257 lines
36 KiB
XML
257 lines
36 KiB
XML
<document id="2F93DBC655C5E33A6C2957E05474E7A8" ID-CLB-Dataset="46709" ID-DOI="10.5281/zenodo.277803" ID-GBIF-Dataset="c7a7345c-c0ba-4a28-bf43-2f2f58cbc35e" ID-ISSN="1175-5326" ID-Zenodo-Dep="277803" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460366400850" checkinUser="plazi" docAuthor="Moradmand, Majid & Jäger, Peter" docDate="2011" docId="03C787F54E34F031EFA0F8832E66FC01" docLanguage="en" docName="zt02910p062.pdf" docOrigin="Zootaxa 2910" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Spariolenus Simon 1880" docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="FFFEFF8D4E35F033EF37FF8F2F35FFED" masterDocTitle="A review on the huntsman spider genus Spariolenus Simon, 1880 (Araneae: Sparassidae: Heteropodinae) in Iran with description of four new species" masterLastPageNumber="62" masterPageNumber="46" pageNumber="47" updateTime="1698295240505" updateUser="ExternalLinkService">
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<mods:title id="65D01117C96B7FDC6FCAB14EC3D85E0F">A review on the huntsman spider genus Spariolenus Simon, 1880 (Araneae: Sparassidae: Heteropodinae) in Iran with description of four new species</mods:title>
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<mods:namePart id="84DCE094FB6452FC28C265D27A0B293B">Moradmand, Majid</mods:namePart>
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<mods:namePart id="713114BB1C48FD751736A743FF2224FC">Jäger, Peter</mods:namePart>
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<mods:title id="E812F74B0CCA7EB753EF5A6E5877388B">Zootaxa</mods:title>
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<mods:part id="EF665B7B77C47C507EE224BA837FA591">
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<mods:date id="6D8BE34D9BC36F6281CC48265B450969">2011</mods:date>
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<treatment id="03C787F54E34F031EFA0F8832E66FC01" ID-DOI="http://doi.org/10.5281/zenodo.6184001" ID-GBIF-Taxon="119410871" ID-Zenodo-Dep="6184001" LSID="urn:lsid:plazi:treatment:03C787F54E34F031EFA0F8832E66FC01" httpUri="http://treatment.plazi.org/id/03C787F54E34F031EFA0F8832E66FC01" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47">
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<subSubSection id="C37465684E34F032EFA0F8832D2AF8CB" box="[151,543,1804,1830]" pageId="1" pageNumber="47" type="nomenclature">
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<paragraph id="8BD136E34E34F032EFA0F8832D2AF8CB" blockId="1.[151,543,1804,1830]" box="[151,543,1804,1830]" pageId="1" pageNumber="47">
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<heading id="D099818F4E34F032EFA0F8832D2AF8CB" bold="true" box="[151,543,1804,1830]" fontSize="11" level="1" pageId="1" pageNumber="47" reason="1">
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<emphasis id="B91AEAF14E34F032EFA0F8832D2AF8CB" bold="true" box="[151,543,1804,1830]" pageId="1" pageNumber="47">
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Genus
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<taxonomicName id="4C6E4D604E34F032EFD9F8832D2AF8CB" ID-CoL="7JYT" ID-ENA="1304664" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[238,543,1804,1830]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF14E34F032EFD9F8832E4AF8CB" bold="true" box="[238,383,1804,1830]" italics="true" pageId="1" pageNumber="47">Spariolenus</emphasis>
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<bibRefCitation id="EFFF4B124E34F032EEB1F8832D2AF8CB" author="Simon" box="[390,543,1804,1830]" pageId="1" pageNumber="47" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465684E34F032EFA0F8DC2EADF86A" pageId="1" pageNumber="47" type="materials_examined">
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<paragraph id="8BD136E34E34F032EFA0F8DC2EADF86A" blockId="1.[151,1436,1874,1989]" pageId="1" pageNumber="47">
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<emphasis id="B91AEAF14E34F032EFA0F8DC2E1DF884" bold="true" box="[151,296,1874,1897]" pageId="1" pageNumber="47">
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<typeStatus id="54D588414E34F032EFA0F8DC2FF9F885" box="[151,204,1875,1896]" pageId="1" pageNumber="47">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C6E4D604E34F032EE06F8DC2D4DF885" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[305,632,1874,1897]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="tigris">
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<emphasis id="B91AEAF14E34F032EE06F8DC2EDDF884" box="[305,488,1875,1897]" italics="true" pageId="1" pageNumber="47">Spariolenus tigris</emphasis>
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<bibRefCitation id="EFFF4B124E34F032EEC6F8DD2D4DF885" author="Simon" box="[497,632,1874,1896]" pageId="1" pageNumber="47" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
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</taxonomicName>
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, by original designation from Calcutta (
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<collectingCountry id="F37976734E34F032EB2BF8DD2B60F885" box="[1052,1109,1874,1896]" name="India" pageId="1" pageNumber="47">India</collectingCountry>
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), redescribed and illustrated by
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||
<bibRefCitation id="EFFF4B124E34F032EFF0F8FD2E75F86A" author="Jager" box="[199,320,1906,1927]" pageId="1" pageNumber="47" refString="Jager, P. (2001) Diversitat der Riesenkrabbenspinnen im Himalaya. Uber eine Radiation zweier Gattungen in den Schneetropen. (Araneae: Sparassidae: Heteropodinae). Courier Forschungsinstitut Senckenberg, 232, 1 - 136." type="journal article" year="2001">Jäger (2001</bibRefCitation>
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,
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<bibRefCitation id="EFFF4B124E34F032EE7DF8FE2EB9F86A" author="Jager" box="[330,396,1905,1927]" pageId="1" pageNumber="47" refString="Jager, P. (2006 a) A new Spariolenus species from caves in Oman - the first representative of the Heteropodinae in the Arabian peninsula (Araneae: Sparassidae). Bulletin of the British Arachnological Society, 13, 309 - 313." type="journal article" year="2006" yearSuffix="a">2006a</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37465684E34F031EFA0F81F2E41FE41" lastPageId="2" lastPageNumber="48" pageId="1" pageNumber="47" type="reference_group">
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<paragraph id="8BD136E34E34F032EFA0F81F2A15F828" blockId="1.[151,1436,1874,1989]" pageId="1" pageNumber="47">
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<treatmentCitationGroup id="AB7E11CD4E34F032EFA0F81F2A15F828" pageId="1" pageNumber="47">
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<taxonomicName id="4C6E4D604E34F032EFA0F81F2EFEF84B" ID-CoL="7JYT" authority="Simon 1880: 281" authorityName="Simon" authorityPageNumber="281" authorityYear="1880" box="[151,459,1936,1958]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF14E34F032EFA0F81F2E25F84B" box="[151,272,1936,1958]" italics="true" pageId="1" pageNumber="47">Spariolenus</emphasis>
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<treatmentCitation id="0ACF10F24E34F032EE2FF81F2EFEF84B" author="Simon" box="[280,459,1936,1958]" page="281" pageId="1" pageNumber="47" year="1880">
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<bibRefCitation id="EFFF4B124E34F032EE2FF81F2EACF84B" author="Simon" box="[280,409,1936,1958]" pageId="1" pageNumber="47" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon 1880</bibRefCitation>
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: 281
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</treatmentCitation>
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||
</taxonomicName>
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–282.
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<treatmentCitation id="0ACF10F24E34F032ED3AF81F2DF7F84B" author="Simon" box="[525,706,1936,1958]" page="56" pageId="1" pageNumber="47" year="1897">
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<bibRefCitation id="EFFF4B124E34F032ED3AF81F2DAFF84B" author="Simon" box="[525,666,1936,1958]" pageId="1" pageNumber="47" refString="Simon, E. (1897 a) Histoire naturelle des araignees. 2. Paris, Roret, 1 - 192." type="journal article" year="1897" yearSuffix="a">Simon 1897a</bibRefCitation>
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: 56
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||
</treatmentCitation>
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;
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<treatmentCitation id="0ACF10F24E34F032EDF8F81F2CA7F84B" author="Simon" box="[719,914,1936,1958]" page="253" pageId="1" pageNumber="47" year="1897">
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<bibRefCitation id="EFFF4B124E34F032EDF8F81F2C68F84B" author="Simon" box="[719,861,1936,1958]" pageId="1" pageNumber="47" refString="Simon, E. (1897 b) Etudes arachnologiques. 27 e Memoire. XLII. Descriptions d'especes nouvelles de l'ordre des Araneae. Annales de la Scociete Entomologique de France, 65, 465 - 510." type="journal article" year="1897" yearSuffix="b">Simon 1897b</bibRefCitation>
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: 253
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</treatmentCitation>
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;
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<treatmentCitation id="0ACF10F24E34F032ECA9F81F2B56F84B" author="Gravely" box="[926,1123,1936,1958]" page="257" pageId="1" pageNumber="47" year="1931">
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||
<bibRefCitation id="EFFF4B124E34F032ECA9F81F2B1BF84B" author="Gravely" box="[926,1070,1936,1958]" pageId="1" pageNumber="47" refString="Gravely, F. H. (1931) Some Indian spiders of the families Ctenidae, Sparassidae, Selenopidae and Clubionidae. Records of the Indian Museum, Calcutta, 33, 211 - 282." type="journal article" year="1931">Gravely 1931</bibRefCitation>
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||
: 257
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||
</treatmentCitation>
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, figs.13C–D;
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<treatmentCitation id="0ACF10F24E34F032EBC2F81F2E20F828" author="Sethi" page="7" pageId="1" pageNumber="47" year="1988">
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<bibRefCitation id="EFFF4B124E34F032EBC2F81F2FC8F829" author="Sethi" pageId="1" pageNumber="47" refString="Sethi, V. D. & Tikader, B. K. (1988) Studies on some giant crab spiders of the family Heteropodidae from India. Records of the zoological survey of India, Miscellanous Publications, Occasional Paper, 93, 1 - 94." type="journal article" year="1988">Sethi & Tikader 1988</bibRefCitation>
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||
: 7
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</treatmentCitation>
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, figs. 19–24;
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<treatmentCitation id="0ACF10F24E34F032EEA8F83F2D1CF829" author="Jager" box="[415,553,1967,1989]" page="9" pageId="1" pageNumber="47" year="2001">
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||
<bibRefCitation id="EFFF4B124E34F032EEA8F83F2D24F829" author="Jager" box="[415,529,1967,1989]" pageId="1" pageNumber="47" refString="Jager, P. (2001) Diversitat der Riesenkrabbenspinnen im Himalaya. Uber eine Radiation zweier Gattungen in den Schneetropen. (Araneae: Sparassidae: Heteropodinae). Courier Forschungsinstitut Senckenberg, 232, 1 - 136." type="journal article" year="2001">Jäger 2001</bibRefCitation>
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: 9
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</treatmentCitation>
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, figs. 2e, m–p, 14d, 15a–b;
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<treatmentCitation id="0ACF10F24E34F032EC74F83F2CECF828" author="Jager" box="[835,985,1967,1989]" page="58" pageId="1" pageNumber="47" year="2002">
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||
<bibRefCitation id="EFFF4B124E34F032EC74F83F2C80F829" author="Jager" box="[835,949,1967,1989]" pageId="1" pageNumber="47" refString="Jager, P. (2002) Heteropodinae: transfers and synonymies (Arachnida: Araneae: Sparassidae). Acta Arachnologica, 51, 33 - 61." type="journal article" year="2002">Jäger 2002</bibRefCitation>
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||
: 58
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||
</treatmentCitation>
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||
–59;
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<treatmentCitation id="0ACF10F24E34F032EB3CF83F2B8EF829" author="Jager" box="[1035,1211,1967,1989]" page="310" pageId="1" pageNumber="47" year="2006">
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||
<bibRefCitation id="EFFF4B124E34F032EB3CF83F2BBDF828" author="Jager" box="[1035,1160,1967,1989]" pageId="1" pageNumber="47" refString="Jager, P. (2006 a) A new Spariolenus species from caves in Oman - the first representative of the Heteropodinae in the Arabian peninsula (Araneae: Sparassidae). Bulletin of the British Arachnological Society, 13, 309 - 313." type="journal article" year="2006" yearSuffix="a">Jäger 2006a</bibRefCitation>
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||
: 310
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</treatmentCitation>
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, f. 12–15.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8BD136E34E37F031EFA0FF172E41FE41" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="48">
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<emphasis id="B91AEAF14E37F031EFA0FF172EB0FF5D" bold="true" box="[151,389,151,176]" pageId="2" pageNumber="48">Extended diagnosis.</emphasis>
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<taxonomicName id="4C6E4D604E37F031EEBBFF172D27FF5D" ID-CoL="7JYT" box="[396,530,152,176]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF14E37F031EEBBFF172D27FF5D" box="[396,530,152,176]" italics="true" pageId="2" pageNumber="48">Spariolenus</emphasis>
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</taxonomicName>
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spp. can be distinguished from all other Heteropodinae by the number of ventral tibial spines:
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<date id="FFD010234E37F031EE1BFF322E4FFF39" box="[300,378,188,212]" pageId="2" pageNumber="48" value="2010-01" valueMax="2010-02-31" valueMin="2010-01-10">I–II 10</date>
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, III 8 and IV 6 (usually
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<date id="FFD010234E37F031EDB1FF322DE5FF39" box="[646,720,188,212]" pageId="2" pageNumber="48">
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I–IV
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||
<quantity id="4C969B064E37F031EDF5FF332DE5FF39" box="[706,720,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="2" pageNumber="48" unit="in" value="6.0">6</quantity>
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</date>
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in other Heteropodinae, exception
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<taxonomicName id="4C6E4D604E37F031EB6EFF332EA2FF1A" ID-CoL="3LC8L" authority="Barrion & Litsinger, 1995" authorityName="Barrion & Litsinger" authorityYear="1995" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="cyperusiria">
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<emphasis id="B91AEAF14E37F031EB6EFF332A51FF39" box="[1113,1380,188,212]" italics="true" pageId="2" pageNumber="48">Heteropoda cyperusiria</emphasis>
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||
Barrion & Litsinger, 1995
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</taxonomicName>
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I–II 12, III 8,
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<date id="FFD010234E37F031ED76FF6F2D4DFF1A" box="[577,632,223,247]" pageId="2" pageNumber="48">IV 6</date>
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). Moreover, they can be recognized by a combination of characters of copulatory organs: male palp with blunt and short RTA in ventral view, conductor membranous (sheath-like in
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<taxonomicName id="4C6E4D604E37F031EA5DFE8A2FC6FEAD" ID-CoL="4WNK" class="Arachnida" family="Sparassidae" genus="Heteropoda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B91AEAF14E37F031EA5DFE8A2FC6FEAD" italics="true" pageId="2" pageNumber="48">Heteropoda</emphasis>
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||
</taxonomicName>
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||
spp.). Female epigynes are characterized by two spirally coiled copulatory openings (similar to those of
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||
<taxonomicName id="4C6E4D604E37F031EFA0FEC32EF2FE89" ID-CoL="5LMB" authority="Jager, 2006" authorityName="Jager" authorityYear="2006" box="[151,455,332,357]" class="Arachnida" family="Sparassidae" genus="Martensopoda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B91AEAF14E37F031EFA0FEC32E0FFE89" box="[151,314,332,356]" italics="true" pageId="2" pageNumber="48">Martensopoda</emphasis>
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||
Jäger, 2006
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||
</taxonomicName>
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||
) but lacking posterior pit. The openings form two large semicircular rims (
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<figureCitation id="13552A664E37F031EA15FEC32A5FFE89" box="[1314,1386,332,356]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,264,1705,1728]" captionTargetBox="[229,1392,221,1634]" captionTargetId="figure@3.[151,1436,195,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Spariolenus aratta spec. nov., holotype female from Iran, Kerman Province, Jebal Barez. 1 Epigyne, ventral; 2 Vulva, lateral; 3 Vulva, dorsal; 4 Schematic course of internal duct system, dorsal; 5 Left female palpal claw, prolateral; 6 Eye arrangement, dorsal; 7 Left chelicera, ventral. Abbreviations: CO—Copulatory opening; CR—Central rim; EF—Epigynal field; EP—Epigynal pit; FD—Fertilisation duct; HGD—Hump-like glandular part of internal duct system; LID—Lumen of internal duct system; MS—Median Slit; SS—Slit sensillum." httpUri="https://zenodo.org/record/277804/files/figure.png" pageId="2" pageNumber="48">Figs 1</figureCitation>
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,
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<figureCitation id="13552A664E37F031EA4DFEC32AA2FE89" box="[1402,1431,332,356]" captionStart="FIGURES 11 – 19" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[219,1364,186,1865]" captionTargetId="figure@6.[219,1365,181,1882]" captionTargetPageId="6" captionText="FIGURES 11 – 19. Spariolenus iranomaximus spec. nov., paratype female. 11 Epigyne, ventral; 12 Vulva, dorsal; 13 Vulva, lateral; 14 Schematic course of internal duct system, dorsal (from left duct of the vulva figure 12, not scaled); 15 Eye arrangement, dorsal; 16 Left chelicera, ventral; 17 Left female palpal claw, retrolateral; 18 – 19 Variation of vulvae, dorsal. AB— Anterior band of epigynal field." httpUri="https://zenodo.org/record/277806/files/figure.png" pageId="2" pageNumber="48">11</figureCitation>
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,
|
||
<figureCitation id="13552A664E37F031EFA0FEE02F83FE6A" box="[151,182,367,391]" captionStart="FIGURES 20 – 26" captionStartId="8.[151,264,1938,1961]" captionTargetBox="[192,1336,215,1915]" captionTargetId="figure@8.[191,1406,163,1921]" captionTargetPageId="8" captionText="FIGURES 20 – 26. Spariolenus manesht spec. nov., holotype female from Iran, Ilam Province, Karezan, Manesht. 20 Epigyne, ventral; 21 Vulva, lateral; 22 Vulva, dorsal; 23 Schematic course of internal duct system, dorsal; 24 Left female palpal claw, retrolateral; 25 Eye arrangement, dorsal; 26 Left chelicera, ventral." httpUri="https://zenodo.org/record/277807/files/figure.png" pageId="2" pageNumber="48">20</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EFF4FEE02FD7FE6A" box="[195,226,367,391]" captionStart="FIGURES 31 – 38" captionStartId="11.[151,264,1950,1973]" captionTargetBox="[213,1374,208,1916]" captionTargetId="figure@11.[212,1380,193,1946]" captionTargetPageId="11" captionText="FIGURES 31 – 38. Spariolenus zagros spec. nov., paratype female. 31 Epigyne, ventral; 32 Vulva, dorsal; 33 Vulva, lateral; 34 Schematic course of internal duct system, dorsal (not scaled from left duct of vulva figure 33); 35 Left female palpal claw, retrolateral; 36 Eye arrangement, dorsal; 37 Left chelicera, ventral; 38 Variation of vulva, dorsal." httpUri="https://zenodo.org/record/277809/files/figure.png" pageId="2" pageNumber="48">31</figureCitation>
|
||
). In dorsal view, vulvae show three distinct windings parts. Glandular pores are situated on the third coils (
|
||
<figureCitation id="13552A664E37F031EFA8FE1B2FD2FE41" box="[159,231,404,428]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,264,1705,1728]" captionTargetBox="[229,1392,221,1634]" captionTargetId="figure@3.[151,1436,195,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Spariolenus aratta spec. nov., holotype female from Iran, Kerman Province, Jebal Barez. 1 Epigyne, ventral; 2 Vulva, lateral; 3 Vulva, dorsal; 4 Schematic course of internal duct system, dorsal; 5 Left female palpal claw, prolateral; 6 Eye arrangement, dorsal; 7 Left chelicera, ventral. Abbreviations: CO—Copulatory opening; CR—Central rim; EF—Epigynal field; EP—Epigynal pit; FD—Fertilisation duct; HGD—Hump-like glandular part of internal duct system; LID—Lumen of internal duct system; MS—Median Slit; SS—Slit sensillum." httpUri="https://zenodo.org/record/277804/files/figure.png" pageId="2" pageNumber="48">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EFC4FE1B2E24FE41" box="[243,273,404,428]" captionStart="FIGURES 11 – 19" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[219,1364,186,1865]" captionTargetId="figure@6.[219,1365,181,1882]" captionTargetPageId="6" captionText="FIGURES 11 – 19. Spariolenus iranomaximus spec. nov., paratype female. 11 Epigyne, ventral; 12 Vulva, dorsal; 13 Vulva, lateral; 14 Schematic course of internal duct system, dorsal (from left duct of the vulva figure 12, not scaled); 15 Eye arrangement, dorsal; 16 Left chelicera, ventral; 17 Left female palpal claw, retrolateral; 18 – 19 Variation of vulvae, dorsal. AB— Anterior band of epigynal field." httpUri="https://zenodo.org/record/277806/files/figure.png" pageId="2" pageNumber="48">12</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EE2AFE1B2E0EFE41" box="[285,315,404,428]" captionStart="FIGURES 20 – 26" captionStartId="8.[151,264,1938,1961]" captionTargetBox="[192,1336,215,1915]" captionTargetId="figure@8.[191,1406,163,1921]" captionTargetPageId="8" captionText="FIGURES 20 – 26. Spariolenus manesht spec. nov., holotype female from Iran, Ilam Province, Karezan, Manesht. 20 Epigyne, ventral; 21 Vulva, lateral; 22 Vulva, dorsal; 23 Schematic course of internal duct system, dorsal; 24 Left female palpal claw, retrolateral; 25 Eye arrangement, dorsal; 26 Left chelicera, ventral." httpUri="https://zenodo.org/record/277807/files/figure.png" pageId="2" pageNumber="48">22</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EE70FE1B2E52FE41" box="[327,359,404,428]" captionStart="FIGURES 31 – 38" captionStartId="11.[151,264,1950,1973]" captionTargetBox="[213,1374,208,1916]" captionTargetId="figure@11.[212,1380,193,1946]" captionTargetPageId="11" captionText="FIGURES 31 – 38. Spariolenus zagros spec. nov., paratype female. 31 Epigyne, ventral; 32 Vulva, dorsal; 33 Vulva, lateral; 34 Schematic course of internal duct system, dorsal (not scaled from left duct of vulva figure 33); 35 Left female palpal claw, retrolateral; 36 Eye arrangement, dorsal; 37 Left chelicera, ventral; 38 Variation of vulva, dorsal." httpUri="https://zenodo.org/record/277809/files/figure.png" pageId="2" pageNumber="48">32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465684E37F031EFF2FE382DC4FC49" pageId="2" pageNumber="48" type="description">
|
||
<paragraph id="8BD136E34E37F031EFF2FE382A07FCF9" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="48">
|
||
<emphasis id="B91AEAF14E37F031EFF2FE382E63FE3D" bold="true" box="[197,342,439,464]" pageId="2" pageNumber="48">Description.</emphasis>
|
||
Medium to very large Heteropodinae (body length 8.5–31.0 mm), prosoma slightly longer than wide; leg length formula: 2143; eyes arranged in two rows and slightly recurved, lateral eyes larger than median eyes, PME larger than AME, PLE slightly larger or equal to ALE (
|
||
<figureCitation id="13552A664E37F031EC4AFE702CF5FDFA" box="[893,960,511,535]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,264,1705,1728]" captionTargetBox="[229,1392,221,1634]" captionTargetId="figure@3.[151,1436,195,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Spariolenus aratta spec. nov., holotype female from Iran, Kerman Province, Jebal Barez. 1 Epigyne, ventral; 2 Vulva, lateral; 3 Vulva, dorsal; 4 Schematic course of internal duct system, dorsal; 5 Left female palpal claw, prolateral; 6 Eye arrangement, dorsal; 7 Left chelicera, ventral. Abbreviations: CO—Copulatory opening; CR—Central rim; EF—Epigynal field; EP—Epigynal pit; FD—Fertilisation duct; HGD—Hump-like glandular part of internal duct system; LID—Lumen of internal duct system; MS—Median Slit; SS—Slit sensillum." httpUri="https://zenodo.org/record/277804/files/figure.png" pageId="2" pageNumber="48">Fig. 6</figureCitation>
|
||
); chelicerae with three anterior and four to seven (
|
||
<taxonomicName id="4C6E4D604E37F031EFD1FDAA2E08FDD1" box="[230,317,549,572]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="tigris">
|
||
<emphasis id="B91AEAF14E37F031EFD1FDAA2E08FDD1" box="[230,317,549,572]" italics="true" pageId="2" pageNumber="48">S. tigris</emphasis>
|
||
</taxonomicName>
|
||
) posterior teeth, intermarginal denticles present (
|
||
<figureCitation id="13552A664E37F031EC56FDAB2C9FFDD1" box="[865,938,548,572]" captionStart="FIGURES 1 – 7" captionStartId="3.[151,264,1705,1728]" captionTargetBox="[229,1392,221,1634]" captionTargetId="figure@3.[151,1436,195,1681]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Spariolenus aratta spec. nov., holotype female from Iran, Kerman Province, Jebal Barez. 1 Epigyne, ventral; 2 Vulva, lateral; 3 Vulva, dorsal; 4 Schematic course of internal duct system, dorsal; 5 Left female palpal claw, prolateral; 6 Eye arrangement, dorsal; 7 Left chelicera, ventral. Abbreviations: CO—Copulatory opening; CR—Central rim; EF—Epigynal field; EP—Epigynal pit; FD—Fertilisation duct; HGD—Hump-like glandular part of internal duct system; LID—Lumen of internal duct system; MS—Median Slit; SS—Slit sensillum." httpUri="https://zenodo.org/record/277804/files/figure.png" pageId="2" pageNumber="48">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EC82FDAB2CE0FDD1" box="[949,981,548,572]" captionStart="FIGURES 20 – 26" captionStartId="8.[151,264,1938,1961]" captionTargetBox="[192,1336,215,1915]" captionTargetId="figure@8.[191,1406,163,1921]" captionTargetPageId="8" captionText="FIGURES 20 – 26. Spariolenus manesht spec. nov., holotype female from Iran, Ilam Province, Karezan, Manesht. 20 Epigyne, ventral; 21 Vulva, lateral; 22 Vulva, dorsal; 23 Schematic course of internal duct system, dorsal; 24 Left female palpal claw, retrolateral; 25 Eye arrangement, dorsal; 26 Left chelicera, ventral." httpUri="https://zenodo.org/record/277807/files/figure.png" pageId="2" pageNumber="48">26</figureCitation>
|
||
); strong male embolus running in prolateral loop partially within groove of tegulum (
|
||
<figureCitation id="13552A664E37F031ED97FDC82DDFFDB2" box="[672,746,583,607]" captionStart="FIGURES 8 – 10" captionStartId="5.[151,264,1478,1501]" captionTargetBox="[217,1434,231,1442]" captionTargetId="figure@5.[151,1436,193,1457]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Spariolenus iranomaximus spec. nov., holotype male from Iran, Ilam Province, Khofash cave. Left palp (8 prolateral, 9 ventral, 10 retrolateral). Abbreviations: BRB—Basal retrolateral bulge; C—Conductor; dRTA—Dorsal part of retrolateral tibial apophysis; E—Embolus; SD—Sperm duct; T—Tegulum; vRTA — Ventral part of retrolateral tibial apophysis." httpUri="https://zenodo.org/record/277805/files/figure.png" pageId="2" pageNumber="48">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031EDCEFDC82C2DFDB2" box="[761,792,583,607]" captionStart="FIGURES 27 – 30" captionStartId="10.[151,264,1360,1383]" captionTargetBox="[220,1412,209,1313]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURES 27 – 30. Spariolenus zagros spec. nov., holotype male from Iran, Kermanshah Province, Kenesht cave. 27 – 29 Left palp (27 prolateral, 28 ventral, 29 retrolateral); 30 Tip of embolus and conductor, ventral." httpUri="https://zenodo.org/record/277808/files/figure.png" pageId="2" pageNumber="48">27</figureCitation>
|
||
), membranous conductor small to well developed, cymbium longer than tibia, cymbium with retrolateral-basal bulge; RTA short and divided in a dorsal and ventral branch, dRTA and vRTA arising from broad conjoint base (
|
||
<figureCitation id="13552A664E37F031EC05FD002C4EFD4A" box="[818,891,655,679]" captionStart="FIGURES 8 – 10" captionStartId="5.[151,264,1478,1501]" captionTargetBox="[217,1434,231,1442]" captionTargetId="figure@5.[151,1436,193,1457]" captionTargetPageId="5" captionText="FIGURES 8 – 10. Spariolenus iranomaximus spec. nov., holotype male from Iran, Ilam Province, Khofash cave. Left palp (8 prolateral, 9 ventral, 10 retrolateral). Abbreviations: BRB—Basal retrolateral bulge; C—Conductor; dRTA—Dorsal part of retrolateral tibial apophysis; E—Embolus; SD—Sperm duct; T—Tegulum; vRTA — Ventral part of retrolateral tibial apophysis." httpUri="https://zenodo.org/record/277805/files/figure.png" pageId="2" pageNumber="48">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="13552A664E37F031ECB0FD002C93FD4A" box="[903,934,655,679]" captionStart="FIGURES 27 – 30" captionStartId="10.[151,264,1360,1383]" captionTargetBox="[220,1412,209,1313]" captionTargetId="figure@10.[151,1436,193,1339]" captionTargetPageId="10" captionText="FIGURES 27 – 30. Spariolenus zagros spec. nov., holotype male from Iran, Kermanshah Province, Kenesht cave. 27 – 29 Left palp (27 prolateral, 28 ventral, 29 retrolateral); 30 Tip of embolus and conductor, ventral." httpUri="https://zenodo.org/record/277808/files/figure.png" pageId="2" pageNumber="48">28</figureCitation>
|
||
); female palpal claw with primary tooth distinctly longer than secondary teeth, 4–8 secondary teeth present; epigyne as in diagnosis with epigynal fields roundish or oval, with or without anterior bands, slit sense organs present, vulva as in diagnosis; colour is generally cream-brown or olive-brown with dark patterns on prosoma and opisthosoma and dark bands on the legs.
|
||
</paragraph>
|
||
<paragraph id="8BD136E34E37F031EFF2FC902DC4FC49" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="48">
|
||
<emphasis id="B91AEAF14E37F031EFF2FC902EA6FCD5" bold="true" box="[197,403,799,824]" pageId="2" pageNumber="48">Species included:</emphasis>
|
||
<taxonomicName id="4C6E4D604E37F031EEADFCAF2DC9FCDA" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[410,764,799,824]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="tigris">
|
||
<emphasis id="B91AEAF14E37F031EEADFCAF2D57FCD5" box="[410,610,800,824]" italics="true" pageId="2" pageNumber="48">Spariolenus tigris</emphasis>
|
||
<bibRefCitation id="EFFF4B124E37F031ED5DFC902DC9FCDA" author="Simon" box="[618,764,799,823]" pageId="2" pageNumber="48" refString="Simon, E. (1880) Revision de la famille des Sparassidae (Arachnides). Actes de la Societe linneenne de Bordeaux, 34, 223 - 351." type="journal article" year="1880">Simon, 1880</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C6E4D604E37F031EC3FFCAF2B23FCDA" authority="Jager, 2006" authorityName="Jager" authorityYear="2006" box="[776,1046,799,825]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="secundus">
|
||
<emphasis id="B91AEAF14E37F031EC3FFCAF2CBFFCD5" box="[776,906,800,824]" italics="true" pageId="2" pageNumber="48">S. secundus</emphasis>
|
||
Jäger, 2006
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C6E4D604E37F031EB16FCAF2BB4FCD5" box="[1057,1153,800,824]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="aratta" status="sp. nov.">
|
||
<emphasis id="B91AEAF14E37F031EB16FCAF2BB4FCD5" box="[1057,1153,800,824]" italics="true" pageId="2" pageNumber="48">S. aratta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91AEAF14E37F031EBBEFCAF2BC9FCD5" bold="true" box="[1161,1276,800,824]" pageId="2" pageNumber="48">
|
||
<taxonomicNameLabel id="A229578A4E37F031EBBEFCAF2BC9FCD5" box="[1161,1276,800,824]" pageId="2" pageNumber="48" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<taxonomicName id="4C6E4D604E37F031EA3CFCAF2FF9FCB1" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="iranomaximus" status="sp. nov.">
|
||
<emphasis id="B91AEAF14E37F031EA3CFCAF2FF9FCB1" italics="true" pageId="2" pageNumber="48">S. iranomaximus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A4E37F031EFE2FCCB2E7FFCB1" box="[213,330,836,860]" pageId="2" pageNumber="48" rank="species">
|
||
<emphasis id="B91AEAF14E37F031EFE2FCCB2E7FFCB1" bold="true" box="[213,330,836,860]" pageId="2" pageNumber="48">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
;
|
||
<taxonomicName id="4C6E4D604E37F031EE6BFCCA2EE2FCB1" box="[348,471,836,860]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="manesht" status="sp. nov.">
|
||
<emphasis id="B91AEAF14E37F031EE6BFCCA2EE2FCB1" box="[348,471,836,860]" italics="true" pageId="2" pageNumber="48">S. manesht</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A4E37F031EED6FCCB2D60FCB1" box="[481,597,836,860]" pageId="2" pageNumber="48" rank="species">
|
||
<emphasis id="B91AEAF14E37F031EED6FCCB2D60FCB1" bold="true" box="[481,597,836,860]" pageId="2" pageNumber="48">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
;
|
||
<taxonomicName id="4C6E4D604E37F031ED50FCCA2DFAFCB1" box="[615,719,837,860]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="zagros" status="sp. nov.">
|
||
<emphasis id="B91AEAF14E37F031ED50FCCA2DFAFCB1" box="[615,719,837,860]" italics="true" pageId="2" pageNumber="48">S. zagros</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A4E37F031EDEEFCCB2C7BFCB1" box="[729,846,836,860]" pageId="2" pageNumber="48" rank="species">
|
||
<emphasis id="B91AEAF14E37F031EDEEFCCB2C7BFCB1" bold="true" box="[729,846,836,860]" pageId="2" pageNumber="48">spec. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
It should be noted that
|
||
<bibRefCitation id="EFFF4B124E37F031EB52FCCB2A21FCB1" author="Platnick" box="[1125,1300,836,860]" pageId="2" pageNumber="48" refString="Platnick, N. I. (2011) The world spider catalog, version 11.5. American Museum of Natural History, New York. Available from http: // research. amnh. org / iz / spiders / catalog. (Accessed: 2 February 2011)" type="journal article" year="2011">Platnick (2011)</bibRefCitation>
|
||
listed three other species for which
|
||
<bibRefCitation id="EFFF4B124E37F031EE94FCE62D05FC92" author="Jager" box="[419,560,871,897]" pageId="2" pageNumber="48" refString="Jager, P. (2002) Heteropodinae: transfers and synonymies (Arachnida: Araneae: Sparassidae). Acta Arachnologica, 51, 33 - 61." type="journal article" year="2002">Jäger (2002)</bibRefCitation>
|
||
ascertained uncertain taxonomic status, they cannot be clearly assigned to the genus
|
||
<taxonomicName id="4C6E4D604E37F031EFE8FC032E50FC49" box="[223,357,908,932]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91AEAF14E37F031EFE8FC032E50FC49" box="[223,357,908,932]" italics="true" pageId="2" pageNumber="48">Spariolenus</emphasis>
|
||
</taxonomicName>
|
||
. For more details see
|
||
<bibRefCitation id="EFFF4B124E37F031ED6BFC022DD9FC49" author="Jager" box="[604,748,908,933]" pageId="2" pageNumber="48" refString="Jager, P. (2002) Heteropodinae: transfers and synonymies (Arachnida: Araneae: Sparassidae). Acta Arachnologica, 51, 33 - 61." type="journal article" year="2002">Jäger (2002)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465684E37F031EFF2FC202E66FC01" pageId="2" pageNumber="48" type="distribution">
|
||
<paragraph id="8BD136E34E37F031EFF2FC202E66FC01" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="48">
|
||
<emphasis id="B91AEAF14E37F031EFF2FC202E6BFC25" bold="true" box="[197,350,943,968]" pageId="2" pageNumber="48">Distribution.</emphasis>
|
||
Indian Peninsula (Calcutta), North-eastern Arabian Peninsula (
|
||
<collectingCountry id="F37976734E37F031EB05FC202B49FC2A" box="[1074,1148,943,967]" name="Oman" pageId="2" pageNumber="48">Oman</collectingCountry>
|
||
), Iranian Plateau (
|
||
<taxonomicName id="4C6E4D604E37F031EA7BFC202AA9FC2A" box="[1356,1436,943,967]" class="Arachnida" family="Sparassidae" genus="Spariolenus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="zagros" status="sp. nov.">Zagros</taxonomicName>
|
||
Mountains,
|
||
<collectingCountry id="F37976734E37F031EE2BFC5B2E7EFC01" box="[284,331,980,1004]" name="Iran" pageId="2" pageNumber="48">Iran</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |