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<mods:title id="C9FB16A01B63FCFAF1761B651ABF4272">Crinoidea and Holothuroidea (Echinodermata) of the abyssal Angola Basin — Results of the DIVA­ 1 expedition of FS &quot; Meteor &quot; (Cruise M 48 / 1)</mods:title>
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<taxonomicName id="4C6E4D344D49D226E43AFA14FD3BF99C" ID-CoL="4PR92" authority="Theel, 1882" authorityName="Theel" authorityYear="1882" box="[264,722,1515,1541]" class="Holothuroidea" family="Psychropotidae" genus="Psychropotes" kingdom="Animalia" order="Elasipodida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="semperiana">
<emphasis id="B91AEAA54D49D226E43AFA14FD3BF99C" bold="true" box="[264,722,1515,1541]" pageId="9" pageNumber="10">
<emphasis id="B91AEAA54D49D226E43AFA14FDD2F99C" bold="true" box="[264,571,1515,1541]" italics="true" pageId="9" pageNumber="10">Psychropotes semperiana</emphasis>
<bibRefCitation id="EFFF4B464D49D226E770FA14FD3BF99C" author="Theel" box="[578,722,1515,1541]" pageId="9" pageNumber="31" refString="Theel, H. (1882) Report on the Holothurioidea dredged by H. M. S. Challenger during the years 1873 - 76. Part I. Report of the scientific results of the voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 4, 176 pp." type="book" year="1882">Théel, 1882</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BD136B74D49D226E43AF9F1FE5CF9B1" blockId="9.[264,722,1515,1576]" box="[264,437,1550,1576]" pageId="9" pageNumber="10">(Figs 2BE, 5)</paragraph>
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<taxonomicName id="4C6E4D344D49D226E43AF9A1FD33F9EC" ID-CoL="4PR92" authority="Theel, 1882: 100" authorityName="Theel" authorityPageNumber="100" authorityYear="1882" box="[264,730,1630,1653]" class="Holothuroidea" family="Psychropotidae" genus="Psychropotes" kingdom="Animalia" order="Elasipodida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="semperiana">
<emphasis id="B91AEAA54D49D226E43AF9A1FDF2F9EC" box="[264,539,1630,1653]" italics="true" pageId="9" pageNumber="10">Psychropotes semperiana</emphasis>
<treatmentCitation id="0ACF10A64D49D226E710F9A1FD33F9EC" author="Theel" box="[546,730,1630,1653]" page="100" pageId="9" pageNumber="10" year="1882">
<bibRefCitation id="EFFF4B464D49D226E710F9A1FD4FF9EC" author="Theel" box="[546,678,1630,1653]" pageId="9" pageNumber="31" refString="Theel, H. (1882) Report on the Holothurioidea dredged by H. M. S. Challenger during the years 1873 - 76. Part I. Report of the scientific results of the voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 4, 176 pp." type="book" year="1882">Théel, 1882</bibRefCitation>
: 100
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101, pl. 34 (figs 1011);
<treatmentCitation id="0ACF10A64D49D226E6C6F9A1FB29F9EC" author="Hansen" box="[1012,1216,1630,1653]" page="102" pageId="9" pageNumber="10" year="1975">
<bibRefCitation id="EFFF4B464D49D226E6C6F9A1FB62F9EC" author="Hansen" box="[1012,1163,1630,1653]" pageId="9" pageNumber="29" refString="Hansen, B. (1975) Systematics and Biology of the Deep-Sea Holothurians. Part 1. Elasipoda. Galathea Report, 13, 262 pp." type="book" year="1975">Hansen, 1975</bibRefCitation>
: 102
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105, figs 4142 [synonymy]
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</paragraph>
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<taxonomicName id="4C6E4D344D49D226E43AF961FD30F92C" ID-CoL="3CS8J" authority="Herouard, 1902: 27" authorityName="Herouard" authorityPageNumber="27" authorityYear="1902" box="[264,729,1694,1717]" class="Holothuroidea" family="Psychropotidae" genus="Psychropotes" kingdom="Animalia" order="Elasipodida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="kerhervei">
<emphasis id="B91AEAA54D49D226E43AF961FDE8F92C" box="[264,513,1694,1717]" italics="true" pageId="9" pageNumber="10">Psychropotes kerhervei</emphasis>
<treatmentCitation id="0ACF10A64D49D226E73AF961FD30F92C" author="Herouard" box="[520,729,1694,1717]" page="27" pageId="9" pageNumber="10" year="1902">
<bibRefCitation id="EFFF4B464D49D226E73AF961FD5DF92C" author="Herouard" box="[520,692,1694,1717]" pageId="9" pageNumber="29" refString="Herouard, E. (1902) Holothuries provenant des campagnes de la Princesse-Alice (1892 - 1897). Resultats des campagnes scientifiques accomplies sur son yatch par Albert Ier, Prince Souverain de Monaco, 21, 61 pp." type="book" year="1902">Hérouard, 1902</bibRefCitation>
: 27
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30, pl. 4 (figs 19)
<taxonomicName id="4C6E4D344D49D226E43AF941FD50F94C" authority="Herouard, 1912: 6" authorityName="Herouard" authorityPageNumber="6" authorityYear="1912" box="[264,697,1726,1749]" class="Holothuroidea" family="Psychropotidae" genus="Euphronides" higherTaxonomySource="CoL" kingdom="Animalia" order="Elasipodida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="anchora">
<emphasis id="B91AEAA54D49D226E43AF941FE06F94C" box="[264,495,1726,1749]" italics="true" pageId="9" pageNumber="10">Euphronides anchora</emphasis>
<treatmentCitation id="0ACF10A64D49D226E4C7F941FD50F94C" author="Herouard" box="[501,697,1726,1749]" page="6" pageId="9" pageNumber="10" year="1912">
<bibRefCitation id="EFFF4B464D49D226E4C7F941FD49F94C" author="Herouard" box="[501,672,1726,1749]" pageId="9" pageNumber="29" refString="Herouard, E. (1912) Holothuries nouvelles des campagnes du yacht &quot; Princesse-Alice &quot;. Bulletin de l'Institut oceanographique, Monaco, 239, 1 - 9." type="journal article" year="1912">Hérouard, 1912</bibRefCitation>
: 6
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7, fig. 5
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</paragraph>
</subSubSection>
<subSubSection id="C374653C4D49D226E43AF8D7FC63F8DA" box="[264,906,1832,1859]" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BD136B74D49D226E43AF8D7FC63F8DA" blockId="9.[264,1323,1832,1899]" box="[264,906,1832,1859]" pageId="9" pageNumber="10">
<emphasis id="B91AEAA54D49D226E43AF8D7FE95F8DB" bold="true" box="[264,380,1832,1858]" pageId="9" pageNumber="10">Material.</emphasis>
M 48/1­321:
<specimenCount id="9D68FD3E4D49D226E72CF8D6FD47F8DA" box="[542,686,1833,1859]" pageId="9" pageNumber="10" type="generic">2 specimens</specimenCount>
(
<collectionCode id="ED7FAE724D49D226E78CF8D6FD15F8DA" box="[702,764,1833,1859]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="9" pageNumber="10">ZSM</collectionCode>
20043075).
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D49D225E40AF8AFFC26FBE7" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
<paragraph id="8BD136B74D49D225E40AF8AFFC74FCFF" blockId="9.[264,1323,1832,1899]" lastBlockId="10.[264,1324,284,1150]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B91AEAA54D49D226E40AF8AFFE39F8F3" bold="true" box="[312,464,1872,1898]" pageId="9" pageNumber="10">Description.</emphasis>
Anteriorly, the specimens are more or less flat, while the posterior part of the body is elevated and gives rise to the unpaired dorsal appendage. The larger specimen is
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long and
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wide (across ventral sole), while the smaller specimen measures
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in length and
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across the ventral sole. Both are of a purple colour (preserved). The mouth and the anus are ventral and confined to the anterior and the posterior end of the body. There are 16 tentacles, with conspicuous, rounded discs. The ventral sole is delimited on both sides by a narrow brim (damaged for the most part), formed by basally fused tube feet. Mid­ventral tube feet are conical and restricted to a double row, which seems to be present (partly damaged) throughout the length of the ventral sole. The unpaired dorsal appendage arises from a dorsal bulge, which is situated one­seventh body length in the large specimen and one­fourth body length in the small specimen from the posterior end of the body. In both specimens, the dorsal appendage is more or less contracted and thus nothing can be said about the original size. The almost complete dorsal appendage of the large specimen ends in two long slender papillae, each about
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long. Anterior to the dorsal bulge of the small specimen, are three pairs of short conical papillae (indistinct in the larger specimen).
</paragraph>
<paragraph id="8BD136B74D4AD225E40AFC8BFC26FBE7" blockId="10.[264,1324,284,1150]" pageId="10" pageNumber="11">
Calcareous deposits of the dorsal body wall are crosses of two
<typeStatus id="54D588154D4AD225E125FC8BFBBCFC17" box="[1047,1109,884,910]" pageId="10" pageNumber="11">types</typeStatus>
(Figs 2BD). The four arms of the larger
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are smooth in their proximal parts and are equipped with irregularly placed spines close to their distal ends (Fig. 2C). The high central apophysis is smooth and ends in three or four downwardly bent hooks (Fig. 2B). Crosses of the second
<typeStatus id="54D588154D4AD225E43AFBEBFED3FBB7" box="[264,314,1044,1070]" pageId="10" pageNumber="11">type</typeStatus>
are considerably smaller, have irregularly placed spines along the arms and a low and also spinous central apophysis (Fig. 2D). The calcareous deposits of the ventral body wall are rods and crosses with irregularly placed spines (Fig. 2E).
</paragraph>
</subSubSection>
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<paragraph id="8BD136B74D4AD225E43AF920FC3CF96E" blockId="10.[264,981,1759,1783]" box="[264,981,1759,1783]" pageId="10" pageNumber="11">
<emphasis id="B91AEAA54D4AD225E43AF920FE63F96E" bold="true" box="[264,394,1759,1783]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
Distribution of
<taxonomicName id="4C6E4D344D4AD225E707F91FFC39F96E" authority="Theel, 1882" authorityName="Theel" authorityYear="1882" box="[565,976,1760,1783]" class="Holothuroidea" family="Psychropotidae" genus="Psychropotes" kingdom="Animalia" order="Elasipodida" pageId="10" pageNumber="11" phylum="Echinodermata" rank="species" species="semperiana">
<emphasis id="B91AEAA54D4AD225E707F91FFCA1F96E" box="[565,840,1760,1783]" italics="true" pageId="10" pageNumber="11">Psychropotes semperiana</emphasis>
Théel, 1882
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.
</paragraph>
</caption>
<subSubSection id="C374653C4D4AD224E40AF8D5FB30FEC7" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BD136B74D4AD224E40AF8D5FB30FEC7" blockId="10.[264,1324,1834,1900]" lastBlockId="11.[264,1325,284,470]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B91AEAA54D4AD225E40AF8D5FE58F8DD" bold="true" box="[312,433,1834,1860]" pageId="10" pageNumber="11">Remarks.</emphasis>
The two specimens described herein conform to the detailed description given by
<bibRefCitation id="EFFF4B464D4AD225E44FF8ADFDC6F8F5" author="Hansen" box="[381,559,1874,1900]" pageId="10" pageNumber="29" refString="Hansen, B. (1975) Systematics and Biology of the Deep-Sea Holothurians. Part 1. Elasipoda. Galathea Report, 13, 262 pp." type="book" year="1975">Hansen (1975)</bibRefCitation>
for this species. Very characteristic are the large cross­shaped deposits of the dorsal body wall with a high and smooth central apophysis, ending in three or four downwardly bent hooks, which are unique within the genus (
<bibRefCitation id="EFFF4B464D4BD224E119FEBBFB22FEC7" author="Hansen" box="[1067,1227,324,350]" pageId="11" pageNumber="29" refString="Hansen, B. (1975) Systematics and Biology of the Deep-Sea Holothurians. Part 1. Elasipoda. Galathea Report, 13, 262 pp." type="book" year="1975">Hansen 1975</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374653C4D4BD224E40AFE94FB4CFE4F" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BD136B74D4BD224E40AFE94FB4CFE4F" blockId="11.[264,1325,284,470]" pageId="11" pageNumber="12">
<emphasis id="B91AEAA54D4BD224E40AFE94FE31FE1C" bold="true" box="[312,472,363,389]" pageId="11" pageNumber="12">Distribution.</emphasis>
(
<figureCitation id="13552A324D4BD224E4DEFE93FDDCFE1F" box="[492,565,364,390]" captionStart="FIGURE 5" captionStartId="10.[264,368,1759,1783]" captionTargetBox="[270,1310,1207,1727]" captionTargetId="figure@10.[264,1323,1206,1735]" captionTargetPageId="10" captionText="FIGURE 5. Distribution of Psychropotes semperiana Théel, 1882." httpUri="https://zenodo.org/record/173341/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
) Northern and southern Atlantic Ocean, northern and western Indian Ocean,
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(
<bibRefCitation id="EFFF4B464D4BD224E74DFE6BFCBCFE37" author="Deichmann" box="[639,853,404,430]" pageId="11" pageNumber="28" refString="Deichmann, E. (1930) The holothurians of the western part of the Atlantic Ocean. Bulletin of the Museum of Comparative Zoology at Harvard College, 71, 43 - 226." type="journal article" year="1930">Deichmann 1930</bibRefCitation>
,
<bibRefCitation id="EFFF4B464D4BD224E655FE6BFC4CFE37" author="Deichmann" box="[871,933,404,430]" pageId="11" pageNumber="28" refString="Deichmann, E. (1940) Report on the Holothurians, collected by the Harvard-Havana Expeditions 1938 and 1939, with a revision of the Molpadonia of the Atlantic Ocean. Memorias de la Sociedad Cubana de Historia Natural ' Felipe Poey', 14, 183 - 240." type="journal article" year="1940">1940</bibRefCitation>
;
<bibRefCitation id="EFFF4B464D4BD224E685FE6BFBB2FE37" author="Hansen" box="[951,1115,404,430]" pageId="11" pageNumber="29" refString="Hansen, B. (1975) Systematics and Biology of the Deep-Sea Holothurians. Part 1. Elasipoda. Galathea Report, 13, 262 pp." type="book" year="1975">Hansen 1975</bibRefCitation>
;
<bibRefCitation id="EFFF4B464D4BD224E15CFE6BFAC1FE37" author="Herouard" box="[1134,1320,404,430]" pageId="11" pageNumber="29" refString="Herouard, E. (1902) Holothuries provenant des campagnes de la Princesse-Alice (1892 - 1897). Resultats des campagnes scientifiques accomplies sur son yatch par Albert Ier, Prince Souverain de Monaco, 21, 61 pp." type="book" year="1902">Hérouard 1902</bibRefCitation>
,
<bibRefCitation id="EFFF4B464D4BD224E43AFE43FEAFFE4F" author="Herouard" box="[264,326,444,470]" pageId="11" pageNumber="29" refString="Herouard, E. (1923) Holothuries provenant des campagnes des yachts PRINCESSE-ALICE et HIRONDELLE II (1898 - 1915). Resultats des campagnes scientifiques accomplies sur son yatch par Albert Ier, Prince Souverain de Monaco, 66, 161 pp." type="book" year="1923">1923</bibRefCitation>
; IFREMER BIOCEAN;
<bibRefCitation id="EFFF4B464D4BD224E759FE43FCE4FE4F" author="Madsen" box="[619,781,444,470]" pageId="11" pageNumber="30" refString="Madsen, F. J. (1953) Holothurioidea. Reports of the Swedish deep-sea expedition 1947 - 1948, 2, 151 - 173." type="journal article" year="1953">Madsen 1953</bibRefCitation>
;
<bibRefCitation id="EFFF4B464D4BD224E62BFE43FC43FE4F" author="Sibuet" box="[793,938,444,470]" pageId="11" pageNumber="30" refString="Sibuet, M. (1977) Distribution and diversity of echinoderms (Holothuroidea, Asteroidea) in the abyssal zone in the Bay of Biscay. Deep-Sea Research, 24, 549 - 564." type="journal article" year="1977">Sibuet 1977</bibRefCitation>
;
<bibRefCitation id="EFFF4B464D4BD224E684FE43FBD7FE4F" author="Theel" box="[950,1086,444,470]" pageId="11" pageNumber="31" refString="Theel, H. (1882) Report on the Holothurioidea dredged by H. M. S. Challenger during the years 1873 - 76. Part I. Report of the scientific results of the voyage of H. M. S. Challenger during the years 1873 - 76. Zoology, 4, 176 pp." type="book" year="1882">Théel 1882</bibRefCitation>
; herein).
</paragraph>
</subSubSection>
</treatment>
</document>