498 lines
56 KiB
XML
498 lines
56 KiB
XML
<document id="DEA520F62C7E9D156E8D0842A1DAF0C0" ID-DOI="10.11646/zootaxa.3722.1.4" ID-GBIF-Dataset="20a6b8ea-cf3b-4863-96f7-31aa49033c5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="224213" ID-ZooBank="C9674D23-8E8E-4197-B389-8EA55016E372" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460690503293" checkinUser="plazi" docAuthor="Unnikrishnan, V." docDate="2013" docId="03C787DD2764FFE9D3A147A02176F895" docLanguage="en" docName="zt03722p060.pdf" docOrigin="Zootaxa 3722 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrobrachium Bate 1868" docType="treatment" docVersion="8" masterDocId="FFFEFFA52765FFE1D336465D2427FF83" masterDocTitle="Description of a new subspecies, Macrobrachium aemulum madhusoodani (Decapoda: Palaemonidae) from Neyyar River, Kerala, S. India" updateTime="1698351779146" updateUser="plazi">
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<mods:mods id="49514F234112F18CCE42EBF82E94FF41" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F519E896572A096099A34B8630342480">
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<mods:title id="EAFF6A6383D0B1EDD77B02E10FC8B51C">Description of a new subspecies, Macrobrachium aemulum madhusoodani (Decapoda: Palaemonidae) from Neyyar River, Kerala, S. India</mods:title>
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<mods:name id="4B3AEC1EC22DC790696E3804F40E4998" type="personal">
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<mods:role id="1C8AB2FE6ACB3D41B5A6F993386E34DB">
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<mods:roleTerm id="0CA0DD7EF45DF80A156DB90AB51FCDE8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0A15263CC2B28FF6AFD989660CBFE561">Unnikrishnan, V.</mods:namePart>
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<mods:typeOfResource id="67C1131301B0E85062F08565C0D263D9">text</mods:typeOfResource>
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<mods:titleInfo id="2997F95DE19094DE72D0C2029D37010C">
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<mods:title id="A17CDF09B44C2B0DC38ADCD3F77B2237">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CF7990CF48898DBAB4A3CEF3A08D280B">
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<mods:date id="C8785DFA12A201B63D8A182996765EE7">2013</mods:date>
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<mods:detail id="042C48C9824676DC445E9FDABA5E4605" type="volume">
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<mods:number id="AD10547F0E661B45F117B52627B7EACE">3722</mods:number>
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</mods:detail>
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<mods:detail id="BD24C8B408E59D4D376A40B0FB2C2D84" type="issue">
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<mods:number id="2056AF420346188C17323F6789FA5A75">1</mods:number>
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</mods:detail>
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</mods:part>
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<mods:classification id="6CF87840D6995A0A9E3067FFA7B9F733">journal volume</mods:classification>
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<mods:identifier id="C1BEDDD2924BA03F695BAEFF9DC65F74" type="DOI">10.11646/zootaxa.3722.1.4</mods:identifier>
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<mods:identifier id="0012D7CCAEAE3BB63F1C3770E4DAD19C" type="GBIF-Dataset">20a6b8ea-cf3b-4863-96f7-31aa49033c5c</mods:identifier>
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<mods:identifier id="E8C9AD78E15F7D645F3020A88669D715" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9887F9EADACF3002500C8BC548E888F2" type="Zenodo-Dep">224213</mods:identifier>
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<mods:identifier id="8053C5B705A4720144F625C36F100D84" type="ZooBank">C9674D23-8E8E-4197-B389-8EA55016E372</mods:identifier>
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</mods:mods>
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<treatment id="03C787DD2764FFE9D3A147A02176F895" ID-DOI="http://doi.org/10.5281/zenodo.5698257" ID-GBIF-Taxon="119579229" ID-Zenodo-Dep="5698257" LSID="urn:lsid:plazi:treatment:03C787DD2764FFE9D3A147A02176F895" httpUri="http://treatment.plazi.org/id/03C787DD2764FFE9D3A147A02176F895" lastPageId="8" pageId="1">
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<subSubSection id="C37465402764FFE0D3A147A02668FD4B" pageId="1" type="nomenclature">
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<paragraph id="8BD136CB2764FFE0D3A147A0261AFD94" blockId="1.[151,573,508,535]" box="[151,573,508,535]" pageId="1">
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<heading id="D09981A72764FFE0D3A147A0261AFD94" bold="true" box="[151,573,508,535]" fontSize="11" level="1" pageId="1" reason="1">
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<emphasis id="B91AEAD92764FFE0D3A147A0261AFD94" bold="true" box="[151,573,508,535]" pageId="1">
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Genus
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<taxonomicName id="4C6E4D482764FFE0D3D847A1261AFD94" ID-CoL="4a489b2a-7bc3-4b4b-8982-82855d710e9d" authority="Bate, 1868" authorityName="Bate" authorityYear="1868" box="[238,573,508,535]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAD92764FFE0D3D847A12593FD95" bold="true" box="[238,436,508,534]" italics="true" pageId="1">Macrobrachium</emphasis>
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Bate, 1868
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD136CB2764FFE0D3A144192668FD4B" blockId="1.[151,1436,580,712]" pageId="1">
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<taxonomicName id="4C6E4D482764FFE0D3A14419256BFDDF" box="[151,332,580,604]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B91AEAD92764FFE0D3A14419256BFDDF" box="[151,332,580,604]" italics="true" pageId="1">Macrobrachium</emphasis>
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</taxonomicName>
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is characterised by the absence of branchio-stegal spine, presence of hepatic spine and mandibular palp, dactyl of last 3 legs simple, second chelate leg more robust, mandibular palp 3- jointed, third maxilliped with arthrobranch and pleurobranch, propodus of fifth pereiopod with transverse rows of setae on posterio-distal margin. Second chelate legs equal or sub equal.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465402764FFE0D3A145462535FCDB" pageId="1" type="description">
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<paragraph id="8BD136CB2764FFE0D3A145462535FCDB" blockId="1.[151,772,795,856]" pageId="1">
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<emphasis id="B91AEAD92764FFE0D3A145462723FCB6" bold="true" box="[151,772,795,821]" pageId="1">
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<taxonomicName id="4C6E4D482764FFE0D3A14546265EFCB7" box="[151,633,795,820]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="1" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
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<emphasis id="B91AEAD92764FFE0D3A14546265EFCB7" bold="true" box="[151,633,795,820]" italics="true" pageId="1">Macrobrachium aemulum madhusoodani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957A22764FFE0D1B645412723FCB6" box="[640,772,796,821]" pageId="1" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="13552A4E2764FFE0D3A94562252EFCDB" box="[159,265,831,856]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 4" captionStartId-0="2.[151,250,1982,2004]" captionStartId-1="6.[151,250,948,970]" captionTargetBox-0="[234,1345,195,1912]" captionTargetBox-1="[178,1404,221,894]" captionTargetId-0="figure@2.[196,1373,195,1931]" captionTargetId-1="figure@6.[150,1436,195,923]" captionTargetPageId-0="2" captionTargetPageId-1="6" captionText-0="FIGURE 1. Macrobrachium aemulum madhusoodani subsp. nov., specimen no. 1, holotype (male). A, entire animal, dorsal view" captionText-1="FIGURE 4. Macrobrachium aemulum madhusoodani subsp. nov., specimen no. 2, paratype. Carapace left lateral view." httpUri-0="https://zenodo.org/record/224214/files/figure.png" httpUri-1="https://zenodo.org/record/224215/files/figure.png" pageId="1">Figs. 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465402764FFE0D3A145DA2000FA1B" pageId="1" type="materials_examined">
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<paragraph id="8BD136CB2764FFE0D3A145DA259BFB1E" blockId="1.[151,1436,903,1901]" pageId="1">
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<emphasis id="B91AEAD92764FFE0D3A145DA25A2FC23" bold="true" box="[151,389,903,928]" pageId="1">Materials examined</emphasis>
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. Analysis of the nine specimens of the new subspecies collected from the middle reaches of the Neyyar River (
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<geoCoordinate id="EE5A500C2764FFE0D25A45F12628FC47" box="[364,527,940,964]" direction="east" orientation="longitude" pageId="1" precision="1" value="77.12523">77°7´30.83´´E</geoCoordinate>
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<geoCoordinate id="EE5A500C2764FFE0D12145F1269BFC46" box="[535,700,940,965]" direction="north" orientation="latitude" pageId="1" precision="1" value="8.4904585">8°29´25.65´´N</geoCoordinate>
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) Kerala, South
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<collectingCountry id="F379765B2764FFE0D04445F12789FC46" box="[882,942,940,965]" name="India" pageId="1">India</collectingCountry>
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, on
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<date id="FFD0100B2764FFE0D0D645F120B6FC47" box="[992,1169,940,965]" pageId="1" value="2007-03-24">24 March 2007</date>
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, showed
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<specimenCount id="9D68FD422764FFE0D7CA45F12173FC46" box="[1276,1364,940,965]" pageId="1" type="male">8 males</specimenCount>
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and
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<specimenCount id="9D68FD422764FFE0D6B845F124C0FC6B" pageId="1" type="female">1 female</specimenCount>
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. One male specimen is described here as the
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<typeStatus id="54D588692764FFE0D1D04592276FFC6B" box="[742,840,975,1000]" pageId="1" type="holotype">holotype</typeStatus>
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(no.1,
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<tableCitation id="C6EC03702764FFE0D0AE459227CCFC6B" box="[920,1003,975,1000]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionTargetBox="[159,1383,234,799]" captionTargetPageId="3" captionText="TABLE 1. Morphometric and meristic features of Macrobrachium aemulum madhusoodani subsp. nov. (length in mm and weight in g.)" httpUri="http://table.plazi.org/id/DF1166432766FFE2D3A146C5256CFF4C" pageId="1" tableUuid="DF1166432766FFE2D3A146C5256CFF4C">Table 1</tableCitation>
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).
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<typeStatus id="54D588692764FFE0D7324592204BFC6B" box="[1028,1132,975,1000]" pageId="1" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="54D588692764FFE0D792458D2121FC6B" box="[1188,1286,976,1000]" pageId="1" type="paratype">Paratype</typeStatus>
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(no. 2,
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<tableCitation id="C6EC03702764FFE0D66845922484FB8F" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionTargetBox="[159,1383,234,799]" captionTargetPageId="3" captionText="TABLE 1. Morphometric and meristic features of Macrobrachium aemulum madhusoodani subsp. nov. (length in mm and weight in g.)" httpUri="http://table.plazi.org/id/DF1166432766FFE2D3A146C5256CFF4C" pageId="1" tableUuid="DF1166432766FFE2D3A146C5256CFF4C">Table 1</tableCitation>
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) are deposited in the faunal depository of the Zoological Survey of
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<collectingCountry id="F379765B2764FFE0D0EF45A92032FB8E" box="[985,1045,1012,1037]" name="India" pageId="1">India</collectingCountry>
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, Western Ghats Regional Centre, Kozhikkode (Reg. nos.- holotype- ZSI/WGRS/IR/INV 2167; paratype- ZSI/WGRS/IR/INV 2168). The
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<typeStatus id="54D588692764FFE0D607424521BBFBB3" box="[1329,1436,1048,1072]" pageId="1" type="paratype">paratypes</typeStatus>
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(nos. 3–9) are currently stored in the Natural History Museum of the Department of Zoology and Research Centre, Mahatma Gandhi College, Thiruvananthapuram (MGC/TVM/NHM-INV/3/2012). All length measurements are in mm. and weight in grams.
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</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F142FA209FFB66" blockId="1.[151,1436,903,1901]" pageId="1">
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<emphasis id="B91AEAD92764FFE0D3F142FA2624FB43" bold="true" box="[199,515,1191,1216]" pageId="1">
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Measurement of
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<typeStatus id="54D588692764FFE0D2A542FA25D9FB43" box="[403,510,1191,1216]" pageId="1" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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total length, 60; carapace length, 18; length of rostrum, 12; length of telson, 7.0; first chelate leg: ischium (i) 3.0; merus (m) 6.0; carpus (c) 7.0; propodus (p) 4.0; dactyl (d) 1.5
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</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F142B22032FA8B" blockId="1.[151,1436,903,1901]" box="[199,1045,1263,1288]" pageId="1">Left second chelate leg (large): i = 8.0, m = 13.0, c = 22.0, p = 30.0, d = 9.0</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F14349200EFAAF" blockId="1.[151,1436,903,1901]" box="[199,1065,1300,1325]" pageId="1">Right second chelate leg (small): i = 7.0, m = 11.0, c = 16.0, p = 23.0, d = 7.0</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F1436A203BFAD3" blockId="1.[151,1436,903,1901]" box="[199,1052,1335,1360]" pageId="1">First non-chelate leg (pereiopod 3): i = 3.0, m = 7.0, c = 4.0, p = 6.0, d = 2.0</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F14301201CFAF7" blockId="1.[151,1436,903,1901]" box="[199,1083,1372,1397]" pageId="1">Second non-chelate leg (pereiopod 4): i = 3.0, m = 7.0, c = 4.0, p = 6.0, d = 2.0</paragraph>
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<paragraph id="8BD136CB2764FFE0D3F143222000FA1B" blockId="1.[151,1436,903,1901]" box="[199,1063,1407,1432]" pageId="1">Third non-chelate leg (pereiopod 5): i = 3.0, m = 7.0, c = 4.0, p = 6.0, d = 2.0</paragraph>
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</subSubSection>
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<subSubSection id="C37465402764FFE0D3F143F92696F883" pageId="1" type="diagnosis">
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<paragraph id="8BD136CB2764FFE0D3F143F92696F883" blockId="1.[151,1436,903,1901]" pageId="1">
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<emphasis id="B91AEAD92764FFE0D3F143F92566FA3E" bold="true" box="[199,321,1444,1469]" pageId="1">Diagnosis.</emphasis>
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Rostral formula 9–13/2 with 4–5 postorbital teeth; carapace smooth; distal end of rostrum directed upwards and upper margin convex, second chelate legs robust, dissimilar in shape and unequal in size. In large second pereiopod, merus shorter than carpus and about half length of chela; carpus shorter than chela, palm longer than fingers; width of palm slightly greater than width of distal carpus; palm of larger chela narrower than smaller chela; fingers stout and equal. Two large denticles and six weak denticles respectively at the proximal and distal parts of movable finger. Similar pattern of dentition is present in the immovable finger also. In small second pereiopod, merus shorter than carpus, about half-length of chela; carpus shorter than chela, palm longer than fingers; width of palm greater than width of distal carpus. Fingers stout, equal, unarmed and wide gap between fingers when closed. Numerous setae present on the inner margin of both fingers. Four pairs of plumose setae present between inner movable spines of telson.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37465402764FFE2D3F1415126D9F9AB" lastPageId="3" pageId="1" type="description">
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<paragraph id="8BD136CB2764FFE0D3F1415127D7F8EE" blockId="1.[151,1436,903,1901]" pageId="1">
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<emphasis id="B91AEAD92764FFE0D3F14151257EF8A6" bold="true" box="[199,345,1804,1829]" pageId="1">Description.</emphasis>
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Rostrum short, extending as far as the middle of distal segment of antennular peduncle; dorsal margin convex with 9–13 teeth, 4–5 postorbital (table 1), distal end directed upwards and ventral margin with two teeth. Small setae present between teeth, on both dorsal and ventral margins.
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</paragraph>
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<caption id="DF1166432767FFE3D3A141E324EFF870" httpUri="https://zenodo.org/record/224214/files/figure.png" pageId="2" targetBox="[234,1345,195,1912]" targetPageId="2">
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<paragraph id="8BD136CB2767FFE3D3A141E324EFF870" blockId="2.[151,1436,1982,2035]" pageId="2">
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<emphasis id="B91AEAD92767FFE3D3A141E32533F850" bold="true" box="[151,276,1982,2004]" pageId="2">FIGURE 1.</emphasis>
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<taxonomicName id="4C6E4D482767FFE3D22A41E326E5F857" box="[284,706,1982,2004]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="2" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
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<emphasis id="B91AEAD92767FFE3D22A41E326E5F857" box="[284,706,1982,2004]" italics="true" pageId="2">Macrobrachium aemulum madhusoodani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957A22767FFE3D1FD41E32765F857" box="[715,834,1982,2004]" pageId="2" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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, specimen no. 1, holotype (male). A, entire animal, dorsal view
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</paragraph>
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</caption>
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<caption id="DF1166432766FFE2D3A146C5256CFF4C" ID-Table-UUID="DF1166432766FFE2D3A146C5256CFF4C" httpUri="http://table.plazi.org/id/DF1166432766FFE2D3A146C5256CFF4C" pageId="3" targetBox="[159,1383,234,799]" targetIsTable="true" targetPageId="3">
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||
<paragraph id="8BD136CB2766FFE2D3A146C5256CFF4C" blockId="3.[151,1436,151,207]" pageId="3">
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||
<emphasis id="B91AEAD92766FFE2D3A146C5252CFF2C" bold="true" box="[151,267,152,175]" pageId="3">TABLE 1.</emphasis>
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Morphometric and meristic features of
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<taxonomicName id="4C6E4D482766FFE2D18B46CA205EFF2C" box="[701,1145,151,175]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
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<emphasis id="B91AEAD92766FFE2D18B46CA205EFF2C" box="[701,1145,151,175]" italics="true" pageId="3">Macrobrachium aemulum madhusoodani</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22957A22766FFE2D7B746C520DDFF2C" box="[1153,1274,152,175]" pageId="3" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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(length in mm and weight in g.)
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||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD136CB2766FFE2D3A946B7216AFC9C" pageId="3">
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<table id="F96EC46B2766001ED3A946B72140FC9C" box="[159,1383,234,799]" gridcols="10" gridrows="14" pageId="3">
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||
<tr id="355E34892766001ED3A946B72140FE83" box="[159,1383,234,256]" gridrow="0" pageId="3">
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<th id="768F5DF52766001ED3A946B725F4FE83" box="[159,467,234,256]" gridcol="0" gridrow="0" pageId="3">Specimen nos.</th>
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<th id="768F5DF52766001ED12146B72663FE83" box="[535,580,234,256]" gridcol="1" gridrow="0" pageId="3">1</th>
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<th id="768F5DF52766001ED14246B72687FE83" box="[628,672,234,256]" gridcol="2" gridrow="0" pageId="3">2</th>
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<th id="768F5DF52766001ED1EB46B7272CFE83" box="[733,779,234,256]" gridcol="3" gridrow="0" pageId="3">3</th>
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<th id="768F5DF52766001ED00146B72743FE83" box="[823,868,234,256]" gridcol="4" gridrow="0" pageId="3">4</th>
|
||
<th id="768F5DF52766001ED09646B727EAFE83" box="[928,973,234,256]" gridcol="5" gridrow="0" pageId="3">5</th>
|
||
<th id="768F5DF52766001ED0C946B7200CFE83" box="[1023,1067,234,256]" gridcol="6" gridrow="0" pageId="3">6</th>
|
||
<th id="768F5DF52766001ED75E46B720B2FE83" box="[1128,1173,234,256]" gridcol="7" gridrow="0" pageId="3">7</th>
|
||
<th id="768F5DF52766001ED7EC46B72121FE83" box="[1242,1286,234,256]" gridcol="8" gridrow="0" pageId="3">8</th>
|
||
<th id="768F5DF52766001ED60C46B72140FE83" box="[1338,1383,234,256]" gridcol="9" gridrow="0" pageId="3">9</th>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A947492140FEA9" box="[159,1383,276,298]" gridrow="1" pageId="3">
|
||
<th id="768F5DF52766001ED3A9474925F4FEA9" box="[159,467,276,298]" gridcol="0" gridrow="1" pageId="3">Total length</th>
|
||
<td id="768F5DF52766001ED12147492663FEA9" box="[535,580,276,298]" gridcol="1" gridrow="1" pageId="3">60</td>
|
||
<td id="768F5DF52766001ED14247492687FEA9" box="[628,672,276,298]" gridcol="2" gridrow="1" pageId="3">75</td>
|
||
<td id="768F5DF52766001ED1EB4749272CFEA9" box="[733,779,276,298]" gridcol="3" gridrow="1" pageId="3">40</td>
|
||
<td id="768F5DF52766001ED00147492743FEA9" box="[823,868,276,298]" gridcol="4" gridrow="1" pageId="3">43</td>
|
||
<td id="768F5DF52766001ED096474927EAFEA9" box="[928,973,276,298]" gridcol="5" gridrow="1" pageId="3">38</td>
|
||
<td id="768F5DF52766001ED0C94749200CFEA9" box="[1023,1067,276,298]" gridcol="6" gridrow="1" pageId="3">36</td>
|
||
<td id="768F5DF52766001ED75E474920B2FEA9" box="[1128,1173,276,298]" gridcol="7" gridrow="1" pageId="3">34</td>
|
||
<td id="768F5DF52766001ED7EC47492121FEA9" box="[1242,1286,276,298]" gridcol="8" gridrow="1" pageId="3">64</td>
|
||
<td id="768F5DF52766001ED60C47492140FEA9" box="[1338,1383,276,298]" gridcol="9" gridrow="1" pageId="3">58</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A947602140FED0" box="[159,1383,317,339]" gridrow="2" pageId="3">
|
||
<th id="768F5DF52766001ED3A9476025F4FED0" box="[159,467,317,339]" gridcol="0" gridrow="2" pageId="3">Carapace length</th>
|
||
<td id="768F5DF52766001ED12147602663FED0" box="[535,580,317,339]" gridcol="1" gridrow="2" pageId="3">18</td>
|
||
<td id="768F5DF52766001ED14247602687FED0" box="[628,672,317,339]" gridcol="2" gridrow="2" pageId="3">20</td>
|
||
<td id="768F5DF52766001ED1EB4760272CFED0" box="[733,779,317,339]" gridcol="3" gridrow="2" pageId="3">11</td>
|
||
<td id="768F5DF52766001ED00147602743FED0" box="[823,868,317,339]" gridcol="4" gridrow="2" pageId="3">13</td>
|
||
<td id="768F5DF52766001ED096476027EAFED0" box="[928,973,317,339]" gridcol="5" gridrow="2" pageId="3">11</td>
|
||
<td id="768F5DF52766001ED0C94760200CFED0" box="[1023,1067,317,339]" gridcol="6" gridrow="2" pageId="3">10</td>
|
||
<td id="768F5DF52766001ED75E476020B2FED0" box="[1128,1173,317,339]" gridcol="7" gridrow="2" pageId="3">10</td>
|
||
<td id="768F5DF52766001ED7EC47602121FED0" box="[1242,1286,317,339]" gridcol="8" gridrow="2" pageId="3">18</td>
|
||
<td id="768F5DF52766001ED60C47602140FED0" box="[1338,1383,317,339]" gridcol="9" gridrow="2" pageId="3">17</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A9473B2140FEFF" box="[159,1383,358,380]" gridrow="3" pageId="3">
|
||
<th id="768F5DF52766001ED3A9473B25F4FEFF" box="[159,467,358,380]" gridcol="0" gridrow="3" pageId="3">Length of rostrum</th>
|
||
<td id="768F5DF52766001ED121473B2663FEFF" box="[535,580,358,380]" gridcol="1" gridrow="3" pageId="3">12</td>
|
||
<td id="768F5DF52766001ED142473B2687FEFF" box="[628,672,358,380]" gridcol="2" gridrow="3" pageId="3">14</td>
|
||
<td id="768F5DF52766001ED1EB473B272CFEFF" box="[733,779,358,380]" gridcol="3" gridrow="3" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED001473B2743FEFF" box="[823,868,358,380]" gridcol="4" gridrow="3" pageId="3">10</td>
|
||
<td id="768F5DF52766001ED096473B27EAFEFF" box="[928,973,358,380]" gridcol="5" gridrow="3" pageId="3">9.0</td>
|
||
<td id="768F5DF52766001ED0C9473B200CFEFF" box="[1023,1067,358,380]" gridcol="6" gridrow="3" pageId="3">8.0</td>
|
||
<td id="768F5DF52766001ED75E473B20B2FEFF" box="[1128,1173,358,380]" gridcol="7" gridrow="3" pageId="3">8.0</td>
|
||
<td id="768F5DF52766001ED7EC473B2121FEFF" box="[1242,1286,358,380]" gridcol="8" gridrow="3" pageId="3">14</td>
|
||
<td id="768F5DF52766001ED60C473B2140FEFF" box="[1338,1383,358,380]" gridcol="9" gridrow="3" pageId="3">11</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A947CD2140FE25" box="[159,1383,400,422]" gridrow="4" pageId="3">
|
||
<th id="768F5DF52766001ED3A947CD25F4FE25" box="[159,467,400,422]" gridcol="0" gridrow="4" pageId="3">Max. width of carapace</th>
|
||
<td id="768F5DF52766001ED12147CD2663FE25" box="[535,580,400,422]" gridcol="1" gridrow="4" pageId="3">10.5</td>
|
||
<td id="768F5DF52766001ED14247CD2687FE25" box="[628,672,400,422]" gridcol="2" gridrow="4" pageId="3">12.0</td>
|
||
<td id="768F5DF52766001ED1EB47CD272CFE25" box="[733,779,400,422]" gridcol="3" gridrow="4" pageId="3">5.5</td>
|
||
<td id="768F5DF52766001ED00147CD2743FE25" box="[823,868,400,422]" gridcol="4" gridrow="4" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED09647CD27EAFE25" box="[928,973,400,422]" gridcol="5" gridrow="4" pageId="3">6.0</td>
|
||
<td id="768F5DF52766001ED0C947CD200CFE25" box="[1023,1067,400,422]" gridcol="6" gridrow="4" pageId="3">6.2</td>
|
||
<td id="768F5DF52766001ED75E47CD20B2FE25" box="[1128,1173,400,422]" gridcol="7" gridrow="4" pageId="3">6.0</td>
|
||
<td id="768F5DF52766001ED7EC47CD2121FE25" box="[1242,1286,400,422]" gridcol="8" gridrow="4" pageId="3">11.0</td>
|
||
<td id="768F5DF52766001ED60C47CD2140FE25" box="[1338,1383,400,422]" gridcol="9" gridrow="4" pageId="3">9.0</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A947E42140FE4C" box="[159,1383,441,463]" gridrow="5" pageId="3">
|
||
<th id="768F5DF52766001ED3A947E425F4FE4C" box="[159,467,441,463]" gridcol="0" gridrow="5" pageId="3">Preorbital teeth</th>
|
||
<td id="768F5DF52766001ED12147E42663FE4C" box="[535,580,441,463]" gridcol="1" gridrow="5" pageId="3">9</td>
|
||
<td id="768F5DF52766001ED14247E42687FE4C" box="[628,672,441,463]" gridcol="2" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED1EB47E4272CFE4C" box="[733,779,441,463]" gridcol="3" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED00147E42743FE4C" box="[823,868,441,463]" gridcol="4" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED09647E427EAFE4C" box="[928,973,441,463]" gridcol="5" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED0C947E4200CFE4C" box="[1023,1067,441,463]" gridcol="6" gridrow="5" pageId="3">5</td>
|
||
<td id="768F5DF52766001ED75E47E420B2FE4C" box="[1128,1173,441,463]" gridcol="7" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED7EC47E42121FE4C" box="[1242,1286,441,463]" gridcol="8" gridrow="5" pageId="3">6</td>
|
||
<td id="768F5DF52766001ED60C47E42140FE4C" box="[1338,1383,441,463]" gridcol="9" gridrow="5" pageId="3">6</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A947BF2140FE7B" box="[159,1383,482,504]" gridrow="6" pageId="3">
|
||
<th id="768F5DF52766001ED3A947BF25F4FE7B" box="[159,467,482,504]" gridcol="0" gridrow="6" pageId="3">Postorbital teeth</th>
|
||
<td id="768F5DF52766001ED12147BF2663FE7B" box="[535,580,482,504]" gridcol="1" gridrow="6" pageId="3">4</td>
|
||
<td id="768F5DF52766001ED14247BF2687FE7B" box="[628,672,482,504]" gridcol="2" gridrow="6" pageId="3">5</td>
|
||
<td id="768F5DF52766001ED1EB47BF272CFE7B" box="[733,779,482,504]" gridcol="3" gridrow="6" pageId="3">4</td>
|
||
<td id="768F5DF52766001ED00147BF2743FE7B" box="[823,868,482,504]" gridcol="4" gridrow="6" pageId="3">5</td>
|
||
<td id="768F5DF52766001ED09647BF27EAFE7B" box="[928,973,482,504]" gridcol="5" gridrow="6" pageId="3">5</td>
|
||
<td id="768F5DF52766001ED0C947BF200CFE7B" box="[1023,1067,482,504]" gridcol="6" gridrow="6" pageId="3">4</td>
|
||
<td id="768F5DF52766001ED75E47BF20B2FE7B" box="[1128,1173,482,504]" gridcol="7" gridrow="6" pageId="3">4</td>
|
||
<td id="768F5DF52766001ED7EC47BF2121FE7B" box="[1242,1286,482,504]" gridcol="8" gridrow="6" pageId="3">5</td>
|
||
<td id="768F5DF52766001ED60C47BF2140FE7B" box="[1338,1383,482,504]" gridcol="9" gridrow="6" pageId="3">4</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A944512140FDA1" box="[159,1383,524,546]" gridrow="7" pageId="3">
|
||
<th id="768F5DF52766001ED3A9445125F4FDA1" box="[159,467,524,546]" gridcol="0" gridrow="7" pageId="3">Rostral formula</th>
|
||
<td id="768F5DF52766001ED12144512663FDA1" box="[535,580,524,546]" gridcol="1" gridrow="7" pageId="3">13/2</td>
|
||
<td id="768F5DF52766001ED14244512687FDA1" box="[628,672,524,546]" gridcol="2" gridrow="7" pageId="3">11/2</td>
|
||
<td id="768F5DF52766001ED1EB4451272CFDA1" box="[733,779,524,546]" gridcol="3" gridrow="7" pageId="3">10/2</td>
|
||
<td id="768F5DF52766001ED00144512743FDA1" box="[823,868,524,546]" gridcol="4" gridrow="7" pageId="3">11/2</td>
|
||
<td id="768F5DF52766001ED096445127EAFDA1" box="[928,973,524,546]" gridcol="5" gridrow="7" pageId="3">11/2</td>
|
||
<td id="768F5DF52766001ED0C94451200CFDA1" box="[1023,1067,524,546]" gridcol="6" gridrow="7" pageId="3">9/2</td>
|
||
<td id="768F5DF52766001ED75E445120B2FDA1" box="[1128,1173,524,546]" gridcol="7" gridrow="7" pageId="3">10/2</td>
|
||
<td id="768F5DF52766001ED7EC44512121FDA1" box="[1242,1286,524,546]" gridcol="8" gridrow="7" pageId="3">11/2</td>
|
||
<td id="768F5DF52766001ED60C44512140FDA1" box="[1338,1383,524,546]" gridcol="9" gridrow="7" pageId="3">10/2</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A944692140FDCE" box="[159,1383,564,589]" gridrow="8" pageId="3">
|
||
<th id="768F5DF52766001ED3A9446925F4FDCE" box="[159,467,564,589]" gridcol="0" gridrow="8" pageId="3">Length of large 2nd chelate leg</th>
|
||
<td id="768F5DF52766001ED12144692663FDCE" box="[535,580,564,589]" gridcol="1" gridrow="8" pageId="3">73.0</td>
|
||
<td id="768F5DF52766001ED14244692687FDCE" box="[628,672,564,589]" gridcol="2" gridrow="8" pageId="3">69.0</td>
|
||
<td id="768F5DF52766001ED1EB4469272CFDCE" box="[733,779,564,589]" gridcol="3" gridrow="8" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED00144692743FDCE" box="[823,868,564,589]" gridcol="4" gridrow="8" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED096446927EAFDCE" box="[928,973,564,589]" gridcol="5" gridrow="8" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED0C94469200CFDCE" box="[1023,1067,564,589]" gridcol="6" gridrow="8" pageId="3">22.0</td>
|
||
<td id="768F5DF52766001ED75E446920B2FDCE" box="[1128,1173,564,589]" gridcol="7" gridrow="8" pageId="3">19.5</td>
|
||
<td id="768F5DF52766001ED7EC44692121FDCE" box="[1242,1286,564,589]" gridcol="8" gridrow="8" pageId="3">69.0</td>
|
||
<td id="768F5DF52766001ED60C44692140FDCE" box="[1338,1383,564,589]" gridcol="9" gridrow="8" pageId="3">-</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A9443D2140FDFA" box="[159,1383,608,633]" gridrow="9" pageId="3">
|
||
<th id="768F5DF52766001ED3A9443D25F4FDFA" box="[159,467,608,633]" gridcol="0" gridrow="9" pageId="3">Length of small 2nd chelate leg</th>
|
||
<td id="768F5DF52766001ED121443D2663FDFA" box="[535,580,608,633]" gridcol="1" gridrow="9" pageId="3">57.0</td>
|
||
<td id="768F5DF52766001ED142443D2687FDFA" box="[628,672,608,633]" gridcol="2" gridrow="9" pageId="3">49.0</td>
|
||
<td id="768F5DF52766001ED1EB443D272CFDFA" box="[733,779,608,633]" gridcol="3" gridrow="9" pageId="3">14.5</td>
|
||
<td id="768F5DF52766001ED001443D2743FDFA" box="[823,868,608,633]" gridcol="4" gridrow="9" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED096443D27EAFDFA" box="[928,973,608,633]" gridcol="5" gridrow="9" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED0C9443D200CFDFA" box="[1023,1067,608,633]" gridcol="6" gridrow="9" pageId="3">-</td>
|
||
<td id="768F5DF52766001ED75E443D20B2FDFA" box="[1128,1173,608,633]" gridcol="7" gridrow="9" pageId="3">18.0</td>
|
||
<td id="768F5DF52766001ED7EC443D2121FDFA" box="[1242,1286,608,633]" gridcol="8" gridrow="9" pageId="3">53.0</td>
|
||
<td id="768F5DF52766001ED60C443D2140FDFA" box="[1338,1383,608,633]" gridcol="9" gridrow="9" pageId="3">49.0</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A944D02140FD20" box="[159,1383,653,675]" gridrow="10" pageId="3">
|
||
<th id="768F5DF52766001ED3A944D025F4FD20" box="[159,467,653,675]" gridcol="0" gridrow="10" pageId="3">Uropod length</th>
|
||
<td id="768F5DF52766001ED12144D02663FD20" box="[535,580,653,675]" gridcol="1" gridrow="10" pageId="3">10.0</td>
|
||
<td id="768F5DF52766001ED14244D02687FD20" box="[628,672,653,675]" gridcol="2" gridrow="10" pageId="3">10.0</td>
|
||
<td id="768F5DF52766001ED1EB44D0272CFD20" box="[733,779,653,675]" gridcol="3" gridrow="10" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED00144D02743FD20" box="[823,868,653,675]" gridcol="4" gridrow="10" pageId="3">9.0</td>
|
||
<td id="768F5DF52766001ED09644D027EAFD20" box="[928,973,653,675]" gridcol="5" gridrow="10" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED0C944D0200CFD20" box="[1023,1067,653,675]" gridcol="6" gridrow="10" pageId="3">6.0</td>
|
||
<td id="768F5DF52766001ED75E44D020B2FD20" box="[1128,1173,653,675]" gridcol="7" gridrow="10" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED7EC44D02121FD20" box="[1242,1286,653,675]" gridcol="8" gridrow="10" pageId="3">9.0</td>
|
||
<td id="768F5DF52766001ED60C44D02140FD20" box="[1338,1383,653,675]" gridcol="9" gridrow="10" pageId="3">8.5</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A944EB2140FD4F" box="[159,1383,694,716]" gridrow="11" pageId="3">
|
||
<th id="768F5DF52766001ED3A944EB25F4FD4F" box="[159,467,694,716]" gridcol="0" gridrow="11" pageId="3">Telson length</th>
|
||
<td id="768F5DF52766001ED12144EB2663FD4F" box="[535,580,694,716]" gridcol="1" gridrow="11" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED14244EB2687FD4F" box="[628,672,694,716]" gridcol="2" gridrow="11" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED1EB44EB272CFD4F" box="[733,779,694,716]" gridcol="3" gridrow="11" pageId="3">5.0</td>
|
||
<td id="768F5DF52766001ED00144EB2743FD4F" box="[823,868,694,716]" gridcol="4" gridrow="11" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED09644EB27EAFD4F" box="[928,973,694,716]" gridcol="5" gridrow="11" pageId="3">5.0</td>
|
||
<td id="768F5DF52766001ED0C944EB200CFD4F" box="[1023,1067,694,716]" gridcol="6" gridrow="11" pageId="3">4.0</td>
|
||
<td id="768F5DF52766001ED75E44EB20B2FD4F" box="[1128,1173,694,716]" gridcol="7" gridrow="11" pageId="3">6.0</td>
|
||
<td id="768F5DF52766001ED7EC44EB2121FD4F" box="[1242,1286,694,716]" gridcol="8" gridrow="11" pageId="3">7.0</td>
|
||
<td id="768F5DF52766001ED60C44EB2140FD4F" box="[1338,1383,694,716]" gridcol="9" gridrow="11" pageId="3">6.5</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A944822140FD76" box="[159,1383,735,757]" gridrow="12" pageId="3">
|
||
<th id="768F5DF52766001ED3A9448225F4FD76" box="[159,467,735,757]" gridcol="0" gridrow="12" pageId="3">Weight</th>
|
||
<td id="768F5DF52766001ED12144822663FD76" box="[535,580,735,757]" gridcol="1" gridrow="12" pageId="3">3.94</td>
|
||
<td id="768F5DF52766001ED14244822687FD76" box="[628,672,735,757]" gridcol="2" gridrow="12" pageId="3">4.74</td>
|
||
<td id="768F5DF52766001ED1EB4482272CFD76" box="[733,779,735,757]" gridcol="3" gridrow="12" pageId="3">0.64</td>
|
||
<td id="768F5DF52766001ED00144822743FD76" box="[823,868,735,757]" gridcol="4" gridrow="12" pageId="3">1.28</td>
|
||
<td id="768F5DF52766001ED096448227EAFD76" box="[928,973,735,757]" gridcol="5" gridrow="12" pageId="3">0.65</td>
|
||
<td id="768F5DF52766001ED0C94482200CFD76" box="[1023,1067,735,757]" gridcol="6" gridrow="12" pageId="3">0.60</td>
|
||
<td id="768F5DF52766001ED75E448220B2FD76" box="[1128,1173,735,757]" gridcol="7" gridrow="12" pageId="3">0.56</td>
|
||
<td id="768F5DF52766001ED7EC44822121FD76" box="[1242,1286,735,757]" gridcol="8" gridrow="12" pageId="3">3.59</td>
|
||
<td id="768F5DF52766001ED60C44822140FD76" box="[1338,1383,735,757]" gridcol="9" gridrow="12" pageId="3">2.53</td>
|
||
</tr>
|
||
<tr id="355E34892766001ED3A945542140FC9C" box="[159,1383,777,799]" gridrow="13" pageId="3">
|
||
<th id="768F5DF52766001ED3A9455425F4FC9C" box="[159,467,777,799]" gridcol="0" gridrow="13" pageId="3">Sex</th>
|
||
<td id="768F5DF52766001ED12145542663FC9C" box="[535,580,777,799]" gridcol="1" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED14245542687FC9C" box="[628,672,777,799]" gridcol="2" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED1EB4554272CFC9C" box="[733,779,777,799]" gridcol="3" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED00145542743FC9C" box="[823,868,777,799]" gridcol="4" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED096455427EAFC9C" box="[928,973,777,799]" gridcol="5" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED0C94554200CFC9C" box="[1023,1067,777,799]" gridcol="6" gridrow="13" pageId="3">♀</td>
|
||
<td id="768F5DF52766001ED75E455420B2FC9C" box="[1128,1173,777,799]" gridcol="7" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED7EC45542121FC9C" box="[1242,1286,777,799]" gridcol="8" gridrow="13" pageId="3">♂</td>
|
||
<td id="768F5DF52766001ED60C45542140FC9C" box="[1338,1383,777,799]" gridcol="9" gridrow="13" pageId="3">♂</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2766FFE2D3F1453920FFFBB3" blockId="3.[151,1436,867,2008]" pageId="3">
|
||
Carapace smooth, both antennal and hepatic spines characteristic of
|
||
<emphasis id="B91AEAD92766FFE2D0F0453920A5FCFF" box="[966,1154,868,892]" italics="true" pageId="3">
|
||
<taxonomicName id="4C6E4D482766FFE2D0F045392059FCFF" box="[966,1150,868,892]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="genus">Macrobrachium</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
latter situated just below and behind level of former (figs. 3A, 4). Abdomen glabrous, pleurae of somites I–III typical, those of IV and V directed backwards, and of VI ending in spine. Telson conical with two pairs of spines on dorsal surface, proximal pair situated midway and distal at about ¾ length of telson. Posterior end of telson with two pairs of movable spines; inner longer, overreaching tip of telson. Four pairs of plumose setae present between inner pair of movable spines (fig. 2A). Antennules typical, outer lateral spine of antennal scale sharp, pointed forwards.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2766FFE2D3F142662177FBFB" blockId="3.[151,1436,867,2008]" pageId="3">First pair of chelate legs slender, tips of chelae over reaching antennal scale; ischium half merus, merus shorter than carpus; carpus almost twice length of chela; palm longer than fingers and devoid of spination (fig. 3B).</paragraph>
|
||
<paragraph id="8BD136CB2766FFE2D3F142DE20FBFA3F" blockId="3.[151,1436,867,2008]" pageId="3">Second chelate legs dissimilar, unequal and covered with spinules (fig. 1). Length range of larger second chelate leg 19.5 to 73.0 mm, 0.57 to 1.22 times total body length. Merus shorter than carpus (fig. 2B), about half length of chela; carpus shorter than chela; palm longer than fingers. Width of palm slightly greater than width of distal carpus; palm of larger chela narrower than smaller (fig. 2B, D). Fingers stout and equal, two large denticles at proximal part of movable finger, six weak denticles at distal part. Similar pattern of dentition present in immovable finger also (fig. 2C). In small second pereiopod, merus shorter than carpus, about half length of chela; carpus shorter than chela, palm longer than fingers; width of palm greater than width of distal carpus. Fingers stout, equal, unarmed, wide gap between fingers when closed. Numerous setae on inner margin of both fingers (fig. 2E). Second large and small, third, fourth and fifth pereiopods densely spinulose, spines distributed uniformly.</paragraph>
|
||
<paragraph id="8BD136CB2766FFE2D3F1439A26D9F9AB" blockId="3.[151,1436,867,2008]" pageId="3">Non-chelate legs and pleopods, normal in structure of the genus (fig. 2F, 3D). Appendix interna less than half length of appendix masculina, spinous setae on inner side (fig. 3C). In the present specimens, telson extends beyond level of outer lateral spine of uropodal exopod.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465402766FFE7D3F1406E2617F839" lastPageId="6" pageId="3" type="discussion">
|
||
<paragraph id="8BD136CB2766FFE2D3F1406E211AF917" blockId="3.[151,1436,867,2008]" pageId="3">
|
||
<emphasis id="B91AEAD92766FFE2D3F1406E251CF9CF" bold="true" box="[199,315,1587,1612]" pageId="3">Remarks.</emphasis>
|
||
The present specimens share most of its characters with
|
||
<taxonomicName id="4C6E4D482766FFE2D0F0406920D9F9CF" authority="Nobili, 1906" authorityName="Nobili" authorityYear="1906" box="[966,1278,1587,1612]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92766FFE2D0F040692073F9CF" box="[966,1108,1588,1612]" italics="true" pageId="3">M. aemulum</emphasis>
|
||
(Nobili, 1906)
|
||
</taxonomicName>
|
||
collected and described from
|
||
<collectingCountry id="F379765B2766FFE2D27B400425D9F9F3" box="[333,510,1623,1648]" name="New Caledonia" pageId="3">New Caledonia</collectingCountry>
|
||
and subsequently recorded from
|
||
<collectingCountry id="F379765B2766FFE2D0B3400A2799F9F3" box="[901,958,1623,1648]" name="India" pageId="3">India</collectingCountry>
|
||
by Jayachandran and Joseph (1988), but there are perceptible differences with these specimens in certain distinctive diagnostic characters (table 2).
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2766FFE2D3F140C22075F85B" blockId="3.[151,1436,867,2008]" pageId="3">
|
||
According to Nobili (1906), in
|
||
<taxonomicName id="4C6E4D482766FFE2D10640FC2699F93B" box="[560,702,1696,1720]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="3" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92766FFE2D10640FC2699F93B" box="[560,702,1696,1720]" italics="true" pageId="3">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
, rostrum short, always a little shorter than antennular peduncle, slightly turned down, but elevated a little towards tip with rostral formula 10–13/1–3 with 4–5 dorsal rostral teeth, carapace smooth. Second chelate legs unequal in size, larger second pereiopod with merus shorter than carpus and about half the size of chela (fig.5), carpus shorter than chela. Basal third of the movable finger with a big tooth, preceded at the articulation by two small teeth; followed by another big tooth in the middle; over which follow one another to the ends 15–16 pairs of blackish tubercles, a little more larger than the tubercles lining the finger. The smaller chelate leg repeats nearly the same provision, but the fingers much shorter and more arched and far apart. Ischium shorter than merus, merus about half size of chela, carpus shorter than chela; palm almost twice length of fingers. Telson bears two pairs of distal spines; inner pair much longer than the outer.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2761FFE5D3A140CF24E2F96B" blockId="4.[151,1436,1682,1768]" pageId="4">
|
||
<emphasis id="B91AEAD92761FFE5D3A140CF2535F924" bold="true" box="[151,274,1682,1704]" pageId="4">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C6E4D482761FFE5D22F40CF269BF92B" box="[281,700,1682,1704]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
|
||
<emphasis id="B91AEAD92761FFE5D22F40CF269BF92B" box="[281,700,1682,1704]" italics="true" pageId="4">Macrobrachium aemulum madhusoodani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957A22761FFE5D1F540CF271FF92B" box="[707,824,1682,1704]" pageId="4" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
, specimen no. 1,
|
||
<typeStatus id="54D588692761FFE5D0D140CF2067F92B" box="[999,1088,1682,1704]" pageId="4" type="holotype">holotype</typeStatus>
|
||
(male). A, telson and uropod; B, 2nd large chelate leg; C, 2nd large chela enlarged; D, 2nd small chelate leg; E, 2nd small chela enlarged; F, 3rd pereiopod, distal part.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2761FFE5D3F141492792F867" blockId="4.[151,1436,1812,2020]" pageId="4">
|
||
Jayachandran and Joseph (1988) recorded
|
||
<taxonomicName id="4C6E4D482761FFE5D19B414B271CF8AE" box="[685,827,1813,1837]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="4" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92761FFE5D19B414B271CF8AE" box="[685,827,1813,1837]" italics="true" pageId="4">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
from Indian waters, and compared the lengths of the podomeres of second chelate leg of specimens from
|
||
<collectingCountry id="F379765B2761FFE5D03641672795F8D1" box="[768,946,1849,1874]" name="New Caledonia" pageId="4">New Caledonia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F379765B2761FFE5D0D94164200EF8D1" box="[1007,1065,1849,1874]" name="India" pageId="4">India</collectingCountry>
|
||
to establish similarities in their morphometric characters and also noticed some other variations in Indian specimens. Width of rostrum just in front of orbit slightly greater, 2nd cheliped unequal, palm width of smaller cheliped greater than that of the larger, gap between fingers when closed wider in smaller cheliped than larger etc. are the variations. Rostral formula 11/2–3 also is within the range 10–13/1–3 seen in New Caledonian specimens.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2760FFE4D3A1410227B7F815" blockId="5.[151,1435,1887,1942]" pageId="5">
|
||
<emphasis id="B91AEAD92760FFE4D3A141022533F8F7" bold="true" box="[151,276,1887,1909]" pageId="5">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C6E4D482760FFE4D22A410226E6F8F6" box="[284,705,1887,1909]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="5" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
|
||
<emphasis id="B91AEAD92760FFE4D22A410226E6F8F6" box="[284,705,1887,1909]" italics="true" pageId="5">Macrobrachium aemulum madhusoodani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957A22760FFE4D1FF41022718F8F6" box="[713,831,1887,1909]" pageId="5" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
, specimen no. 1,
|
||
<typeStatus id="54D588692760FFE4D0C24102206AF8F6" box="[1012,1101,1887,1909]" pageId="5" type="holotype">holotype</typeStatus>
|
||
(male). A, carapace-right lateral view; B, 1st pereiopod, distal part; C, 2nd pleopod of male; D, 3rd pleopod.
|
||
</paragraph>
|
||
<caption id="DF1166432763FFE7D3A145E92172FC49" box="[151,1365,948,970]" httpUri="https://zenodo.org/record/224215/files/figure.png" pageId="6" targetBox="[178,1404,221,894]" targetPageId="6">
|
||
<paragraph id="8BD136CB2763FFE7D3A145E92172FC49" blockId="6.[151,1365,948,970]" box="[151,1365,948,970]" pageId="6">
|
||
<emphasis id="B91AEAD92763FFE7D3A145E92535FC4A" bold="true" box="[151,274,948,970]" pageId="6">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C6E4D482763FFE7D22F45E9269BFC49" box="[281,700,948,970]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="subSpecies" species="aemulum" status="subsp. nov." subSpecies="madhusoodani">
|
||
<emphasis id="B91AEAD92763FFE7D22F45E9269BFC49" box="[281,700,948,970]" italics="true" pageId="6">Macrobrachium aemulum madhusoodani</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22957A22763FFE7D1F445E9271FFC49" box="[706,824,948,970]" pageId="6" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
, specimen no. 2, paratype. Carapace left lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD136CB2763FFE7D3F145AB260DFA8E" blockId="6.[151,1436,1014,1978]" pageId="6">
|
||
There are similarities and differences between the new subspecies and the
|
||
<taxonomicName id="4C6E4D482763FFE7D75C45A520D8FB8C" box="[1130,1279,1015,1039]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D75C45A520D8FB8C" box="[1130,1279,1015,1039]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
recorded by Jayachandran and Joseph. In
|
||
<emphasis id="B91AEAD92763FFE7D2D742412653FBB0" box="[481,628,1051,1075]" italics="true" pageId="6">
|
||
<taxonomicName id="4C6E4D482763FFE7D2D742412657FBB0" box="[481,624,1051,1075]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">M. aemulum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
rostrum short, extends up to the middle of distal segment of antennular peduncle, tip very sharp, directed slightly upwards. In the new subspecies also the same characters could be seen. Jayachandran recorded 11 dorsal rostral teeth in
|
||
<taxonomicName id="4C6E4D482763FFE7D18C42392760FBF8" box="[698,839,1123,1147]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D18C42392760FBF8" box="[698,839,1123,1147]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
whereas in the new subspecies, it ranges from 9–13. However, number of dorsal rostral postorbital teeth is 4–5 (interval between first four teeth greater than the remaining teeth), which is similar to the situation in the new subspecies. Moreover ventral margin of rostrum in
|
||
<taxonomicName id="4C6E4D482763FFE7D64B42F124D9FB69" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D64B42F124D9FB69" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
has 2–3 teeth (situated in between 9th and 11th dorsal teeth), but in new subspecies, it is 2 only, whereas carapace smooth in both specimens.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2763FFE7D3F14347200EF919" blockId="6.[151,1436,1014,1978]" pageId="6">
|
||
In first pereiopod of
|
||
<emphasis id="B91AEAD92763FFE7D2834347266DFAB1" box="[437,586,1306,1330]" italics="true" pageId="6">
|
||
<taxonomicName id="4C6E4D482763FFE7D28343472661FAB1" box="[437,582,1306,1330]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">M. aemulum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
ischium about half size of merus; merus shorter than carpus, carpus twice size of chela, palm longer than fingers and the new subspecies also agree with these characters. In
|
||
<emphasis id="B91AEAD92763FFE7D7ED4363214AFAD6" box="[1243,1389,1341,1365]" italics="true" pageId="6">
|
||
<taxonomicName id="4C6E4D482763FFE7D7ED4363214EFAD6" box="[1243,1385,1341,1365]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">M. aemulum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and it’s new subspecies, second pereiopods are strong, distinctly unequal and dissimilar in shape. Merus shorter than carpus, about half length of chela in larger second pereiopod of
|
||
<taxonomicName id="4C6E4D482763FFE7D04B43DB202CFA1E" box="[893,1035,1413,1437]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D04B43DB202CFA1E" box="[893,1035,1413,1437]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
; carpus shorter than chela (fig. 6). Width of palm slightly greater than that of distal carpus, palm of larger chela, narrower than smaller. Proximal part of cutting edges of movable finger with two denticles, fixed finger with one, rest of cutting edge entire. Whole leg scabrous due to close-set spinules. The new subspecies, share characters with
|
||
<taxonomicName id="4C6E4D482763FFE7D73643AF20ABF989" box="[1024,1164,1522,1546]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D73643AF20ABF989" box="[1024,1164,1522,1546]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
, like merus shorter than carpus, about half-length of chela, carpus shorter than chela, palm longer than fingers; width of palm slightly greater than that of distal carpus. Palm of larger chelate leg narrower than the smaller chelate. Fingers are stout and equal, but in the new subspecies movable finger has two large denticles in the proximal part and six weak denticles in the distal part. Similar pattern of dentition is present on immovable finger also.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2763FFE7D3F140FB273EF8CD" blockId="6.[151,1436,1014,1978]" pageId="6">
|
||
In smaller second pereiopod of
|
||
<emphasis id="B91AEAD92763FFE7D11340FB269FF93E" box="[549,696,1701,1725]" italics="true" pageId="6">
|
||
<taxonomicName id="4C6E4D482763FFE7D11340FB2693F93E" box="[549,692,1701,1725]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">M. aemulum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
ischium shorter than merus, merus about half size of chela, carpus shorter than chela; palm more than twice length of fingers. Fingers stout, equal, distinct gap when closed; wider than that between fingers of larger chela. Cutting edges of fingers with 1–2 weak denticles in proximal part, stiff setae at cutting edges. The new subspecies also show similar characters except that cutting edges of fingers unarmed, numerous stiff setae at cutting edges of fingers.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB2763FFE7D3F141072617F839" blockId="6.[151,1436,1014,1978]" pageId="6">
|
||
In
|
||
<taxonomicName id="4C6E4D482763FFE7D3D041072555F8F1" box="[230,370,1882,1906]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D3D041072555F8F1" box="[230,370,1882,1906]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
telson reaches up to the level of the outer lateral spine of the uropodal exopod, whereas in the present specimens it extends beyond. In both
|
||
<taxonomicName id="4C6E4D482763FFE7D19F41232710F816" box="[681,823,1917,1941]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="6" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92763FFE7D19F41232710F816" box="[681,823,1917,1941]" italics="true" pageId="6">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
and new subspecies, there are four pairs of plumose setae between inner spines of telson.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37465402762FFE9D3A141FC266CF94D" lastPageId="8" pageId="7" type="description">
|
||
<paragraph id="8BD136CB2762FFE6D3A141FC2650F856" blockId="7.[151,1437,1949,2005]" pageId="7">
|
||
<emphasis id="B91AEAD92762FFE6D3A141FC2534F835" bold="true" box="[151,275,1953,1975]" pageId="7">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C6E4D482762FFE6D22D41FC269AF834" authority="Nobili, 1906" authorityName="Nobili" authorityYear="1906" box="[283,701,1953,1975]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="7" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD92762FFE6D22D41FC2603F834" box="[283,548,1953,1975]" italics="true" pageId="7">Macrobrachium aemulum</emphasis>
|
||
(Nobili, 1906)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F379765B2762FFE6D1CB41FF27B9F834" box="[765,926,1953,1975]" name="New Caledonia" pageId="7">New Caledonia</collectingCountry>
|
||
. A, 2nd small chelate leg; B, 2nd large chelate leg. (Courtesy, Bull. Mus. Histoire Naturelle, Paris)
|
||
</paragraph>
|
||
<paragraph id="8BD136CB276DFFE9D3A142B427B5FAA3" blockId="8.[151,1435,1257,1312]" pageId="8">
|
||
<emphasis id="B91AEAD9276DFFE9D3A142B42534FB7D" bold="true" box="[151,275,1257,1279]" pageId="8">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C6E4D48276DFFE9D22F42B42690FB7C" authority="Nobili, 1906" authorityName="Nobili" authorityYear="1906" box="[281,695,1257,1279]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD9276DFFE9D22F42B42607FB7C" box="[281,544,1257,1279]" italics="true" pageId="8">Macrobrachium aemulum</emphasis>
|
||
(Nobili, 1906)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F379765B276DFFE9D1C342B4270BFB7C" box="[757,812,1257,1279]" name="India" pageId="8">India</collectingCountry>
|
||
, Jayachandran and Joseph, 1988. A, 2nd small chelate leg; B, 2nd large chelate leg. (Courtesy, Oxford and IBH Publ. Company Pvt. Ltd.).
|
||
</paragraph>
|
||
<paragraph id="8BD136CB276DFFE9D3F14313276EFA75" blockId="8.[151,1436,1356,1814]" pageId="8">
|
||
In the specimens of Nobili (1906) and Roux (1926), fingers of second chelate legs are slender. However, in
|
||
<taxonomicName id="4C6E4D48276DFFE9D64B431324D9FA0A" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD9276DFFE9D64B431324D9FA0A" italics="true" pageId="8">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
reported by Jayachandran and Joseph and new subspecies, fingers of second chelate legs are stout. The new subspecies has two large denticles at proximal part of movable finger and six weak denticles at the distal part. Similar pattern of dentition is present in immovable finger also. But the specimens of Nobili and Roux have a strong triangular tooth at about half the length of fixed finger.
|
||
</paragraph>
|
||
<paragraph id="8BD136CB276DFFE9D3F1405D266CF94D" blockId="8.[151,1436,1356,1814]" pageId="8">
|
||
The collections during the pre monsoon consisted of
|
||
<taxonomicName id="4C6E4D48276DFFE9D07D405C27FAF99A" box="[843,989,1537,1561]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD9276DFFE9D07D405C27FAF99A" box="[843,989,1537,1561]" italics="true" pageId="8">M. aemulum</emphasis>
|
||
</taxonomicName>
|
||
(11.8%) and new subspecies (7.3%). However, during the monsoon and post monsoon seasons, representation of
|
||
<taxonomicName id="4C6E4D48276DFFE9D726407B2169F9BD" authority="Nobili, 1906" authorityName="Nobili" authorityYear="1906" box="[1040,1358,1573,1598]" class="Malacostraca" family="Palaemonidae" genus="Macrobrachium" kingdom="Animalia" order="Decapoda" pageId="8" phylum="Arthropoda" rank="species" species="aemulum">
|
||
<emphasis id="B91AEAD9276DFFE9D726407B2087F9BE" box="[1040,1184,1573,1597]" italics="true" pageId="8">M. aemulum</emphasis>
|
||
(Nobili, 1906)
|
||
</taxonomicName>
|
||
in the collections was 11.8 % and 6.4% respectively whereas new subspecies could not be found. Even though the new subspecies described here share some characters with the specimens of Nobili, whether it can be designated as new species has to be confirmed only after gene sequencing. This was not attempted as prolonged formalin preservation often fails to produce precision results.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3746540276DFFE9D3F140852176F895" pageId="8" type="etymology">
|
||
<paragraph id="8BD136CB276DFFE9D3F140852176F895" blockId="8.[151,1436,1356,1814]" pageId="8">
|
||
<emphasis id="B91AEAD9276DFFE9D3F140852569F972" bold="true" box="[199,334,1752,1777]" pageId="8">Etymology.</emphasis>
|
||
The subspecies name is in honor of P. Madhusoodanan Pillai, former Professor, Mahatma Gandhi College, Thiruvananthapuram and teacher of the junior author. The name thus is a noun in genitive singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |