430 lines
54 KiB
XML
430 lines
54 KiB
XML
<document id="3CB71CB03E0D22BA9B2A07699E1F8A85" ID-DOI="10.5281/zenodo.281232" ID-GBIF-Dataset="cd359e84-c028-4f73-a84f-841017d87263" ID-ISSN="1175-5326" ID-Zenodo-Dep="281232" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460485267434" checkinUser="plazi" docAuthor="Skartveit, John & Nel, André" docDate="2012" docId="03C787F53A7CFE6918EEFC33FE3FFA3D" docLanguage="en" docName="zt03329p063.pdf" docOrigin="Zootaxa 3329" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bibio Geoffroy 1762" docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="FFFEFF8D3A7DFE6E1879FFB9FFA1FFA1" masterDocTitle="Fossil Bibionidae (Diptera: Bibionomorpha) from the late Oligocene of Bes-Konak, Anatolia, Turkey" masterLastPageNumber="63" masterPageNumber="51" pageNumber="52" updateTime="1698308914814" updateUser="plazi">
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<mods:titleInfo id="DDB2B4B828741F8715225C059905BEF5">
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<mods:title id="B0157DE2DB9A1D20003834B284D0EEBA">Fossil Bibionidae (Diptera: Bibionomorpha) from the late Oligocene of Bes-Konak, Anatolia, Turkey</mods:title>
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<mods:name id="6E439A8784C60AF8DE2E9CBCF7B92B6E" type="personal">
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<mods:namePart id="5C230BF3136B59C430D6643008CBA764">Skartveit, John</mods:namePart>
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<mods:namePart id="4897213D710E3D53C5355570DEAA0735">Nel, André</mods:namePart>
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<mods:title id="586A480B39C52DD8936E7809B560F86E">Zootaxa</mods:title>
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<mods:date id="6DD00E3BDB2F767423C53B400F5F9025">2012</mods:date>
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<mods:detail id="B260764FC88ED9ACA950A42D2F989785" type="volume">
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<mods:number id="5E3EC7462994664B18DA3A33D357CBC0">3329</mods:number>
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<mods:identifier id="D6956475D0D750350668965E5235577F" type="DOI">10.5281/zenodo.281232</mods:identifier>
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<treatment id="03C787F53A7CFE6918EEFC33FE3FFA3D" ID-DOI="http://doi.org/10.5281/zenodo.6181705" ID-GBIF-Taxon="119518101" ID-Zenodo-Dep="6181705" LSID="urn:lsid:plazi:treatment:03C787F53A7CFE6918EEFC33FE3FFA3D" httpUri="http://treatment.plazi.org/id/03C787F53A7CFE6918EEFC33FE3FFA3D" lastPageId="7" lastPageNumber="58" pageId="1" pageNumber="52">
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<subSubSection id="C37465683A7CFE6F18EEFC33FE4CFC05" box="[151,493,906,932]" pageId="1" pageNumber="52" type="nomenclature">
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<paragraph id="8BD136E33A7CFE6F18EEFC33FE4CFC05" blockId="1.[151,493,906,932]" box="[151,493,906,932]" pageId="1" pageNumber="52">
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<heading id="D099818F3A7CFE6F18EEFC33FE4CFC05" bold="true" box="[151,493,906,932]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
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<emphasis id="B91AEAF13A7CFE6F18EEFC33FE4CFC05" bold="true" box="[151,493,906,932]" pageId="1" pageNumber="52">
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||
Genus
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<taxonomicName id="4C6E4D603A7CFE6F1897FC33FE4CFC05" ID-CoL="9CJF7" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[238,493,906,932]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF13A7CFE6F1897FC33FE8EFC05" bold="true" box="[238,303,906,932]" italics="true" pageId="1" pageNumber="52">Bibio</emphasis>
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<bibRefCitation id="EFFF4B123A7CFE6F194FFC33FE4CFC05" author="Geoffroy" box="[310,493,906,932]" pageId="1" pageNumber="52" refString="Geoffroy, E. L. (1762) Histoire abregee des insectes qui se trouvent aux environs de Paris. 2. Paris, 690 pp." type="book" year="1762">Geoffroy, 1762</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37465683A7CFE6F18EEFC68FD85F8AF" pageId="1" pageNumber="52" type="reference_group">
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||
<paragraph id="8BD136E33A7CFE6F18EEFC68FDB2FBA7" blockId="1.[151,1436,977,1030]" pageId="1" pageNumber="52">
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<treatmentCitationGroup id="AB7E11CD3A7CFE6F18EEFC68FDB2FBA7" pageId="1" pageNumber="52">
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||
<taxonomicName id="4C6E4D603A7CFE6F18EEFC68FE06FC46" ID-CoL="9CJF7" authority="Goeffroy, 1762: 568" authorityName="Goeffroy" authorityPageNumber="568" authorityYear="1762" box="[151,423,977,999]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF13A7CFE6F18EEFC68FF6FFC46" box="[151,206,977,999]" italics="true" pageId="1" pageNumber="52">Bibio</emphasis>
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Goeffroy, 1762: 568
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</taxonomicName>
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.
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<typeStatus id="54D588413A7CFE6F19C8FC6BFE42FC46" box="[433,483,978,999]" pageId="1" pageNumber="52">Type</typeStatus>
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species,
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<taxonomicName id="4C6E4D603A7CFE6F1A3BFC68FC33FC46" ID-CoL="8Z2M9" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[578,914,977,999]" class="Insecta" family="Bibionidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="hortulana">
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<emphasis id="B91AEAF13A7CFE6F1A3BFC68FD4FFC46" box="[578,750,977,999]" italics="true" pageId="1" pageNumber="52">Tipula hortulana</emphasis>
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||
<bibRefCitation id="EFFF4B123A7CFE6F1A8DFC68FC33FC46" author="Linnaeus" box="[756,914,977,999]" pageId="1" pageNumber="52" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. L. Salvii, Holmiae [= Stockholm]. 824 p." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
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</taxonomicName>
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(by subsequent designation of
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<treatmentCitation id="0ACF10F23A7CFE6F1CABFC68FA39FC46" author="Latreille" box="[1234,1432,977,999]" page="442" pageId="1" pageNumber="52" year="1810">
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<bibRefCitation id="EFFF4B123A7CFE6F1CABFC68FAC4FC46" author="Latreille" box="[1234,1381,977,999]" pageId="1" pageNumber="52" refString="Latreille, P. A. (1810) Considerations generales sur l'ordre naturel des animaux composant les classes des crustaces, des arachnides, et des insectes; avec un tableau methodique de leurs genres, disposes en familles. F. Schoell, Paris. 444 pp." type="book" year="1810">Latreille 1810</bibRefCitation>
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: 442
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</treatmentCitation>
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, validated by I.C.Z.N., 1957: 86).
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</treatmentCitationGroup>
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</paragraph>
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<caption id="DF11666B3A7CFE6F18EEF936FCC0F904" box="[151,865,1679,1702]" pageId="1" pageNumber="52">
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<paragraph id="8BD136E33A7CFE6F18EEF936FCC0F904" blockId="1.[151,865,1679,1702]" box="[151,865,1679,1702]" pageId="1" pageNumber="52">
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<emphasis id="B91AEAF13A7CFE6F18EEF936FEB2F905" bold="true" box="[151,275,1679,1702]" pageId="1" pageNumber="52">FIGURE 1.</emphasis>
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<taxonomicName id="4C6E4D603A7CFE6F1960F936FE1EF904" box="[281,447,1679,1701]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
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<emphasis id="B91AEAF13A7CFE6F1960F936FE1EF904" box="[281,447,1679,1701]" italics="true" pageId="1" pageNumber="52">Bibio anatolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229578A3A7CFE6F19BFF929FE56F904" box="[454,503,1680,1701]" pageId="1" pageNumber="52" rank="species">sp.n.</taxonomicNameLabel>
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, male, holotype (MNHN B 47826).
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</paragraph>
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</caption>
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<paragraph id="8BD136E33A7CFE6F18EEF94FFD85F8AF" blockId="1.[151,548,1782,1807]" box="[151,548,1782,1807]" pageId="1" pageNumber="52">
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<emphasis id="B91AEAF13A7CFE6F18EEF94FFE3BF8AF" bold="true" box="[151,410,1782,1807]" pageId="1" pageNumber="52">
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<taxonomicName id="4C6E4D603A7CFE6F18EEF94FFEF5F8AE" box="[151,340,1782,1807]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
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<emphasis id="B91AEAF13A7CFE6F18EEF94FFEF5F8AE" bold="true" box="[151,340,1782,1807]" italics="true" pageId="1" pageNumber="52">Bibio anatolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229578A3A7CFE6F1922F94FFE3BF8AF" box="[347,410,1782,1806]" pageId="1" pageNumber="52" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(figs. 1–14)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465683A7CFE6D18EEF885FEBCF83A" lastPageId="3" lastPageNumber="54" pageId="1" pageNumber="52" type="etymology">
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<paragraph id="8BD136E33A7CFE6F18EEF885FC5AF8F5" blockId="1.[151,1436,1852,2020]" box="[151,1019,1852,1876]" pageId="1" pageNumber="52">
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Etymology: Named after
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<collectingCountry id="F37976733A7CFE6F19CDF885FDB6F8F5" box="[436,535,1852,1876]" name="Turkey" pageId="1" pageNumber="52">Anatolia</collectingCountry>
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, home province of the Bes-Konak outcrop.
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</paragraph>
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<paragraph id="8BD136E33A7CFE6F18B0F8D9FC19F861" blockId="1.[151,1436,1852,2020]" pageId="1" pageNumber="52">
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Diagnosis: A moderately large
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<taxonomicName id="4C6E4D603A7CFE6F1A5EF8D8FDC2F8D8" box="[551,611,1889,1913]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAF13A7CFE6F1A5EF8D8FDC2F8D8" box="[551,611,1889,1913]" italics="true" pageId="1" pageNumber="52">Bibio</emphasis>
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</taxonomicName>
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species. Wing light brown fumose, rather narrow, crossvein r-m approximately half as long as Rs. Antennal flagellum seven- or eight-segmented.
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<collectingCountry id="F37976733A7CFE6F1B98F83DFBB8F83D" box="[993,1049,1924,1948]" name="India" pageId="1" pageNumber="52">Hind</collectingCountry>
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legs moderately clavate with first tarsomere not swollen. Lobes of epandrium short and widely separated.
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</paragraph>
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<paragraph id="8BD136E33A7CFE6C18B0F875FF49FF56" blockId="1.[151,1436,1852,2020]" lastBlockId="2.[151,1436,151,247]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
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Material:
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<typeStatus id="54D588413A7CFE6F1940F875FE00F845" box="[313,417,1996,2020]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
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(male) MNHN B 47826, fig.1. The specimen is preserved lying on its side, but with the abdomen twisted so that the terminalia are seen from a dorsal view. Preservation of wings, body and hind legs quite good, of head and terminalia fair. The two front pairs of legs are poorly preserved. Pile of head and body not preserved.
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</paragraph>
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<caption id="DF11666B3A7FFE6C18EEF836FF73F843" httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="2" pageNumber="53" targetBox="[156,1427,307,1908]" targetPageId="2">
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<paragraph id="8BD136E33A7FFE6C18EEF836FF73F843" blockId="2.[151,1436,1935,2018]" pageId="2" pageNumber="53">
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<emphasis id="B91AEAF13A7FFE6C18EEF836FE8FF804" bold="true" box="[151,302,1935,1958]" pageId="2" pageNumber="53">FIGURE 2–6.</emphasis>
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<taxonomicName id="4C6E4D603A7FFE6C194FF836FE7FF804" box="[310,478,1935,1957]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
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<emphasis id="B91AEAF13A7FFE6C194FF836FE7FF804" box="[310,478,1935,1957]" italics="true" pageId="2" pageNumber="53">Bibio anatolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229578A3A7FFE6C199EF829FDBBF804" box="[487,538,1936,1957]" pageId="2" pageNumber="53" rank="species">
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<emphasis id="B91AEAF13A7FFE6C199EF829FDBBF804" bold="true" box="[487,538,1936,1957]" pageId="2" pageNumber="53">sp.n.</emphasis>
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</taxonomicNameLabel>
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, male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1mm.
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</paragraph>
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</caption>
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<caption id="DF11666B3A7EFE6D18EEFB23FCC7FB10" box="[151,870,1178,1201]" httpUri="https://zenodo.org/record/281234/files/figure.png" pageId="3" pageNumber="54" targetBox="[151,1436,193,1156]" targetPageId="3">
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<paragraph id="8BD136E33A7EFE6D18EEFB23FCC7FB10" blockId="3.[151,870,1178,1201]" box="[151,870,1178,1201]" pageId="3" pageNumber="54">
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<emphasis id="B91AEAF13A7EFE6D18EEFB23FEB2FB11" bold="true" box="[151,275,1178,1201]" pageId="3" pageNumber="54">FIGURE 7.</emphasis>
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<taxonomicName id="4C6E4D603A7EFE6D1960FB23FE1EFB11" box="[281,447,1178,1200]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
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<emphasis id="B91AEAF13A7EFE6D1960FB23FE1EFB11" box="[281,447,1178,1200]" italics="true" pageId="3" pageNumber="54">Bibio anatolicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A229578A3A7EFE6D19BFFB22FE59FB11" box="[454,504,1179,1200]" pageId="3" pageNumber="54" rank="species">sp.n.</taxonomicNameLabel>
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, female, habitus (MNHN B 47789).
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</paragraph>
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</caption>
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<paragraph id="8BD136E33A7EFE6D18B0FB6EFBB6FA96" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
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Other material, all in MNHN: five males (B 47751, B 47833, B 48011, IY 262, 1 unlabelled)
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<specimenCount id="9D68FD6A3A7EFE6D1C90FB6EFAC5FB4E" box="[1257,1380,1239,1263]" pageId="3" pageNumber="54" type="female">20 females</specimenCount>
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(BK 32, BK 92, BK 345, BK 605, BK 642, B 47578, B 47603, B 47728, B 47747, B 47771, B 47776, B 47789, B 47820, B 47884, B 47906, B 47926, B 47932, B 47972, B 48014, 1 unlabelled).
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</paragraph>
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<paragraph id="8BD136E33A7EFE6D18B0FAFAFB2DFA02" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
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Description:
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<emphasis id="B91AEAF13A7EFE6D1924FAFDFE38FAFD" bold="true" box="[349,409,1348,1372]" pageId="3" pageNumber="54">Male</emphasis>
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: A moderately large species, body 8.0–
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<quantity id="4C969B063A7EFE6D1B12FAFAFC62FAFA" box="[875,963,1347,1371]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="54" unit="mm" value="8.7">8.7 mm</quantity>
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long (N = 4), black with brownish legs. Head (
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<figureCitation id="13552A663A7EFE6D189DFADEFE8BFADE" box="[228,298,1383,1407]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="3" pageNumber="54">Fig. 2</figureCitation>
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)
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<quantity id="4C969B063A7EFE6D1945FADEFE43FADE" box="[316,482,1383,1407]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="3" pageNumber="54" unit="mm" value="1.525" valueMax="1.6" valueMin="1.45">1.45–1.60 mm</quantity>
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long (N = 3), plus rostrum and proboscis
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<quantity id="4C969B063A7EFE6D1BA8FADEFB99FADE" box="[977,1080,1383,1407]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="54" unit="mm" value="0.75">0.75 mm</quantity>
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(N = 1). Flagellum apparently seven-segmented, but not well preserved in any of the male specimens. Pile not preserved.
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</paragraph>
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<paragraph id="8BD136E33A7EFE6D18B0FA16FC3CFA66" blockId="3.[151,1437,1239,2020]" box="[201,925,1455,1480]" pageId="3" pageNumber="54">
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Thorax black. Pile not preserved. Length
|
||
<quantity id="4C969B063A7EFE6D1AE3FA16FC9CFA69" box="[666,829,1455,1480]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.25" metricValueMin="2.75" pageId="3" pageNumber="54" unit="mm" value="3.0" valueMax="3.25" valueMin="2.75">2.75–3.25 mm</quantity>
|
||
(N = 3).
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7EFE6D18B0FA6DFDAAF901" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
|
||
Legs: Brownish. Fore femur with long, black setae, tibia
|
||
<quantity id="4C969B063A7EFE6D1B34FA6AFC13FA4A" box="[845,946,1491,1515]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" pageId="3" pageNumber="54" unit="mm" value="1.85">1.85 mm</quantity>
|
||
long (N = 1). Shape of mucron and spur of fore tibia not possible to see well in any of the male specimens.
|
||
<collectingCountry id="F37976733A7EFE6D1B24FA4EFC34F9AE" box="[861,917,1527,1551]" name="India" pageId="3" pageNumber="54">Hind</collectingCountry>
|
||
femur and tibia (
|
||
<figureCitation id="13552A663A7EFE6D1C2AFA4EFB34F9AE" box="[1107,1173,1527,1551]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="3" pageNumber="54">Fig. 3</figureCitation>
|
||
) strongly clavate.
|
||
<collectingCountry id="F37976733A7EFE6D1D1DFA4EFA3DF9AE" box="[1380,1436,1527,1551]" name="India" pageId="3" pageNumber="54">Hind</collectingCountry>
|
||
femur dark basally, light yellowish-brown in apical half, 2.75–3.0 mm (N = 3) long,
|
||
<quantity id="4C969B063A7EFE6D1C25F9A2FAA1F992" box="[1116,1280,1563,1587]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="3" pageNumber="54" unit="mm" value="0.22" valueMax="0.25" valueMin="0.19">0.19–0.25 mm</quantity>
|
||
(N = 3) wide basally,
|
||
<quantity id="4C969B063A7EFE6D188BF986FE35F9F9" box="[242,404,1599,1624]" metricMagnitude="-4" metricUnit="m" metricValue="5.249999999999999" metricValueMax="5.699999999999999" metricValueMin="4.8" pageId="3" pageNumber="54" unit="mm" value="0.5249999999999999" valueMax="0.57" valueMin="0.48">0.48–0.57 mm</quantity>
|
||
(N = 4) apically.
|
||
<collectingCountry id="F37976733A7EFE6D1A2EF986FD2EF9F6" box="[599,655,1599,1623]" name="India" pageId="3" pageNumber="54">Hind</collectingCountry>
|
||
tibia yellowish-brown basally, darkened apically,
|
||
<quantity id="4C969B063A7EFE6D1CC7F986FAE5F9F9" box="[1214,1348,1599,1624]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="3" pageNumber="54" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
|
||
(N = 4) long,
|
||
<quantity id="4C969B063A7EFE6D18A1F9DAFEDCF9DA" box="[216,381,1635,1659]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="3" pageNumber="54" unit="mm" value="0.25" valueMax="0.28" valueMin="0.22">0.22–0.28 mm</quantity>
|
||
(N = 4) wide basally,
|
||
<quantity id="4C969B063A7EFE6D1AF8F9DAFC87F9DA" box="[641,806,1635,1659]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.2" metricValueMin="4.8" pageId="3" pageNumber="54" unit="mm" value="0.55" valueMax="0.62" valueMin="0.48">0.48–0.62 mm</quantity>
|
||
(N = 4) apically. First tarsomere slightly swollen,
|
||
<quantity id="4C969B063A7EFE6D1D12F9DAFF62F901" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="3" pageNumber="54" unit="mm" value="0.96">0.96 mm</quantity>
|
||
long,
|
||
<quantity id="4C969B063A7EFE6D1970F931FECCF901" box="[265,365,1672,1696]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="54" unit="mm" value="0.32">0.32 mm</quantity>
|
||
wide (N = 1).
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7EFE6D18B0F912FCBFF8AA" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
|
||
Wing (
|
||
<figureCitation id="13552A663A7EFE6D196EF912FEFCF962" box="[279,349,1707,1731]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="3" pageNumber="54">Fig. 4</figureCitation>
|
||
)
|
||
<quantity id="4C969B063A7EFE6D1917F912FE59F962" box="[366,504,1707,1731]" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.6" metricValueMin="6.3" pageId="3" pageNumber="54" unit="mm" value="6.949999999999999" valueMax="7.6" valueMin="6.3">6.3–7.6 mm</quantity>
|
||
long (N = 5),
|
||
<quantity id="4C969B063A7EFE6D1ADAF912FC9BF962" box="[675,826,1707,1731]" metricMagnitude="-3" metricUnit="m" metricValue="2.625" metricValueMax="2.75" metricValueMin="2.5" pageId="3" pageNumber="54" unit="mm" value="2.625" valueMax="2.75" valueMin="2.5">2.5–2.75 mm</quantity>
|
||
wide (N = 2). Light brown fumose, veins distinctive throughout, R-veins brown, more posterior veins more or less colourless. Pterostigma brown and distinctive. Crossvein r-m is roughly half as long as first sector of Rs.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7EFE6D18B0F8A1FEBCF83A" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
|
||
Abdomen 4.4–5.0 mm long (N = 3), relatively slender, somewhat conical, tergites short and transverse. No trace of pile in the specimens at hand. Epandrium (
|
||
<figureCitation id="13552A663A7EFE6D1AA9F882FCB3F8F2" box="[720,786,1851,1875]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="3" pageNumber="54">Fig. 5</figureCitation>
|
||
) with two short, rounded lobes separated by a wide mesal cleft. Gonocoxosternite (
|
||
<figureCitation id="13552A663A7EFE6D19D7F8D9FE52F8D9" box="[430,499,1888,1912]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="3" pageNumber="54">Fig. 6</figureCitation>
|
||
) with a wide and shallow, mesal indentation. Gonostyli sickle-shaped, rather small and slender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465683A7EFE6918B0F811FE3FFA3D" lastPageId="7" lastPageNumber="58" pageId="3" pageNumber="54" type="description">
|
||
<paragraph id="8BD136E33A7EFE6D18B0F811FBFDF845" blockId="3.[151,1437,1239,2020]" pageId="3" pageNumber="54">
|
||
<emphasis id="B91AEAF13A7EFE6D18B0F811FE81F861" bold="true" box="[201,288,1960,1984]" pageId="3" pageNumber="54">Female</emphasis>
|
||
(fig. 7): Body
|
||
<quantity id="4C969B063A7EFE6D19ACF811FDD8F861" box="[469,633,1960,1984]" metricMagnitude="-2" metricUnit="m" metricValue="1.145" metricValueMax="1.27" metricValueMin="1.02" pageId="3" pageNumber="54" unit="mm" value="11.45" valueMax="12.7" valueMin="10.2">10.2–12.7 mm</quantity>
|
||
long (N = 5). Head (
|
||
<figureCitation id="13552A663A7EFE6D1B15F811FC11F861" box="[876,944,1960,1984]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="3" pageNumber="54">Fig. 8</figureCitation>
|
||
) 1.5–2.0 mm long (N = 4), black, antennal flagellum (
|
||
<figureCitation id="13552A663A7EFE6D1968F875FEF4F845" box="[273,341,1996,2020]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="3" pageNumber="54">Fig. 9</figureCitation>
|
||
) rather short and stout, eight-segmented. Ocellar tubercle prominent.
|
||
</paragraph>
|
||
<caption id="DF11666B3A79FE6A18EEF830FD07F87D" httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="4" pageNumber="55" targetBox="[198,1395,215,1896]" targetPageId="4">
|
||
<paragraph id="8BD136E33A79FE6A18EEF830FD07F87D" blockId="4.[151,1436,1929,2013]" pageId="4" pageNumber="55">
|
||
<emphasis id="B91AEAF13A79FE6A18EEF830FE98F83E" bold="true" box="[151,313,1929,1952]" pageId="4" pageNumber="55">FIGURE 8–14.</emphasis>
|
||
<taxonomicName id="4C6E4D603A79FE6A1946F833FE47F801" box="[319,486,1930,1952]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="anatolicus" status="sp. nov.">
|
||
<emphasis id="B91AEAF13A79FE6A1946F833FE47F801" box="[319,486,1930,1952]" italics="true" pageId="4" pageNumber="55">Bibio anatolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A3A79FE6A1994F832FDBFF801" box="[493,542,1931,1952]" pageId="4" pageNumber="55" rank="species">sp.n.</taxonomicNameLabel>
|
||
, female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD136E33A78FE6B18B0FF2EFE4CFEBD" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="56">
|
||
Thorax 3.8–4.0 mm long (N = 3), yellowish-brown to dark brown dorsally, pleura darker. Legs brown, fore tibiae black, mid- and hind tibiae and all tarsal segments darker apically; fore tibia (
|
||
<figureCitation id="13552A663A78FE6B1C65FF05FBCCFF75" box="[1052,1133,188,212]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="5" pageNumber="56">Fig. 10</figureCitation>
|
||
) quite stout, spur short and curved, less than half as long as mucron. Mid- and hind femora clavate, rather thick, hind tibia slender, all tarsi slender.
|
||
<collectingCountry id="F37976733A78FE6B188DFEBDFE8DFEBD" box="[244,300,260,284]" name="India" pageId="5" pageNumber="56">Hind</collectingCountry>
|
||
leg as in
|
||
<figureCitation id="13552A663A78FE6B19EEFEBDFE48FEBD" box="[407,489,260,284]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="5" pageNumber="56">Fig. 11</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18B0FE9EFDDEFEC5" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="56">
|
||
Wing (
|
||
<figureCitation id="13552A663A78FE6B196EFE9EFECAFE9E" box="[279,363,295,319]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="5" pageNumber="56">Fig. 12</figureCitation>
|
||
) 8.0–
|
||
<quantity id="4C969B063A78FE6B19D4FE9EFDA4FE9E" box="[429,517,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="56" unit="mm" value="9.5">9.5 mm</quantity>
|
||
long (N = 5), light brown fumose, venation as in male, veins brown, distinctive throughout. Pterostigma brown, distinctive.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18B0FED6FB03FE26" blockId="5.[151,1437,151,501]" box="[201,1186,367,391]" pageId="5" pageNumber="56">
|
||
Abdomen yellowish-brown to dark brown. Terminalia as in
|
||
<figureCitation id="13552A663A78FE6B1B10FED6FC50FE26" box="[873,1009,367,391]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="5" pageNumber="56">Figs. 13–14</figureCitation>
|
||
. Cerci rounded.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18B0FE2DFA93FE55" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="56">
|
||
Remark: The antenna is well preserved in just one of the specimens at hand (female, MNHN B 47906,
|
||
<figureCitation id="13552A663A78FE6B1D33FE2DFA2CFE0D" box="[1354,1421,404,428]" captionStart="FIGURE 8 – 14" captionStartId="4.[151,249,1929,1952]" captionTargetBox="[198,1395,215,1896]" captionTargetId="figure@4.[188,1399,193,1908]" captionTargetPageId="4" captionText="FIGURE 8 – 14. Bibio anatolicus sp. n., female. 8 head (MNHN BK 223). 9. antenna (MNHN B 47906). 10. fore tibia (MNHN BK 223). 11. hind leg (MNHN BK 223). 12. wing (MNHN B 47789). 13. terminalia, dorsal view (MNHN B 47789). 14. terminalia, lateral view (MNHN B 47906). Scale = 1 mm." httpUri="https://zenodo.org/record/281235/files/figure.png" pageId="5" pageNumber="56">Fig. 9</figureCitation>
|
||
). In this case the flagellum consists of eight short, transverse segments. In several other specimens (e.g.,
|
||
<figureCitation id="13552A663A78FE6B1D58FE0EFAC5FE6E" box="[1313,1380,439,463]" captionStart="FIGURE 2 – 6" captionStartId="2.[151,250,1935,1958]" captionTargetBox="[156,1427,307,1908]" captionTargetId="figure@2.[151,1436,301,1914]" captionTargetPageId="2" captionText="FIGURE 2 – 6. Bibio anatolicus sp. n., male. 2. head (MNHN B 47754). 3. hind femur and tibia (MNHN B 48011). 4. wing (MNHN B 47826). 5. terminalia, dorsal view (MNHN B 47890). 6. gonocoxosternite, ventral view (MNHN B 47826). Scale = 1 mm." httpUri="https://zenodo.org/record/281233/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
|
||
), the flagella appear to have just six or seven segments; they are, however, too fragmentary for a certain count.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18EEFDFEFD8DFDFE" blockId="5.[151,556,583,608]" box="[151,556,583,608]" pageId="5" pageNumber="56">
|
||
<emphasis id="B91AEAF13A78FE6B18EEFDFEFE34FDC1" bold="true" box="[151,405,583,608]" pageId="5" pageNumber="56">
|
||
<taxonomicName id="4C6E4D603A78FE6B18EEFDFEFEEEFDC1" box="[151,335,583,608]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="nigricosta" status="sp. nov.">
|
||
<emphasis id="B91AEAF13A78FE6B18EEFDFEFEEEFDC1" bold="true" box="[151,335,583,608]" italics="true" pageId="5" pageNumber="56">Bibio nigricosta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A3A78FE6B192FFDF1FE34FDC1" box="[342,405,584,608]" pageId="5" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(figs. 15–21)
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18EEFD36FC15FD6D" blockId="5.[151,1436,655,860]" pageId="5" pageNumber="56">
|
||
Etymology: The species has a conspicuously darkened costal cell in the wing and is named accordingly. From Latin
|
||
<emphasis id="B91AEAF13A78FE6B18EEFD0CFF70FD6D" box="[151,209,693,716]" italics="true" pageId="5" pageNumber="56">niger</emphasis>
|
||
, black, and
|
||
<emphasis id="B91AEAF13A78FE6B1920FD0CFE35FD6D" box="[345,404,693,716]" italics="true" pageId="5" pageNumber="56">costa</emphasis>
|
||
, rib. The species epithet is a noun in opposition.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A78FE6B18B0FD6EFE9DFCFD" blockId="5.[151,1436,655,860]" pageId="5" pageNumber="56">Diagnosis: A large, robust species. Femorae red with black tips, tibiae and tarsi black. Wing rather short and broad, fumose with costal cell darkened, pterostigma black. Crossvein r-m half as long as to subequal to first sector of Rs. Legs robust, first hind tarsomere moderately swollen. Male gonostyli robust, epandrium rather small with rounded lobes.</paragraph>
|
||
<caption id="DF11666B3A78FE6B18EEF801FCC5F86F" box="[151,868,1976,1999]" httpUri="https://zenodo.org/record/281236/files/figure.png" pageId="5" pageNumber="56" targetBox="[151,1436,913,1954]" targetPageId="5">
|
||
<paragraph id="8BD136E33A78FE6B18EEF801FCC5F86F" blockId="5.[151,868,1976,1999]" box="[151,868,1976,1999]" pageId="5" pageNumber="56">
|
||
<emphasis id="B91AEAF13A78FE6B18EEF801FEBEF86F" bold="true" box="[151,287,1976,1999]" pageId="5" pageNumber="56">FIGURE 15.</emphasis>
|
||
<taxonomicName id="4C6E4D603A78FE6B195CF801FE6BF86E" box="[293,458,1976,1999]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="nigricosta" status="sp. nov.">
|
||
<emphasis id="B91AEAF13A78FE6B195CF801FE6BF86E" box="[293,458,1976,1999]" italics="true" pageId="5" pageNumber="56">Bibio nigricosta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A3A78FE6B19A9F800FDA0F86F" box="[464,513,1977,1998]" pageId="5" pageNumber="56" rank="species">sp.n.</taxonomicNameLabel>
|
||
, male, holotype (MHNH BK 450).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF11666B3A7BFE6818EEF8C7FD19F870" httpUri="https://zenodo.org/record/281237/files/figure.png" pageId="6" pageNumber="57" targetBox="[155,1436,202,1891]" targetPageId="6">
|
||
<paragraph id="8BD136E33A7BFE6818EEF8C7FD19F870" blockId="6.[151,1437,1918,2002]" pageId="6" pageNumber="57">
|
||
<emphasis id="B91AEAF13A7BFE6818EEF8C7FEE4F835" bold="true" box="[151,325,1918,1941]" pageId="6" pageNumber="57">FIGURE 16–20.</emphasis>
|
||
<taxonomicName id="4C6E4D603A7BFE681932F8C6FE4FF834" box="[331,494,1919,1941]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="nigricosta" status="sp. nov.">
|
||
<emphasis id="B91AEAF13A7BFE681932F8C6FE4FF834" box="[331,494,1919,1941]" italics="true" pageId="6" pageNumber="57">Bibio nigricosta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A229578A3A7BFE68198DF8C6FD87F835" box="[500,550,1919,1940]" pageId="6" pageNumber="57" rank="species">sp.n.</taxonomicNameLabel>
|
||
, male. 16. hind leg (MNHN BK 450). 17. wing. (MNHN BK 450) Position of crossvein m-cu not possible to ascertain in this specimen. 18. terminalia, dorsal view (MNHN BK 450). 19. habitus (MNHN B 47847). 20. female (?), wing (MNHN B 47961). Scale = 1mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD136E33A7AFE6918B0FF2EFDA6FE0D" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Remark. The species appears to be similar to
|
||
<taxonomicName id="4C6E4D603A7AFE691ABFFF21FC5AFF0E" authority="Heer, 1849" authorityName="Heer" authorityYear="1849" box="[710,1019,151,176]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="B91AEAF13A7AFE691ABFFF21FCD7FF11" box="[710,886,152,176]" italics="true" pageId="7" pageNumber="58">Bibio giganteus</emphasis>
|
||
<bibRefCitation id="EFFF4B123A7AFE691B04FF21FC5AFF0E" author="Heer" box="[893,1019,151,175]" pageId="7" pageNumber="58" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von OEningen und von Radoboj in Croatien. Zweite Abtheilung: Heuschrecken, Florfliegen, Aderfluger, Schmetterlinge und Fliegen. Leipzig, Engelmann, W. (ed.). Neue Denkschriften Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, Zurich, 11, iv + 264 pp." type="book" year="1849">Heer, 1849</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6E4D603A7AFE691C7FFF21FA84FF0E" authority="Heer, 1849" authorityName="Heer" authorityYear="1849" box="[1030,1317,151,176]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="linearis">
|
||
<emphasis id="B91AEAF13A7AFE691C7FFF21FB01FF11" box="[1030,1184,152,176]" italics="true" pageId="7" pageNumber="58">Bibio linearis</emphasis>
|
||
<bibRefCitation id="EFFF4B123A7AFE691CDEFF21FA84FF0E" author="Heer" box="[1191,1317,151,175]" pageId="7" pageNumber="58" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von OEningen und von Radoboj in Croatien. Zweite Abtheilung: Heuschrecken, Florfliegen, Aderfluger, Schmetterlinge und Fliegen. Leipzig, Engelmann, W. (ed.). Neue Denkschriften Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, Zurich, 11, iv + 264 pp." type="book" year="1849">Heer, 1849</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6E4D603A7AFE691D26FF21FE20FF75" authority="Heer, 1849" authorityName="Heer" authorityYear="1849" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="partschii">
|
||
<emphasis id="B91AEAF13A7AFE691D26FF21FF5AFF75" italics="true" pageId="7" pageNumber="58">Bibio partschii</emphasis>
|
||
<bibRefCitation id="EFFF4B123A7AFE69197DFF04FE20FF75" author="Heer" box="[260,385,188,212]" pageId="7" pageNumber="58" refString="Heer, O. (1849) Die Insektenfauna der Tertiargebilde von OEningen und von Radoboj in Croatien. Zweite Abtheilung: Heuschrecken, Florfliegen, Aderfluger, Schmetterlinge und Fliegen. Leipzig, Engelmann, W. (ed.). Neue Denkschriften Allgemeinen Schweizerischen Gesellschaft fur die Gesammten Naturwissenschaften, Zurich, 11, iv + 264 pp." type="book" year="1849">Heer, 1849</bibRefCitation>
|
||
</taxonomicName>
|
||
described from the Early Miocene of Radoboj (
|
||
<collectingCountry id="F37976733A7AFE691BD3FF05FBA2FF75" box="[938,1027,188,212]" name="Croatia" pageId="7" pageNumber="58">Croatia</collectingCountry>
|
||
). It can be distinguished from these species by the much broader wing (length/width approximately 2.1 versus
|
||
<quantity id="4C969B063A7AFE691BA2FF66FBBAFF56" box="[987,1051,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" pageId="7" pageNumber="58" unit="in" value="2.7">2.7 in</quantity>
|
||
<taxonomicName id="4C6E4D603A7AFE691C5BFF59FB73FF59" box="[1058,1234,224,248]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="B91AEAF13A7AFE691C5BFF59FB73FF59" box="[1058,1234,224,248]" italics="true" pageId="7" pageNumber="58">Bibio giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6E4D603A7AFE691D70FF58FA2DFF59" box="[1289,1420,224,248]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="partschii">
|
||
<emphasis id="B91AEAF13A7AFE691D70FF58FA2DFF59" box="[1289,1420,224,248]" italics="true" pageId="7" pageNumber="58">B. partschii</emphasis>
|
||
</taxonomicName>
|
||
). The latter species also have crossvein r-m markedly shorter than half the length of first sector of Rs. The comparison is somewhat complicated by the fact that Heer’s three species were described based on females, while the
|
||
<typeStatus id="54D588413A7AFE691D15FE91FA3DFEE1" box="[1388,1436,296,320]" pageId="7" pageNumber="58">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6E4D603A7AFE6918CFFEF4FEE7FEC5" box="[182,326,333,356]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="nigricosta">
|
||
<emphasis id="B91AEAF13A7AFE6918CFFEF4FEE7FEC5" box="[182,326,333,356]" italics="true" pageId="7" pageNumber="58">B. nigricosta</emphasis>
|
||
</taxonomicName>
|
||
is a male. However, female bibionids tend to have slightly wider, not narrower, wings than males. We did not find the
|
||
<typeStatus id="54D588413A7AFE691901FEC9FE09FE29" box="[376,424,368,392]" pageId="7" pageNumber="58">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C6E4D603A7AFE6919B6FEC9FDC9FE29" box="[463,616,368,392]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="linearis">
|
||
<emphasis id="B91AEAF13A7AFE6919B6FEC9FDC9FE29" box="[463,616,368,392]" italics="true" pageId="7" pageNumber="58">Bibio linearis</emphasis>
|
||
</taxonomicName>
|
||
, but judging from Heer’s description we believe that it is very similar to
|
||
<taxonomicName id="4C6E4D603A7AFE6918EEFE2DFEE6FE0D" box="[151,327,404,428]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="giganteus">
|
||
<emphasis id="B91AEAF13A7AFE6918EEFE2DFEE6FE0D" box="[151,327,404,428]" italics="true" pageId="7" pageNumber="58">Bibio giganteus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6E4D603A7AFE691907FE2CFDA0FE0D" box="[382,513,404,428]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="partschii">
|
||
<emphasis id="B91AEAF13A7AFE691907FE2CFDA0FE0D" box="[382,513,404,428]" italics="true" pageId="7" pageNumber="58">B. partschii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FE0EFC86FD9D" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Material.
|
||
<typeStatus id="54D588413A7AFE69194CFE0EFE3CFE6E" box="[309,413,439,463]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
(male) BK 450, fig. 15. The specimen is preserved in a spread position, dorsal side up. The head is poorly preserved but body, hind legs and wings are in a fairly good state of preservation. The two anterior pairs of legs are poorly preserved.
|
||
<typeStatus id="54D588413A7AFE691A5EFDB9FD2CFDB9" box="[551,653,512,536]" pageId="7" pageNumber="58" type="paratype">Paratype</typeStatus>
|
||
: one female (B 47847), fig.19. Other material: partial specimen, sex uncertain (BK 21), one wing, presumed female (B 47961).
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FDFEFC38FD06" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Description:
|
||
<typeStatus id="54D588413A7AFE691923FDFEFE63FDFE" box="[346,450,583,607]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
(male) BK 450. Total length of
|
||
<typeStatus id="54D588413A7AFE691B50FDFEFC2AFDFE" box="[809,907,583,607]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
<quantity id="4C969B063A7AFE691BEBFDFEFC58FDFE" box="[914,1017,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="7" pageNumber="58" unit="mm" value="13.1">13.1 mm</quantity>
|
||
; colour black except for reddish femora. Body pile apparently short and fine. Head
|
||
<quantity id="4C969B063A7AFE691AD8FDD5FCA4FD25" box="[673,773,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="7" pageNumber="58" unit="mm" value="1.95">1.95 mm</quantity>
|
||
wide,
|
||
<quantity id="4C969B063A7AFE691B36FDD5FC12FD25" box="[847,947,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="7" pageNumber="58" unit="mm" value="2.05">2.05 mm</quantity>
|
||
long (including rostrum), poorly preserved. Antennal flagellum poorly preserved but probably seven-segmented.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FD0DFDC7FD6D" blockId="7.[151,1437,151,1436]" box="[201,614,692,716]" pageId="7" pageNumber="58">
|
||
Thorax
|
||
<quantity id="4C969B063A7AFE691959FD0DFED7FD6D" box="[288,374,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="58" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long,
|
||
<quantity id="4C969B063A7AFE6919C4FD0DFD80FD6D" box="[445,545,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="58" unit="mm" value="2.15">2.15 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FD6EFC10FC98" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Wing (
|
||
<figureCitation id="13552A663A7AFE69196CFD6EFEC7FD4E" box="[277,358,727,751]" captionStart="FIGURE 16 – 20" captionStartId="6.[151,250,1918,1941]" captionTargetBox="[155,1436,202,1891]" captionTargetId="figure@6.[151,1449,193,1089]" captionTargetPageId="6" captionText="FIGURE 16 – 20. Bibio nigricosta sp. n., male. 16. hind leg (MNHN BK 450). 17. wing. (MNHN BK 450) Position of crossvein m-cu not possible to ascertain in this specimen. 18. terminalia, dorsal view (MNHN BK 450). 19. habitus (MNHN B 47847). 20. female (?), wing (MNHN B 47961). Scale = 1 mm" httpUri="https://zenodo.org/record/281237/files/figure.png" pageId="7" pageNumber="58">Fig. 17</figureCitation>
|
||
)
|
||
<quantity id="4C969B063A7AFE69190CFD6EFE6AFD4E" box="[373,459,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="58" unit="mm" value="7.1">7.1 mm</quantity>
|
||
long,
|
||
<quantity id="4C969B063A7AFE691A68FD6EFDD4FD4E" box="[529,629,727,751]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="7" pageNumber="58" unit="mm" value="3.35">3.35 mm</quantity>
|
||
wide, brown fumose, costal cell from pterostigma and two thirds of distance basad dark. Pterostigma brownish-black, distinctive. Veins distinctive throughout. Crossvein r-m approximately half as long as radial sector. Crossvein m-cu not visible in
|
||
<typeStatus id="54D588413A7AFE691B00FC98FC0CFC98" box="[889,941,801,825]" pageId="7" pageNumber="58">type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FCFDFC2CFC05" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
<collectingCountry id="F37976733A7AFE6918B0FCFDFEA0FCFD" box="[201,257,836,860]" name="India" pageId="7" pageNumber="58">Hind</collectingCountry>
|
||
leg (
|
||
<figureCitation id="13552A663A7AFE691943FCFDFE2FFCFD" box="[314,398,836,860]" captionStart="FIGURE 16 – 20" captionStartId="6.[151,250,1918,1941]" captionTargetBox="[155,1436,202,1891]" captionTargetId="figure@6.[151,1449,193,1089]" captionTargetPageId="6" captionText="FIGURE 16 – 20. Bibio nigricosta sp. n., male. 16. hind leg (MNHN BK 450). 17. wing. (MNHN BK 450) Position of crossvein m-cu not possible to ascertain in this specimen. 18. terminalia, dorsal view (MNHN BK 450). 19. habitus (MNHN B 47847). 20. female (?), wing (MNHN B 47961). Scale = 1 mm" httpUri="https://zenodo.org/record/281237/files/figure.png" pageId="7" pageNumber="58">Fig. 16</figureCitation>
|
||
): Femur
|
||
<quantity id="4C969B063A7AFE69198BFCFDFDE9FCFD" box="[498,584,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="58" unit="mm" value="2.6">2.6 mm</quantity>
|
||
long,
|
||
<quantity id="4C969B063A7AFE691AF6FCFDFD52FCFD" box="[655,755,836,860]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="7" pageNumber="58" unit="mm" value="0.68">0.68 mm</quantity>
|
||
wide, reddish (probably bright red in life), clavate. Tibia 3.0 mm long,
|
||
<quantity id="4C969B063A7AFE691973FCDEFECFFC21" box="[266,366,871,896]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="7" pageNumber="58" unit="mm" value="0.64">0.64 mm</quantity>
|
||
wide, black, truncate apically. First tarsomere
|
||
<quantity id="4C969B063A7AFE691B05FCDEFC40FC21" box="[892,993,871,896]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="58" unit="mm" value="1.24">1.24 mm</quantity>
|
||
long,
|
||
<quantity id="4C969B063A7AFE691C51FCDEFB2CFC21" box="[1064,1165,871,896]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="58" unit="mm" value="0.5">0.50 mm</quantity>
|
||
wide, moderately swollen. Length of second to fifth tarsomeres 0.64, 0.46, 0.32,
|
||
<quantity id="4C969B063A7AFE691B5BFC35FC28FC05" box="[802,905,908,932]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="58" unit="mm" value="0.42">0.42 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FC16FA3AFC66" blockId="7.[151,1437,151,1436]" box="[201,1435,943,968]" pageId="7" pageNumber="58">
|
||
Abdominal tergites quite broad, clearly separated, in this respect similar to the recent species
|
||
<taxonomicName id="4C6E4D603A7AFE691CA6FC09FAC3FC69" box="[1247,1378,944,968]" class="Insecta" family="Bibionidae" genus="Bibio" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="marci">
|
||
<emphasis id="B91AEAF13A7AFE691CA6FC09FAC3FC69" box="[1247,1378,944,968]" italics="true" pageId="7" pageNumber="58">Bibio marci</emphasis>
|
||
</taxonomicName>
|
||
(L.).
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FC6DFEA6FBAE" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Terminalia (
|
||
<figureCitation id="13552A663A7AFE69192BFC6DFE06FC4D" box="[338,423,980,1004]" captionStart="FIGURE 16 – 20" captionStartId="6.[151,250,1918,1941]" captionTargetBox="[155,1436,202,1891]" captionTargetId="figure@6.[151,1449,193,1089]" captionTargetPageId="6" captionText="FIGURE 16 – 20. Bibio nigricosta sp. n., male. 16. hind leg (MNHN BK 450). 17. wing. (MNHN BK 450) Position of crossvein m-cu not possible to ascertain in this specimen. 18. terminalia, dorsal view (MNHN BK 450). 19. habitus (MNHN B 47847). 20. female (?), wing (MNHN B 47961). Scale = 1 mm" httpUri="https://zenodo.org/record/281237/files/figure.png" pageId="7" pageNumber="58">Fig. 18</figureCitation>
|
||
): gonostyli quite robust. Lobes of epandrium rounded, indentation between them broadly V-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FBA5FD65FB01" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Female (specimen B 47847), fig. 19: body and head black, femora reddish. Thorax length
|
||
<quantity id="4C969B063A7AFE691CB0FBA5FA82FB95" box="[1225,1315,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="58" unit="mm" value="3.7">3.7 mm</quantity>
|
||
. Fore tibia rather thick, black (only basal part visible). Wings as in male but larger (only basal half preserved). One large wing (BK 47961,
|
||
<figureCitation id="13552A663A7AFE69195DFBDDFED4FBDD" box="[292,373,1124,1148]" captionStart="FIGURE 16 – 20" captionStartId="6.[151,250,1918,1941]" captionTargetBox="[155,1436,202,1891]" captionTargetId="figure@6.[151,1449,193,1089]" captionTargetPageId="6" captionText="FIGURE 16 – 20. Bibio nigricosta sp. n., male. 16. hind leg (MNHN BK 450). 17. wing. (MNHN BK 450) Position of crossvein m-cu not possible to ascertain in this specimen. 18. terminalia, dorsal view (MNHN BK 450). 19. habitus (MNHN B 47847). 20. female (?), wing (MNHN B 47961). Scale = 1 mm" httpUri="https://zenodo.org/record/281237/files/figure.png" pageId="7" pageNumber="58">Fig. 20</figureCitation>
|
||
) is presumably female. The basal and apical parts are not preserved, but the visible part is
|
||
<quantity id="4C969B063A7AFE691D00FBDDFF62FB01" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="58" unit="mm" value="9.1">9.1 mm</quantity>
|
||
long, longer than the entire wing of the male.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FB15FF69FAAD" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
A partial specimen (BK 21) consists of most of the thorax, the basal part of both wings and the posterior margin of the head including the ocellar tubercle. It is certainly conspecific with the
|
||
<typeStatus id="54D588413A7AFE691C51FB69FB2BFB49" box="[1064,1162,1232,1256]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
but the gender is uncertain.
|
||
</paragraph>
|
||
<paragraph id="8BD136E33A7AFE6918B0FAA1FE3FFA3D" blockId="7.[151,1437,151,1436]" pageId="7" pageNumber="58">
|
||
Remark: It is not possible to see a crossvein m-cu in the wings of the
|
||
<typeStatus id="54D588413A7AFE691BAEFAA1FB98FA91" box="[983,1081,1304,1328]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
male; this may be partly due to the somewhat rough structure of the matrix. Nothing suggests that the vein should be absent in this species and the female specimen figured has a clear m-cu. This vein is probably present in all
|
||
<taxonomicName id="4C6E4D603A7AFE691C73FAD9FB29FAD9" box="[1034,1160,1376,1400]" class="Insecta" family="Bibionidae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="family">Bibionidae</taxonomicName>
|
||
, but is often inconspicuous in fossilized wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |