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<mods:title id="A0198FA3A96E599B2A738A98666BFED8">First record of the genus Chorinaeus Holmgren, 1858 (Hymenoptera: Ichneumonidae) from India and Thailand with the description of two new species</mods:title>
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<mods:namePart id="5CCE260CC498400D900FC2ED9FF2B81B">Ranjith, A. P.</mods:namePart>
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<mods:affiliation id="07BA60A27F2A8C7DC33F1474EDB59E4C">Integrative Insect Ecology Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
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<mods:namePart id="38C22D1A5A77391FAF9A0A2BFE975061">Butcher, Buntika A.</mods:namePart>
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<mods:affiliation id="ADE7C78ABEF13E0F5B1E4718F5F31118">Integrative Insect Ecology Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, Thailand</mods:affiliation>
<mods:nameIdentifier id="1E72EAF34E559CCACA07AC94614EDF3C" type="email">buntika.a@chula.ac.th</mods:nameIdentifier>
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<heading id="D09981C4FFFDFFB1579DA446CB6F377E" bold="true" box="[151,875,1036,1063]" fontSize="11" level="1" pageId="1" pageNumber="284" reason="1">
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Key to the Oriental species of
<taxonomicName id="4C6E4D2BFFFDFFB15504A447CB6F377E" ID-CoL="3P4P" ID-ENA="494203" authority="Holmgren, 1858" authorityName="Holmgren" authorityYear="1858" box="[526,875,1036,1063]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEABAFFFDFFB15504A447CA98377F" bold="true" box="[526,668,1036,1063]" italics="true" pageId="1" pageNumber="284">Chorinaeus</emphasis>
Holmgren, 1858
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<subSubSection id="C3746523FFFDFFB1579DA419CD983559" pageId="1" pageNumber="284" type="key">
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<paragraph id="8BD136A8FFFDFFB1579DA419CD9837DC" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA419CD9837DC" pageId="1" pageNumber="284">
1. Propodeal spiracle more close to pleural carina; malar space 2.03.0 × as long as basal width of mandible..........................................................................................
<taxonomicName id="4C6E4D2BFFFDFFB15361A424CD9837DC" ID-CoL="5YC5P" authority="Chao, 1981" authorityName="Chao" authorityYear="1981" box="[1131,1436,1135,1157]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="facialis">
<emphasis id="B91AEABAFFFDFFB15361A424CD2C37DC" box="[1131,1320,1135,1156]" italics="true" pageId="1" pageNumber="284">Chorinaeus facialis</emphasis>
Chao, 1981
</taxonomicName>
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</paragraph>
<paragraph id="8BD136A8FFFDFFB1579DA4C6CD9837E7" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA4C6CD9837E7" pageId="1" pageNumber="284">
Propodeal spiracle more close to lateral carina (
<figureCitation id="13552A2DFFFDFFB155A0A4C7CAFD37F9" box="[682,761,1164,1186]" captionStart="FIGURE 2" captionStartId="3.[152,255,1673,1697]" captionTargetBox="[169,1414,187,1640]" captionTargetId="figure-24@3.[159,1428,178,1652]" captionTargetPageId="3" captionText="FIGURE 2. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) scutellum and propodeum, dorsal view; B) propodeum and first metasomal tergite, dorso-lateral view; C) metasoma, lateral view; D) metasoma, dorsal view, E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.11611071" httpUri="https://zenodo.org/record/11611071/files/figure.png" pageId="1" pageNumber="284">Figs 2B</figureCitation>
,
<figureCitation id="13552A2DFFFDFFB15409A4C7CB2637F9" box="[771,802,1164,1185]" captionStart="FIGURE 4" captionStartId="6.[152,255,1676,1700]" captionTargetBox="[169,1413,189,1641]" captionTargetId="figure-29@6.[160,1427,178,1654]" captionTargetPageId="6" captionText="FIGURE 4. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) scutellum and propodeum, dorsal view; B) propodeum and first metasomal tergite, dorso-lateral view; C) metasoma, lateral view; D) metasoma, dorsal view, E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.11611082" httpUri="https://zenodo.org/record/11611082/files/figure.png" pageId="1" pageNumber="284">4B</figureCitation>
); malar space less than 2.0 × as long as basal width of mandible. ................................................................................................... 2
</keyLead>
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<keyStep id="309A2E0DFFFDFFB1579DA48CCD9837A1" pageId="1" pageNumber="284">
<paragraph id="8BD136A8FFFDFFB1579DA48CCD983784" blockId="1.[151,1437,1105,1538]" box="[151,1436,1223,1245]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA48CCD983784" box="[151,1436,1223,1245]" pageId="1" pageNumber="284">
2. Face, clypeus and malar space yellow (
<figureCitation id="13552A2DFFFDFFB15553A48CCAC03784" box="[601,708,1223,1245]" captionStart="FIGURE 3" captionStartId="5.[152,255,1876,1900]" captionTargetBox="[171,1416,388,1840]" captionTargetId="figure-119@5.[161,1425,378,1855]" captionTargetPageId="5" captionText="FIGURE 3. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611075" httpUri="https://zenodo.org/record/11611075/files/figure.png" pageId="1" pageNumber="284">Figs 3B, D</figureCitation>
).......................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BD136A8FFFDFFB1579DA4AECD9837A1" blockId="1.[151,1437,1105,1538]" box="[151,1436,1252,1274]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA4AECD9837A1" box="[151,1436,1252,1274]" pageId="1" pageNumber="284">
Face brown or at least with brown patch (
<figureCitation id="13552A2DFFFDFFB1557AA4AFCAD937A1" box="[624,733,1252,1274]" captionStart="FIGURE 1" captionStartId="2.[152,255,1982,2006]" captionTargetBox="[171,1415,495,1946]" captionTargetId="figure-180@2.[162,1425,486,1961]" captionTargetPageId="2" captionText="FIGURE 1. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611069" httpUri="https://zenodo.org/record/11611069/files/figure.png" pageId="1" pageNumber="284">Figs 1B, D</figureCitation>
), clypeus and malar space brown (
<figureCitation id="13552A2DFFFDFFB15322A4AFCC9737A1" box="[1064,1171,1252,1274]" captionStart="FIGURE 1" captionStartId="2.[152,255,1982,2006]" captionTargetBox="[171,1415,495,1946]" captionTargetId="figure-180@2.[162,1425,486,1961]" captionTargetPageId="2" captionText="FIGURE 1. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611069" httpUri="https://zenodo.org/record/11611069/files/figure.png" pageId="1" pageNumber="284">Figs 1B, D</figureCitation>
).................... 4
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<paragraph id="8BD136A8FFFDFFB1579DA549CD98366C" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA549CD98366C" pageId="1" pageNumber="284">
3. Antenna with 38 antennomeres; second metasomal tergite with short sublateral longitudinal carina basally; hind femur black; hind tibia black in apical half, remainder yellow............................
<taxonomicName id="4C6E4D2BFFFDFFB1530EA554CD98366C" ID-CoL="5YBTC" authority="Kusigemati, 1987" authorityName="Kusigemati" authorityYear="1987" box="[1028,1436,1311,1333]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="ryukyuensis">
<emphasis id="B91AEABAFFFDFFB1530EA554CCEF366C" box="[1028,1259,1311,1332]" italics="true" pageId="1" pageNumber="284">Chorinaeus ryukyuensis</emphasis>
Kusigemati, 1987
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD136A8FFFDFFB1579DA576CD9836D4" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA576CD9836D4" pageId="1" pageNumber="284">
Antenna with 25 antennomeres (
<figureCitation id="13552A2DFFFDFFB15514A577CA683609" box="[542,620,1340,1362]" captionStart="FIGURE 3" captionStartId="5.[152,255,1876,1900]" captionTargetBox="[171,1416,388,1840]" captionTargetId="figure-119@5.[161,1425,378,1855]" captionTargetPageId="5" captionText="FIGURE 3. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611075" httpUri="https://zenodo.org/record/11611075/files/figure.png" pageId="1" pageNumber="284">Fig. 3A</figureCitation>
); second metasomal tergite without sublateral longitudinal carina (
<figureCitation id="13552A2DFFFDFFB153F8A577CD5B3609" box="[1266,1375,1340,1362]" captionStart="FIGURE 4" captionStartId="6.[152,255,1676,1700]" captionTargetBox="[169,1413,189,1641]" captionTargetId="figure-29@6.[160,1427,178,1654]" captionTargetPageId="6" captionText="FIGURE 4. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) scutellum and propodeum, dorsal view; B) propodeum and first metasomal tergite, dorso-lateral view; C) metasoma, lateral view; D) metasoma, dorsal view, E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.11611082" httpUri="https://zenodo.org/record/11611082/files/figure.png" pageId="1" pageNumber="284">Figs 4B, C</figureCitation>
); hind femur brown medially, yellow basally and apically (
<figureCitation id="13552A2DFFFDFFB155DCA512CB273637" box="[726,803,1369,1391]" captionStart="FIGURE 3" captionStartId="5.[152,255,1876,1900]" captionTargetBox="[171,1416,388,1840]" captionTargetId="figure-119@5.[161,1425,378,1855]" captionTargetPageId="5" captionText="FIGURE 3. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611075" httpUri="https://zenodo.org/record/11611075/files/figure.png" pageId="1" pageNumber="284">Fig. 3A</figureCitation>
); hind tibia completely yellow (
<figureCitation id="13552A2DFFFDFFB1535FA512CCA43637" box="[1109,1184,1369,1391]" captionStart="FIGURE 3" captionStartId="5.[152,255,1876,1900]" captionTargetBox="[171,1416,388,1840]" captionTargetId="figure-119@5.[161,1425,378,1855]" captionTargetPageId="5" captionText="FIGURE 3. Chorinaeus femiae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611075" httpUri="https://zenodo.org/record/11611075/files/figure.png" pageId="1" pageNumber="284">Fig. 3A</figureCitation>
)..................................................................................
<taxonomicName id="4C6E4D2BFFFDFFB154D6A53CCD4E36D5" authority="Ranjith &amp; Butcher" authorityName="Ranjith &amp; Butcher" authorityYear="2024" box="[988,1354,1399,1421]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="femiae" status="sp. nov.">
<emphasis id="B91AEABAFFFDFFB154D6A53CCC9636D4" box="[988,1170,1399,1420]" italics="true" pageId="1" pageNumber="284">Chorinaeus femiae</emphasis>
Ranjith &amp; Butcher
</taxonomicName>
<emphasis id="B91AEABAFFFDFFB1525AA53CCD9836D4" bold="true" box="[1360,1436,1399,1420]" pageId="1" pageNumber="284">
<taxonomicNameLabel id="A22957C1FFFDFFB1525AA53CCD9836D4" box="[1360,1436,1399,1420]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
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<paragraph id="8BD136A8FFFDFFB1579DA5DFCD98369F" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA5DFCD98369F" pageId="1" pageNumber="284">
4. Lateral carina of scutellum very weak or obsolete; first flagellomere 2.5 × as long as wide; first metasomal tergite smooth without punctures....................................................
<taxonomicName id="4C6E4D2BFFFDFFB15318A5FACD98369F" ID-CoL="5YBTG" authority="Kusigemati, 1967" authorityName="Kusigemati" authorityYear="1967" box="[1042,1436,1457,1479]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="aizanensis">
<emphasis id="B91AEABAFFFDFFB15318A5FACCE8369F" box="[1042,1260,1457,1479]" italics="true" pageId="1" pageNumber="284">Chorinaeus aizanensis</emphasis>
Kusigemati, 1967
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD136A8FFFDFFB1579DA584CD983559" blockId="1.[151,1437,1105,1538]" pageId="1" pageNumber="284">
<keyLead id="309F959DFFFDFFB1579DA584CD983559" pageId="1" pageNumber="284">
Lateral carina of scutellum strong and complete (
<figureCitation id="13552A2DFFFDFFB155B0A584CB0736BC" box="[698,771,1487,1509]" captionStart="FIGURE 1" captionStartId="2.[152,255,1982,2006]" captionTargetBox="[171,1415,495,1946]" captionTargetId="figure-180@2.[162,1425,486,1961]" captionTargetPageId="2" captionText="FIGURE 1. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) habitus, lateral view; B) head, anterior view; C) head, dorsal view; D) head, antero-ventral view; E) head and mesosoma, lateral view; F) mesosoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11611069" httpUri="https://zenodo.org/record/11611069/files/figure.png" pageId="1" pageNumber="284">Figs 1F</figureCitation>
,
<figureCitation id="13552A2DFFFDFFB15404A584CB2B36BC" box="[782,815,1487,1508]" captionStart="FIGURE 2" captionStartId="3.[152,255,1673,1697]" captionTargetBox="[169,1414,187,1640]" captionTargetId="figure-24@3.[159,1428,178,1652]" captionTargetPageId="3" captionText="FIGURE 2. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) scutellum and propodeum, dorsal view; B) propodeum and first metasomal tergite, dorso-lateral view; C) metasoma, lateral view; D) metasoma, dorsal view, E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.11611071" httpUri="https://zenodo.org/record/11611071/files/figure.png" pageId="1" pageNumber="284">2A</figureCitation>
); first flagellomere 2.8 × as long as wide; first metasomal tergite smooth anterior half, rugose-punctate posterior half (
<figureCitation id="13552A2DFFFDFFB155D4A5A7CB2D3559" box="[734,809,1516,1538]" captionStart="FIGURE 2" captionStartId="3.[152,255,1673,1697]" captionTargetBox="[169,1414,187,1640]" captionTargetId="figure-24@3.[159,1428,178,1652]" captionTargetPageId="3" captionText="FIGURE 2. Chorinaeus amrithae Ranjith &amp; Butcher sp. nov., holotype, female A) scutellum and propodeum, dorsal view; B) propodeum and first metasomal tergite, dorso-lateral view; C) metasoma, lateral view; D) metasoma, dorsal view, E) fore wing." figureDoi="http://doi.org/10.5281/zenodo.11611071" httpUri="https://zenodo.org/record/11611071/files/figure.png" pageId="1" pageNumber="284">Fig. 2D</figureCitation>
)............
<taxonomicName id="4C6E4D2BFFFDFFB154CCA5A7CD4E355A" authority="Ranjith &amp; Butcher" authorityName="Ranjith &amp; Butcher" authorityYear="2024" box="[966,1354,1516,1538]" class="Insecta" family="Ichneumonidae" genus="Chorinaeus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="amrithae" status="sp. nov.">
<emphasis id="B91AEABAFFFDFFB154CCA5A7CC973559" box="[966,1171,1516,1537]" italics="true" pageId="1" pageNumber="284">Chorinaeus amrithae</emphasis>
Ranjith &amp; Butcher
</taxonomicName>
<emphasis id="B91AEABAFFFDFFB1525AA5A7CD983559" bold="true" box="[1360,1436,1516,1537]" pageId="1" pageNumber="284">
<taxonomicNameLabel id="A22957C1FFFDFFB1525AA5A7CD983559" box="[1360,1436,1516,1537]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>