treatments-xml/data/03/C7/87/03C787A14D4DD220E43AFAC1FE41FBE7.xml

<document id="2BB209F596E7A44BDD0F8B45958F9DE9" ID-DOI="10.5281/zenodo.173337" ID-GBIF-Dataset="773475fd-f7cb-4f71-993a-bc7424e1ea52" ID-ISSN="1175­5326" ID-Zenodo-Dep="173337" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459773031704" checkinUser="plazi" docAuthor="Bohn, Jens Michael" docDate="2006" docId="03C787A14D4DD220E43AFAC1FE41FBE7" docLanguage="en" docName="zt01276p031.pdf" docOrigin="Zootaxa 1276" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Achlyonice longicornis Bohn, 2006, spec. nov." docType="treatment" docVersion="7" lastPageNumber="16" masterDocId="FFFEFFD94D40D22FE532FFFFFFE9FF99" masterDocTitle="Crinoidea and Holothuroidea (Echinodermata) of the abyssal Angola Basin — Results of the DIVA­ 1 expedition of FS &quot; Meteor &quot; (Cruise M 48 / 1)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="14" updateTime="1698217476750" updateUser="plazi">
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<mods:title id="09AFA41E84A6713BADE9229C6C006ED3">Crinoidea and Holothuroidea (Echinodermata) of the abyssal Angola Basin — Results of the DIVA­ 1 expedition of FS &quot; Meteor &quot; (Cruise M 48 / 1)</mods:title>
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<mods:namePart id="CA0941A3C93C87871360338CF9C904E7">Bohn, Jens Michael</mods:namePart>
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<mods:date id="44EFB8E25C32CA2200B1F60E4CC2CC76">2006</mods:date>
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<treatment id="03C787A14D4DD220E43AFAC1FE41FBE7" ID-DOI="http://doi.org/10.5281/zenodo.6255864" ID-GBIF-Taxon="119347430" ID-Zenodo-Dep="6255864" LSID="urn:lsid:plazi:treatment:03C787A14D4DD220E43AFAC1FE41FBE7" httpUri="http://treatment.plazi.org/id/03C787A14D4DD220E43AFAC1FE41FBE7" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<paragraph id="8BD136B74D4DD222E43AFAC1FD77FAC1" blockId="13.[264,670,1342,1403]" box="[264,670,1342,1368]" pageId="13" pageNumber="14">
<heading id="D09981DB4D4DD222E43AFAC1FD77FAC1" bold="true" box="[264,670,1342,1368]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B91AEAA54D4DD222E43AFAC1FD77FAC1" bold="true" box="[264,670,1342,1368]" pageId="13" pageNumber="14">
<taxonomicName id="4C6E4D344D4DD222E43AFAC1FDF6FAC1" ID-CoL="9869" box="[264,543,1342,1368]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="longicornis" status="sp. nov.">
<emphasis id="B91AEAA54D4DD222E43AFAC1FDF6FAC1" bold="true" box="[264,543,1342,1368]" italics="true" pageId="13" pageNumber="14">Achlyonice longicornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957DE4D4DD222E715FAC1FD77FAC1" box="[551,670,1342,1368]" pageId="13" pageNumber="14" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD136B74D4DD222E43AFA9EFE5FFAE2" blockId="13.[264,670,1342,1403]" box="[264,438,1377,1403]" pageId="13" pageNumber="14">
(
<figureCitation id="13552A324D4DD222E422FA9EFEB2FAE2" box="[272,347,1377,1403]" captionStart="FIGURE 7" captionStartId="12.[264,367,1779,1803]" captionTargetBox="[270,1309,1228,1745]" captionTargetId="figure@12.[264,1323,1226,1755]" captionTargetPageId="12" captionText="FIGURE 7. Distribution of Peniagone purpurea (Théel, 1882) and Achlyonice longicornis spec. nov." httpUri="https://zenodo.org/record/173343/files/figure.png" pageId="13" pageNumber="14">Figs 7</figureCitation>
, 
<figureCitation id="13552A324D4DD222E45BFA9EFE9FFAE2" box="[361,374,1377,1403]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="13" pageNumber="14">8</figureCitation>
A–E)
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4DD222E43AFA4EFEA2F982" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BD136B74D4DD222E43AFA4EFEA2F982" blockId="13.[264,1326,1457,1883]" pageId="13" pageNumber="14">
<emphasis id="B91AEAA54D4DD222E43AFA4EFE95FA52" bold="true" box="[264,380,1457,1483]" pageId="13" pageNumber="14">Material.</emphasis>
<typeStatus id="54D588154D4DD222E4BCFA4EFE12FA52" box="[398,507,1457,1483]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED7FAE724D4DD222E724FA4EFDBDFA52" box="[534,596,1457,1483]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="13" pageNumber="14">ZSM</collectionCode>
20043073), FS &quot;Meteor&quot;, DIVA­1, station M 48/1­351, 
<geoCoordinate id="EE5A50704D4DD222E43AFA26FE6EFA6A" box="[264,391,1497,1523]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="92" value="-16.42">16°25.2' S</geoCoordinate>
, 
<geoCoordinate id="EE5A50704D4DD222E4A5FA26FDE2FA6A" box="[407,523,1497,1523]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="92" value="5.451667">5°27.1' E</geoCoordinate>
, 
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to 
<geoCoordinate id="EE5A50704D4DD222E791FA26FCCBFA6A" box="[675,802,1497,1523]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="92" value="-16.553333">16°33.2' S</geoCoordinate>
, 
<geoCoordinate id="EE5A50704D4DD222E601FA26FC4CFA6A" box="[819,933,1497,1523]" direction="east" orientation="longitude" pageId="13" pageNumber="14" precision="92" value="5.455">5°27.3' E</geoCoordinate>
, 
<quantity id="4C969B524D4DD222E687FA26FBFCFA6A" box="[949,1045,1497,1523]" metricMagnitude="3" metricUnit="m" metricValue="5.385" pageId="13" pageNumber="14" unit="m" value="5385.0">5385 m</quantity>
, Agassiz trawl, 
<date id="FFD010774D4DD222E1E9FA26FAC0FA6A" box="[1243,1321,1497,1523]" pageId="13" pageNumber="14" value="1930-07">30 Jul</date>
, 2000.
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4DD221E40AF9D6FE5DFCD9" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BD136B74D4DD221E40AF9D6FB61FE67" blockId="13.[264,1326,1457,1883]" lastBlockId="14.[264,1324,284,1272]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B91AEAA54D4DD222E40AF9D6FE26F9DA" bold="true" box="[312,463,1577,1603]" pageId="13" pageNumber="14">Description.</emphasis>
The 
<typeStatus id="54D588154D4DD222E726F9D6FD92F9DA" box="[532,635,1577,1603]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
is more or less ovoid (
<figureCitation id="13552A324D4DD222E6ADF9D6FC02F9DA" box="[927,1003,1577,1603]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="13" pageNumber="14">Figs 8</figureCitation>
A–B), 
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long, about 
<quantity id="4C969B524D4DD222E43AF9AEFEB4F9F2" box="[264,349,1617,1643]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="mm" value="10.0">10 mm</quantity>
wide (across ventral sole), and of a dirty­white colour (preserved). The mouth is ventral and the anus is terminal, dorsally of the ventral sole. Ten tentacles encompass the mouth, eight (?) of which are connected by a membrane, while at least the two posteriormost tentacles are free. Terminal discs of tentacles with numerous minute processes. Nine pairs of tube feet border the posterior two­thirds of the ventral sole (
<figureCitation id="13552A324D4DD222E16AF90EFB4CF892" box="[1112,1189,1777,1803]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="13" pageNumber="14">Figs 8</figureCitation>
A–B). The anterior five pairs are conical and bear a terminal disc consisting of four lobes (
<figureCitation id="13552A324D4DD222E189F8E6FAE8F8AA" box="[1211,1281,1817,1843]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="13" pageNumber="14">Fig. 8</figureCitation>
D). They are well separated from each other and slightly decrease in size from anterior to posterior. The remaining four pairs, all of about equal size, are clustered brim­like at the posterior end of the ventral sole. A conspicuous dorsal velum, about as long as the body, is situated close to the anterior end of the specimen (
<figureCitation id="13552A324D4ED221E647FE93FC2BFE1F" box="[885,962,364,390]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="14" pageNumber="15">Figs 8</figureCitation>
A, C). It is composed of four papillae, of which the long median pair are fully fused along their length, while the outer papillae are much shorter and separate in their distal part, forming small lateral lobes. A third pair of small and free papillae is present close to the base of the velum.
</paragraph>
<paragraph id="8BD136B74D4ED221E40AFDF3FEADFD07" blockId="14.[264,1324,284,1272]" pageId="14" pageNumber="15">A calcareous ring seems to be missing. The single polian vesicle is large. The unpaired left gonad consists of richly branched short tubules opening into a well developed common duct. The anterior part of the intestine is straight, while the posterior forms a loop.</paragraph>
<paragraph id="8BD136B74D4ED221E40AFD51FE5DFCD9" blockId="14.[264,1324,284,1272]" pageId="14" pageNumber="15">
Calcareous deposits are straight or slightly curved spinous rods (
<figureCitation id="13552A324D4ED221E118FD51FB87FD51" box="[1066,1134,686,712]" captionStart="FIGURE 8" captionStartId="14.[264,368,1806,1830]" captionTargetBox="[279,1315,1342,1776]" captionTargetId="figure@14.[264,1323,1328,1782]" captionTargetPageId="14" captionText="FIGURE 8. Achlyonice longicornis spec. nov. (A) Right lateral view (schematic). (B) Ventral view (schematic). (C) Velum. (D) Distal end of tubefoot. (E) Rods from dorsal papillae. Figs A – C to the same scale." httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
E), up to 470 long, which are present in the tentacles the dorsal papillae and the tube feet; but none were found in the body wall (some of the deposits are in the process of dissolving, due to acidic fixation fluid).
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4ED220E40AFCB2FDA5FC47" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BD136B74D4ED221E40AFCB2FDDFFC79" blockId="14.[264,1324,284,1272]" pageId="14" pageNumber="15">
<emphasis id="B91AEAA54D4ED221E40AFCB2FE58FCFE" bold="true" box="[312,433,845,871]" pageId="14" pageNumber="15">Remarks.</emphasis>
<taxonomicName id="4C6E4D344D4ED221E48BFCB1FD2EFCFE" box="[441,711,846,871]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4ED221E48BFCB1FD2EFCFE" box="[441,711,846,871]" italics="true" pageId="14" pageNumber="15">Achlyonice longicornis</emphasis>
</taxonomicName>
differs from all other known elpidiid species by the combined occurrence of the following two characteristics: the peculiar triangular velum with two small lateral lobes and the large spinous rods present in the tentacles, the dorsal papillae and the tube feet.
</paragraph>
<paragraph id="8BD136B74D4ED221E40AFC11FC2AFB61" blockId="14.[264,1324,284,1272]" pageId="14" pageNumber="15">
Only one species is known to possess a somewhat similar velum, the single specimen of 
<taxonomicName id="4C6E4D344D4ED221E41CFBE9FCB8FBA9" authority="Koehler and Vaney, 1905" authorityName="Koehler and Vaney" authorityYear="1905" box="[302,849,1046,1072]" class="Holothuroidea" family="Elpidiidae" genus="Peniagone" kingdom="Animalia" order="Elasipodida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="stabilis">
<emphasis id="B91AEAA54D4ED221E41CFBE9FDE7FBB6" box="[302,526,1046,1071]" italics="true" pageId="14" pageNumber="15">Peniagone stabilis</emphasis>
<bibRefCitation id="EFFF4B464D4ED221E72EFBE9FCB8FBA9" author="Koehler" box="[540,849,1046,1072]" pageId="14" pageNumber="29" refString="Koehler, R. &amp; Vaney, C. (1905) An account of the deep-sea Holothurioidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, 126 pp." type="book" year="1905">Koehler and Vaney, 1905</bibRefCitation>
</taxonomicName>
collected in the Bay of Bengal. It is characterised by a triangular velum consisting of one large and two small papillae and a posterior lobated border of fused tube feet (
<bibRefCitation id="EFFF4B464D4ED221E605FB99FBB4FB19" author="Koehler" box="[823,1117,1126,1152]" pageId="14" pageNumber="29" refString="Koehler, R. &amp; Vaney, C. (1905) An account of the deep-sea Holothurioidea collected by the Royal Indian Marine Survey Ship Investigator. Echinoderma of the Indian Museum, 126 pp." type="book" year="1905">Koehler &amp; Vaney 1905</bibRefCitation>
). It differs from 
<taxonomicName id="4C6E4D344D4ED221E43AFB71FDF3FB3E" box="[264,538,1166,1191]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4ED221E43AFB71FDF3FB3E" box="[264,538,1166,1191]" italics="true" pageId="14" pageNumber="15">Achlyonice longicornis</emphasis>
</taxonomicName>
by the much shorter velum, which is only 
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high (with both specimens being of similar size) and by its calcareous deposits, which are four­armed and have one central apophysis and one apophysis on each arm.
</paragraph>
<caption id="DF11663F4D4ED221E43AF8F1FE6BF8F2" httpUri="https://zenodo.org/record/173344/files/figure.png" pageId="14" pageNumber="15" targetBox="[279,1315,1342,1776]" targetPageId="14">
<paragraph id="8BD136B74D4ED221E43AF8F1FE6BF8F2" blockId="14.[264,1323,1806,1899]" pageId="14" pageNumber="15">
<emphasis id="B91AEAA54D4ED221E43AF8F1FE62F8BF" bold="true" box="[264,395,1806,1830]" pageId="14" pageNumber="15">FIGURE 8.</emphasis>
<taxonomicName id="4C6E4D344D4ED221E4A3F8F1FD6EF8BC" box="[401,647,1806,1829]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="longicornis" status="sp. nov.">
<emphasis id="B91AEAA54D4ED221E4A3F8F1FD6EF8BC" box="[401,647,1806,1829]" italics="true" pageId="14" pageNumber="15">Achlyonice longicornis</emphasis>
</taxonomicName>
<emphasis id="B91AEAA54D4ED221E7BFF8F1FCCCF8BF" bold="true" box="[653,805,1806,1830]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A22957DE4D4ED221E7BFF8F1FD13F8BF" box="[653,762,1806,1830]" pageId="14" pageNumber="15" rank="species">spec. nov.</taxonomicNameLabel>
(A)
</emphasis>
Right lateral view (schematic). 
<emphasis id="B91AEAA54D4ED221E14BF8F1FB75F8BF" bold="true" box="[1145,1180,1806,1830]" pageId="14" pageNumber="15">(B)</emphasis>
Ventral view (schematic). 
<emphasis id="B91AEAA54D4ED221E4A0F8CFFE5EF8D1" bold="true" box="[402,439,1840,1864]" pageId="14" pageNumber="15">(C)</emphasis>
Velum. 
<emphasis id="B91AEAA54D4ED221E723F8CFFDDFF8D1" bold="true" box="[529,566,1840,1864]" pageId="14" pageNumber="15">(D)</emphasis>
Distal end of tubefoot. 
<emphasis id="B91AEAA54D4ED221E606F8CFFCBEF8D1" bold="true" box="[820,855,1840,1864]" pageId="14" pageNumber="15">(E)</emphasis>
Rods from dorsal papillae. Figs A–C to the same scale.
</paragraph>
</caption>
<paragraph id="8BD136B74D4FD220E40AFEE3FC02FDD7" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
The generic assignment of the new species is somewhat ambiguous. Three out of five species of the genus 
<taxonomicName id="4C6E4D344D4FD220E727FEBBFCD1FEC7" authority="Theel, 1879" authorityName="Theel" authorityYear="1879" box="[533,824,324,350]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E727FEBBFD7EFEC4" box="[533,663,324,349]" italics="true" pageId="15" pageNumber="16">Achlyonice</emphasis>
<bibRefCitation id="EFFF4B464D4FD220E797FEBBFCD1FEC7" author="Theel" box="[677,824,324,350]" pageId="15" pageNumber="30" refString="Theel, H. (1879). Preliminary report on the Holothuridae of the Exploring voyage of H. M. S. &quot; Challenger &quot;, I. Bihang till Kungliga Svenska Vetenskaps-Akademiens Handlingar, 5, 1 - 20." type="journal article" year="1879">Théel, 1879</bibRefCitation>
</taxonomicName>
and all species of the genus 
<taxonomicName id="4C6E4D344D4FD220E187FEBBFE2DFE1F" authority="Herouard, 1923" authorityName="Herouard" authorityYear="1923" class="Holothuroidea" family="Elpidiidae" genus="Ellipinion" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E187FEBBFAC2FEC4" box="[1205,1323,324,349]" italics="true" pageId="15" pageNumber="16">Ellipinion</emphasis>
<bibRefCitation id="EFFF4B464D4FD220E43AFE93FE2DFE1F" author="Herouard" box="[264,452,364,390]" pageId="15" pageNumber="29" refString="Herouard, E. (1923) Holothuries provenant des campagnes des yachts PRINCESSE-ALICE et HIRONDELLE II (1898 - 1915). Resultats des campagnes scientifiques accomplies sur son yatch par Albert Ier, Prince Souverain de Monaco, 66, 161 pp." type="book" year="1923">Hérouard, 1923</bibRefCitation>
</taxonomicName>
are known to possess rod­shaped, often spinous deposits and a velum. 
<taxonomicName id="4C6E4D344D4FD220E43AFE6BFE94FE34" box="[264,381,404,429]" class="Holothuroidea" family="Elpidiidae" genus="Ellipinion" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E43AFE6BFE94FE34" box="[264,381,404,429]" italics="true" pageId="15" pageNumber="16">Ellipinion</emphasis>
</taxonomicName>
species differ from 
<taxonomicName id="4C6E4D344D4FD220E74AFE6BFD13FE34" box="[632,762,404,429]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E74AFE6BFD13FE34" box="[632,762,404,429]" italics="true" pageId="15" pageNumber="16">Achlyonice</emphasis>
</taxonomicName>
by the additional presence of small C­shaped deposits and constantly 10 tentacles. 
<bibRefCitation id="EFFF4B464D4FD220E7F6FE43FC9DFE4F" author="Hansen" box="[708,884,444,470]" pageId="15" pageNumber="29" refString="Hansen, B. (1975) Systematics and Biology of the Deep-Sea Holothurians. Part 1. Elasipoda. Galathea Report, 13, 262 pp." type="book" year="1975">Hansen (1975)</bibRefCitation>
characterises 
<taxonomicName id="4C6E4D344D4FD220E12FFE43FB76FE4C" box="[1053,1183,444,469]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E12FFE43FB76FE4C" box="[1053,1183,444,469]" italics="true" pageId="15" pageNumber="16">Achlyonice</emphasis>
</taxonomicName>
as follows: tentacles 10–12; deposits tripartite, rod­shaped or absent; calcareous ring consisting of five isolated pieces, each having a varying number of arms. Absence of C­shaped deposits assigns the new species to 
<taxonomicName id="4C6E4D344D4FD220E773FDCBFD2AFDD4" box="[577,707,564,589]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E773FDCBFD2AFDD4" box="[577,707,564,589]" italics="true" pageId="15" pageNumber="16">Achlyonice</emphasis>
</taxonomicName>
rather than to 
<taxonomicName id="4C6E4D344D4FD220E65CFDCBFC0AFDD4" box="[878,995,564,589]" class="Holothuroidea" family="Elpidiidae" genus="Ellipinion" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="genus">
<emphasis id="B91AEAA54D4FD220E65CFDCBFC0AFDD4" box="[878,995,564,589]" italics="true" pageId="15" pageNumber="16">Ellipinion</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD136B74D4FD220E40AFDA3FDA5FC47" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
Within this genus there are two species, 
<taxonomicName id="4C6E4D344D4FD220E62FFDA3FB1EFDEF" authority="Ohshima, 1915" authorityName="Ohshima" authorityYear="1915" box="[797,1271,604,630]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="monactinica">
<emphasis id="B91AEAA54D4FD220E62FFDA3FBD0FDEC" box="[797,1081,604,629]" italics="true" pageId="15" pageNumber="16">Achlyonice monactinica</emphasis>
<bibRefCitation id="EFFF4B464D4FD220E170FDA3FB1EFDEF" author="Ohshima" box="[1090,1271,604,630]" pageId="15" pageNumber="30" refString="Ohshima, H. (1915) Report on the holothurians collected by the United States Fisheries Steamer &quot; Albatross &quot; in the northwestern Pacific during the summer of 1906. Proceedings of the United States National Museum, 48, 213 - 291." type="journal article" year="1915">Ohshima, 1915</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C6E4D344D4FD220E43AFD7BFD55FD07" authority="Gebruk, 1997" authorityName="Gebruk" authorityYear="1997" box="[264,700,644,670]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="myriamae">
<emphasis id="B91AEAA54D4FD220E43AFD7BFDE3FD04" box="[264,522,644,669]" italics="true" pageId="15" pageNumber="16">Achlyonice myriamae</emphasis>
<bibRefCitation id="EFFF4B464D4FD220E727FD7BFD55FD07" author="Gebruk" box="[533,700,644,670]" pageId="15" pageNumber="29" refString="Gebruk, A. V. (1997) New species of the deep-sea holothurian family Elpidiidae Theel, 1879. Zoosystema, 19, 211 - 217." type="journal article" year="1997">Gebruk, 1997</bibRefCitation>
</taxonomicName>
, which share some similarities with 
<taxonomicName id="4C6E4D344D4FD220E148FD7BFACAFD04" box="[1146,1315,644,669]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4FD220E148FD7BFACAFD04" box="[1146,1315,644,669]" italics="true" pageId="15" pageNumber="16">A. longicornis</emphasis>
</taxonomicName>
. All three have the anterior tentacles connected by a membrane and the calcareous deposits are rods (
<bibRefCitation id="EFFF4B464D4FD220E44EFD2BFDD9FD77" author="Ohshima" box="[380,560,724,750]" pageId="15" pageNumber="30" refString="Ohshima, H. (1915) Report on the holothurians collected by the United States Fisheries Steamer &quot; Albatross &quot; in the northwestern Pacific during the summer of 1906. Proceedings of the United States National Museum, 48, 213 - 291." type="journal article" year="1915">Ohshima 1915</bibRefCitation>
; 
<bibRefCitation id="EFFF4B464D4FD220E772FD2BFD08FD77" author="Gebruk" box="[576,737,724,750]" pageId="15" pageNumber="29" refString="Gebruk, A. V. (1997) New species of the deep-sea holothurian family Elpidiidae Theel, 1879. Zoosystema, 19, 211 - 217." type="journal article" year="1997">Gebruk 1997</bibRefCitation>
). Most obviously, 
<taxonomicName id="4C6E4D344D4FD220E6F1FD2BFB84FD74" box="[963,1133,724,749]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4FD220E6F1FD2BFB84FD74" box="[963,1133,724,749]" italics="true" pageId="15" pageNumber="16">A. longicornis</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C6E4D344D4FD220E023FD2BFE94FC8C" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="myriamae">
<emphasis id="B91AEAA54D4FD220E023FD2BFACAFD74" box="[1297,1315,724,749]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E43AFD03FE94FC8C" box="[264,381,764,789]" italics="true" pageId="15" pageNumber="16">myriamae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C6E4D344D4FD220E4F4FD03FD69FC8C" box="[454,640,764,790]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="monactinica">
<emphasis id="B91AEAA54D4FD220E4F4FD03FE31FC8C" box="[454,472,764,789]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E4DDFD03FD69FC8C" box="[495,640,764,789]" italics="true" pageId="15" pageNumber="16">monactinica</emphasis>
</taxonomicName>
by: different tentacle numbers, both species have 12 tentacles as opposed to 
<taxonomicName id="4C6E4D344D4FD220E72EFCDBFD28FCA4" box="[540,705,804,830]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4FD220E72EFCDBFDC7FCA4" box="[540,558,804,829]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E70EFCDBFD28FCA4" box="[572,705,804,829]" italics="true" pageId="15" pageNumber="16">longicornis</emphasis>
</taxonomicName>
, which has only ten; its peculiar dominant triangular velum; the restriction of rod deposits to the tentacles, the dorsal papillae and the tube feet, which are present throughout the body wall in the other species. Furthermore, 
<taxonomicName id="4C6E4D344D4FD220E1A7FC8BFAC3FC14" box="[1173,1322,884,910]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="myriamae">
<emphasis id="B91AEAA54D4FD220E1A7FC8BFB4EFC14" box="[1173,1191,884,909]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E187FC8BFAC3FC14" box="[1205,1322,884,909]" italics="true" pageId="15" pageNumber="16">myriamae</emphasis>
</taxonomicName>
has in addition to rod deposits also tripartite deposits, which are lacking in 
<taxonomicName id="4C6E4D344D4FD220E14BFC63FAC2FC2C" box="[1145,1323,924,950]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="monactinica">
<emphasis id="B91AEAA54D4FD220E14BFC63FB62FC2C" box="[1145,1163,924,949]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E1A8FC63FAC2FC2C" box="[1178,1323,924,949]" italics="true" pageId="15" pageNumber="16">monactinica</emphasis>
</taxonomicName>
as well as in 
<taxonomicName id="4C6E4D344D4FD220E4ADFC3BFDACFC44" box="[415,581,964,990]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4FD220E4ADFC3BFE58FC44" box="[415,433,964,989]" italics="true" pageId="15" pageNumber="16">A</emphasis>
. 
<emphasis id="B91AEAA54D4FD220E4F2FC3BFDACFC44" box="[448,581,964,989]" italics="true" pageId="15" pageNumber="16">longicornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4FD220E40AFC13FCF6FBB7" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BD136B74D4FD220E40AFC13FCF6FBB7" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
<emphasis id="B91AEAA54D4FD220E40AFC13FE31FB9F" bold="true" box="[312,472,1004,1030]" pageId="15" pageNumber="16">Distribution.</emphasis>
(
<figureCitation id="13552A324D4FD220E4DDFC13FDD5FB9F" box="[495,572,1004,1030]" captionStart="FIGURE 7" captionStartId="12.[264,367,1779,1803]" captionTargetBox="[270,1309,1228,1745]" captionTargetId="figure@12.[264,1323,1226,1755]" captionTargetPageId="12" captionText="FIGURE 7. Distribution of Peniagone purpurea (Théel, 1882) and Achlyonice longicornis spec. nov." httpUri="https://zenodo.org/record/173343/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
) So far, this species is only known from the 
<typeStatus id="54D588154D4FD220E1BFFC13FB29FB9F" box="[1165,1216,1004,1030]" pageId="15" pageNumber="16">type</typeStatus>
locality, Atlantic Ocean, 
<collectingCountry id="F37976274D4FD220E4F5FBEBFDF4FBB7" box="[455,541,1044,1070]" name="Angola" pageId="15" pageNumber="16">Angola</collectingCountry>
Basin, 
<quantity id="4C969B524D4FD220E744FBEBFCF2FBB7" box="[630,795,1044,1070]" metricMagnitude="3" metricUnit="m" metricValue="5.386" metricValueMax="5.387" metricValueMin="5.385" pageId="15" pageNumber="16" unit="m" value="5386.0" valueMax="5387.0" valueMin="5385.0">5385–5387 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4FD220E40AFBC3FE41FBE7" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BD136B74D4FD220E40AFBC3FE41FBE7" blockId="15.[264,1323,284,1150]" pageId="15" pageNumber="16">
<emphasis id="B91AEAA54D4FD220E40AFBC3FE2DFBCF" bold="true" box="[312,452,1084,1110]" pageId="15" pageNumber="16">Etymology.</emphasis>
The name, 
<taxonomicName id="4C6E4D344D4FD220E763FBC3FD3FFBCC" box="[593,726,1084,1109]" class="Holothuroidea" family="Elpidiidae" genus="Achlyonice" kingdom="Animalia" order="Elasipodida" pageId="15" pageNumber="16" phylum="Echinodermata" rank="species" species="longicornis">
<emphasis id="B91AEAA54D4FD220E763FBC3FD3FFBCC" box="[593,726,1084,1109]" italics="true" pageId="15" pageNumber="16">longicornis</emphasis>
</taxonomicName>
, refers to the characteristic, horn­like shape of the dorsal velum.
</paragraph>
</subSubSection>
</treatment>
</document>