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<document id="6E125AEB3992AEF9549135F4185209F1" ID-DOI="10.5281/zenodo.281232" ID-GBIF-Dataset="cd359e84-c028-4f73-a84f-841017d87263" ID-ISSN="1175-5326" ID-Zenodo-Dep="281232" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460485267434" checkinUser="plazi" docAuthor="Skartveit, John &amp; Nel, André" docDate="2012" docId="03C787F53A7AFE6618EEFA56FE4DF84F" docLanguage="en" docName="zt03329p063.pdf" docOrigin="Zootaxa 3329" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Penthetria Meigen 1803" docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="FFFEFF8D3A7DFE6E1879FFB9FFA1FFA1" masterDocTitle="Fossil Bibionidae (Diptera: Bibionomorpha) from the late Oligocene of Bes-Konak, Anatolia, Turkey" masterLastPageNumber="63" masterPageNumber="51" pageNumber="58" updateTime="1698308914814" updateUser="plazi">
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<mods:title id="C42A981A04C9F535D8B3B85A387CAE3A">Fossil Bibionidae (Diptera: Bibionomorpha) from the late Oligocene of Bes-Konak, Anatolia, Turkey</mods:title>
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<mods:namePart id="751D9F873EEC524CAC73E3348357F908">Skartveit, John</mods:namePart>
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<mods:namePart id="F31A2EFB0D4B5F4667FBEEF4BED03E6D">Nel, André</mods:namePart>
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<mods:typeOfResource id="602A644075E3BB9E7D99F8531EC29892">text</mods:typeOfResource>
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<mods:date id="CC13A6757CD6D8AA09594F261734803E">2012</mods:date>
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<treatment id="03C787F53A7AFE6618EEFA56FE4DF84F" ID-DOI="http://doi.org/10.5281/zenodo.6181707" ID-GBIF-Taxon="119518102" ID-Zenodo-Dep="6181707" LSID="urn:lsid:plazi:treatment:03C787F53A7AFE6618EEFA56FE4DF84F" httpUri="http://treatment.plazi.org/id/03C787F53A7AFE6618EEFA56FE4DF84F" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<subSubSection id="C37465683A7AFE6918EEFA56FDBAF9A8" box="[151,539,1519,1545]" pageId="7" pageNumber="58" type="nomenclature">
<paragraph id="8BD136E33A7AFE6918EEFA56FDBAF9A8" blockId="7.[151,539,1519,1545]" box="[151,539,1519,1545]" pageId="7" pageNumber="58">
<heading id="D099818F3A7AFE6918EEFA56FDBAF9A8" bold="true" box="[151,539,1519,1545]" fontSize="11" level="1" pageId="7" pageNumber="58" reason="1">
<emphasis id="B91AEAF13A7AFE6918EEFA56FDBAF9A8" bold="true" box="[151,539,1519,1545]" pageId="7" pageNumber="58">
Genus
<taxonomicName id="4C6E4D603A7AFE691897FA56FDBAF9A8" ID-CoL="8YQLD" ID-ENA="560770" authority="Meigen, 1803" authorityName="Meigen" authorityYear="1803" box="[238,539,1519,1545]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF13A7AFE691897FA56FECCF9A8" bold="true" box="[238,365,1519,1545]" italics="true" pageId="7" pageNumber="58">Penthetria</emphasis>
<bibRefCitation id="EFFF4B123A7AFE69190DFA56FDBAF9A8" author="Meigen" box="[372,539,1519,1545]" pageId="7" pageNumber="58" refString="Meigen, J. W. (1803) Versuch einer neuer GattungsEintheilung der europaischen zweiflugeligen Insecten. Magazin fur Insektenkunde (Illiger), 2, 259 - 281." type="journal article" year="1803">Meigen, 1803</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37465683A7AFE6918EEF98EFD39F96F" pageId="7" pageNumber="58" type="reference_group">
<paragraph id="8BD136E33A7AFE6918EEF98EFEA2F9CA" blockId="7.[151,1436,1590,1643]" pageId="7" pageNumber="58">
<treatmentCitationGroup id="AB7E11CD3A7AFE6918EEF98EFEA2F9CA" pageId="7" pageNumber="58">
<taxonomicName id="4C6E4D603A7AFE6918EEF98EFE73F9ED" ID-CoL="8YQLD" authority="Meigen, 1803: 264" authorityName="Meigen" authorityPageNumber="264" authorityYear="1803" box="[151,466,1590,1613]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF13A7AFE6918EEF98EFEA3F9EC" box="[151,258,1591,1613]" italics="true" pageId="7" pageNumber="58">Penthetria</emphasis>
<treatmentCitation id="0ACF10F23A7AFE691973F98FFE73F9ED" author="Meigen" box="[266,466,1590,1612]" page="264" pageId="7" pageNumber="58" year="1803">
<bibRefCitation id="EFFF4B123A7AFE691973F98FFE3DF9ED" author="Meigen" box="[266,412,1590,1612]" pageId="7" pageNumber="58" refString="Meigen, J. W. (1803) Versuch einer neuer GattungsEintheilung der europaischen zweiflugeligen Insecten. Magazin fur Insektenkunde (Illiger), 2, 259 - 281." type="journal article" year="1803">Meigen, 1803</bibRefCitation>
: 264
</treatmentCitation>
</taxonomicName>
. Type-species:
<taxonomicName id="4C6E4D603A7AFE691A08F98EFBA5F9ED" ID-CoL="6V23R" authority="Meigen, 1804: 104" authorityName="Meigen" authorityPageNumber="104" authorityYear="1804" box="[625,1028,1590,1613]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="B91AEAF13A7AFE691A08F98EFC96F9EC" box="[625,823,1591,1613]" italics="true" pageId="7" pageNumber="58">Penthetria funebris</emphasis>
<treatmentCitation id="0ACF10F23A7AFE691B46F98FFBA5F9ED" author="Meigen" box="[831,1028,1590,1612]" page="104" pageId="7" pageNumber="58" year="1804">
<bibRefCitation id="EFFF4B123A7AFE691B46F98FFC72F9ED" author="Meigen" box="[831,979,1590,1612]" pageId="7" pageNumber="58" refString="Meigen, J. W. (1804) Klassifikation und Beschreibung der europaischen zweiflugeligen Insekten. (Diptera Linn.). Erster Band. Abt. I. Reichard, Braunschweig." type="book" year="1804">Meigen, 1804</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
(by subsequent designation and monotypy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD136E33A7AFE6918EEF90FFD39F96F" blockId="7.[151,664,1718,1743]" box="[151,664,1718,1743]" pageId="7" pageNumber="58">
<taxonomicName id="4C6E4D603A7AFE6918EEF90FFE1FF96E" box="[151,446,1718,1743]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="beskonakensis" status="sp. nov.">
<emphasis id="B91AEAF13A7AFE6918EEF90FFE1FF96E" bold="true" box="[151,446,1718,1743]" italics="true" pageId="7" pageNumber="58">Penthetria beskonakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229578A3A7AFE6919BDF90EFDA1F96F" box="[452,512,1719,1742]" pageId="7" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
(figs. 2128)
</paragraph>
</subSubSection>
<subSubSection id="C37465683A7AFE6618EEF947FAB5FE74" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58" type="diagnosis">
<paragraph id="8BD136E33A7AFE6918EEF947FBD3F8C0" blockId="7.[151,1437,1790,2033]" pageId="7" pageNumber="58">Diagnosis. Wing rather long and narrow, R2+3 long, subparallel to R4+5. Antenna robust and relatively long, flagellum 12-segmented. Male terminalia with robust gonostyli, epandrium bilobate, lobes rather narrowly separated. Female cerci two-segmented, basal segment large and triangular, distal segment slender.</paragraph>
<paragraph id="8BD136E33A7AFE6918B0F8D4FAFCF850" blockId="7.[151,1437,1790,2033]" pageId="7" pageNumber="58">
Material:
<typeStatus id="54D588413A7AFE69194FF8D4FE39F824" box="[310,408,1901,1925]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
(male) Labelled 8C, fig. 21. The
<typeStatus id="54D588413A7AFE691B77F8D4FCD1F824" box="[782,880,1901,1925]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
is smaller than the other specimens available (
<quantity id="4C969B063A7AFE691D01F8D4FF62F808" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="7" pageNumber="58" unit="mm" value="7.1">7.1 mm</quantity>
vs.
<quantity id="4C969B063A7AFE691889F828FED9F808" box="[240,376,1937,1961]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="7" pageNumber="58" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">8.59.5 mm</quantity>
long) but otherwise is quite similar to them. The colours of the
<typeStatus id="54D588413A7AFE691C3DF828FB07F808" box="[1092,1190,1937,1961]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
appear faded but otherwise the state of preservation is very good. The apical parts of the wings are covered by rock. Other material: five males (BK 257, BK 442, B 47593, B 47598, B 47748), four females (B 47737, B 47793, B 47981, B 48072).
</paragraph>
<paragraph id="8BD136E33A75FE6618B0FF2EFC1BFF75" blockId="8.[151,1436,151,469]" pageId="8" pageNumber="59">
Description:
<emphasis id="B91AEAF13A75FE661925FF21FE38FF11" bold="true" box="[348,409,152,176]" pageId="8" pageNumber="59">Male</emphasis>
: body, legs and wings all black. Total length
<quantity id="4C969B063A75FE661BDFFF2EFB8EFF0E" box="[934,1071,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.5" metricValueMin="7.1" pageId="8" pageNumber="59" unit="mm" value="8.3" valueMax="9.5" valueMin="7.1">7.19.5 mm</quantity>
(N = 3). Head width
<quantity id="4C969B063A75FE661D55FF2EFF62FF75" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.08" metricValueMin="0.95" pageId="8" pageNumber="59" unit="mm" value="1.015" valueMax="1.08" valueMin="0.95">0.951.08 mm</quantity>
(N = 2). Thorax length 2.0
<quantity id="4C969B063A75FE661A79FF05FDF7FF75" box="[512,598,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="59" unit="mm" value="3.2">3.2 mm</quantity>
(N = 3), width
<quantity id="4C969B063A75FE661B7DFF05FCFBFF75" box="[772,858,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="59" unit="mm" value="2.1">2.1 mm</quantity>
(N = 1).
</paragraph>
<paragraph id="8BD136E33A75FE6618B0FF59FE2DFEBD" blockId="8.[151,1436,151,469]" pageId="8" pageNumber="59">
Legs: All parts slender. Length of fore tibia
<quantity id="4C969B063A75FE661AC7FF66FCC3FF56" box="[702,866,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.05" metricValueMin="2.25" pageId="8" pageNumber="59" unit="mm" value="2.65" valueMax="3.05" valueMin="2.25">2.253.05 mm</quantity>
(N = 3), mid tibia 1.62.0 mm (N = 2), hind tibia
<quantity id="4C969B063A75FE6618EEFEBDFE8DFEBD" box="[151,300,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="3.2" metricValueMin="2.55" pageId="8" pageNumber="59" unit="mm" value="2.875" valueMax="3.2" valueMin="2.55">2.553.2 mm</quantity>
(N = 3).
</paragraph>
<paragraph id="8BD136E33A75FE6618B0FE9EFBACFEC6" blockId="8.[151,1436,151,469]" pageId="8" pageNumber="59">
Wing (
<figureCitation id="13552A663A75FE66196FFE9EFEC9FE9E" box="[278,360,295,319]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 22</figureCitation>
) 5.0
<quantity id="4C969B063A75FE6619D0FE9EFE5EFE9E" box="[425,511,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="59" unit="mm" value="6.7">6.7 mm</quantity>
long (N = 3), rather narrow, pterostigma distinctive. Vein R2+3 long, subparallel to R4+
<quantity id="4C969B063A75FE6618C3FEE7FF7EFEC6" box="[186,223,335,364]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="8" pageNumber="59" unit="in" value="5.0">5 in</quantity>
basal half. Crossvein r-m vertical, length approximately one sixth of Rs.
</paragraph>
<paragraph id="8BD136E33A75FE6618B0FECCFBBEFE2C" blockId="8.[151,1436,151,469]" box="[201,1055,373,397]" pageId="8" pageNumber="59">Abdomen slender, slightly conical. Pile not visible in the specimens at hand.</paragraph>
<paragraph id="8BD136E33A75FE6618B0FE20FAB5FE74" blockId="8.[151,1436,151,469]" pageId="8" pageNumber="59">
Terminalia (
<figureCitation id="13552A663A75FE66192BFE20FE06FE10" box="[338,423,409,433]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 23</figureCitation>
): epandrium strongly bilobed, the two lobes slightly pointed, separated by a deep but narrow, U-shaped cleft. Gonostyli slender, mesally straight, distinctly curved only in apical third, pointed.
</paragraph>
</subSubSection>
<caption id="DF11666B3A75FE6618EEFA78FC47FA79" box="[151,998,1473,1496]" httpUri="https://zenodo.org/record/281238/files/figure.png" pageId="8" pageNumber="59" targetBox="[151,1436,506,1460]" targetPageId="8">
<paragraph id="8BD136E33A75FE6618EEFA78FC47FA79" blockId="8.[151,998,1473,1496]" box="[151,998,1473,1496]" pageId="8" pageNumber="59">
<emphasis id="B91AEAF13A75FE6618EEFA78FEBEFA76" bold="true" box="[151,287,1473,1496]" pageId="8" pageNumber="59">FIGURE 21.</emphasis>
<taxonomicName id="4C6E4D603A75FE66195CFA7BFD89FA79" box="[293,552,1474,1496]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="beskonakensis" status="sp. nov.">
<emphasis id="B91AEAF13A75FE66195CFA7BFD89FA79" box="[293,552,1474,1496]" italics="true" pageId="8" pageNumber="59">Penthetria beskonakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229578A3A75FE661A57FA7AFDFEFA79" box="[558,607,1475,1496]" pageId="8" pageNumber="59" rank="species">sp.n.</taxonomicNameLabel>
, male, holotype (MHNH BK 223 8C).
</paragraph>
</caption>
<subSubSection id="C37465683A75FE6618B0F9B9FE4DF84F" pageId="8" pageNumber="59" type="description">
<paragraph id="8BD136E33A75FE6618B0F9B9FE78F9FE" blockId="8.[151,1436,1535,2030]" pageId="8" pageNumber="59">
<emphasis id="B91AEAF13A75FE6618B0F9B9FE81F9B9" bold="true" box="[201,288,1536,1560]" pageId="8" pageNumber="59">Female</emphasis>
: Habitus as in fig. 24, total length
<quantity id="4C969B063A75FE661ADEFA46FC9CF9B6" box="[679,829,1535,1559]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.27" metricValueMin="0.95" pageId="8" pageNumber="59" unit="mm" value="11.1" valueMax="12.7" valueMin="9.5">9.512.7 mm</quantity>
(N = 3). Head width
<quantity id="4C969B063A75FE661C56FA46FB35F9B6" box="[1071,1172,1535,1559]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="59" unit="mm" value="1.05">1.05 mm</quantity>
(N = 1). Head and thorax as in
<figureCitation id="13552A663A75FE661884F99AFEF1F99A" box="[253,336,1571,1595]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 25</figureCitation>
. Antenna (
<figureCitation id="13552A663A75FE6619B3F99AFDBAF99A" box="[458,539,1571,1595]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 26</figureCitation>
) robust with 12-segmented flagellum, the segments strongly transverse. Thorax length
<quantity id="4C969B063A75FE66189DF9FEFED9F9FE" box="[228,376,1607,1631]" metricMagnitude="-3" metricUnit="m" metricValue="2.475" metricValueMax="2.7" metricValueMin="2.25" pageId="8" pageNumber="59" unit="mm" value="2.475" valueMax="2.7" valueMin="2.25">2.252.7 mm</quantity>
(N = 2).
</paragraph>
<paragraph id="8BD136E33A75FE6618B0F9D5FCC8F922" blockId="8.[151,1436,1535,2030]" box="[201,873,1643,1668]" pageId="8" pageNumber="59">
Legs: as in male. Length of hind tibia
<quantity id="4C969B063A75FE661A0DF9D2FCA8F922" box="[628,777,1643,1667]" metricMagnitude="-3" metricUnit="m" metricValue="3.325" metricValueMax="3.5" metricValueMin="3.15" pageId="8" pageNumber="59" unit="mm" value="3.325" valueMax="3.5" valueMin="3.15">3.153.5 mm</quantity>
(N = 2).
</paragraph>
<paragraph id="8BD136E33A75FE6618B0F936FB64F96A" blockId="8.[151,1436,1535,2030]" pageId="8" pageNumber="59">
Wing (
<figureCitation id="13552A663A75FE66196FF936FEC9F906" box="[278,360,1679,1703]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 27</figureCitation>
) length 10.0 mm (N = 1), width 4.0 mm. Wider than in male. Pterostigma apparently indistinctive, but this may be due to fading. R2+3 long, rather straight. Crossvein r-m longer than in male.
</paragraph>
<paragraph id="8BD136E33A75FE6618B0F963FDDBF8B7" blockId="8.[151,1436,1535,2030]" pageId="8" pageNumber="59">
Abdomen length
<quantity id="4C969B063A75FE6619F7F963FDB6F953" box="[398,535,1754,1778]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="8" pageNumber="59" unit="mm" value="8.0" valueMax="9.5" valueMin="6.5">6.59.5 mm</quantity>
(N = 2), rather large, wider than thorax in the specimens at hand. Tergites and sternites with rather dense, short, dark pile.
</paragraph>
<paragraph id="8BD136E33A75FE6618B0F89BFD50F8FF" blockId="8.[151,1436,1535,2030]" pageId="8" pageNumber="59">
Terminalia (
<figureCitation id="13552A663A75FE66192AF89BFE08F89B" box="[339,425,1826,1850]" captionStart="FIGURE 22 28" captionStartId="9.[151,250,1961,1984]" captionTargetBox="[255,1326,153,1949]" captionTargetId="figure@9.[258,1328,149,1246]" captionTargetPageId="9" captionText="FIGURE 22 28. Penthetria beskonakensis sp. n. 22. male, wing (MNHN B 47593). 23. male, terminalia, dorsal view (MNHN BK 223 8 C). 24. female, habitus (MNHN B 47793). 25. female, head and thorax (MNHN B 47981). 26. female, antenna (MNHN BK 614). 27. female, wing (MNHN BK 614). 28. female, terminalia (MNHN BK 614). Scale = 1 mm." httpUri="https://zenodo.org/record/281239/files/figure.png" pageId="8" pageNumber="59">Fig. 28</figureCitation>
): cerci two-segmented, basal segment wide and triangular, terminal segment narrow and pointed. Tergite IX with slightly concave hind border.
</paragraph>
<paragraph id="8BD136E33A75FE6618B0F8D3FE4DF84F" blockId="8.[151,1436,1535,2030]" pageId="8" pageNumber="59">
Remark: The male and female have conspicuously differently shaped wings, but a marked sexual dimorphism in wing shape is common also among extant
<taxonomicName id="4C6E4D603A75FE661AE7F837FCB5F807" box="[670,788,1934,1958]" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAF13A75FE661AE7F837FCB5F807" box="[670,788,1934,1958]" italics="true" pageId="8" pageNumber="59">Penthetria</emphasis>
</taxonomicName>
species, being most extreme in the European
<taxonomicName id="4C6E4D603A75FE661D5FF837FE3CF86B" authority="Meigen, 1804" authorityName="Meigen" authorityYear="1804" class="Insecta" family="Bibionidae" genus="Penthetria" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="funebris">
<emphasis id="B91AEAF13A75FE661D5FF837FF52F86B" italics="true" pageId="8" pageNumber="59">Penthetria funebris</emphasis>
<bibRefCitation id="EFFF4B123A75FE661884F80BFE3CF86B" author="Meigen" box="[253,413,1970,1994]" pageId="8" pageNumber="59" refString="Meigen, J. W. (1804) Klassifikation und Beschreibung der europaischen zweiflugeligen Insekten. (Diptera Linn.). Erster Band. Abt. I. Reichard, Braunschweig." type="book" year="1804">Meigen, 1804</bibRefCitation>
</taxonomicName>
where the male is strongly brachypterous and flightless while the female has normally shaped, although short, wings.
</paragraph>
</subSubSection>
</treatment>
</document>