">Taxonomic decisions</emphasis>
. After the previous information, it must be emphasized that the taxonomic criteria ruling the assignation of specimens to known or undescribed species has depended on a certain combination of morphological features. When a different set of different data is available, as molecular indicators, a different set of conclusions can be reached upon. Of course, some of the conclusions of the future studies might disagree with the current situation, but that is just the way that Science proceeds. Large scale sampling for molecular purposes, especially for those species having a large or very large distribution such as 
<emphasis id="B91AFD14FF8DD764FCB1786C279DFC6B" box="[854,1179,871,897]" italics="true" pageId="123" pageNumber="124">
<taxonomicName id="4C6E5A85FF8DD764FCB1786C20B5FC6A" authorityName=": de Quatrefages" authorityYear="1866" box="[854,947,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="species" species="flava">C. flava</taxonomicName>
, 
<taxonomicName id="4C6E5A85FF8DD764FC58786C2718FC6A" authorityName="sensu Potts" authorityYear="1909" box="[959,1054,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="species" species="fusca">C. fusca</taxonomicName>
, 
<taxonomicName id="4C6E5A85FF8DD764FBCD786C279DFC6B" authorityName="de Quatrefages" authorityYear="1866" box="[1066,1179,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="species" species="incerta">C. incerta</taxonomicName>
</emphasis>
, or 
<taxonomicName id="4C6E5A85FF8DD764FB2E786C2655FC6A" authorityName="Baird" authorityYear="1868" box="[1225,1363,871,897]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="species" species="pulchella">
<emphasis id="B91AFD14FF8DD764FB2E786C2655FC6A" box="[1225,1363,871,897]" italics="true" pageId="123" pageNumber="124">C. pulchella</emphasis>
</taxonomicName>
might be especially revealing. Of course, this is not the final study on 
<taxonomicName id="4C6E5A85FF8DD764FC8A788020C0FC4E" authorityName="Savigny" authorityYear="1818" box="[877,966,907,933]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FF8DD764FC8A788020C0FC4E" box="[877,966,907,933]" italics="true" pageId="123" pageNumber="124">Chloeia</emphasis>
</taxonomicName>
, and the conclusions presented above are just working taxonomic hypothesis.
</paragraph>
<paragraph id="8BD12106FF8DD764FF2078D9268CFB2E" blockId="123.[151,1437,583,1221]" pageId="123" pageNumber="124">
<emphasis id="B91AFD14FF8DD764FF2078D92239FC07" bold="true" box="[199,319,978,1004]" pageId="123" pageNumber="124">Symbiosis</emphasis>
. The only species that has been recorded as associated with a sessile invertebrate is 
<taxonomicName id="4C6E5A85FF8DD764FB1878D8265BFC07" authorityName=": Fauvel" authorityYear="1932" box="[1279,1373,979,1005]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="species" species="rosea">
<emphasis id="B91AFD14FF8DD764FB1878D8265BFC07" box="[1279,1373,979,1005]" italics="true" pageId="123" pageNumber="124">C. rosea</emphasis>
</taxonomicName>
from the Indian Ocean, being found on a subtidal soft coral. The species is also unique by having pinnate, not bippinate, branchiae, but it has not been collected after its original description. It would be useful for future studies that collectors make notes about where the 
<taxonomicName id="4C6E5A85FF8DD764FDEB7F342163FBB2" authorityName="Savigny" authorityYear="1818" box="[524,613,1087,1113]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FF8DD764FDEB7F342163FBB2" box="[524,613,1087,1113]" italics="true" pageId="123" pageNumber="124">Chloeia</emphasis>
</taxonomicName>
specimens were found; most have been regarded as free living, and they are depicted in a few photos or short videos in internet, as indicated above for the different species, but some species might have a closer affinity to other invertebrates. This is another interesting perspective for continuing collecting 
<taxonomicName id="4C6E5A85FF8DD764FF707FA023F6FB2E" authorityName="Savigny" authorityYear="1818" box="[151,240,1195,1221]" class="Polychaeta" family="Amphinomidae" genus="Chloeia" kingdom="Animalia" order="Amphinomida" pageId="123" pageNumber="124" phylum="Annelida" rank="genus">
<emphasis id="B91AFD14FF8DD764FF707FA023F6FB2E" box="[151,240,1195,1221]" italics="true" pageId="123" pageNumber="124">Chloeia</emphasis>
</taxonomicName>
specimens, and for making better observations of what specimens do and what could be their symbionts.
</paragraph>
</subSubSection>
</treatment>
</document>