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<document id="D00493596A1EF23B6787430038919BCE" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C78785462DFFE04161409932ABFD84" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Costulodonta trilamellaris Herbert 2020, gen. et comb. nov." docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="34" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E3271A146457F97385ECEC243EEDC718">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:namePart id="7D4E7EF1831F6CCE01EFA4429A2BF318">Herbert, David G.</mods:namePart>
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<mods:affiliation id="EA255D3D238CD8D0AFEBC9983BA8A599">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="285DE25D5B9D4AD8F4361FFCEAA3FDCF" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="B2B1AFB75D2FEF40F38BFAB0158F5C4C">European Journal of Taxonomy</mods:title>
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<mods:part id="65B8C1E0C11F89AC30098A5504CC4AEF">
<mods:date id="93BC92E7374491220C80148C73772F35">2020</mods:date>
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<mods:number id="21AC62366E8BE3DAEA6F633D0BB37DE1">2020-04-17</mods:number>
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<mods:number id="F4B406209763E2C8CC3FC7F362F5D135">629</mods:number>
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<treatment id="03C78785462DFFE04161409932ABFD84" ID-DOI="http://doi.org/10.5281/zenodo.3804760" ID-GBIF-Taxon="163732373" ID-Zenodo-Dep="3804760" LSID="urn:lsid:plazi:treatment:03C78785462DFFE04161409932ABFD84" httpUri="http://treatment.plazi.org/id/03C78785462DFFE04161409932ABFD84" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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<paragraph id="8BD13693462DFFE34161409935E9FC0A" blockId="33.[355,1233,952,1015]" box="[355,1233,952,979]" pageId="33" pageNumber="34">
<heading id="D09981FF462DFFE34161409935E9FC0A" box="[355,1233,952,979]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="4C6E4D10462DFFE34161409932CAFC0A" ID-CoL="7QTRZ" authority="(Melvill &amp; Ponsonby, 1908)" authorityName="Herbert" authorityYear="2020" baseAuthorityName="Melvill &amp; Ponsonby" baseAuthorityYear="1908" box="[355,1010,952,979]" class="Gastropoda" family="Charopidae" genus="Costulodonta" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="trilamellaris" status="gen. et comb. nov.">
<emphasis id="B91AEA81462DFFE3416140993398FC0A" bold="true" box="[355,672,952,979]" italics="true" pageId="33" pageNumber="34">Costulodonta trilamellaris</emphasis>
(
<bibRefCitation id="EFFF4B62462DFFE342B2409832D2FC0A" author="Melvill J. C. &amp; Ponsonby J. H." box="[688,1002,953,979]" pageId="33" pageNumber="34" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill &amp; Ponsonby, 1908</bibRefCitation>
)
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<taxonomicNameLabel id="A22957FA462DFFE343FB409835E9FC0A" box="[1017,1233,953,979]" pageId="33" pageNumber="34" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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<subSubSection id="C3746518462DFFE3428E40FC329FFC2E" box="[652,935,989,1015]" pageId="33" pageNumber="34" type="description">
<paragraph id="8BD13693462DFFE3428E40FC329FFC2E" blockId="33.[355,1233,952,1015]" box="[652,935,989,1015]" pageId="33" pageNumber="34">
<figureCitation id="13552A16462DFFE3428E40FC33DCFC2E" box="[652,740,989,1015]" captionStart="Fig" captionStartId="29.[189,232,1896,1922]" captionTargetBox="[302,1285,1093,1856]" captionTargetId="figure@29.[302,1285,1093,1856]" captionTargetPageId="29" captionText="Fig. 11. Distribution of Costulodonta gen. nov. species. C. acinaces (Connolly, 1933) gen. et comb. nov. (green square), C. bidens gen. et sp. nov. (turquoise squares), C. burnupi (Connolly, 1933) gen. et comb. nov. (orange square) C. pluridens gen. et sp. nov. (pink squares), C. trilamellaris (Melvill &amp; Ponsonby, 1908) gen. et comb. nov. (royal blue square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762302" httpUri="https://zenodo.org/record/3762302/files/figure.png" pageId="33" pageNumber="34">Figs 11</figureCitation>
,
<figureCitation id="13552A16462DFFE342F340FC3235FC2E" box="[753,781,989,1015]" captionStart="Fig" captionStartId="32.[189,232,1932,1958]" captionTargetBox="[190,1395,1119,1884]" captionTargetId="figure@32.[189,1399,1113,1892]" captionTargetPageId="32" captionText="Fig. 12. Shells of Costulodonta gen. nov. species. A–D. C. pluridens gen. et sp. nov., holotype, diameter 1.8 mm (NMSA V5920/T4267). E–G. C. trilamellaris (Melvill &amp; Ponsonby, 1908) gen. et comb. nov., paralectotype, Dargle, KwaZulu-Natal, diameter 1.45 mm (NMSA 2181/T624)." figureDoi="http://doi.org/10.5281/zenodo.3762304" httpUri="https://zenodo.org/record/3762304/files/figure.png" pageId="33" pageNumber="34">12</figureCitation>
E–G, 18Q–R
</paragraph>
</subSubSection>
<subSubSection id="C3746518462DFFE340BF470133EAFB32" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="8BD13693462DFFE340BF4701350EFBE3" blockId="33.[189,1078,1056,1083]" box="[189,1078,1056,1083]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7E11BD462DFFE340BF4701350EFBE3" box="[189,1078,1056,1083]" pageId="33" pageNumber="34">
<treatmentCitation id="0ACF1082462DFFE340BF47013272FBE3" author="Melvill J. C. &amp; Ponsonby J. H." box="[189,842,1056,1082]" page="134" pageId="33" pageNumber="34" year="1908">
<taxonomicName id="4C6E4D10462DFFE340BF47013272FBE3" authority="Melvill &amp; Ponsonby, 1908: 134" authorityName="Melvill &amp; Ponsonby" authorityPageNumber="134" authorityYear="1908" box="[189,842,1056,1082]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="trilamellaris">
<emphasis id="B91AEA81462DFFE340BF470130F5FBE3" box="[189,461,1056,1082]" italics="true" pageId="33" pageNumber="34">Afrodonta trilamellaris</emphasis>
<bibRefCitation id="EFFF4B62462DFFE341D647013272FBE3" author="Melvill J. C. &amp; Ponsonby J. H." box="[468,842,1056,1082]" pageId="33" pageNumber="34" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill &amp; Ponsonby, 1908: 134</bibRefCitation>
</taxonomicName>
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, pl. 7, figs 7, 7a, 7b.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD13693462DFFE340BF474232C7FBA4" blockId="33.[189,1399,1123,1260]" box="[189,1023,1123,1150]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7E11BD462DFFE340BF474232C7FBA4" box="[189,1023,1123,1150]" pageId="33" pageNumber="34">
<taxonomicName id="4C6E4D10462DFFE340BF47423006FBA4" ID-CoL="7P3F9" authorityName="Albers" authorityYear="1850" box="[189,318,1123,1149]" class="Gastropoda" family="Endodontidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA81462DFFE340BF47423006FBA4" box="[189,318,1123,1149]" italics="true" pageId="33" pageNumber="34">Endodonta</emphasis>
</taxonomicName>
[
<taxonomicName id="4C6E4D10462DFFE3414D4742335AFBA7" authority="" box="[335,610,1123,1150]" class="Gastropoda" family="Endodontidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="subGenus" subGenus="Afrodonta">
<emphasis id="B91AEA81462DFFE3414D474230E8FBA4" box="[335,464,1123,1149]" italics="true" pageId="33" pageNumber="34">Endodonta</emphasis>
(
<emphasis id="B91AEA81462DFFE341E34742336FFBA4" box="[481,599,1123,1149]" italics="true" pageId="33" pageNumber="34">Afrodonta</emphasis>
)
</taxonomicName>
]
<taxonomicName id="4C6E4D10462DFFE342734742323CFBA4" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[625,772,1123,1149]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="trilamellaris">
<emphasis id="B91AEA81462DFFE342734742323CFBA4" box="[625,772,1123,1149]" italics="true" pageId="33" pageNumber="34">trilamellaris</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACF1082462DFFE34323474532C3FBA4" author="Burnup H. C." box="[801,1019,1123,1150]" page="336" pageId="33" pageNumber="34" year="1912">
<bibRefCitation id="EFFF4B62462DFFE34323474532C3FBA4" author="Burnup H. C." box="[801,1019,1123,1150]" pageId="33" pageNumber="34" pagination="333 - 343" refId="ref26463" refString="Burnup H. C. 1912. On Afrodonta Melv. and Pons., with descriptions of new species. Annals of the Natal Museum 2 (3): 333 - 343." type="journal article" year="1912">Burnup 1912: 336</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD13693462DFFE340BF47A93240FB7B" blockId="33.[189,1399,1123,1260]" box="[189,888,1160,1186]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7E11BD462DFFE340BF47A93240FB7B" box="[189,888,1160,1186]" pageId="33" pageNumber="34">
<taxonomicName id="4C6E4D10462DFFE340BF47A93351FB7B" box="[189,617,1160,1186]" class="Gastropoda" family="Endodontidae" genus="Endodonta" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="trilamellaris" subGenus="Afrodonta">
<emphasis id="B91AEA81462DFFE340BF47A93006FB7B" box="[189,318,1160,1186]" italics="true" pageId="33" pageNumber="34">Endodonta</emphasis>
(
<emphasis id="B91AEA81462DFFE3414D47A930FDFB7B" box="[335,453,1160,1186]" italics="true" pageId="33" pageNumber="34">Afrodonta</emphasis>
)
<emphasis id="B91AEA81462DFFE341D447A93351FB7B" box="[470,617,1160,1186]" italics="true" pageId="33" pageNumber="34">trilamellaris</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACF1082462DFFE3428447A9324CFB7B" author="Connolly M." box="[646,884,1160,1186]" page="128" pageId="33" pageNumber="34" year="1912">
<bibRefCitation id="EFFF4B62462DFFE3428447A9324CFB7B" author="Connolly M." box="[646,884,1160,1186]" pageId="33" pageNumber="34" pagination="59 - 306" refId="ref26546" refString="Connolly M. 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306. https: // doi. org / 10.5962 / bhl. part. 5071" type="journal article" year="1912">Connolly 1912: 128</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD13693462DFFE340BF478C33EAFB32" blockId="33.[189,1399,1123,1260]" pageId="33" pageNumber="34">
<treatmentCitationGroup id="AB7E11BD462DFFE340BF478C33EAFB32" pageId="33" pageNumber="34">
<taxonomicName id="4C6E4D10462DFFE340BF478C30F5FB1E" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[189,461,1197,1223]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="trilamellaris">
<emphasis id="B91AEA81462DFFE340BF478C30F5FB1E" box="[189,461,1197,1223]" italics="true" pageId="33" pageNumber="34">Afrodonta trilamellaris</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFFF4B62462DFFE341E8478C33A5FB1E" author="Connolly M." box="[490,669,1197,1223]" pageId="33" pageNumber="34" pagination="145 - 150" refId="ref26638" refString="Connolly M. 1933. On South African Endodontidae. Part 2. Annals of the Natal Museum 7 (2): 145 - 150." type="journal article" year="1933">Connolly 1933</bibRefCitation>
: text-fig. 1(2); 1939: 252, text-fig. 19(2). —
<treatmentCitation id="0ACF1082462DFFE344A5478C344BFB1E" author="Solem A." box="[1191,1395,1196,1223]" page="357" pageId="33" pageNumber="34" year="1970">
<bibRefCitation id="EFFF4B62462DFFE344A5478C344BFB1E" author="Solem A." box="[1191,1395,1196,1223]" pageId="33" pageNumber="34" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">Solem 1970: 357</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ACF1082462DFFE3410D47F03352FB35" author="Herbert D. &amp; Kilburn D." box="[271,618,1233,1260]" page="250" pageId="33" pageNumber="34" year="2004">
<bibRefCitation id="EFFF4B62462DFFE3410D47F03352FB35" author="Herbert D. &amp; Kilburn D." box="[271,618,1233,1260]" pageId="33" pageNumber="34" refId="ref26755" refString="Herbert D. &amp; Kilburn D. 2004. Field Guide to the Land Snails and Slugs of Eastern South Africa. Natal Museum, Pietermaritzburg." type="book" year="2004">Herbert &amp; Kilburn 2004: 250</bibRefCitation>
</treatmentCitation>
, text-fig.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3746518462DFFE340BF46353318F9CF" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BD13693462DFFE340BF4635300EFAF7" blockId="33.[189,310,1300,1326]" box="[189,310,1300,1326]" pageId="33" pageNumber="34">
<heading id="D09981FF462DFFE340BF4635300EFAF7" bold="true" box="[189,310,1300,1326]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B91AEA81462DFFE340BF4635300EFAF7" bold="true" box="[189,310,1300,1326]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462DFFE340BF46653318F9CF" blockId="33.[189,1399,1348,1558]" pageId="33" pageNumber="34">
Shell very small, spire slightly raised; protoconch with close-set axial riblets (diameter ±415 μm); teleoconch sculpture of compound axial riblets with 2–4 intermediary axial threads; spiral sculpture of close-set, spiral threads, strongest below suture, indistinct elsewhere; parietal region with a low, narrow, in-running ridge; baso-columellar region with a broad, low, in-running ridge; palatal region with a strong, broad, ridge-like denticle just below mid-whorl; umbilicus relatively wide. Shell buff to pale ochre; diameter up to
<quantity id="4C969B76462DFFE341BC46DA3323F9CF" box="[446,539,1531,1558]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="34" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462DFFE340BF451F3403F832" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BD13693462DFFE340BF451F3092F981" blockId="33.[189,426,1598,1624]" box="[189,426,1598,1624]" pageId="33" pageNumber="34">
<heading id="D09981FF462DFFE340BF451F3092F981" bold="true" box="[189,426,1598,1624]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B91AEA81462DFFE340BF451F3092F981" bold="true" box="[189,426,1598,1624]" pageId="33" pageNumber="34">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462DFFE340E8454E3235F950" blockId="33.[189,1228,1647,1710]" box="[234,781,1647,1673]" pageId="33" pageNumber="34">
<materialsCitation id="3B063CCE462DFFE340E8454E3235F950" ID-GBIF-Occurrence="2605305657" box="[234,781,1647,1673]" collectionCode="NHMUK" collectorName="H. C. Burnup" country="South Africa" location="KwaZulu-Natal" pageId="33" pageNumber="34" specimenCode="NHMUK 1908.12.14.42" specimenCount="1" stateProvince="KwaZulu-Natal" typeStatus="lectotype">
<emphasis id="B91AEA81462DFFE340E8454E305EF950" bold="true" box="[234,358,1647,1673]" pageId="33" pageNumber="34">
<typeStatus id="54D58831462DFFE340E8454E305EF950" box="[234,358,1647,1673]" pageId="33" pageNumber="34" type="lectotype">Lectotype</typeStatus>
</emphasis>
(by inference,
<bibRefCitation id="EFFF4B62462DFFE3421A454E323BF950" author="Connolly M." box="[536,771,1647,1673]" pageId="33" pageNumber="34" pagination="59 - 306" refId="ref26546" refString="Connolly M. 1912. A revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum 11: 59 - 306. https: // doi. org / 10.5962 / bhl. part. 5071" type="journal article" year="1912">Connolly 1912: 129</bibRefCitation>
)
</materialsCitation>
</paragraph>
<paragraph id="8BD13693462DFFE340BF45B235F4F974" blockId="33.[189,1228,1647,1710]" box="[189,1228,1683,1710]" pageId="33" pageNumber="34">
<collectingCountry id="F3797603462DFFE340BF45B230B6F974" box="[189,398,1683,1709]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
•
<collectingRegion id="49AAF871462DFFE341A545B2335AF974" box="[423,610,1683,1709]" country="South Africa" name="KwaZulu-Natal" pageId="33" pageNumber="34">KwaZulu-Natal</collectingRegion>
, Dargle;
<collectorName id="269B5345462DFFE342CF45B2325CF977" box="[717,868,1683,1710]" pageId="33" pageNumber="34">H.C. Burnup</collectorName>
leg.;
<specimenCode id="DBC89EE8462DFFE343A445B535F0F974" box="[934,1224,1683,1710]" pageId="33" pageNumber="34">NHMUK 1908.12.14.42</specimenCode>
.
</paragraph>
<paragraph id="8BD13693462DFFE340E845F6309AF928" blockId="33.[189,1399,1751,1887]" box="[234,418,1751,1777]" pageId="33" pageNumber="34">
<emphasis id="B91AEA81462DFFE340E845F6309AF928" bold="true" box="[234,418,1751,1777]" pageId="33" pageNumber="34">
<typeStatus id="54D58831462DFFE340E845F6309AF928" box="[234,418,1751,1777]" pageId="33" pageNumber="34" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD13693462DFFE340BF45DA322FF887" blockId="33.[189,1399,1751,1887]" pageId="33" pageNumber="34">
<materialsCitation id="3B063CCE462DFFE340BF45DA31C0F8E3" ID-GBIF-Occurrence="2605305740" collectionCode="NHMUK" country="South Africa" location="SOUTH AFRICA" pageId="33" pageNumber="34" specimenCode="NHMUK 1937.12.30.2793 to 2794" specimenCount="2" typeStatus="paralectotype">
<collectingCountry id="F3797603462DFFE340BF45DA30A8F8CC" box="[189,400,1787,1813]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
•
<specimenCount id="9D68FD1A462DFFE341AF45DA337BF8CC" box="[429,579,1787,1813]" pageId="33" pageNumber="34" type="generic">2 specimens</specimenCount>
; same collection data as for lectotype;
<specimenCode id="DBC89EE8462DFFE3441A45DD31C0F8E3" pageId="33" pageNumber="34">NHMUK 1937.12.30.2793 to 2794</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B063CCE462DFFE3411A4401354AF8E3" ID-GBIF-Occurrence="2605305671" box="[280,1138,1823,1850]" collectionCode="NMSA" country="South Africa" pageId="33" pageNumber="34" specimenCode="NMSA 2175/T624" specimenCount="4" typeStatus="paralectotype">
<specimenCount id="9D68FD1A462DFFE3411A44013088F8E3" box="[280,432,1824,1850]" pageId="33" pageNumber="34" type="generic">4 specimens</specimenCount>
; same collection data as for lectotype;
<specimenCode id="DBC89EE8462DFFE343914401354AF8E3" box="[915,1138,1823,1850]" pageId="33" pageNumber="34">NMSA 2175/T624</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B063CCE462DFFE344914401322CF887" ID-GBIF-Occurrence="2605305700" collectionCode="NMSA" country="South Africa" pageId="33" pageNumber="34" specimenCode="NMSA 2181/T624" specimenCount="3" typeStatus="paralectotype">
<specimenCount id="9D68FD1A462DFFE3449144013413F8E3" box="[1171,1323,1824,1850]" pageId="33" pageNumber="34" type="generic">3 specimens</specimenCount>
; same collection data as for lectotype;
<specimenCode id="DBC89EE8462DFFE342374465322CF887" box="[565,788,1860,1886]" pageId="33" pageNumber="34">NMSA 2181/T624</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD13693462DFFE340E844A6309EF878" blockId="33.[189,1398,1927,2027]" box="[234,422,1927,1953]" pageId="33" pageNumber="34">
<emphasis id="B91AEA81462DFFE340E844A6309EF878" bold="true" box="[234,422,1927,1953]" pageId="33" pageNumber="34">Other material</emphasis>
</paragraph>
<paragraph id="8BD13693462DFFE340BF448D3403F832" blockId="33.[189,1398,1927,2027]" pageId="33" pageNumber="34">
<materialsCitation id="3B063CCE462DFFE340BF448D3035F832" ID-GBIF-Occurrence="2605305656" collectionCode="NMSA" country="South Africa" location="SOUTH AFRICA" pageId="33" pageNumber="34" specimenCode="V3552" specimenCount="4">
<collectingCountry id="F3797603462DFFE340BF448D30AAF81F" box="[189,402,1964,1990]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
•
<specimenCount id="9D68FD1A462DFFE341B3448D3370F81F" box="[433,584,1964,1990]" pageId="33" pageNumber="34" type="generic">4 specimens</specimenCount>
; same collection data as for lectotype;
<emphasis id="B91AEA81462DFFE34424448C3578F81F" box="[1062,1088,1965,1990]" italics="true" pageId="33" pageNumber="34">ex</emphasis>
Albany Museum;
<collectionCode id="ED7FAE56462DFFE34521448D344EF81F" box="[1315,1398,1964,1990]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="33" pageNumber="34" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462DFFE340BF44F13035F832" box="[189,269,2000,2027]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="33" pageNumber="34" type="Museum">V3552</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B063CCE462DFFE3412444F0340FF832" ID-GBIF-Occurrence="2605305736" box="[294,1335,2001,2027]" collectionCode="NMSA" country="South Africa" location="Transvaal Museum" pageId="33" pageNumber="34" specimenCode="W462" specimenCount="4">
<specimenCount id="9D68FD1A462DFFE3412444F03082F832" box="[294,442,2001,2027]" pageId="33" pageNumber="34" type="generic">4 specimens</specimenCount>
; same collection data as for lectotype;
<emphasis id="B91AEA81462DFFE3438344F332A3F832" box="[897,923,2002,2027]" italics="true" pageId="33" pageNumber="34">ex</emphasis>
Transvaal Museum;
<collectionCode id="ED7FAE56462DFFE3449344F035DCF832" box="[1169,1252,2001,2027]" collectionName="South Africa, Kwa-Zulu Natal, Pietermaritzburg, Natal Museum" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="33" pageNumber="34" type="Museum">NMSA</collectionCode>
<specimenCode id="DBC89EE8462DFFE344E944F0340FF832" box="[1259,1335,2001,2027]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="33" pageNumber="34" type="Museum">W462</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462EFFE040BF422A3338FE1C" pageId="34" pageNumber="35" type="distribution">
<paragraph id="8BD13693462EFFE040BF422A330CFEFC" blockId="34.[189,564,267,293]" box="[189,564,267,293]" pageId="34" pageNumber="35">
<heading id="D09981FF462EFFE040BF422A330CFEFC" bold="true" box="[189,564,267,293]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B91AEA81462EFFE040BF422A330CFEFC" bold="true" box="[189,564,267,293]" pageId="34" pageNumber="35">Distribution and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462EFFE040BF421A3338FE1C" blockId="34.[189,1399,315,453]" pageId="34" pageNumber="35">
A narrow-range endemic (
<figureCitation id="13552A16462EFFE041FF421A336EFE8C" box="[509,598,315,341]" captionStart="Fig" captionStartId="29.[189,232,1896,1922]" captionTargetBox="[302,1285,1093,1856]" captionTargetId="figure@29.[302,1285,1093,1856]" captionTargetPageId="29" captionText="Fig. 11. Distribution of Costulodonta gen. nov. species. C. acinaces (Connolly, 1933) gen. et comb. nov. (green square), C. bidens gen. et sp. nov. (turquoise squares), C. burnupi (Connolly, 1933) gen. et comb. nov. (orange square) C. pluridens gen. et sp. nov. (pink squares), C. trilamellaris (Melvill &amp; Ponsonby, 1908) gen. et comb. nov. (royal blue square). Contour at 1000 m." figureDoi="http://doi.org/10.5281/zenodo.3762302" httpUri="https://zenodo.org/record/3762302/files/figure.png" pageId="34" pageNumber="35">Fig. 11</figureCitation>
), known only from the Dargle area,
<collectingRegion id="49AAF871462EFFE0441D421A35E2FE8C" box="[1055,1242,315,341]" country="South Africa" name="KwaZulu-Natal" pageId="34" pageNumber="35">KwaZulu-Natal</collectingRegion>
, at ±
<quantity id="4C969B76462EFFE04518421A344FFE8F" box="[1306,1399,315,342]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="34" pageNumber="35" unit="m" value="1150.0">1150 m</quantity>
a.s.l.; presumably in leaf-litter of southern mistbelt forest. Forested habitats in nature reserves in the Bulwer-Dargle-Karkloof area need to be surveyed in the hope of finding extant colonies of this species in formally protected areas.
</paragraph>
</subSubSection>
<subSubSection id="C3746518462EFFE040BF42CF32ABFD84" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BD13693462EFFE040BF42CF3017FDD1" blockId="34.[189,303,494,520]" box="[189,303,494,520]" pageId="34" pageNumber="35">
<heading id="D09981FF462EFFE040BF42CF3017FDD1" bold="true" box="[189,303,494,520]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B91AEA81462EFFE040BF42CF3017FDD1" bold="true" box="[189,303,494,520]" pageId="34" pageNumber="35">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693462EFFE040BF413F32ABFD84" blockId="34.[189,1399,542,606]" pageId="34" pageNumber="35">
Despite the
<collectingRegion id="49AAF871462EFFE04146413F30C4FDE1" box="[324,508,542,568]" country="South Africa" name="KwaZulu-Natal" pageId="34" pageNumber="35">KwaZulu-Natal</collectingRegion>
<collectingRegion id="49AAF871462EFFE04203413F3348FDE1" box="[513,624,542,568]" country="Zimbabwe" name="Midlands" pageId="34" pageNumber="35">Midlands</collectingRegion>
being a relatively well-sampled area, the original samples collected by Henry Burnup remain the only ones known of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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