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<document id="8022ECCDBCD1391B8EA05EF25C8FB24A" ID-DOI="10.1649/0010-065X(2003)057[0017:FDFLSO]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10103378" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630161078303" checkinUser="tatiana" docAuthor="McHugh, Joseph V. &amp; Kiselyova, Tatiana G." docDate="2003" docId="03C787EA015DFFEC9DE79421FC0C482D" docLanguage="en" docName="0010-065X_2003_057_0017_FDFLSO_2.0.CO_2.pdf.imf" docOrigin="The Coleopterists Bulletin 57 (1)" docSource="http://dx.doi.org/10.1649/0010-065x(2003)057[0017:fdflso]2.0.co;2" docStyle="DocumentStyle:07711681861DB98305E87147CAB6DAD5.4:TheColeopBull.2003-2009.journal_article.0cover" docStyleId="07711681861DB98305E87147CAB6DAD5" docStyleName="TheColeopBull.2003-2009.journal_article.0cover" docStyleVersion="4" docTitle="Eurysphindus LeConte" docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="FFFEFF92015CFFEF9C6B951EFFDE4D5B" masterDocTitle="First Descriptions for Larval Stages of Eurysphindus (Coleoptera: Cucujoidea: Sphindidae)" masterLastPageNumber="25" masterPageNumber="17" pageNumber="18" updateTime="1699625600604" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1D084D910B5757FB28CAD4D2733965CE">First Descriptions for Larval Stages of Eurysphindus (Coleoptera: Cucujoidea: Sphindidae)</mods:title>
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<mods:namePart id="F506A9D9F6E5C8103D65548778E5C700">McHugh, Joseph V.</mods:namePart>
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<mods:title id="00258289F375C85E0992BC62D6B7DDF3">The Coleopterists Bulletin</mods:title>
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<treatment id="03C787EA015DFFEC9DE79421FC0C482D" LSID="urn:lsid:plazi:treatment:03C787EA015DFFEC9DE79421FC0C482D" httpUri="http://treatment.plazi.org/id/03C787EA015DFFEC9DE79421FC0C482D" lastPageId="3" lastPageNumber="20" pageId="1" pageNumber="18">
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<paragraph id="8BD136FC015DFFEE9DE79421FD064C0F" blockId="1.[396,728,319,341]" box="[396,728,319,341]" pageId="1" pageNumber="18">
<heading id="D0998190015DFFEE9DE79421FD064C0F" box="[396,728,319,341]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="18" reason="2">
<emphasis id="B91AEAEE015DFFEE9DE79421FD064C0F" bold="true" box="[396,728,319,341]" pageId="1" pageNumber="18">
Larva of
<taxonomicName id="4C6E4D7F015DFFEE9D84945EFD064C0F" ID-CoL="4GYQ" authority="LeConte" authorityName="LeConte" box="[495,728,319,341]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAEE015DFFEE9D84945EFDA74C0E" bold="true" box="[495,633,320,341]" italics="true" pageId="1" pageNumber="18">Eurysphindus</emphasis>
LeConte
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3746577015DFFEE9CC29477FE434982" pageId="1" pageNumber="18" type="description">
<paragraph id="8BD136FC015DFFEE9CC29477FD784CAF" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
<emphasis id="B91AEAEE015DFFEE9CC29477FEC94C27" bold="true" box="[169,279,361,380]" pageId="1" pageNumber="18">Description.</emphasis>
Mature larvae 2.93.4 mm. Body (
<figureCitation id="13552A79015DFFEE9E549477FDA74C27" box="[575,633,361,380]" captionStart="Fig" captionStartId="2.[197,228,1408,1427]" captionText="Fig. 1. Eurysphindus comatulus, habitus, dorsal." pageId="1" pageNumber="18">Figs. 1</figureCitation>
,
<figureCitation id="13552A79015DFFEE9EE89477FD464C27" box="[643,664,361,380]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">15</figureCitation>
) elongate, fusiform, straight, slightly compressed. Dorsum of thorax and abdomen with narrow transverse bands of weakly sclerotized tergal plates. Integument smooth with some small granules, especially on terga (
<figureCitation id="13552A79015DFFEE9F329487FC514CF7" box="[857,911,409,428]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Fig. 7</figureCitation>
). Body lightly pigmented, with darker head and a few weakly pigmented posterior abdominal terga. Vestiture consisting of erect setae, most setae on head simple. Most setae on dorsum of thorax and abdomen spiraled at apex and minutely capitate (
<figureCitation id="13552A79015DFFEE9E5794FFFDA74CAF" box="[572,633,481,500]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Figs. 7</figureCitation>
,
<figureCitation id="13552A79015DFFEE9EEF94FFFD454CAF" box="[644,667,481,500]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">15</figureCitation>
).
</paragraph>
<paragraph id="8BD136FC015DFFEE9CC294E7FF0E4EB1" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
Head: Protracted and prognathous, moderately broad, slightly flattened, lightly sclerotized. Antenna (
<figureCitation id="13552A79015DFFEE9C8D970FFEFF4F7F" box="[230,289,529,548]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Figs. 5</figureCitation>
,
<figureCitation id="13552A79015DFFEE9D41970FFE9E4F7F" box="[298,320,529,548]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">10</figureCitation>
) well-developed, 3-segmented, inserted laterally, anterior to stemmata, arising from large swollen membranous base; antennomere ratio AI:II:III 5 4:13:3; AII broadest at midlength, bearing sensorium apically and AIII subapically, sensorium large, 4/5 length AIII, conical, gradually curved toward AIII; AIII bearing 1 long and 2 short apical setae. Stemmata 6 per side (
<figureCitation id="13552A79015DFFEE9CF1976EFF044FD8" box="[154,218,624,643]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">Fig. 12</figureCitation>
), arranged in 2 subparallel rows of 3. Epicranial stem short, wide (
<figureCitation id="13552A79015DFFEE9F4A976EFC8B4FD8" box="[801,853,624,643]" captionStart="Fig" captionStartId="2.[197,228,1408,1427]" captionText="Fig. 1. Eurysphindus comatulus, habitus, dorsal." pageId="1" pageNumber="18">Fig. 1</figureCitation>
); frontal arms lyriform and contiguous at base. Median endocarina absent. Frontoclypeal suture absent (
<figureCitation id="13552A79015DFFEE9FEC9796FC184FC0" box="[903,966,648,667]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">Fig. 10</figureCitation>
); clypeolabral suture incomplete; labrum free. Mandibles (
<figureCitation id="13552A79015DFFEE9E1297BEFD3C4FE8" box="[633,738,672,691]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Figs. 3, 4, 6</figureCitation>
,
<figureCitation id="13552A79015DFFEE9E8697BEFCDC4FE8" box="[749,770,672,691]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">11</figureCitation>
) symmetrical to weakly asymmetrical, bidentate, with prominent ventral accessory process; mola well-developed, tuberculate, prostheca narrow, fixed, hyaline, acute at apex, abruptly curved mesad at about 1/3 length. Ventral mouthparts (
<figureCitation id="13552A79015DFFEE9DE397F6FE024FA0" box="[392,476,744,763]" captionStart="Figs" captionStartId="4.[197,236,1073,1092]" captionTargetBox="[286,894,195,1052]" captionTargetId="figure-201@4.[286,894,194,1052]" captionTargetPageId="4" captionText="Figs. 89. Eurysphindus comatulus larval maxillolabial complex. 7) Dorsal; 8) ventral." figureDoi="http://doi.org/10.5281/zenodo.10103382" httpUri="https://zenodo.org/record/10103382/files/figure.png" pageId="1" pageNumber="18">Figs. 8, 9</figureCitation>
) retracted. Maxilla with transverse cardo, elongate stipes, well developed articulating area, 3-segmented palp, well developed palpifer; galea and lacinia completely fused to form broad, obtuse mala, malar apex covered with moderately dense, curled setae, mesal margin bearing a single longitudinal row of 89 long stout setae and 2 small, straight teeth near apex of mesal margin. Labium free to base of mentum; ligula short and broad; labial palps short, 2-segmented, widely separated. Hypopharynx (
<figureCitation id="13552A79015DFFEE9EFE967EFD174E28" box="[661,713,864,883]" captionStart="Figs" captionStartId="4.[197,236,1073,1092]" captionTargetBox="[286,894,195,1052]" captionTargetId="figure-201@4.[286,894,194,1052]" captionTargetPageId="4" captionText="Figs. 89. Eurysphindus comatulus larval maxillolabial complex. 7) Dorsal; 8) ventral." figureDoi="http://doi.org/10.5281/zenodo.10103382" httpUri="https://zenodo.org/record/10103382/files/figure.png" pageId="1" pageNumber="18">Fig. 8</figureCitation>
) bilobed at apex, with 6 stout setae arranged in a V shape in median area; lateral lobes covered with very long, seta-like asperities; hypopharyngeal sclerome a curved transverse bar with median acute projection; epipharynx membranous, with long, seta-like asperities laterally and a transverse row of approximately 25 placoid sensilla near mid-length (
<figureCitation id="13552A79015DFFEE9E2196A1FDA14E89" box="[586,639,959,978]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Fig. 2</figureCitation>
). Ventral epicranial ridges absent. Gula absent.
</paragraph>
<paragraph id="8BD136FC015DFFEE9CC296F1FE9F49F2" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
Thorax and Abdomen: Thoracic and abdominal terga IVIII with paired, weakly sclerotized plates (
<figureCitation id="13552A79015DFFEE9CBD9119FED24941" box="[214,268,1031,1050]" captionStart="Fig" captionStartId="2.[197,228,1408,1427]" captionText="Fig. 1. Eurysphindus comatulus, habitus, dorsal." pageId="1" pageNumber="18">Fig. 1</figureCitation>
); anterior tergites unpigmented, posterior abdominal plates weakly pigmented. Thoracic tergites only weakly projecting laterally. Legs well-developed, 5-segmented; tarsungulus with 2 setae lying one distal to the other (
<figureCitation id="13552A79015DFFEE9D979129FDE24911" box="[508,572,1079,1098]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">Fig. 14</figureCitation>
); coxae widely separated. Segments TIIII and AIVIII each with a membranous, pleural, mound-like projection bearing 812 long setae (
<figureCitation id="13552A79015DFFEE9FFA9151FC124939" box="[913,972,1103,1122]" captionStart="Fig" captionStartId="2.[197,228,1408,1427]" captionText="Fig. 1. Eurysphindus comatulus, habitus, dorsal." pageId="1" pageNumber="18">Figs. 1</figureCitation>
,
<figureCitation id="13552A79015DFFEE9CFF9178FF754922" box="[148,171,1126,1145]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">15</figureCitation>
). Abdominal segments AIVIII each also bearing a more ventral pleural, mound-like projection (
<figureCitation id="13552A79015DFFEE9CF19160FF0449CA" box="[154,218,1150,1169]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">Fig. 15</figureCitation>
), with 45 long setae. Tergum A9 well-developed, simple. Segment A10 short, circular, posteriorly oriented.
</paragraph>
<paragraph id="8BD136FC015DFFEE9CC291B0FE434982" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
Spiracles: small, annular, projecting slightly from body wall (
<figureCitation id="13552A79015DFFEE9EDF91B0FD2D499A" box="[692,755,1198,1217]" captionStart="Figs" captionStartId="5.[169,208,1187,1206]" captionTargetBox="[148,980,194,1166]" captionTargetId="figure-112@5.[148,980,194,1166]" captionTargetPageId="5" captionText="Figs. 1015. Eurysphindus comatulus larval anatomy. 10) Head, dorsal; 11) mandible, right, dorsal; 12) stemmata, right side, lateral; 13) AIII spiracle, right, lateral; 14) TII tarsungulus, right, anterior; 15) final instar larva, habitus, right side, lateral." figureDoi="http://doi.org/10.5281/zenodo.10103384" httpUri="https://zenodo.org/record/10103384/files/figure.png" pageId="1" pageNumber="18">Fig. 13</figureCitation>
), one pair on mesothorax, 8 abdominal pairs on AIVIII.
</paragraph>
</subSubSection>
<subSubSection id="C3746577015DFFED9CC791F6FC334B79" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="discussion">
<paragraph id="8BD136FC015DFFEE9CC791F6FD7B48D8" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
<emphasis id="B91AEAEE015DFFEE9CC791F6FEC149A6" bold="true" box="[172,287,1256,1277]" pageId="1" pageNumber="18">Comments.</emphasis>
<taxonomicName id="4C6E4D7F015DFFEE9D4791F7FE7149A5" box="[300,431,1257,1278]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAEE015DFFEE9D4791F7FE7149A5" box="[300,431,1257,1278]" italics="true" pageId="1" pageNumber="18">Eurysphindus</emphasis>
</taxonomicName>
differs from all known sphindid larvae by the following combination of characters (see Chiao and McHugh 2000): Urogomphi absent; maxillary palp 3-segmented; labial palp 2-segmented; AX lacking pair of ventral membranous lobes each bearing a single curved, stout seta; Tarsungulus bearing 2 setae; Mandible bidentate; Antennomere II 3.54.5 3 length AIII. Larvae of the two North American species may be diagnosed as follows.
</paragraph>
<paragraph id="8BD136FC015DFFEE9CC79096FF1048B6" blockId="1.[148,978,361,1570]" pageId="1" pageNumber="18">
<emphasis id="B91AEAEE015DFFEE9CC79096FE6C48C6" bold="true" box="[172,434,1416,1437]" pageId="1" pageNumber="18">
<taxonomicName id="4C6E4D7F015DFFEE9CC79096FE7248C6" box="[172,428,1416,1437]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015DFFEE9CC79096FE7248C6" bold="true" box="[172,428,1416,1437]" italics="true" pageId="1" pageNumber="18">Eurysphindus comatulus</emphasis>
</taxonomicName>
.
</emphasis>
Mature larva 3.33.4 mm; mandibles symmetrical, preapical tooth smooth- feebly serrate on distal edge, tubercles of mola broad, rounded (
<figureCitation id="13552A79015DFFEE9C8490A3FEF74889" box="[239,297,1469,1490]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="1" pageNumber="18">Fig. 6</figureCitation>
); lacinial teeth very small, inconspicuous; integument of nota and terga shiny.
</paragraph>
<paragraph id="8BD136FC015DFFED9CC790ECFC334B79" blockId="1.[148,978,361,1570]" lastBlockId="2.[175,1005,1469,1570]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
<emphasis id="B91AEAEE015DFFEE9CC790ECFEA14B5C" bold="true" box="[172,383,1522,1543]" pageId="1" pageNumber="18">
<taxonomicName id="4C6E4D7F015DFFEE9CC790ECFEA74B5C" authorityName="LeConte" authorityYear="1878" box="[172,377,1522,1543]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B91AEAEE015DFFEE9CC790ECFEA74B5C" bold="true" box="[172,377,1522,1543]" italics="true" pageId="1" pageNumber="18">Eurysphindus hirtus</emphasis>
</taxonomicName>
.
</emphasis>
Mature larva 2.93.0 mm; head smaller and narrower than in
<taxonomicName id="4C6E4D7F015DFFEE9CD89313FEE94B79" box="[179,311,1549,1570]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015DFFEE9CD89313FEE94B79" box="[179,311,1549,1570]" italics="true" pageId="1" pageNumber="18">E. comatulus</emphasis>
</taxonomicName>
; antennae proportionally longer, legs longer with more strongly projecting coxae; mandibles slightly asymmetrical, apices acute, somewhat falcate; preapical tooth with blunt denticles on distal edge, 4 denticles on right mandible 3 on left; tubercles on mola oblong, rounded, becoming acute on the ventral proximal margin (
<figureCitation id="13552A79015EFFED9C949313FE844B79" box="[255,346,1549,1570]" captionStart="Figs" captionStartId="3.[169,208,1080,1099]" captionTargetBox="[149,978,195,1062]" captionTargetId="figure-153@3.[148,980,194,1062]" captionTargetPageId="3" captionText="Figs. 27. Eurysphindus larval anatomy. 2) E. comatulus, labrum, ventral; 34) E. hirtus, mandible, ventral; 3) right; 4) left; 56) E. comatulus; 5) antenna, left, ventral; 6) mandible, left, ventral; 7) E. hirtus, TII notum, right side, dorsal." figureDoi="http://doi.org/10.5281/zenodo.10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="2" pageNumber="19">Figs. 3, 4</figureCitation>
); lacinial teeth more prominent; integument of nota and terga opaque.
</paragraph>
</subSubSection>
<caption id="DF116674015EFFED9CAE909EFDAB48C8" box="[197,629,1408,1427]" pageId="2" pageNumber="19" startId="2.[197,228,1408,1427]">
<paragraph id="8BD136FC015EFFED9CAE909EFDAB48C8" blockId="2.[197,629,1408,1427]" box="[197,629,1408,1427]" pageId="2" pageNumber="19">
<emphasis id="B91AEAEE015EFFED9CAE909EFF2148C8" bold="true" box="[197,255,1408,1427]" pageId="2" pageNumber="19">Fig. 1.</emphasis>
<taxonomicName id="4C6E4D7F015EFFED9D7F909EFE3848C8" box="[276,486,1408,1427]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015EFFED9D7F909EFE3848C8" box="[276,486,1408,1427]" italics="true" pageId="2" pageNumber="19">Eurysphindus comatulus</emphasis>
</taxonomicName>
, habitus, dorsal.
</paragraph>
</caption>
<caption id="DF116674015FFFEC9CC29126FD9B4920" ID-DOI="http://doi.org/10.5281/zenodo.10103380" ID-Zenodo-Dep="10103380" httpUri="https://zenodo.org/record/10103380/files/figure.png" pageId="3" pageNumber="20" startId="3.[169,208,1080,1099]" targetBox="[149,978,195,1062]" targetPageId="3">
<paragraph id="8BD136FC015FFFEC9CC29126FD9B4920" blockId="3.[148,978,1080,1147]" pageId="3" pageNumber="20">
<emphasis id="B91AEAEE015FFFEC9CC29126FEDC4910" bold="true" box="[169,258,1080,1099]" pageId="3" pageNumber="20">Figs. 27.</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9D7C9127FE554917" box="[279,395,1081,1100]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEAEE015FFFEC9D7C9127FE554917" box="[279,395,1081,1100]" italics="true" pageId="3" pageNumber="20">Eurysphindus</emphasis>
</taxonomicName>
larval anatomy.
<emphasis id="B91AEAEE015FFFEC9E489126FDEA4910" bold="true" box="[547,564,1080,1099]" pageId="3" pageNumber="20">2)</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9E559127FD714917" box="[574,687,1081,1100]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015FFFEC9E559127FD714917" box="[574,687,1081,1100]" italics="true" pageId="3" pageNumber="20">E. comatulus</emphasis>
</taxonomicName>
, labrum, ventral;
<emphasis id="B91AEAEE015FFFEC9F3B9126FCA84910" bold="true" box="[848,886,1080,1099]" pageId="3" pageNumber="20">34)</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9F149127FC154917" authorityName="LeConte" authorityYear="1878" box="[895,971,1081,1100]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B91AEAEE015FFFEC9F149127FC154917" box="[895,971,1081,1100]" italics="true" pageId="3" pageNumber="20">E. hirtus</emphasis>
</taxonomicName>
, mandible, ventral;
<emphasis id="B91AEAEE015FFFEC9D5F914EFE984938" bold="true" box="[308,326,1104,1123]" pageId="3" pageNumber="20">3)</emphasis>
right;
<emphasis id="B91AEAEE015FFFEC9DEB914EFE4C4938" bold="true" box="[384,402,1104,1123]" pageId="3" pageNumber="20">4)</emphasis>
left;
<emphasis id="B91AEAEE015FFFEC9DAA914EFE364938" bold="true" box="[449,488,1104,1123]" pageId="3" pageNumber="20">56)</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9D85914EFD804938" box="[494,606,1104,1123]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015FFFEC9D85914EFD804938" box="[494,606,1104,1123]" italics="true" pageId="3" pageNumber="20">E. comatulus</emphasis>
</taxonomicName>
;
<emphasis id="B91AEAEE015FFFEC9E01914EFDA54938" bold="true" box="[618,635,1104,1123]" pageId="3" pageNumber="20">5)</emphasis>
antenna, left, ventral;
<emphasis id="B91AEAEE015FFFEC9F56914EFC8E4938" bold="true" box="[829,848,1104,1123]" pageId="3" pageNumber="20">6)</emphasis>
mandible, left, ventral;
<emphasis id="B91AEAEE015FFFEC9CB09176FF324920" bold="true" box="[219,236,1128,1147]" pageId="3" pageNumber="20">7)</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9C989176FEBB4920" authority=", TII" authorityName="TII" box="[243,357,1128,1147]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B91AEAEE015FFFEC9C989176FEE34920" box="[243,317,1128,1147]" italics="true" pageId="3" pageNumber="20">E. hirtus</emphasis>
, TII
</taxonomicName>
notum, right side, dorsal.
</paragraph>
</caption>
<subSubSection id="C3746577015FFFEC9CC791B9FC0C482D" pageId="3" pageNumber="20" type="materials_examined">
<paragraph id="8BD136FC015FFFEC9CC791B9FC0C482D" blockId="3.[148,978,1191,1398]" pageId="3" pageNumber="20">
<emphasis id="B91AEAEE015FFFEC9CC791B9FEA449E7" bold="true" box="[172,378,1191,1212]" pageId="3" pageNumber="20">Material Examined.</emphasis>
<taxonomicName id="4C6E4D7F015FFFEC9DED91B9FDAC49E7" box="[390,626,1191,1212]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015FFFEC9DED91B9FDAC49E7" box="[390,626,1191,1212]" italics="true" pageId="3" pageNumber="20">Eurysphindus comatulus</emphasis>
</taxonomicName>
:
<collectingCountry id="F379766C015FFFEC9E1491B9FD1A49E7" box="[639,708,1191,1212]" name="United States of America" pageId="3" pageNumber="20">U.S.A.</collectingCountry>
:
<collectingRegion id="49AAF81E015FFFEC9EA591B9FCE949E7" box="[718,823,1191,1212]" country="United States of America" name="New York" pageId="3" pageNumber="20">New York</collectingRegion>
: Tompkins Co., Brooktondale, ex
<taxonomicName id="4C6E4D7F015FFFEC9D2E91DFFDD8498D" box="[325,518,1217,1238]" class="Myxomycetes" family="Didymiaceae" genus="Mucilago" kingdom="Protozoa" order="Physarales" pageId="3" pageNumber="20" phylum="Mycetozoa" rank="species" species="crustacea">
<emphasis id="B91AEAEE015FFFEC9D2E91DFFDD8498D" box="[325,518,1217,1238]" italics="true" pageId="3" pageNumber="20">Mucilago crustacea</emphasis>
</taxonomicName>
,
<date id="FFD0103C015FFFEC9E7C91DFFD56498D" box="[535,648,1217,1238]" pageId="3" pageNumber="20" value="1992-08-03">3.VIII.1992</date>
, J.
<collectionCode id="ED7FAE39015FFFEC9ECE91DFFD64498D" box="[677,698,1217,1238]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="20" type="Museum">V</collectionCode>
. McHugh (.50 early instar larvae, 13 mature larvae,
<specimenCount id="9D68FD75015FFFEC9DF091C2FE2449AA" box="[411,506,1244,1265]" pageId="3" pageNumber="20" type="pupa">10 pupae</specimenCount>
,.2,000 adults).
<taxonomicName id="4C6E4D7F015FFFEC9EDA91C2FCA849AA" box="[689,886,1244,1265]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B91AEAEE015FFFEC9EDA91C2FCA849AA" box="[689,886,1244,1265]" italics="true" pageId="3" pageNumber="20">Eurysphindus hirtus</emphasis>
</taxonomicName>
:
<collectingCountry id="F379766C015FFFEC9FE291C2FC1049AA" box="[905,974,1244,1265]" name="United States of America" pageId="3" pageNumber="20">U.S.A.</collectingCountry>
:
<collectingCountry id="F379766C015FFFEC9CFF91E9FF3A4857" box="[148,228,1271,1292]" name="Georgia" pageId="3" pageNumber="20">Georgia</collectingCountry>
: Clarke Co., Athens, Memorial Park, ex
<taxonomicName id="4C6E4D7F015FFFEC9EE391E9FC904857" box="[648,846,1271,1292]" class="Myxomycetes" family="Didymiaceae" genus="Diachea" kingdom="Protozoa" order="Physarales" pageId="3" pageNumber="20" phylum="Mycetozoa" rank="species" species="leucopodia">
<emphasis id="B91AEAEE015FFFEC9EE391E9FC904857" box="[648,846,1271,1292]" italics="true" pageId="3" pageNumber="20">Diachea leucopodia</emphasis>
</taxonomicName>
,
<date id="FFD0103C015FFFEC9F3591E9FC124857" box="[862,972,1271,1292]" pageId="3" pageNumber="20" value="1998-06-15">
15.
<collectionCode id="ED7FAE39015FFFEC9F1291E9FC484857" box="[889,918,1271,1292]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="20">VI</collectionCode>
.1998
</date>
, J.
<collectionCode id="ED7FAE39015FFFEC9CC2900FFF63487D" box="[169,189,1297,1318]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="20" type="Museum">V</collectionCode>
. McHugh (1 early instar larva, 4 mature larvae,
<specimenCount id="9D68FD75015FFFEC9EEA900FFD05487D" box="[641,731,1297,1318]" pageId="3" pageNumber="20" type="adult">26 adults</specimenCount>
). Descriptions and figures of mouthparts are based on a single dissected specimen of
<taxonomicName id="4C6E4D7F015FFFEC9EA09032FCC0481A" box="[715,798,1324,1345]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B91AEAEE015FFFEC9EA09032FCC0481A" box="[715,798,1324,1345]" italics="true" pageId="3" pageNumber="20">E. hirtus</emphasis>
</taxonomicName>
and five dissected specimens of
<taxonomicName id="4C6E4D7F015FFFEC9D499058FE7B4800" box="[290,421,1350,1371]" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="comatulus">
<emphasis id="B91AEAEE015FFFEC9D499058FE7B4800" box="[290,421,1350,1371]" italics="true" pageId="3" pageNumber="20">E. comatulus</emphasis>
</taxonomicName>
. Immature and adult specimens of both
<taxonomicName id="4C6E4D7F015FFFEC9F259058FF06482D" class="Insecta" family="Sphindidae" genus="Eurysphindus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91AEAEE015FFFEC9F259058FC0F4800" box="[846,977,1350,1371]" italics="true" pageId="3" pageNumber="20">Eurysphindus</emphasis>
species
</taxonomicName>
are deposited in the University of
<collectingCountry id="F379766C015FFFEC9E40907FFDA9482D" box="[555,631,1377,1398]" name="Georgia" pageId="3" pageNumber="20">Georgia</collectingCountry>
Collection of Arthropods (
<collectionCode id="ED7FAE39015FFFEC9F15907FFC19482D" box="[894,967,1377,1398]" pageId="3" pageNumber="20">UGCA</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>