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<document id="344B25FE66E71D41085E9B73EF5B8FD9" ID-CLB-Dataset="32101" ID-DOI="10.11646/zootaxa.4320.1.5" ID-GBIF-Dataset="b2a9ea86-343f-4690-8bec-0bef8965d17a" ID-ISSN="1175-5326" ID-Zenodo-Dep="891584" ID-ZooBank="C6Aa0B10-E6Cb-4B49-8Efb-Bc183B69354B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505452152722" checkinUser="plazi" docAuthor="Morse, John C." docDate="2017" docId="03C787EF7500FFD9FF39CB2FFBDCDB5B" docLanguage="en" docName="zooaxa.4320.1.5.pdf.imf" docOrigin="Zootaxa 4320 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marilia simulans Forsslund 1935" docType="treatment" docVersion="7" lastPageNumber="98" masterDocId="FFFEFF977510FFC8FFAECE52FFD8D943" masterDocTitle="Lannapsyche and Marilia species of China (Trichoptera: Odontoceridae)" masterLastPageNumber="99" masterPageNumber="81" pageNumber="97" updateTime="1698468448558" updateUser="ExternalLinkService">
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<mods:title id="06E8F6B5E1E68DA0E9358DE5895F7CAF">Lannapsyche and Marilia species of China (Trichoptera: Odontoceridae)</mods:title>
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<treatment id="03C787EF7500FFD9FF39CB2FFBDCDB5B" ID-DOI="http://doi.org/10.5281/zenodo.6032279" ID-GBIF-Taxon="134089641" ID-Zenodo-Dep="6032279" LSID="urn:lsid:plazi:treatment:03C787EF7500FFD9FF39CB2FFBDCDB5B" httpUri="http://treatment.plazi.org/id/03C787EF7500FFD9FF39CB2FFBDCDB5B" lastPageId="17" lastPageNumber="98" pageId="16" pageNumber="97">
<subSubSection id="C37465727500FFD8FF39CB2FFF20DCFB" pageId="16" pageNumber="97" type="nomenclature">
<paragraph id="8BD136F97500FFD8FF39CB2FFDF6DCD4" blockId="16.[151,558,1405,1464]" box="[151,558,1405,1431]" pageId="16" pageNumber="97">
<heading id="D09981957500FFD8FF39CB2FFDF6DCD4" bold="true" box="[151,558,1405,1431]" fontSize="11" level="1" pageId="16" pageNumber="97" reason="1">
<taxonomicName id="4C6E4D7A7500FFD8FF39CB2FFDF6DCD4" ID-CoL="3Y6JY" authority="Forsslund 1935" authorityName="Forsslund" authorityYear="1935" box="[151,558,1405,1431]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7500FFD8FF39CB2FFDF6DCD4" bold="true" box="[151,558,1405,1431]" pageId="16" pageNumber="97">
<emphasis id="B91AEAEB7500FFD8FF39CB2FFEBEDCD4" bold="true" box="[151,358,1405,1431]" italics="true" pageId="16" pageNumber="97">Marilia simulans</emphasis>
<bibRefCitation id="EFFF4B087500FFD8FEC0CB2FFDF6DCD4" author="Forsslund" box="[366,558,1405,1431]" pageId="16" pageNumber="97" refString="Forsslund, K. - H. (1935) Swedisch-chinesische wissenshaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin and Prof. Su Ping-Chang. Insecten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 50: Trichoptera. Archiv for Zoologi, 27 A (31), 1 - 21." type="journal article" year="1935">Forsslund 1935</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BD136F97500FFD8FF39CBCDFF20DCFB" blockId="16.[151,558,1405,1464]" box="[151,248,1439,1464]" pageId="16" pageNumber="97">
(
<figureCitation id="13552A7C7500FFD8FF31CBCDFF29DCFB" box="[159,241,1439,1464]" captionStart="FIGURE 12" captionStartId="17.[151,250,1533,1555]" captionTargetBox="[163,1427,603,1498]" captionTargetId="figure@17.[151,1436,591,1511]" captionTargetPageId="17" captionText="FIGURE 12. Marilia simulans Forsslund 1935, male genitalia, head, and wings. 12 A 12 B, male genitalia (after Forsslund 1935): 12 A, right lateral; 12 B, ventral; 12 C, right forewing, dorsal (after Forsslund 1935). 12 E 12 F, male head and wings (original drawings from holotype): 12 E, head, dorsal; 12 Fa 12 Fp, left wings, ventral: 12 Fa, forewing; 12 Fp, hind wing." httpUri="https://zenodo.org/record/891604/files/figure.png" pageId="16" pageNumber="97">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37465727500FFD8FF39CBBAFB9CDCBD" box="[151,1092,1512,1534]" pageId="16" pageNumber="97" type="reference_group">
<paragraph id="8BD136F97500FFD8FF39CBBAFB9CDCBD" blockId="16.[151,1092,1512,1534]" box="[151,1092,1512,1534]" pageId="16" pageNumber="97">
<materialsCitation id="3B063CA47500FFD8FF39CBBAFB9CDCBD" ID-GBIF-Occurrence="1638471353" box="[151,1092,1512,1534]" country="China" location="Si-chuan" pageId="16" pageNumber="97" specimenCount="1" typeStatus="holotype">
<treatmentCitationGroup id="AB7E11D77500FFD8FF39CBBAFB9CDCBD" box="[151,1092,1512,1534]" pageId="16" pageNumber="97">
<taxonomicName id="4C6E4D7A7500FFD8FF39CBBAFDD7DCBD" ID-CoL="3Y6JY" authority="Forsslund 1935: 27" authorityName="Forsslund" authorityPageNumber="27" authorityYear="1935" box="[151,527,1512,1534]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7500FFD8FF39CBBAFE9DDCBD" box="[151,325,1512,1534]" italics="true" pageId="16" pageNumber="97">Marilia simulans</emphasis>
<treatmentCitation id="0ACF10E87500FFD8FEE5CBBAFDD7DCBD" author="Forsslund" box="[331,527,1512,1534]" page="27" pageId="16" pageNumber="97" year="1935">
<bibRefCitation id="EFFF4B087500FFD8FEE5CBBAFE33DCBD" author="Forsslund" box="[331,491,1512,1534]" pageId="16" pageNumber="97" refString="Forsslund, K. - H. (1935) Swedisch-chinesische wissenshaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin and Prof. Su Ping-Chang. Insecten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 50: Trichoptera. Archiv for Zoologi, 27 A (31), 1 - 21." type="journal article" year="1935">Forsslund 1935</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
A no 31, 78, f 6a6c.
<typeStatus id="54D5885B7500FFD8FD53CBBBFCF7DCBD" box="[765,815,1513,1534]" pageId="16" pageNumber="97">Type</typeStatus>
locality:
<collectingCountry id="F37976697500FFD8FC21CBBAFC13DCBD" box="[911,971,1512,1534]" name="China" pageId="16" pageNumber="97">China</collectingCountry>
(
<location id="8EB160227500FFD8FC77CBBAFBE1DCBD" LSID="urn:lsid:plazi:treatment:03C787EF7500FFD9FF39CB2FFBDCDB5B:8EB160227500FFD8FC77CBBAFBE1DCBD" box="[985,1081,1512,1534]" country="China" name="Si-chuan" pageId="16" pageNumber="97">Si-chuan</location>
).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37465727500FFD8FF39C878FD6FDEC4" pageId="16" pageNumber="97" type="materials_examined">
<paragraph id="8BD136F97500FFD8FF39C878FD6FDEC4" blockId="16.[151,1437,1578,2035]" pageId="16" pageNumber="97">
<emphasis id="B91AEAEB7500FFD8FF39C878FEA7DF00" bold="true" box="[151,383,1578,1603]" pageId="16" pageNumber="97">Material examined</emphasis>
. Holotype male with the following labels: Pencil label handwritten with cursive writing appearing to be the following: “ror 7” Pencil label, handwritten with upper and lower case block letters: “Sven Hedins exp. [indecipherable] CenterAsien Leg. Dr. Hummel.” Printed label: “N.O. Szechuan: Ufer von Kia-ling-ho swischen Naupo Und Pao-ning-fu
<date id="FFD010397500FFD8FDB0C8C4FD48DFEC" box="[542,656,1686,1711]" pageId="16" pageNumber="97" value="1930-05-16">16.V.1930</date>
[Hummel leg.]” Pencil label, handwritten with upper and lower case block letters: “
<taxonomicName id="4C6E4D7A7500FFD8FE90C8E8FD8EDF90" authority="Forssl." authorityName="Forsslund" authorityYear="1935" box="[318,598,1722,1747]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="simulans">Marilia simulans Forssl.</taxonomicName>
Typus ♂
<emphasis id="B91AEAEB7500FFD8FD65C8E8FCD4DF90" bold="true" box="[715,780,1722,1747]" pageId="16" pageNumber="97">n. sp.</emphasis>
” Printed label: “NHRS-TOBI 000000856” Printed label: “Loan 0 0 1/2016 NHRS-TOBI 000000856” This holotype specimen is missing the following body parts: Left flagellum beyond segment 3; Maxillary and labial palps; Left foreleg; Right wings; Abdominal segments VIX. The remaining specimen has been cleared and is mostly transparent. The left wings remain, but are frayed, without setae other than the bristles on the forewing anal vein, and the dorsal and ventral cuticular layers have separated, so that parts of these wings could not be illustrated.
</paragraph>
</subSubSection>
<subSubSection id="C37465727500FFD9FF69C9C0FEB5D8B6" lastPageId="17" lastPageNumber="98" pageId="16" pageNumber="97" type="discussion">
<paragraph id="8BD136F97500FFD9FF69C9C0FEB5D8B6" blockId="16.[151,1437,1578,2035]" lastBlockId="17.[151,1437,151,536]" lastPageId="17" lastPageNumber="98" pageId="16" pageNumber="97">
<emphasis id="B91AEAEB7500FFD8FF69C9C0FEE3DEE8" bold="true" box="[199,315,1938,1963]" pageId="16" pageNumber="97">Remarks.</emphasis>
The wing venation of this species closely resembles that of
<taxonomicName id="4C6E4D7A7500FFD8FC78C9C1FB87DEE8" box="[982,1119,1939,1963]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="97" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7500FFD8FC78C9C1FC35DEE9" box="[982,1005,1939,1962]" italics="true" pageId="16" pageNumber="97">M</emphasis>
.
<emphasis id="B91AEAEB7500FFD8FC55C9C1FB87DEE8" box="[1019,1119,1939,1963]" italics="true" pageId="16" pageNumber="97">discaulis</emphasis>
</taxonomicName>
. Each hind wing has a small posterobasal lobe (not shown in
<figureCitation id="13552A7C7500FFD8FDA7C9E4FD81DE8C" box="[521,601,1974,1999]" captionStart="FIGURE 12" captionStartId="17.[151,250,1533,1555]" captionTargetBox="[163,1427,603,1498]" captionTargetId="figure@17.[151,1436,591,1511]" captionTargetPageId="17" captionText="FIGURE 12. Marilia simulans Forsslund 1935, male genitalia, head, and wings. 12 A 12 B, male genitalia (after Forsslund 1935): 12 A, right lateral; 12 B, ventral; 12 C, right forewing, dorsal (after Forsslund 1935). 12 E 12 F, male head and wings (original drawings from holotype): 12 E, head, dorsal; 12 Fa 12 Fp, left wings, ventral: 12 Fa, forewing; 12 Fp, hind wing." httpUri="https://zenodo.org/record/891604/files/figure.png" pageId="16" pageNumber="97">Fig. 12</figureCitation>
Fp); any long basal brush of setae is missing, but the lobe has many setal alveoli, suggesting that the brush was present in the undamaged specimen. Also, the compound eyes are separated from each other and the intervening setal warts are poorly defined, as in
<taxonomicName id="4C6E4D7A7501FFD9FC11CECBFB9FD9F3" box="[959,1095,152,176]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7501FFD9FC11CECBFC0ED9F3" box="[959,982,153,176]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FC4ACECAFB9FD9F3" box="[996,1095,152,176]" italics="true" pageId="17" pageNumber="98">discaulis</emphasis>
</taxonomicName>
. The genitalia were illustrated in lateral and ventral views by
<bibRefCitation id="EFFF4B087501FFD9FDA6CEEEFD08D996" author="Forsslund" box="[520,720,188,213]" pageId="17" pageNumber="98" refString="Forsslund, K. - H. (1935) Swedisch-chinesische wissenshaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin and Prof. Su Ping-Chang. Insecten gesammelt vom schwedischen Arzt der Expedition Dr. David Hummel 1927 - 1930. 50: Trichoptera. Archiv for Zoologi, 27 A (31), 1 - 21." type="journal article" year="1935">Forsslund (1935)</bibRefCitation>
.
<taxonomicName id="4C6E4D7A7501FFD9FD70CEEEFC7BD997" authorityName="Forsslund" authorityYear="1935" box="[734,931,188,212]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7501FFD9FD70CEEEFC7BD997" box="[734,931,188,212]" italics="true" pageId="17" pageNumber="98">Marilia simulans</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C6E4D7A7501FFD9FBEDCEEFFB08D997" box="[1091,1232,188,212]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7501FFD9FBEDCEEFFB82D997" box="[1091,1114,189,212]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FBC2CEEEFB08D997" box="[1132,1232,188,212]" italics="true" pageId="17" pageNumber="98">discaulis</emphasis>
</taxonomicName>
by the following characters: (1) the lateral view of tergum X was depicted as a slender and acute structure in
<taxonomicName id="4C6E4D7A7501FFD9FB6FCEB3FA94D9BB" authorityName="Forsslund" authorityYear="1935" box="[1217,1356,224,248]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7501FFD9FB6FCEB3FA94D9BB" box="[1217,1356,224,248]" italics="true" pageId="17" pageNumber="98">M. simulans</emphasis>
</taxonomicName>
, but is nearly rounded in
<taxonomicName id="4C6E4D7A7501FFD9FECACF57FE34D85F" box="[356,492,260,284]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7501FFD9FECACF57FEA3D85F" box="[356,379,261,284]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FE27CF56FE34D85F" box="[393,492,260,284]" italics="true" pageId="17" pageNumber="98">discaulis</emphasis>
</taxonomicName>
(
<figureCitation id="13552A7C7501FFD9FE52CF56FDBCD85F" box="[508,612,260,285]" captionStart="FIGURE 10" captionStartId="14.[151,250,1388,1410]" captionTargetBox="[166,1418,452,1355]" captionTargetId="figure@14.[151,1436,447,1366]" captionTargetPageId="14" captionText="FIGURE 10. Marilia discaulis new species, male genitalia, head, and wings. 10 A 10 D, male genitalia: 10 A, left lateral; 10 B, dorsal; 10 Bv, tergum X, ventral; 10 C, left inferior appendage (paired), ventral; 10 D, phallus, left lateral; 10 E, head, dorsal. 10 El, left antennal base, lateral. 10 Fa 10 Fp, right wings, dorsal: 10 Fa, forewing; 10 Fp, hind wing." httpUri="https://zenodo.org/record/891600/files/figure.png" pageId="17" pageNumber="98">Fig. 10A</figureCitation>
); 2) the harpago of each inferior appendage is about 3 times as long as its width and is almost 1/3 as long as the coxopodite in lateral or ventral views in
<taxonomicName id="4C6E4D7A7501FFD9FBADCF7BFB54D803" authorityName="Forsslund" authorityYear="1935" box="[1027,1164,296,320]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7501FFD9FBADCF7BFB54D803" box="[1027,1164,296,320]" italics="true" pageId="17" pageNumber="98">M. simulans</emphasis>
</taxonomicName>
, but it is only 2 times as long as its width and is not quite 1/5 as long as the coxopodite in lateral or ventral views in
<taxonomicName id="4C6E4D7A7501FFD9FB20CF1FFACFD827" box="[1166,1303,332,356]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7501FFD9FB20CF1FFACFD827" box="[1166,1303,332,356]" italics="true" pageId="17" pageNumber="98">M. discaulis</emphasis>
</taxonomicName>
(
<figureCitation id="13552A7C7501FFD9FA88CF1EFF11D8CB" captionStart="FIGURE 10" captionStartId="14.[151,250,1388,1410]" captionTargetBox="[166,1418,452,1355]" captionTargetId="figure@14.[151,1436,447,1366]" captionTargetPageId="14" captionText="FIGURE 10. Marilia discaulis new species, male genitalia, head, and wings. 10 A 10 D, male genitalia: 10 A, left lateral; 10 B, dorsal; 10 Bv, tergum X, ventral; 10 C, left inferior appendage (paired), ventral; 10 D, phallus, left lateral; 10 E, head, dorsal. 10 El, left antennal base, lateral. 10 Fa 10 Fp, right wings, dorsal: 10 Fa, forewing; 10 Fp, hind wing." httpUri="https://zenodo.org/record/891600/files/figure.png" pageId="17" pageNumber="98">Figs. 10A, 10C</figureCitation>
). Despite these apparent differences, until modern specimens of
<taxonomicName id="4C6E4D7A7501FFD9FC05CF23FBEDD8CB" authorityName="Forsslund" authorityYear="1935" box="[939,1077,368,392]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7501FFD9FC05CF23FC1AD8CB" box="[939,962,369,392]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FC7FCF22FBEDD8CB" box="[977,1077,368,392]" italics="true" pageId="17" pageNumber="98">simulans</emphasis>
</taxonomicName>
can be found, we cannot know with confidence whether
<taxonomicName id="4C6E4D7A7501FFD9FE1ECFC7FDE1D8EF" authorityName="Forsslund" authorityYear="1935" box="[432,569,404,428]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="B91AEAEB7501FFD9FE1ECFC7FE1FD8EF" box="[432,455,405,428]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FE7BCFC6FDE1D8EF" box="[469,569,404,428]" italics="true" pageId="17" pageNumber="98">simulans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D7A7501FFD9FDC0CFC7FD2DD8EF" box="[622,757,404,428]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="98" phylum="Arthropoda" rank="species" species="discaulis">
<emphasis id="B91AEAEB7501FFD9FDC0CFC7FD5DD8EF" box="[622,645,405,428]" italics="true" pageId="17" pageNumber="98">M</emphasis>
.
<emphasis id="B91AEAEB7501FFD9FD3CCFC6FD2DD8EF" box="[658,757,404,428]" italics="true" pageId="17" pageNumber="98">discaulis</emphasis>
</taxonomicName>
are different species. Because of the need to be able to speak about readily identifiable species, we choose to consider them distinct species unless and until better evidence becomes available.
</paragraph>
</subSubSection>
<subSubSection id="C37465727501FFD9FF69CFADFBDCDB5B" box="[199,1028,511,536]" pageId="17" pageNumber="98" type="distribution">
<paragraph id="8BD136F97501FFD9FF69CFADFBDCDB5B" blockId="17.[151,1437,151,536]" box="[199,1028,511,536]" pageId="17" pageNumber="98">
<emphasis id="B91AEAEB7501FFD9FF69CFADFEB8DB5B" bold="true" box="[199,352,511,536]" pageId="17" pageNumber="98">Distribution.</emphasis>
Oriental Biogeographic Region, southern
<collectingCountry id="F37976697501FFD9FC92CFADFCA7DB5B" box="[828,895,511,536]" name="China" pageId="17" pageNumber="98">China</collectingCountry>
(Si-chuan).
</paragraph>
</subSubSection>
</treatment>
</document>