treatments-xml/data/03/C7/87/03C78781FFDCEB3574CAF8CFFD9BFF51.xml
2024-06-21 12:22:17 +02:00

1199 lines
140 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="74E32F73BB3E53FD330DA6E5BF72FD59" ID="10.11646/zootaxa.4171.1.6" ID-DOI="10.11646/zootaxa.4171.1.6" ID-GBIF-Dataset="6790e7b7-9ef7-4609-8b4f-92ad38e27fd9" ID-ISSN="1175-5326" ID-Zenodo-Dep="268260" ID-ZooBank="FA8A04EB-BFB0-4320-8E6D-CE28857BEB51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474963706347" checkinUser="plazi" docAuthor="Mcdonald, Keith R., Rowley, Jodi J. L., Richards, Stephen J. &amp; Frankham, Greta J." docDate="2016" docId="03C78781FFDCEB3574CAF8CFFD9BFF51" docLanguage="en" docName="zootaxa.4171.1.6.pdf" docOrigin="Zootaxa 4171 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litoria bella Mcdonald, Rowley, Richards &amp; Frankham, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="166" masterDocId="FFFEFFF9FFDAEB38745DFFA1FF83FF85" masterDocTitle="A new species of treefrog (Litoria) from Cape York Peninsula, Australia" masterLastPageNumber="169" masterPageNumber="153" pageNumber="159" updateTime="1698387458670" updateUser="plazi">
<mods:mods id="368BD14DC591615411FE1E59F8BF71D2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5819DCAE744F6B3DC6C6E262F838249A">
<mods:title id="9DBA59638ECA6D0B91709B76DE22E4F8">A new species of treefrog (Litoria) from Cape York Peninsula, Australia</mods:title>
</mods:titleInfo>
<mods:name id="35C84D15D79D436F8451672601CCA947" type="personal">
<mods:role id="1EF1E935ADBD4940095A516C31E03190">
<mods:roleTerm id="07B8487862C5B57470C0C6E24786C782">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B19C32A0BC5E5D4CBCDBEFBAD4A7ACF8">Mcdonald, Keith R.</mods:namePart>
</mods:name>
<mods:name id="2F52C17D45B61BE3C04F4B7399120C62" type="personal">
<mods:role id="0AD56E57ED5A46C46CDA15B7755188C3">
<mods:roleTerm id="7D40033EEB06451DE26FF4E5C578DD13">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67B127E1101D95BE675D3319693F5533">Rowley, Jodi J. L.</mods:namePart>
</mods:name>
<mods:name id="A9A8C5EF74F44E02F3B54ACA42174CDB" type="personal">
<mods:role id="C6A54D23EC146238633BE0499B314B0E">
<mods:roleTerm id="89F8F384C421D64CB28759C0C28DC30D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7CFC6A69281A4BA5E51E276F158BFDDE">Richards, Stephen J.</mods:namePart>
</mods:name>
<mods:name id="9142255CDCFE3F8E53B3764DE69E31CF" type="personal">
<mods:role id="555B3BEE3327B39A2ADA774FED105791">
<mods:roleTerm id="683723F4C0E7E7D3D3CA9DF56305DED3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A466FDF4E32244A7DA1538E09114FE7E">Frankham, Greta J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A9A4F1950FFF90E7A2578F954A40FEEC">text</mods:typeOfResource>
<mods:relatedItem id="006E2459BEF3164BEF4C40ABC61A86CC" type="host">
<mods:titleInfo id="109A291008B16542C3E5F4065A4CA587">
<mods:title id="893662AC28C6FF13AA2094EDCFF01046">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A514B978B9F14B90C7E030027458C90F">
<mods:date id="ECD12DCFDD7EE5D673FE4180903B3183">2016</mods:date>
<mods:detail id="B44147E17D2B79E92F43B848A3685D2C" type="volume">
<mods:number id="968A20167E64231FFE1C32849E715180">4171</mods:number>
</mods:detail>
<mods:detail id="A158C0250CE679A72120010D56EEC3EC" type="issue">
<mods:number id="B4ECE8EF1A0243AF13C74ECB9E51D09B">1</mods:number>
</mods:detail>
<mods:extent id="DC06DDABCD1F0CDD89F161AEB65C16CD" unit="page">
<mods:start id="03D3B107E934BD2DDE044B1259C8A1EA">153</mods:start>
<mods:end id="896DB5311A52A67402D1859E329F08AB">169</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="87971BFEB57565F6958AEEF063503602">journal article</mods:classification>
<mods:identifier id="ACB8889FB43E65F31D7043A5BA996198" type="DOI">10.11646/zootaxa.4171.1.6</mods:identifier>
<mods:identifier id="A64B50959D4FC10DE007C49645B8D817" type="GBIF-Dataset">6790e7b7-9ef7-4609-8b4f-92ad38e27fd9</mods:identifier>
<mods:identifier id="DC426DD79956C7DFF1F1F4583786558C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4C10B0602C156F3855C86C4C866FDE8C" type="Zenodo-Dep">268260</mods:identifier>
<mods:identifier id="D056918EE7EC7BC90B90A0DD36613706" type="ZooBank">FA8A04EB-BFB0-4320-8E6D-CE28857BEB51</mods:identifier>
</mods:mods>
<treatment id="03C78781FFDCEB3574CAF8CFFD9BFF51" ID-DOI="http://doi.org/10.5281/zenodo.5631613" ID-GBIF-Taxon="124553119" ID-Zenodo-Dep="5631613" LSID="urn:lsid:plazi:treatment:03C78781FFDCEB3574CAF8CFFD9BFF51" httpUri="http://treatment.plazi.org/id/03C78781FFDCEB3574CAF8CFFD9BFF51" lastPageId="13" lastPageNumber="166" pageId="6" pageNumber="159">
<subSubSection id="C374651CFFDCEB3E74CAF8CFFE13F80D" box="[151,400,1902,1928]" pageId="6" pageNumber="159" type="nomenclature">
<paragraph id="8BD13697FFDCEB3E74CAF8CFFE13F80D" blockId="6.[151,400,1902,1928]" box="[151,400,1902,1928]" pageId="6" pageNumber="159">
<heading id="D09981FBFFDCEB3E74CAF8CFFE13F80D" bold="true" box="[151,400,1902,1928]" fontSize="11" level="1" pageId="6" pageNumber="159" reason="1">
<emphasis id="B91AEA85FFDCEB3E74CAF8CFFE13F80D" bold="true" box="[151,400,1902,1928]" pageId="6" pageNumber="159">
<taxonomicName id="4C6E4D14FFDCEB3E74CAF8CFFEAFF80D" box="[151,300,1902,1928]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="6" pageNumber="159" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFDCEB3E74CAF8CFFEAFF80D" bold="true" box="[151,300,1902,1928]" italics="true" pageId="6" pageNumber="159">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFDCEB3E756EF8CFFE13F80D" box="[307,400,1902,1928]" pageId="6" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFDCEB3F74CAF814FBB3FAB9" lastPageId="7" lastPageNumber="160" pageId="6" pageNumber="159" type="materials_examined">
<paragraph id="8BD13697FFDCEB3E74CAF814FB47F874" blockId="6.[151,1436,1973,2034]" pageId="6" pageNumber="159">
<materialsCitation id="3B063CCAFFDCEB3E74CAF814FB47F874" ID-GBIF-Occurrence="1315183437" collectingDate="2000-10-23" country="Australia" elevation="540" latitude="13.8869" location="Lankelly Creek" longLatPrecision="7" longitude="143.2673" municipality="Cape York Peninsula" pageId="6" pageNumber="159" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="54D58835FFDCEB3E74CAF814FE87F84B" box="[151,260,1973,1998]" pageId="6" pageNumber="159" type="holotype">
<emphasis id="B91AEA85FFDCEB3E74CAF814FE87F84B" bold="true" box="[151,260,1973,1998]" pageId="6" pageNumber="159">Holotype</emphasis>
</typeStatus>
: QM
<accessionNumber id="943DAB74FFDCEB3E7514F814FE1EF84B" box="[329,413,1973,1998]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74476" pageId="6" pageNumber="159" type="EnaNcbi">J74476</accessionNumber>
, adult male, from
<location id="8EB1604CFFDCEB3E7633F814FCA3F84B" LSID="urn:lsid:plazi:treatment:03C78781FFDCEB3574CAF8CFFD9BFF51:8EB1604CFFDCEB3E7633F814FCA3F84B" box="[622,800,1973,1998]" country="Australia" latitude="13.8869" longLatPrecision="7" longitude="143.2673" municipality="Cape York Peninsula" name="Lankelly Creek" pageId="6" pageNumber="159" stateProvince="Queensland">Lankelly Creek</location>
,
<locationDeviation id="9A632FB5FFDCEB3E7771F817FB84F84B" box="[812,1031,1973,1998]" pageId="6" pageNumber="159">
<quantity id="4C969B72FFDCEB3E7771F817FCF7F84B" box="[812,884,1973,1998]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="159" unit="km" value="10.0">10 km</quantity>
NE of Coen
</locationDeviation>
,
<collectingMunicipality id="6BB5ACEDFFDCEB3E7049F817FA86F84B" box="[1044,1285,1973,1998]" pageId="6" pageNumber="159">Cape York Peninsula</collectingMunicipality>
,
<collectingRegion id="49AAF875FFDCEB3E714CF814FA1BF84B" box="[1297,1432,1973,1998]" country="Australia" name="Queensland" pageId="6" pageNumber="159">Queensland</collectingRegion>
,
<collectingCountry id="F3797607FFDCEB3E74CAF878FF7DF877" box="[151,254,2009,2034]" name="Australia" pageId="6" pageNumber="159">Australia</collectingCountry>
(
<geoCoordinate id="EE5A5050FFDCEB3E7551F878FE0CF874" box="[268,399,2009,2034]" direction="north" orientation="latitude" pageId="6" pageNumber="159" precision="5" value="13.8869">13.8869º N</geoCoordinate>
;
<geoCoordinate id="EE5A5050FFDCEB3E75C1F878FDA2F877" box="[412,545,2009,2034]" direction="east" orientation="longitude" pageId="6" pageNumber="159" precision="5" value="143.2673">143.2673ºE</geoCoordinate>
,
<elevation id="0043D1A4FFDCEB3E7671F878FDF0F874" box="[556,627,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="159" unit="m" value="540.0">
<quantity id="4C969B72FFDCEB3E7671F878FDF0F874" box="[556,627,2009,2034]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="159" unit="m" value="540.0">540 m</quantity>
</elevation>
asl;
<figureCitation id="13552A12FFDCEB3E76F5F878FD6BF877" box="[680,744,2009,2034]" captionStart="FIGURE 1" captionStartId="1.[151,250,1642,1664]" captionTargetBox="[154,1432,520,1615]" captionTargetId="figure@1.[151,1436,518,1617]" captionTargetPageId="1" captionText="FIGURE 1. Localities of specimens and tissues of Litoria auae (pale yellow), Litoria bella sp. nov. (dark yellow), Litoria gracilenta (dark orange), and L. chloris (dark red) used in this study. DNA only = circle, specimen only = diamond, DNA and specimen = square. The type localities of the first three species are indicated by stars." httpUri="https://zenodo.org/record/268261/files/figure.png" pageId="6" pageNumber="159">Fig. 1</figureCitation>
). Collected on
<collectingDate id="EF94E9BFFFDCEB3E77CBF87BFBD1F877" box="[918,1106,2009,2034]" pageId="6" pageNumber="159" value="2000-10-23">
<date id="FFD01057FFDCEB3E77CBF87BFBD1F877" box="[918,1106,2009,2034]" pageId="6" pageNumber="159" value="2000-10-23">23 October 2000</date>
</collectingDate>
by KRM.
</materialsCitation>
</paragraph>
<caption id="DF11661FFFDDEB3F74CAFBB2FC9BFBCD" httpUri="https://zenodo.org/record/268263/files/figure.png" pageId="7" pageNumber="160" targetBox="[157,1436,193,1022]" targetPageId="7">
<paragraph id="8BD13697FFDDEB3F74CAFBB2FC9BFBCD" blockId="7.[151,1436,1043,1096]" pageId="7" pageNumber="160">
<emphasis id="B91AEA85FFDDEB3F74CAFBB2FE97FBAD" bold="true" box="[151,276,1043,1065]" pageId="7" pageNumber="160">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="4C6E4D14FFDDEB3F75C3FBB5FD9CFBAC" box="[414,543,1043,1065]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFDDEB3F75C3FBB5FD9CFBAC" box="[414,543,1043,1065]" italics="true" pageId="7" pageNumber="160">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFDDEB3F767BFBB5FDFBFBAC" box="[550,632,1044,1065]" pageId="7" pageNumber="160" rank="species">
<emphasis id="B91AEA85FFDDEB3F767BFBB5FDFBFBAC" bold="true" box="[550,632,1044,1065]" pageId="7" pageNumber="160">sp. nov.</emphasis>
</taxonomicNameLabel>
(QM
<accessionNumber id="943DAB74FFDDEB3F76E5FBB2FC80FBAC" box="[696,771,1043,1065]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74476" pageId="7" pageNumber="160" type="EnaNcbi">J74476</accessionNumber>
) in preservative (A) dorsal surface, (B) lateral view of head, (C) palmar surface of left hand, and (D) plantar surface of left foot.
</paragraph>
</caption>
<paragraph id="8BD13697FFDDEB3F749AFBCEFBB3FAB9" blockId="7.[151,1437,1135,2024]" pageId="7" pageNumber="160">
<materialsCitation id="3B063CCAFFDDEB3F749AFBCEFBB9FB55" ID-GBIF-Occurrence="1315183471" collectingDate="1978-12-18" country="Australia" elevation="180" location="Little Stewart Creek" longitude="143.3138" municipality="Cape York Peninsula" pageId="7" pageNumber="160" specimenCount="2" stateProvince="Queensland" typeStatus="paratype">
<typeStatus id="54D58835FFDDEB3F749AFBCEFEBCFB0D" box="[199,319,1135,1160]" pageId="7" pageNumber="160" type="paratype">
<emphasis id="B91AEA85FFDDEB3F749AFBCEFEBCFB0D" bold="true" box="[199,319,1135,1160]" pageId="7" pageNumber="160">Paratypes</emphasis>
</typeStatus>
: QM
<accessionNumber id="943DAB74FFDDEB3F75DCFBCEFE55FB0D" box="[385,470,1135,1160]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74466" pageId="7" pageNumber="160" type="EnaNcbi">J74466</accessionNumber>
8, 74470, 744735, 74478, eight adult males from same locality and date as
<typeStatus id="54D58835FFDDEB3F716EFBCEFA1AFB0D" box="[1331,1433,1135,1160]" pageId="7" pageNumber="160" type="holotype">holotype</typeStatus>
. QM
<accessionNumber id="943DAB74FFDDEB3F748CFB35FEA6FB29" box="[209,293,1172,1196]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J38289" pageId="7" pageNumber="160" type="EnaNcbi">J38289</accessionNumber>
, adult male, from
<location id="8EB1604CFFDDEB3F7655FB32FD79FB29" LSID="urn:lsid:plazi:treatment:03C78781FFDCEB3574CAF8CFFD9BFF51:8EB1604CFFDDEB3F7655FB32FD79FB29" box="[520,762,1171,1196]" country="Australia" longitude="143.3138" municipality="Cape York Peninsula" name="Little Stewart Creek" pageId="7" pageNumber="160" stateProvince="Queensland">Little Stewart Creek</location>
,
<locationDeviation id="9A632FB5FFDDEB3F7756FB35FC76FB29" box="[779,1013,1171,1196]" pageId="7" pageNumber="160">
<quantity id="4C969B72FFDDEB3F7756FB35FCDBFB29" box="[779,856,1171,1196]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="160" unit="km" value="19.0">19 km</quantity>
SE of Coen
</locationDeviation>
,
<collectingMunicipality id="6BB5ACEDFFDDEB3F705BFB35FA83FB29" box="[1030,1280,1171,1196]" pageId="7" pageNumber="160">Cape York Peninsula</collectingMunicipality>
,
<collectingRegion id="49AAF875FFDDEB3F714CFB32FA1BFB29" box="[1297,1432,1171,1196]" country="Australia" name="Queensland" pageId="7" pageNumber="160">Queensland</collectingRegion>
,
<collectingCountry id="F3797607FFDDEB3F74CAFB16FF7DFB55" box="[151,254,1207,1232]" name="Australia" pageId="7" pageNumber="160">Australia</collectingCountry>
(
<geoCoordinate id="EE5A5050FFDDEB3F7553FB16FE04FB55" box="[270,391,1207,1232]" direction="east" orientation="longitude" pageId="7" pageNumber="160" precision="5" value="14.0776">14.0776ºE</geoCoordinate>
,
<geoCoordinate id="EE5A5050FFDDEB3F75C8FB19FDA7FB4A" box="[405,548,1208,1232]" direction="east" orientation="longitude" pageId="7" pageNumber="160" precision="5" value="143.3138">143.3138º E</geoCoordinate>
: ~
<elevation id="0043D1A4FFDDEB3F761DFB19FD09FB4A" box="[576,650,1208,1232]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="160" unit="m" value="180.0">
<quantity id="4C969B72FFDDEB3F761DFB19FD09FB4A" box="[576,650,1208,1232]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="160" unit="m" value="180.0">180 m</quantity>
</elevation>
asl), collected on
<collectingDate id="EF94E9BFFFDDEB3F7703FB19FBB9FB55" box="[862,1082,1207,1232]" pageId="7" pageNumber="160" value="1978-12-18">
<date id="FFD01057FFDDEB3F7703FB19FBB9FB55" box="[862,1082,1207,1232]" pageId="7" pageNumber="160" value="1978-12-18">18 December 1978</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B063CCAFFDDEB3F701AFB19FDEBFA9D" ID-GBIF-Occurrence="1315183469" collectingDate="1981-11-22" country="Australia" county="Cape York Peninsula" elevation="40" latitude="-13.0082" location="Coconut Creek" longLatPrecision="7" longitude="141.7991" municipality="Aurukun" pageId="7" pageNumber="160" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
QM
<accessionNumber id="943DAB74FFDDEB3F7020FB16FB52FB55" box="[1149,1233,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J39656" pageId="7" pageNumber="160" type="EnaNcbi">J39656</accessionNumber>
, adult male, from
<location id="8EB1604CFFDDEB3F74CAFB7DFEC0FB71" LSID="urn:lsid:plazi:treatment:03C78781FFDCEB3574CAF8CFFD9BFF51:8EB1604CFFDDEB3F74CAFB7DFEC0FB71" box="[151,323,1243,1268]" country="Australia" county="Cape York Peninsula" latitude="-13.0082" longLatPrecision="7" longitude="141.7991" municipality="Aurukun" name="Coconut Creek" pageId="7" pageNumber="160" stateProvince="Queensland">Coconut Creek</location>
, ~
<quantity id="4C969B72FFDDEB3F7500FB7DFE24FB71" box="[349,423,1243,1268]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="160" unit="km" value="40.0">40 km</quantity>
North of
<collectingMunicipality id="6BB5ACEDFFDDEB3F764AFB7AFDFDFB71" box="[535,638,1243,1268]" pageId="7" pageNumber="160">Aurukun</collectingMunicipality>
,
<collectingCounty id="62B04E1BFFDDEB3F76D4FB7DFCFAFB71" box="[649,889,1243,1268]" pageId="7" pageNumber="160">Cape York Peninsula</collectingCounty>
,
<collectingRegion id="49AAF875FFDDEB3F77D8FB7AFB8FFB71" box="[901,1036,1243,1268]" country="Australia" name="Queensland" pageId="7" pageNumber="160">Queensland</collectingRegion>
,
<collectingCountry id="F3797607FFDDEB3F7045FB7AFBFCFB71" box="[1048,1151,1243,1268]" name="Australia" pageId="7" pageNumber="160">Australia</collectingCountry>
(
<geoCoordinate id="EE5A5050FFDDEB3F70D2FB7DFA85FB71" box="[1167,1286,1244,1268]" direction="south" orientation="latitude" pageId="7" pageNumber="160" precision="5" value="-13.0082">13.0082ºS</geoCoordinate>
,
<geoCoordinate id="EE5A5050FFDDEB3F714EFB7DFA1BFB71" box="[1299,1432,1244,1268]" direction="east" orientation="longitude" pageId="7" pageNumber="160" precision="5" value="141.7991">141.7991ºE</geoCoordinate>
, ~
<elevation id="0043D1A4FFDDEB3F74F8FAA1FF63FA92" box="[165,224,1280,1304]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="160" unit="m" value="40.0">
<quantity id="4C969B72FFDDEB3F74F8FAA1FF63FA92" box="[165,224,1280,1304]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="160" unit="m" value="40.0">40 m</quantity>
</elevation>
asl), collected
<collectingDate id="EF94E9BFFFDDEB3F75D6FAA1FDEBFA9D" box="[395,616,1279,1304]" pageId="7" pageNumber="160" value="1981-11-22">
<date id="FFD01057FFDDEB3F75D6FAA1FDEBFA9D" box="[395,616,1279,1304]" pageId="7" pageNumber="160" value="1981-11-22">22 November 1981</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B063CCAFFDDEB3F7629FAA1FBB3FAB9" ID-GBIF-Occurrence="1315183466" country="Australia" county="Cape York Peninsula" elevation="50" latitude="-11.9666" location="Double Point Road" longLatPrecision="7" longitude="142.9163" municipality="Shelburne Station" pageId="7" pageNumber="160" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
QM
<accessionNumber id="943DAB74FFDDEB3F76F4FB5EFD7FFA9D" box="[681,764,1279,1304]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J46363" pageId="7" pageNumber="160" type="EnaNcbi">J46363</accessionNumber>
4, two adult males,
<location id="8EB1604CFFDDEB3F77BBFB5EFB42FA9D" LSID="urn:lsid:plazi:treatment:03C78781FFDCEB3574CAF8CFFD9BFF51:8EB1604CFFDDEB3F77BBFB5EFB42FA9D" box="[998,1217,1279,1304]" country="Australia" county="Cape York Peninsula" latitude="-11.9666" longLatPrecision="7" longitude="142.9163" municipality="Shelburne Station" name="Double Point Road" pageId="7" pageNumber="160" stateProvince="Queensland">Double Point Road</location>
,
<collectingMunicipality id="6BB5ACEDFFDDEB3F7090FB5EFA1AFA9D" box="[1229,1433,1279,1304]" pageId="7" pageNumber="160">Shelburne Station</collectingMunicipality>
,
<collectingCounty id="62B04E1BFFDDEB3F74CAFA85FE06FAB9" box="[151,389,1315,1340]" pageId="7" pageNumber="160">Cape York Peninsula</collectingCounty>
,
<collectingRegion id="49AAF875FFDDEB3F75CDFA82FD94FAB9" box="[400,535,1315,1340]" country="Australia" name="Queensland" pageId="7" pageNumber="160">Queensland</collectingRegion>
,
<collectingCountry id="F3797607FFDDEB3F767FFA82FD0BFAB9" box="[546,648,1315,1340]" name="Australia" pageId="7" pageNumber="160">Australia</collectingCountry>
(
<geoCoordinate id="EE5A5050FFDDEB3F76C5FA82FC8DFAB9" box="[664,782,1315,1340]" direction="south" orientation="latitude" pageId="7" pageNumber="160" precision="5" value="-11.9666">11.9666ºS</geoCoordinate>
,
<geoCoordinate id="EE5A5050FFDDEB3F7744FA82FC26FABE" box="[793,933,1315,1340]" direction="east" orientation="longitude" pageId="7" pageNumber="160" precision="5" value="142.9163">142.9163º E</geoCoordinate>
, &lt;
<elevation id="0043D1A4FFDDEB3F779DFA82FC79FABE" box="[960,1018,1315,1340]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="160" unit="m" value="50.0">
<quantity id="4C969B72FFDDEB3F779DFA82FC79FABE" box="[960,1018,1315,1340]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="160" unit="m" value="50.0">50 m</quantity>
</elevation>
asl).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFDDEB3F749AFAE6FCD1FA2D" pageId="7" pageNumber="160" type="etymology">
<paragraph id="8BD13697FFDDEB3F749AFAE6FDCAFA01" blockId="7.[151,1437,1135,2024]" pageId="7" pageNumber="160">
<emphasis id="B91AEA85FFDDEB3F749AFAE6FECDFAE5" bold="true" box="[199,334,1351,1376]" pageId="7" pageNumber="160">Etymology.</emphasis>
The specific name “
<taxonomicName id="4C6E4D14FFDDEB3F766EFAE6FDEEFAE5" authority="Latin" authorityName="Latin" box="[563,621,1351,1376]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="bella">bella</taxonomicName>
” (Latin), meaning pretty or lovely, is used as an adjective in reference to the beautiful coloration of this species.
</paragraph>
<paragraph id="8BD13697FFDDEB3F749AFA2EFCD1FA2D" blockId="7.[151,1437,1135,2024]" box="[199,850,1423,1448]" pageId="7" pageNumber="160">
<emphasis id="B91AEA85FFDDEB3F749AFA2EFE7FFA2D" bold="true" box="[199,508,1423,1448]" pageId="7" pageNumber="160">Suggested common name.</emphasis>
Cape York Graceful Treefrog.
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFDDEB3F749AFA12FCCCF969" pageId="7" pageNumber="160" type="diagnosis">
<paragraph id="8BD13697FFDDEB3F749AFA12FCCCF969" blockId="7.[151,1437,1135,2024]" pageId="7" pageNumber="160">
<emphasis id="B91AEA85FFDDEB3F749AFA12FEC2FA49" bold="true" box="[199,321,1459,1484]" pageId="7" pageNumber="160">Diagnosis.</emphasis>
The new species is assigned to the genus
<taxonomicName id="4C6E4D14FFDDEB3F7779FA15FCF1FA4E" box="[804,882,1460,1483]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="7" pageNumber="160" phylum="Chordata" rank="genus">
<emphasis id="B91AEA85FFDDEB3F7779FA15FCF1FA4E" box="[804,882,1460,1483]" italics="true" pageId="7" pageNumber="160">Litoria</emphasis>
</taxonomicName>
based on the presence of a horizontally oriented contracted pupil and the absence of a palpebral reticulum (Zweifel 1958;
<bibRefCitation id="EFFF4B66FFDDEB3F77A2FA76FB7FFA75" author="Tyler" box="[1023,1276,1495,1520]" pageId="7" pageNumber="160" refString="Tyler, M. J. &amp; Davies, M. (1978) Species-groups within the Australopapuan hulid frog genus Litoria Tschudi. Australian Journal of Zoology, 26, 1 - 47. http: // dx. doi. org / 10.1071 / AJZS 063" type="journal article" year="1978">Tyler &amp; Davies 1978</bibRefCitation>
). Based upon molecular and morphological data the new species is closely related to
<taxonomicName id="4C6E4D14FFDDEB3F77F3FA5DFBBBF996" box="[942,1080,1532,1555]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFDDEB3F77F3FA5DFBBBF996" box="[942,1080,1532,1555]" italics="true" pageId="7" pageNumber="160">Litoria auae</emphasis>
</taxonomicName>
, and it exhibits other characters typical of the
<taxonomicName id="4C6E4D14FFDDEB3F7569F981FE45F9B2" box="[308,454,1567,1591]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="7" pageNumber="160" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFDDEB3F7569F981FE45F9B2" box="[308,454,1567,1591]" italics="true" pageId="7" pageNumber="160">L. gracilenta</emphasis>
</taxonomicName>
group: moderately small size (&lt;
<quantity id="4C969B72FFDDEB3F7769F9BEFC06F9B2" box="[820,901,1567,1592]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="160" unit="mm" value="50.0">50 mm</quantity>
SVL), green dorsum and a male advertisement call resembling a low-pitched growl (
<bibRefCitation id="EFFF4B66FFDDEB3F7663F9E2FD62F9D9" author="Menzies" box="[574,737,1603,1628]" pageId="7" pageNumber="160" refString="Menzies, J. (2006) The frogs of New Guinea and the Solomon Islands. Pensoft Publishers, 345 pp." type="book" year="2006">Menzies 2006</bibRefCitation>
;
<bibRefCitation id="EFFF4B66FFDDEB3F76B3F9E2FC72F9D9" author="Menzies" box="[750,1009,1603,1628]" pageId="7" pageNumber="160" refString="Menzies, J. I. &amp; Tyler, M. J. (2004) Litoria gracilenta (Anura: Hylidae) and related species in New Guinea. Australian Journal of Zoology, 52, 191 - 214. http: // dx. doi. org / 10.1071 / ZO 03008" type="journal article" year="2004">Menzies &amp; Tyler 2004</bibRefCitation>
). However it is distinguished from all congeners by the following combination of characters: size moderate (male body size
<quantity id="4C969B72FFDDEB3F70DAF9C6FAB1F905" box="[1159,1330,1639,1664]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="7" pageNumber="160" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
), dorsum near-immaculate green, venter orange, digits and webbing bright orange, lateral surfaces of the thigh bluish purple, no pale canthal stripe, bones white, and male advertisement call a finely pulsed single note with a pulse rate of 56 64 pulses/s and dominant frequency of 2.62.8 kHz at 21.4ºC.
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFDDEB32749AF956FAACF8DA" lastPageId="10" lastPageNumber="163" pageId="7" pageNumber="160" type="description">
<paragraph id="8BD13697FFDDEB30749AF956FCCCFE55" blockId="7.[151,1437,1135,2024]" lastBlockId="8.[151,1436,151,464]" lastPageId="8" lastPageNumber="161" pageId="7" pageNumber="160">
<emphasis id="B91AEA85FFDDEB3F749AF956FE64F895" bold="true" box="[199,487,1783,1808]" pageId="7" pageNumber="160">
Description of
<typeStatus id="54D58835FFDDEB3F7524F956FE61F895" box="[377,482,1783,1808]" pageId="7" pageNumber="160" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, moderate body size (
<quantity id="4C969B72FFDDEB3F7734F956FC53F88A" box="[873,976,1783,1808]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="7" pageNumber="160" unit="mm" value="38.5">38.5 mm</quantity>
SVL;
<figureCitation id="13552A12FFDDEB3F7040F956FBE3F895" box="[1053,1120,1783,1808]" captionStart="FIGURE 3" captionStartId="7.[151,250,1043,1065]" captionTargetBox="[157,1436,193,1022]" captionTargetId="figure@7.[151,1436,193,1022]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Litoria bella sp. nov. (QM J 74476) in preservative (A) dorsal surface, (B) lateral view of head, (C) palmar surface of left hand, and (D) plantar surface of left foot." httpUri="https://zenodo.org/record/268263/files/figure.png" pageId="7" pageNumber="160">Fig. 3</figureCitation>
); head length equal to head width (101%); snout obtusely rounded in dorsal view and truncate in profile; canthus rostralis poorly defined; loreal region flat; interorbital region flat; nostrils oval, closer to tip of snout than eye; eyes moderately large (10% SVL), pupil oval, horizontal, tympanic rim slightly elevated relative to skin of temporal region, top margin obscured by supratympanic skin fold, tympanum 64% of eye diameter; vomeropalatines with two patches of teeth (one on each side) between internal nares (
<figureCitation id="13552A12FFDDEB3F7624F80AFD3DF841" box="[633,702,1963,1988]" captionStart="FIGURE 3" captionStartId="7.[151,250,1043,1065]" captionTargetBox="[157,1436,193,1022]" captionTargetId="figure@7.[151,1436,193,1022]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Litoria bella sp. nov. (QM J 74476) in preservative (A) dorsal surface, (B) lateral view of head, (C) palmar surface of left hand, and (D) plantar surface of left foot." httpUri="https://zenodo.org/record/268263/files/figure.png" pageId="7" pageNumber="160">Fig. 3</figureCitation>
). Vocal sac present; slightly raised supratympanic fold extending to just beyond level of axilla. Finger webbing does not extend to disks (
<figureCitation id="13552A12FFDDEB3F77E2F86EFB81F86D" box="[959,1026,1999,2024]" captionStart="FIGURE 3" captionStartId="7.[151,250,1043,1065]" captionTargetBox="[157,1436,193,1022]" captionTargetId="figure@7.[151,1436,193,1022]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Litoria bella sp. nov. (QM J 74476) in preservative (A) dorsal surface, (B) lateral view of head, (C) palmar surface of left hand, and (D) plantar surface of left foot." httpUri="https://zenodo.org/record/268263/files/figure.png" pageId="7" pageNumber="160">Fig. 3</figureCitation>
C), relative length of fingers I &lt;II &lt;IV &lt;III; tips of all fingers with well-developed disks with distinct circummarginal grooves, disks relatively wide compared to penultimate phalanx width (third finger disk 143% third finger width); subarticular tubercles indistinct; nuptial pads present on posteriodorsal surface of finger I. Relative length of toes I &lt;II &lt;V &lt;III &lt;IV; tips of toes with well-developed disks with distinct circummarginal grooves; disks slightly smaller than those of fingers (fourth toe disk 131% fourth penultimate phalanx width); toe webbing on fourth and fifth toe reaches base of disk, webbing between first and second toe reaches edge of terminal subarticular tubercle (
<figureCitation id="13552A12FFD2EB3070F7FEEDFB6FFEE1" box="[1194,1260,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,250,1043,1065]" captionTargetBox="[157,1436,193,1022]" captionTargetId="figure@7.[151,1436,193,1022]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Litoria bella sp. nov. (QM J 74476) in preservative (A) dorsal surface, (B) lateral view of head, (C) palmar surface of left hand, and (D) plantar surface of left foot." httpUri="https://zenodo.org/record/268263/files/figure.png" pageId="8" pageNumber="161">Fig. 3</figureCitation>
D); subarticular tubercles indistinct, rounded; inner metatarsal tubercle low, oval; outer metatarsal tubercle absent. Tibia length moderately long (55% SVL). Skin of dorsal surfaces finely granular; ventral surfaces of body and thighs coarsely granular, finer on chest and under arms, smooth under shanks.
</paragraph>
<caption id="DF11661FFFD2EB3074CAF9AEFA93F9C1" httpUri="https://zenodo.org/record/268264/files/figure.png" pageId="8" pageNumber="161" targetBox="[151,1436,501,1530]" targetPageId="8">
<paragraph id="8BD13697FFD2EB3074CAF9AEFA93F9C1" blockId="8.[151,1435,1551,1604]" pageId="8" pageNumber="161">
<emphasis id="B91AEA85FFD2EB3074CAF9AEFE96F9A1" bold="true" box="[151,277,1551,1573]" pageId="8" pageNumber="161">FIGURE 4.</emphasis>
<taxonomicName id="4C6E4D14FFD2EB307540F9B0FE1CF9A0" box="[285,415,1551,1574]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="8" pageNumber="161" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFD2EB307540F9B0FE1CF9A0" box="[285,415,1551,1574]" italics="true" pageId="8" pageNumber="161">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFD2EB3075F5F9B1FE79F9A0" box="[424,506,1552,1573]" pageId="8" pageNumber="161" rank="species">
<emphasis id="B91AEA85FFD2EB3075F5F9B1FE79F9A0" bold="true" box="[424,506,1552,1573]" pageId="8" pageNumber="161">sp. nov.</emphasis>
</taxonomicNameLabel>
in life. (A) female, (B) male showing ventral coloration, an (C) male showing thigh and webbing colouration from Iron Range, QLD, and (D) male QM
<accessionNumber id="943DAB74FFD2EB307742F98FFCEBF9C1" box="[799,872,1582,1604]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J65778" pageId="8" pageNumber="161" type="EnaNcbi">J65778</accessionNumber>
from McIlwraith Range, QLD, Australia.
</paragraph>
</caption>
<paragraph id="8BD13697FFD2EB30749AF9CAFCB0F949" blockId="8.[151,1437,1643,2028]" pageId="8" pageNumber="161">
<emphasis id="B91AEA85FFD2EB30749AF9CAFDEEF901" bold="true" box="[199,621,1643,1668]" pageId="8" pageNumber="161">
Colour of
<typeStatus id="54D58835FFD2EB30751EF9CAFE29F901" box="[323,426,1643,1668]" pageId="8" pageNumber="161" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsal ground colour uniform greyish green. Venter, upper arms, inner surface of lower arms, hands and feet including webbing pale yellowish orange. Upper thighs pale yellow with purplish brown wash. Iris pale yellow with black periphery.
</paragraph>
<paragraph id="8BD13697FFD2EB30749AF976FE7CF891" blockId="8.[151,1437,1643,2028]" pageId="8" pageNumber="161">
<emphasis id="B91AEA85FFD2EB30749AF976FE46F975" bold="true" box="[199,453,1751,1776]" pageId="8" pageNumber="161">Measurements (mm).</emphasis>
<typeStatus id="54D58835FFD2EB307591F976FDBAF975" box="[460,569,1751,1776]" pageId="8" pageNumber="161" type="holotype">Holotype</typeStatus>
: SVL 38.5, TIB 21.2, HL 13.8, HW 13.7, EYE 3.7, TMP 2.4, TEY 1.2, END 3.6, IND 3.4, 3FD 2.0, 4TD 1.7.
</paragraph>
<paragraph id="8BD13697FFD2EB30749AF8BEFCCFF869" blockId="8.[151,1437,1643,2028]" pageId="8" pageNumber="161">
<emphasis id="B91AEA85FFD2EB30749AF8BEFEF1F8BD" bold="true" box="[199,370,1823,1848]" pageId="8" pageNumber="161">Colour in life.</emphasis>
Recorded from images of the species from the Iron and McIlwraith Ranges (
<figureCitation id="13552A12FFD2EB3070AFF8BEFAB9F8BD" box="[1266,1338,1823,1848]" captionStart="FIGURE 4" captionStartId="8.[151,250,1551,1573]" captionTargetBox="[151,1436,501,1530]" captionTargetId="figure@8.[151,1436,501,1530]" captionTargetPageId="8" captionText="FIGURE 4. Litoria bella sp. nov. in life. (A) female, (B) male showing ventral coloration, an (C) male showing thigh and webbing colouration from Iron Range, QLD, and (D) male QM J 65778 from McIlwraith Range, QLD, Australia." httpUri="https://zenodo.org/record/268264/files/figure.png" pageId="8" pageNumber="161">Fig. 4</figureCitation>
). Dorsal surface bright green, with bright orange on the dorsal surface of the upper arms, fingers IIII, toes IIV and distal end of finger IV and toe V, including webbing and toe-disks. Blueish purple inner and outer thighs with only thin strip of green along centre. Pale canthal stripe absent; paler yellowish along bottom of upper lip and along flanks. Ventral surfaces are orange, more yellowish along outer edges of limbs and belly. Iris bright yellowish gold with a network of fine darker gold reticulations; iris periphery black.
</paragraph>
<paragraph id="8BD13697FFD3EB31749AFF36FF2FFEC5" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="162">
<emphasis id="B91AEA85FFD3EB31749AFF36FEB8FF35" bold="true" box="[199,315,151,176]" pageId="9" pageNumber="162">Variation.</emphasis>
The
<typeStatus id="54D58835FFD3EB317520FF39FE2EFF35" box="[381,429,152,176]" pageId="9" pageNumber="162">type</typeStatus>
series exhibits little variation in colour or morphology, and specimens observed in life consistently exhibited the colours described above. The large variation in extent of dorsal spotting that is apparent in many other members of the
<taxonomicName id="4C6E4D14FFD3EB3175B1FF40FDFFFF7D" box="[492,636,224,248]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="162" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD3EB3175B1FF40FDFFFF7D" box="[492,636,224,248]" italics="true" pageId="9" pageNumber="162">L. gracilenta</emphasis>
</taxonomicName>
group is not apparent in
<taxonomicName id="4C6E4D14FFD3EB3177CFFF40FC64FF7D" box="[914,999,224,248]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="162" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD3EB3177CFFF40FC64FF7D" box="[914,999,224,248]" italics="true" pageId="9" pageNumber="162">L. bella</emphasis>
</taxonomicName>
. Adult males vary in size from ~
<quantity id="4C969B72FFD3EB317108FF41FF40FE99" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="9" pageNumber="162" unit="mm" value="37.5" valueMax="41.0" valueMin="34.0">3441 mm</quantity>
but variation in morphological features is otherwise slight: measurements of the
<typeStatus id="54D58835FFD3EB317011FEA4FBFFFE98" box="[1100,1148,261,285]" pageId="9" pageNumber="162">type</typeStatus>
series are shown in
<tableCitation id="C6EC032CFFD3EB317103FEA5FF2BFEC5" captionStart="TABLE 4" captionStartId="9.[151,239,368,391]" captionTargetBox="[159,1400,418,986]" captionTargetPageId="9" captionText="TABLE 4. Measurements (mm) of Litoria bella sp. nov. Abbreviations defined in text." httpUri="http://table.plazi.org/id/DF11661FFFD3EB3174CAFED1FBBAFE02" pageId="9" pageNumber="162" tableUuid="DF11661FFFD3EB3174CAFED1FBBAFE02">Table 4</tableCitation>
.
</paragraph>
<caption id="DF11661FFFD3EB3174CAFED1FBBAFE02" ID-Table-UUID="DF11661FFFD3EB3174CAFED1FBBAFE02" box="[151,1081,367,391]" httpUri="http://table.plazi.org/id/DF11661FFFD3EB3174CAFED1FBBAFE02" pageId="9" pageNumber="162" targetBox="[159,1400,418,986]" targetIsTable="true" targetPageId="9">
<paragraph id="8BD13697FFD3EB3174CAFED1FBBAFE02" blockId="9.[151,1081,367,391]" box="[151,1081,367,391]" pageId="9" pageNumber="162">
<emphasis id="B91AEA85FFD3EB3174CAFED1FE80FE02" bold="true" box="[151,259,368,391]" pageId="9" pageNumber="162">TABLE 4</emphasis>
. Measurements (mm) of
<taxonomicName id="4C6E4D14FFD3EB31764CFED1FD14FE02" box="[529,663,367,391]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="162" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFD3EB31764CFED1FD14FE02" box="[529,663,367,391]" italics="true" pageId="9" pageNumber="162">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFD3EB3176C3FED1FD70FE02" box="[670,755,368,391]" pageId="9" pageNumber="162" rank="species">
<emphasis id="B91AEA85FFD3EB3176C3FED1FD6FFE02" bold="true" box="[670,748,368,391]" pageId="9" pageNumber="162">sp. nov</emphasis>
.
</taxonomicNameLabel>
Abbreviations defined in text.
</paragraph>
</caption>
<paragraph id="8BD13697FFD3EB317548FE03FA9CFC5F" pageId="9" pageNumber="162">
<table id="F96EC437FFD314C774C2FE03FAFBFC5F" box="[159,1400,418,986]" gridcols="7" gridrows="14" pageId="9" pageNumber="162" rowsContinueIn="9.[159,1374,1074,1642]">
<tr id="355E34D5FFD314C774C2FE03FAFBFE3D" box="[159,1400,418,440]" gridrow="0" pageId="9" pageNumber="162" rowspan-0="1">
<th id="768F5DA9FFD314C77548FE03FDA8FE3D" box="[277,555,418,440]" gridcol="1" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317519FE03FE13FE3D" box="[324,400,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74476" pageId="9" pageNumber="162" type="EnaNcbi">J74476</accessionNumber>
* QM
<accessionNumber id="943DAB74FFD3EB3175BFFE03FDA8FE3D" box="[482,555,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J46363" pageId="9" pageNumber="162" type="EnaNcbi">J46363</accessionNumber>
</th>
<th id="768F5DA9FFD314C77606FE03FD57FE3D" box="[603,724,418,440]" gridcol="2" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB3176D6FE03FD57FE3D" box="[651,724,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J39656" pageId="9" pageNumber="162" type="EnaNcbi">J39656</accessionNumber>
</th>
<th id="768F5DA9FFD314C77759FE03FCFEFE3D" box="[772,893,418,440]" gridcol="3" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB31776EFE03FCFEFE3D" box="[819,893,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J38289" pageId="9" pageNumber="162" type="EnaNcbi">J38289</accessionNumber>
</th>
<th id="768F5DA9FFD314C777F0FE03FBA5FE3D" box="[941,1062,418,440]" gridcol="4" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317781FE03FBA5FE3D" box="[988,1062,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74470" pageId="9" pageNumber="162" type="EnaNcbi">J74470</accessionNumber>
</th>
<th id="768F5DA9FFD314C7700BFE03FB4CFE3D" box="[1110,1231,418,440]" gridcol="5" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB3170D8FE03FB4CFE3D" box="[1157,1231,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74773" pageId="9" pageNumber="162" type="EnaNcbi">J74773</accessionNumber>
</th>
<th id="768F5DA9FFD314C770A2FE03FAFBFE3D" box="[1279,1400,418,440]" gridcol="6" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317173FE03FAFBFE3D" box="[1326,1400,418,440]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74474" pageId="9" pageNumber="162" type="EnaNcbi">J74474</accessionNumber>
</th>
</tr>
<tr id="355E34D5FFD314C774C2FE6DFAFBFE67" box="[159,1400,460,482]" gridrow="1" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FE6DFF50FE67" box="[159,211,460,482]" gridcol="0" gridrow="1" pageId="9" pageNumber="162">SVL</th>
<td id="768F5DA9FFD314C77548FE6DFDA8FE67" box="[277,555,460,482]" gridcol="1" gridrow="1" pageId="9" pageNumber="162">38.5 36.9</td>
<td id="768F5DA9FFD314C77606FE6DFD57FE67" box="[603,724,460,482]" gridcol="2" gridrow="1" pageId="9" pageNumber="162">41.0</td>
<td id="768F5DA9FFD314C77759FE6DFCFEFE67" box="[772,893,460,482]" gridcol="3" gridrow="1" pageId="9" pageNumber="162">40.3</td>
<td id="768F5DA9FFD314C777F0FE6DFBA5FE67" box="[941,1062,460,482]" gridcol="4" gridrow="1" pageId="9" pageNumber="162">37.6</td>
<td id="768F5DA9FFD314C7700BFE6DFB4CFE67" box="[1110,1231,460,482]" gridcol="5" gridrow="1" pageId="9" pageNumber="162">41.8</td>
<td id="768F5DA9FFD314C770A2FE6DFAFBFE67" box="[1279,1400,460,482]" gridcol="6" gridrow="1" pageId="9" pageNumber="162">37.9</td>
</tr>
<tr id="355E34D5FFD314C774C2FE57FAFBFD89" box="[159,1400,502,524]" gridrow="2" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FE57FF50FD89" box="[159,211,502,524]" gridcol="0" gridrow="2" pageId="9" pageNumber="162">TIB</th>
<td id="768F5DA9FFD314C77548FE57FDA8FD89" box="[277,555,502,524]" gridcol="1" gridrow="2" pageId="9" pageNumber="162">21.2 19.5</td>
<td id="768F5DA9FFD314C77606FE57FD57FD89" box="[603,724,502,524]" gridcol="2" gridrow="2" pageId="9" pageNumber="162">22.1</td>
<td id="768F5DA9FFD314C77759FE57FCFEFD89" box="[772,893,502,524]" gridcol="3" gridrow="2" pageId="9" pageNumber="162">21.2</td>
<td id="768F5DA9FFD314C777F0FE57FBA5FD89" box="[941,1062,502,524]" gridcol="4" gridrow="2" pageId="9" pageNumber="162">22.3</td>
<td id="768F5DA9FFD314C7700BFE57FB4CFD89" box="[1110,1231,502,524]" gridcol="5" gridrow="2" pageId="9" pageNumber="162">23.6</td>
<td id="768F5DA9FFD314C770A2FE57FAFBFD89" box="[1279,1400,502,524]" gridcol="6" gridrow="2" pageId="9" pageNumber="162">21.7</td>
</tr>
<tr id="355E34D5FFD314C774C2FD81FAFBFDB3" box="[159,1400,544,566]" gridrow="3" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FD81FF50FDB3" box="[159,211,544,566]" gridcol="0" gridrow="3" pageId="9" pageNumber="162">HL</th>
<td id="768F5DA9FFD314C77548FD81FDA8FDB3" box="[277,555,544,566]" gridcol="1" gridrow="3" pageId="9" pageNumber="162">13.8 12.0</td>
<td id="768F5DA9FFD314C77606FD81FD57FDB3" box="[603,724,544,566]" gridcol="2" gridrow="3" pageId="9" pageNumber="162">14.2</td>
<td id="768F5DA9FFD314C77759FD81FCFEFDB3" box="[772,893,544,566]" gridcol="3" gridrow="3" pageId="9" pageNumber="162">13.2</td>
<td id="768F5DA9FFD314C777F0FD81FBA5FDB3" box="[941,1062,544,566]" gridcol="4" gridrow="3" pageId="9" pageNumber="162">13.5</td>
<td id="768F5DA9FFD314C7700BFD81FB4CFDB3" box="[1110,1231,544,566]" gridcol="5" gridrow="3" pageId="9" pageNumber="162">13.5</td>
<td id="768F5DA9FFD314C770A2FD81FAFBFDB3" box="[1279,1400,544,566]" gridcol="6" gridrow="3" pageId="9" pageNumber="162">13.6</td>
</tr>
<tr id="355E34D5FFD314C774C2FDEBFAFBFDE5" box="[159,1400,586,608]" gridrow="4" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FDEBFF50FDE5" box="[159,211,586,608]" gridcol="0" gridrow="4" pageId="9" pageNumber="162">HW</th>
<td id="768F5DA9FFD314C77548FDEBFDA8FDE5" box="[277,555,586,608]" gridcol="1" gridrow="4" pageId="9" pageNumber="162">13.7 12.1</td>
<td id="768F5DA9FFD314C77606FDEBFD57FDE5" box="[603,724,586,608]" gridcol="2" gridrow="4" pageId="9" pageNumber="162">15.4</td>
<td id="768F5DA9FFD314C77759FDEBFCFEFDE5" box="[772,893,586,608]" gridcol="3" gridrow="4" pageId="9" pageNumber="162">14.6</td>
<td id="768F5DA9FFD314C777F0FDEBFBA5FDE5" box="[941,1062,586,608]" gridcol="4" gridrow="4" pageId="9" pageNumber="162">14.0</td>
<td id="768F5DA9FFD314C7700BFDEBFB4CFDE5" box="[1110,1231,586,608]" gridcol="5" gridrow="4" pageId="9" pageNumber="162">14.9</td>
<td id="768F5DA9FFD314C770A2FDEBFAFBFDE5" box="[1279,1400,586,608]" gridcol="6" gridrow="4" pageId="9" pageNumber="162">13.9</td>
</tr>
<tr id="355E34D5FFD314C774C2FDD5FAFBFD0F" box="[159,1400,628,650]" gridrow="5" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FDD5FF50FD0F" box="[159,211,628,650]" gridcol="0" gridrow="5" pageId="9" pageNumber="162">EYE</th>
<td id="768F5DA9FFD314C77548FDD5FDA8FD0F" box="[277,555,628,650]" gridcol="1" gridrow="5" pageId="9" pageNumber="162">3.7 4.0</td>
<td id="768F5DA9FFD314C77606FDD5FD57FD0F" box="[603,724,628,650]" gridcol="2" gridrow="5" pageId="9" pageNumber="162">4.3</td>
<td id="768F5DA9FFD314C77759FDD5FCFEFD0F" box="[772,893,628,650]" gridcol="3" gridrow="5" pageId="9" pageNumber="162">4.0</td>
<td id="768F5DA9FFD314C777F0FDD5FBA5FD0F" box="[941,1062,628,650]" gridcol="4" gridrow="5" pageId="9" pageNumber="162">3.9</td>
<td id="768F5DA9FFD314C7700BFDD5FB4CFD0F" box="[1110,1231,628,650]" gridcol="5" gridrow="5" pageId="9" pageNumber="162">4.1</td>
<td id="768F5DA9FFD314C770A2FDD5FAFBFD0F" box="[1279,1400,628,650]" gridcol="6" gridrow="5" pageId="9" pageNumber="162">4.2</td>
</tr>
<tr id="355E34D5FFD314C774C2FD3FFAFBFD31" box="[159,1400,670,692]" gridrow="6" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FD3FFF50FD31" box="[159,211,670,692]" gridcol="0" gridrow="6" pageId="9" pageNumber="162">TMP</th>
<td id="768F5DA9FFD314C77548FD3FFDA8FD31" box="[277,555,670,692]" gridcol="1" gridrow="6" pageId="9" pageNumber="162">2.4 2.5</td>
<td id="768F5DA9FFD314C77606FD3FFD57FD31" box="[603,724,670,692]" gridcol="2" gridrow="6" pageId="9" pageNumber="162">2.4</td>
<td id="768F5DA9FFD314C77759FD3FFCFEFD31" box="[772,893,670,692]" gridcol="3" gridrow="6" pageId="9" pageNumber="162">2.8</td>
<td id="768F5DA9FFD314C777F0FD3FFBA5FD31" box="[941,1062,670,692]" gridcol="4" gridrow="6" pageId="9" pageNumber="162">2.3</td>
<td id="768F5DA9FFD314C7700BFD3FFB4CFD31" box="[1110,1231,670,692]" gridcol="5" gridrow="6" pageId="9" pageNumber="162">2.5</td>
<td id="768F5DA9FFD314C770A2FD3FFAFBFD31" box="[1279,1400,670,692]" gridcol="6" gridrow="6" pageId="9" pageNumber="162">2.3</td>
</tr>
<tr id="355E34D5FFD314C774C2FD69FAFBFD5B" box="[159,1400,712,734]" gridrow="7" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FD69FF50FD5B" box="[159,211,712,734]" gridcol="0" gridrow="7" pageId="9" pageNumber="162">END</th>
<td id="768F5DA9FFD314C77548FD69FDA8FD5B" box="[277,555,712,734]" gridcol="1" gridrow="7" pageId="9" pageNumber="162">3.6 3.4</td>
<td id="768F5DA9FFD314C77606FD69FD57FD5B" box="[603,724,712,734]" gridcol="2" gridrow="7" pageId="9" pageNumber="162">4.0</td>
<td id="768F5DA9FFD314C77759FD69FCFEFD5B" box="[772,893,712,734]" gridcol="3" gridrow="7" pageId="9" pageNumber="162">4.0</td>
<td id="768F5DA9FFD314C777F0FD69FBA5FD5B" box="[941,1062,712,734]" gridcol="4" gridrow="7" pageId="9" pageNumber="162">3.9</td>
<td id="768F5DA9FFD314C7700BFD69FB4CFD5B" box="[1110,1231,712,734]" gridcol="5" gridrow="7" pageId="9" pageNumber="162">3.7</td>
<td id="768F5DA9FFD314C770A2FD69FAFBFD5B" box="[1279,1400,712,734]" gridcol="6" gridrow="7" pageId="9" pageNumber="162">2.7</td>
</tr>
<tr id="355E34D5FFD314C774C2FD53FAFBFC8D" box="[159,1400,754,776]" gridrow="8" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FD53FF50FC8D" box="[159,211,754,776]" gridcol="0" gridrow="8" pageId="9" pageNumber="162">IND</th>
<td id="768F5DA9FFD314C77548FD53FDA8FC8D" box="[277,555,754,776]" gridcol="1" gridrow="8" pageId="9" pageNumber="162">3.4 2.7</td>
<td id="768F5DA9FFD314C77606FD53FD57FC8D" box="[603,724,754,776]" gridcol="2" gridrow="8" pageId="9" pageNumber="162">3.0</td>
<td id="768F5DA9FFD314C77759FD53FCFEFC8D" box="[772,893,754,776]" gridcol="3" gridrow="8" pageId="9" pageNumber="162">3.0</td>
<td id="768F5DA9FFD314C777F0FD53FBA5FC8D" box="[941,1062,754,776]" gridcol="4" gridrow="8" pageId="9" pageNumber="162">3.3</td>
<td id="768F5DA9FFD314C7700BFD53FB4CFC8D" box="[1110,1231,754,776]" gridcol="5" gridrow="8" pageId="9" pageNumber="162">3.4</td>
<td id="768F5DA9FFD314C770A2FD53FAFBFC8D" box="[1279,1400,754,776]" gridcol="6" gridrow="8" pageId="9" pageNumber="162">2.9</td>
</tr>
<tr id="355E34D5FFD314C774C2FCBDFAFBFCB7" box="[159,1400,796,818]" gridrow="9" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FCBDFF50FCB7" box="[159,211,796,818]" gridcol="0" gridrow="9" pageId="9" pageNumber="162">TEY</th>
<td id="768F5DA9FFD314C77548FCBDFDA8FCB7" box="[277,555,796,818]" gridcol="1" gridrow="9" pageId="9" pageNumber="162">1.2 1.4</td>
<td id="768F5DA9FFD314C77606FCBDFD57FCB7" box="[603,724,796,818]" gridcol="2" gridrow="9" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C77759FCBDFCFEFCB7" box="[772,893,796,818]" gridcol="3" gridrow="9" pageId="9" pageNumber="162">1.2</td>
<td id="768F5DA9FFD314C777F0FCBDFBA5FCB7" box="[941,1062,796,818]" gridcol="4" gridrow="9" pageId="9" pageNumber="162">1.2</td>
<td id="768F5DA9FFD314C7700BFCBDFB4CFCB7" box="[1110,1231,796,818]" gridcol="5" gridrow="9" pageId="9" pageNumber="162">1.3</td>
<td id="768F5DA9FFD314C770A2FCBDFAFBFCB7" box="[1279,1400,796,818]" gridcol="6" gridrow="9" pageId="9" pageNumber="162">1.4</td>
</tr>
<tr id="355E34D5FFD314C774C2FCE7FAFBFCD9" box="[159,1400,838,860]" gridrow="10" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FCE7FF50FCD9" box="[159,211,838,860]" gridcol="0" gridrow="10" pageId="9" pageNumber="162">3FD</th>
<td id="768F5DA9FFD314C77548FCE7FDA8FCD9" box="[277,555,838,860]" gridcol="1" gridrow="10" pageId="9" pageNumber="162">2.0 1.8</td>
<td id="768F5DA9FFD314C77606FCE7FD57FCD9" box="[603,724,838,860]" gridcol="2" gridrow="10" pageId="9" pageNumber="162">2.6</td>
<td id="768F5DA9FFD314C77759FCE7FCFEFCD9" box="[772,893,838,860]" gridcol="3" gridrow="10" pageId="9" pageNumber="162">2.3</td>
<td id="768F5DA9FFD314C777F0FCE7FBA5FCD9" box="[941,1062,838,860]" gridcol="4" gridrow="10" pageId="9" pageNumber="162">2.1</td>
<td id="768F5DA9FFD314C7700BFCE7FB4CFCD9" box="[1110,1231,838,860]" gridcol="5" gridrow="10" pageId="9" pageNumber="162">2.1</td>
<td id="768F5DA9FFD314C770A2FCE7FAFBFCD9" box="[1279,1400,838,860]" gridcol="6" gridrow="10" pageId="9" pageNumber="162">2.3</td>
</tr>
<tr id="355E34D5FFD314C774C2FCD1FAFBFC03" box="[159,1400,880,902]" gridrow="11" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FCD1FF50FC03" box="[159,211,880,902]" gridcol="0" gridrow="11" pageId="9" pageNumber="162">3F</th>
<td id="768F5DA9FFD314C77548FCD1FDA8FC03" box="[277,555,880,902]" gridcol="1" gridrow="11" pageId="9" pageNumber="162">1.4 1.2</td>
<td id="768F5DA9FFD314C77606FCD1FD57FC03" box="[603,724,880,902]" gridcol="2" gridrow="11" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C77759FCD1FCFEFC03" box="[772,893,880,902]" gridcol="3" gridrow="11" pageId="9" pageNumber="162">1.3</td>
<td id="768F5DA9FFD314C777F0FCD1FBA5FC03" box="[941,1062,880,902]" gridcol="4" gridrow="11" pageId="9" pageNumber="162">1.3</td>
<td id="768F5DA9FFD314C7700BFCD1FB4CFC03" box="[1110,1231,880,902]" gridcol="5" gridrow="11" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C770A2FCD1FAFBFC03" box="[1279,1400,880,902]" gridcol="6" gridrow="11" pageId="9" pageNumber="162">1.4</td>
</tr>
<tr id="355E34D5FFD314C774C2FC3BFAFBFC35" box="[159,1400,922,944]" gridrow="12" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FC3BFF50FC35" box="[159,211,922,944]" gridcol="0" gridrow="12" pageId="9" pageNumber="162">4TD</th>
<td id="768F5DA9FFD314C77548FC3BFDA8FC35" box="[277,555,922,944]" gridcol="1" gridrow="12" pageId="9" pageNumber="162">1.7 1.7</td>
<td id="768F5DA9FFD314C77606FC3BFD57FC35" box="[603,724,922,944]" gridcol="2" gridrow="12" pageId="9" pageNumber="162">2.2</td>
<td id="768F5DA9FFD314C77759FC3BFCFEFC35" box="[772,893,922,944]" gridcol="3" gridrow="12" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C777F0FC3BFBA5FC35" box="[941,1062,922,944]" gridcol="4" gridrow="12" pageId="9" pageNumber="162">2.3</td>
<td id="768F5DA9FFD314C7700BFC3BFB4CFC35" box="[1110,1231,922,944]" gridcol="5" gridrow="12" pageId="9" pageNumber="162">2.1</td>
<td id="768F5DA9FFD314C770A2FC3BFAFBFC35" box="[1279,1400,922,944]" gridcol="6" gridrow="12" pageId="9" pageNumber="162">1.8</td>
</tr>
<tr id="355E34D5FFD314C774C2FC65FAFBFC5F" box="[159,1400,964,986]" gridrow="13" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FC65FF50FC5F" box="[159,211,964,986]" gridcol="0" gridrow="13" pageId="9" pageNumber="162">4F</th>
<td id="768F5DA9FFD314C77548FC65FDA8FC5F" box="[277,555,964,986]" gridcol="1" gridrow="13" pageId="9" pageNumber="162">1.3 1.3</td>
<td id="768F5DA9FFD314C77606FC65FD57FC5F" box="[603,724,964,986]" gridcol="2" gridrow="13" pageId="9" pageNumber="162">1.6</td>
<td id="768F5DA9FFD314C77759FC65FCFEFC5F" box="[772,893,964,986]" gridcol="3" gridrow="13" pageId="9" pageNumber="162">1.2</td>
<td id="768F5DA9FFD314C777F0FC65FBA5FC5F" box="[941,1062,964,986]" gridcol="4" gridrow="13" pageId="9" pageNumber="162">1.4</td>
<td id="768F5DA9FFD314C7700BFC65FB4CFC5F" box="[1110,1231,964,986]" gridcol="5" gridrow="13" pageId="9" pageNumber="162">1.2</td>
<td id="768F5DA9FFD314C770A2FC65FAFBFC5F" box="[1279,1400,964,986]" gridcol="6" gridrow="13" pageId="9" pageNumber="162">1.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD13697FFD3EB3174CAFC5EFE8BFB93" blockId="9.[151,264,1023,1046]" box="[151,264,1023,1046]" pageId="9" pageNumber="162">continued.</paragraph>
<paragraph id="8BD13697FFD3EB317578FB93FA85F9EF" pageId="9" pageNumber="162">
<table id="F96EC437FFD314C774C2FB93FADDF9EF" box="[159,1374,1074,1642]" gridcols="7" gridrows="14" pageId="9" pageNumber="162" rowsContinueFrom="9.[159,1400,418,986]">
<tr id="355E34D5FFD314C774C2FB93FADDFBCD" box="[159,1374,1074,1096]" gridrow="0" pageId="9" pageNumber="162" rowspan-0="1">
<th id="768F5DA9FFD314C77578FB93FE1DFBCD" box="[293,414,1074,1096]" gridcol="1" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317509FB93FE1DFBCD" box="[340,414,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74467" pageId="9" pageNumber="162" type="EnaNcbi">J74467</accessionNumber>
</th>
<th id="768F5DA9FFD314C775B8FB93FDDDFBCD" box="[485,606,1074,1096]" gridcol="2" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317649FB93FDDDFBCD" box="[532,606,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J46364" pageId="9" pageNumber="162" type="EnaNcbi">J46364</accessionNumber>
</th>
<th id="768F5DA9FFD314C776F8FB93FC9DFBCD" box="[677,798,1074,1096]" gridcol="3" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317688FB93FC9DFBCD" box="[725,798,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74478" pageId="9" pageNumber="162" type="EnaNcbi">J74478</accessionNumber>
</th>
<th id="768F5DA9FFD314C77738FB93FC5DFBCD" box="[869,990,1074,1096]" gridcol="4" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB3177C8FB93FC5DFBCD" box="[917,990,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74466" pageId="9" pageNumber="162" type="EnaNcbi">J74466</accessionNumber>
</th>
<th id="768F5DA9FFD314C77078FB93FB1DFBCD" box="[1061,1182,1074,1096]" gridcol="5" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317008FB93FB1DFBCD" box="[1109,1182,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74475" pageId="9" pageNumber="162" type="EnaNcbi">J74475</accessionNumber>
</th>
<th id="768F5DA9FFD314C770BBFB93FADDFBCD" box="[1254,1374,1074,1096]" gridcol="6" gridrow="0" pageId="9" pageNumber="162">
QM
<accessionNumber id="943DAB74FFD3EB317148FB93FADDFBCD" box="[1301,1374,1074,1096]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74468" pageId="9" pageNumber="162" type="EnaNcbi">J74468</accessionNumber>
</th>
</tr>
<tr id="355E34D5FFD314C774C2FBFDFADDFBF7" box="[159,1374,1116,1138]" gridrow="1" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FBFDFF50FBF7" box="[159,211,1116,1138]" gridcol="0" gridrow="1" pageId="9" pageNumber="162">SVL</th>
<td id="768F5DA9FFD314C77578FBFDFE1DFBF7" box="[293,414,1116,1138]" gridcol="1" gridrow="1" pageId="9" pageNumber="162">35.1</td>
<td id="768F5DA9FFD314C775B8FBFDFDDDFBF7" box="[485,606,1116,1138]" gridcol="2" gridrow="1" pageId="9" pageNumber="162">34.5</td>
<td id="768F5DA9FFD314C776F8FBFDFC9DFBF7" box="[677,798,1116,1138]" gridcol="3" gridrow="1" pageId="9" pageNumber="162">37.0</td>
<td id="768F5DA9FFD314C77738FBFDFC5DFBF7" box="[869,990,1116,1138]" gridcol="4" gridrow="1" pageId="9" pageNumber="162">35.6</td>
<td id="768F5DA9FFD314C77078FBFDFB1DFBF7" box="[1061,1182,1116,1138]" gridcol="5" gridrow="1" pageId="9" pageNumber="162">38.3</td>
<td id="768F5DA9FFD314C770BBFBFDFADDFBF7" box="[1254,1374,1116,1138]" gridcol="6" gridrow="1" pageId="9" pageNumber="162">37.2</td>
</tr>
<tr id="355E34D5FFD314C774C2FB27FADDFB19" box="[159,1374,1158,1180]" gridrow="2" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FB27FF50FB19" box="[159,211,1158,1180]" gridcol="0" gridrow="2" pageId="9" pageNumber="162">TIB</th>
<td id="768F5DA9FFD314C77578FB27FE1DFB19" box="[293,414,1158,1180]" gridcol="1" gridrow="2" pageId="9" pageNumber="162">19.1</td>
<td id="768F5DA9FFD314C775B8FB27FDDDFB19" box="[485,606,1158,1180]" gridcol="2" gridrow="2" pageId="9" pageNumber="162">18.3</td>
<td id="768F5DA9FFD314C776F8FB27FC9DFB19" box="[677,798,1158,1180]" gridcol="3" gridrow="2" pageId="9" pageNumber="162">21.6</td>
<td id="768F5DA9FFD314C77738FB27FC5DFB19" box="[869,990,1158,1180]" gridcol="4" gridrow="2" pageId="9" pageNumber="162">21.0</td>
<td id="768F5DA9FFD314C77078FB27FB1DFB19" box="[1061,1182,1158,1180]" gridcol="5" gridrow="2" pageId="9" pageNumber="162">22.0</td>
<td id="768F5DA9FFD314C770BBFB27FADDFB19" box="[1254,1374,1158,1180]" gridcol="6" gridrow="2" pageId="9" pageNumber="162">21.7</td>
</tr>
<tr id="355E34D5FFD314C774C2FB11FADDFB43" box="[159,1374,1200,1222]" gridrow="3" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FB11FF50FB43" box="[159,211,1200,1222]" gridcol="0" gridrow="3" pageId="9" pageNumber="162">HL</th>
<td id="768F5DA9FFD314C77578FB11FE1DFB43" box="[293,414,1200,1222]" gridcol="1" gridrow="3" pageId="9" pageNumber="162">11.8</td>
<td id="768F5DA9FFD314C775B8FB11FDDDFB43" box="[485,606,1200,1222]" gridcol="2" gridrow="3" pageId="9" pageNumber="162">12.5</td>
<td id="768F5DA9FFD314C776F8FB11FC9DFB43" box="[677,798,1200,1222]" gridcol="3" gridrow="3" pageId="9" pageNumber="162">13.2</td>
<td id="768F5DA9FFD314C77738FB11FC5DFB43" box="[869,990,1200,1222]" gridcol="4" gridrow="3" pageId="9" pageNumber="162">12.5</td>
<td id="768F5DA9FFD314C77078FB11FB1DFB43" box="[1061,1182,1200,1222]" gridcol="5" gridrow="3" pageId="9" pageNumber="162">13.2</td>
<td id="768F5DA9FFD314C770BBFB11FADDFB43" box="[1254,1374,1200,1222]" gridcol="6" gridrow="3" pageId="9" pageNumber="162">13.2</td>
</tr>
<tr id="355E34D5FFD314C774C2FB7BFADDFB75" box="[159,1374,1242,1264]" gridrow="4" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FB7BFF50FB75" box="[159,211,1242,1264]" gridcol="0" gridrow="4" pageId="9" pageNumber="162">HW</th>
<td id="768F5DA9FFD314C77578FB7BFE1DFB75" box="[293,414,1242,1264]" gridcol="1" gridrow="4" pageId="9" pageNumber="162">12.2</td>
<td id="768F5DA9FFD314C775B8FB7BFDDDFB75" box="[485,606,1242,1264]" gridcol="2" gridrow="4" pageId="9" pageNumber="162">12.8</td>
<td id="768F5DA9FFD314C776F8FB7BFC9DFB75" box="[677,798,1242,1264]" gridcol="3" gridrow="4" pageId="9" pageNumber="162">14.0</td>
<td id="768F5DA9FFD314C77738FB7BFC5DFB75" box="[869,990,1242,1264]" gridcol="4" gridrow="4" pageId="9" pageNumber="162">13.4</td>
<td id="768F5DA9FFD314C77078FB7BFB1DFB75" box="[1061,1182,1242,1264]" gridcol="5" gridrow="4" pageId="9" pageNumber="162">13.7</td>
<td id="768F5DA9FFD314C770BBFB7BFADDFB75" box="[1254,1374,1242,1264]" gridcol="6" gridrow="4" pageId="9" pageNumber="162">13.5</td>
</tr>
<tr id="355E34D5FFD314C774C2FAA5FADDFA9F" box="[159,1374,1284,1306]" gridrow="5" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FAA5FF50FA9F" box="[159,211,1284,1306]" gridcol="0" gridrow="5" pageId="9" pageNumber="162">EYE</th>
<td id="768F5DA9FFD314C77578FAA5FE1DFA9F" box="[293,414,1284,1306]" gridcol="1" gridrow="5" pageId="9" pageNumber="162">4.0</td>
<td id="768F5DA9FFD314C775B8FAA5FDDDFA9F" box="[485,606,1284,1306]" gridcol="2" gridrow="5" pageId="9" pageNumber="162">3.8</td>
<td id="768F5DA9FFD314C776F8FAA5FC9DFA9F" box="[677,798,1284,1306]" gridcol="3" gridrow="5" pageId="9" pageNumber="162">4.2</td>
<td id="768F5DA9FFD314C77738FAA5FC5DFA9F" box="[869,990,1284,1306]" gridcol="4" gridrow="5" pageId="9" pageNumber="162">4.0</td>
<td id="768F5DA9FFD314C77078FAA5FB1DFA9F" box="[1061,1182,1284,1306]" gridcol="5" gridrow="5" pageId="9" pageNumber="162">4.5</td>
<td id="768F5DA9FFD314C770BBFAA5FADDFA9F" box="[1254,1374,1284,1306]" gridcol="6" gridrow="5" pageId="9" pageNumber="162">4.2</td>
</tr>
<tr id="355E34D5FFD314C774C2FA8FFADDFAC1" box="[159,1374,1326,1348]" gridrow="6" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FA8FFF50FAC1" box="[159,211,1326,1348]" gridcol="0" gridrow="6" pageId="9" pageNumber="162">TMP</th>
<td id="768F5DA9FFD314C77578FA8FFE1DFAC1" box="[293,414,1326,1348]" gridcol="1" gridrow="6" pageId="9" pageNumber="162">2.4</td>
<td id="768F5DA9FFD314C775B8FA8FFDDDFAC1" box="[485,606,1326,1348]" gridcol="2" gridrow="6" pageId="9" pageNumber="162">2.6</td>
<td id="768F5DA9FFD314C776F8FA8FFC9DFAC1" box="[677,798,1326,1348]" gridcol="3" gridrow="6" pageId="9" pageNumber="162">2.3</td>
<td id="768F5DA9FFD314C77738FA8FFC5DFAC1" box="[869,990,1326,1348]" gridcol="4" gridrow="6" pageId="9" pageNumber="162">2.5</td>
<td id="768F5DA9FFD314C77078FA8FFB1DFAC1" box="[1061,1182,1326,1348]" gridcol="5" gridrow="6" pageId="9" pageNumber="162">2.4</td>
<td id="768F5DA9FFD314C770BBFA8FFADDFAC1" box="[1254,1374,1326,1348]" gridcol="6" gridrow="6" pageId="9" pageNumber="162">2.4</td>
</tr>
<tr id="355E34D5FFD314C774C2FAF9FADDFAEB" box="[159,1374,1368,1390]" gridrow="7" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FAF9FF50FAEB" box="[159,211,1368,1390]" gridcol="0" gridrow="7" pageId="9" pageNumber="162">END</th>
<td id="768F5DA9FFD314C77578FAF9FE1DFAEB" box="[293,414,1368,1390]" gridcol="1" gridrow="7" pageId="9" pageNumber="162">3.1</td>
<td id="768F5DA9FFD314C775B8FAF9FDDDFAEB" box="[485,606,1368,1390]" gridcol="2" gridrow="7" pageId="9" pageNumber="162">3.7</td>
<td id="768F5DA9FFD314C776F8FAF9FC9DFAEB" box="[677,798,1368,1390]" gridcol="3" gridrow="7" pageId="9" pageNumber="162">3.4</td>
<td id="768F5DA9FFD314C77738FAF9FC5DFAEB" box="[869,990,1368,1390]" gridcol="4" gridrow="7" pageId="9" pageNumber="162">3.5</td>
<td id="768F5DA9FFD314C77078FAF9FB1DFAEB" box="[1061,1182,1368,1390]" gridcol="5" gridrow="7" pageId="9" pageNumber="162">3.7</td>
<td id="768F5DA9FFD314C770BBFAF9FADDFAEB" box="[1254,1374,1368,1390]" gridcol="6" gridrow="7" pageId="9" pageNumber="162">3.4</td>
</tr>
<tr id="355E34D5FFD314C774C2FA23FADDFA1D" box="[159,1374,1410,1432]" gridrow="8" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FA23FF50FA1D" box="[159,211,1410,1432]" gridcol="0" gridrow="8" pageId="9" pageNumber="162">IND</th>
<td id="768F5DA9FFD314C77578FA23FE1DFA1D" box="[293,414,1410,1432]" gridcol="1" gridrow="8" pageId="9" pageNumber="162">2.9</td>
<td id="768F5DA9FFD314C775B8FA23FDDDFA1D" box="[485,606,1410,1432]" gridcol="2" gridrow="8" pageId="9" pageNumber="162">2.6</td>
<td id="768F5DA9FFD314C776F8FA23FC9DFA1D" box="[677,798,1410,1432]" gridcol="3" gridrow="8" pageId="9" pageNumber="162">2.8</td>
<td id="768F5DA9FFD314C77738FA23FC5DFA1D" box="[869,990,1410,1432]" gridcol="4" gridrow="8" pageId="9" pageNumber="162">2.7</td>
<td id="768F5DA9FFD314C77078FA23FB1DFA1D" box="[1061,1182,1410,1432]" gridcol="5" gridrow="8" pageId="9" pageNumber="162">3.6</td>
<td id="768F5DA9FFD314C770BBFA23FADDFA1D" box="[1254,1374,1410,1432]" gridcol="6" gridrow="8" pageId="9" pageNumber="162">2.9</td>
</tr>
<tr id="355E34D5FFD314C774C2FA0DFADDFA47" box="[159,1374,1452,1474]" gridrow="9" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FA0DFF50FA47" box="[159,211,1452,1474]" gridcol="0" gridrow="9" pageId="9" pageNumber="162">TEY</th>
<td id="768F5DA9FFD314C77578FA0DFE1DFA47" box="[293,414,1452,1474]" gridcol="1" gridrow="9" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C775B8FA0DFDDDFA47" box="[485,606,1452,1474]" gridcol="2" gridrow="9" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C776F8FA0DFC9DFA47" box="[677,798,1452,1474]" gridcol="3" gridrow="9" pageId="9" pageNumber="162">1.3</td>
<td id="768F5DA9FFD314C77738FA0DFC5DFA47" box="[869,990,1452,1474]" gridcol="4" gridrow="9" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C77078FA0DFB1DFA47" box="[1061,1182,1452,1474]" gridcol="5" gridrow="9" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C770BBFA0DFADDFA47" box="[1254,1374,1452,1474]" gridcol="6" gridrow="9" pageId="9" pageNumber="162">1.4</td>
</tr>
<tr id="355E34D5FFD314C774C2FA77FADDFA69" box="[159,1374,1494,1516]" gridrow="10" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2FA77FF50FA69" box="[159,211,1494,1516]" gridcol="0" gridrow="10" pageId="9" pageNumber="162">3FD</th>
<td id="768F5DA9FFD314C77578FA77FE1DFA69" box="[293,414,1494,1516]" gridcol="1" gridrow="10" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C775B8FA77FDDDFA69" box="[485,606,1494,1516]" gridcol="2" gridrow="10" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C776F8FA77FC9DFA69" box="[677,798,1494,1516]" gridcol="3" gridrow="10" pageId="9" pageNumber="162">2.0</td>
<td id="768F5DA9FFD314C77738FA77FC5DFA69" box="[869,990,1494,1516]" gridcol="4" gridrow="10" pageId="9" pageNumber="162">1.8</td>
<td id="768F5DA9FFD314C77078FA77FB1DFA69" box="[1061,1182,1494,1516]" gridcol="5" gridrow="10" pageId="9" pageNumber="162">2.1</td>
<td id="768F5DA9FFD314C770BBFA77FADDFA69" box="[1254,1374,1494,1516]" gridcol="6" gridrow="10" pageId="9" pageNumber="162">1.9</td>
</tr>
<tr id="355E34D5FFD314C774C2F9A1FADDF993" box="[159,1374,1536,1558]" gridrow="11" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2F9A1FF50F993" box="[159,211,1536,1558]" gridcol="0" gridrow="11" pageId="9" pageNumber="162">3F</th>
<td id="768F5DA9FFD314C77578F9A1FE1DF993" box="[293,414,1536,1558]" gridcol="1" gridrow="11" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C775B8F9A1FDDDF993" box="[485,606,1536,1558]" gridcol="2" gridrow="11" pageId="9" pageNumber="162">1.0</td>
<td id="768F5DA9FFD314C776F8F9A1FC9DF993" box="[677,798,1536,1558]" gridcol="3" gridrow="11" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C77738F9A1FC5DF993" box="[869,990,1536,1558]" gridcol="4" gridrow="11" pageId="9" pageNumber="162">1.2</td>
<td id="768F5DA9FFD314C77078F9A1FB1DF993" box="[1061,1182,1536,1558]" gridcol="5" gridrow="11" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C770BBF9A1FADDF993" box="[1254,1374,1536,1558]" gridcol="6" gridrow="11" pageId="9" pageNumber="162">1.2</td>
</tr>
<tr id="355E34D5FFD314C774C2F98BFADDF9C5" box="[159,1374,1578,1600]" gridrow="12" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2F98BFF50F9C5" box="[159,211,1578,1600]" gridcol="0" gridrow="12" pageId="9" pageNumber="162">4TD</th>
<td id="768F5DA9FFD314C77578F98BFE1DF9C5" box="[293,414,1578,1600]" gridcol="1" gridrow="12" pageId="9" pageNumber="162">1.6</td>
<td id="768F5DA9FFD314C775B8F98BFDDDF9C5" box="[485,606,1578,1600]" gridcol="2" gridrow="12" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C776F8F98BFC9DF9C5" box="[677,798,1578,1600]" gridcol="3" gridrow="12" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C77738F98BFC5DF9C5" box="[869,990,1578,1600]" gridcol="4" gridrow="12" pageId="9" pageNumber="162">1.7</td>
<td id="768F5DA9FFD314C77078F98BFB1DF9C5" box="[1061,1182,1578,1600]" gridcol="5" gridrow="12" pageId="9" pageNumber="162">1.9</td>
<td id="768F5DA9FFD314C770BBF98BFADDF9C5" box="[1254,1374,1578,1600]" gridcol="6" gridrow="12" pageId="9" pageNumber="162">1.6</td>
</tr>
<tr id="355E34D5FFD314C774C2F9F5FADDF9EF" box="[159,1374,1620,1642]" gridrow="13" pageId="9" pageNumber="162">
<th id="768F5DA9FFD314C774C2F9F5FF50F9EF" box="[159,211,1620,1642]" gridcol="0" gridrow="13" pageId="9" pageNumber="162">4F</th>
<td id="768F5DA9FFD314C77578F9F5FE1DF9EF" box="[293,414,1620,1642]" gridcol="1" gridrow="13" pageId="9" pageNumber="162">1.1</td>
<td id="768F5DA9FFD314C775B8F9F5FDDDF9EF" box="[485,606,1620,1642]" gridcol="2" gridrow="13" pageId="9" pageNumber="162">1.0</td>
<td id="768F5DA9FFD314C776F8F9F5FC9DF9EF" box="[677,798,1620,1642]" gridcol="3" gridrow="13" pageId="9" pageNumber="162">1.4</td>
<td id="768F5DA9FFD314C77738F9F5FC5DF9EF" box="[869,990,1620,1642]" gridcol="4" gridrow="13" pageId="9" pageNumber="162">1.4</td>
<td id="768F5DA9FFD314C77078F9F5FB1DF9EF" box="[1061,1182,1620,1642]" gridcol="5" gridrow="13" pageId="9" pageNumber="162">1.5</td>
<td id="768F5DA9FFD314C770BBF9F5FADDF9EF" box="[1254,1374,1620,1642]" gridcol="6" gridrow="13" pageId="9" pageNumber="162">1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD13697FFD3EB31749AF90EFC44F841" blockId="9.[151,1437,1711,1988]" pageId="9" pageNumber="162">
<emphasis id="B91AEA85FFD3EB31749AF90EFE30F94D" bold="true" box="[199,435,1711,1736]" pageId="9" pageNumber="162">Advertisement call.</emphasis>
Call descriptions are based on the calls of two topoparatypes (QM
<accessionNumber id="943DAB74FFD3EB317083F90EFAACF94D" box="[1246,1327,1711,1736]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74467" pageId="9" pageNumber="162" type="EnaNcbi">J74467</accessionNumber>
and QM
<accessionNumber id="943DAB74FFD3EB3174CAF972FF6FF969" box="[151,236,1747,1772]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74468" pageId="9" pageNumber="162" type="EnaNcbi">J74468</accessionNumber>
), recorded at an air temperature of 24.1ºC within
<quantity id="4C969B72FFD3EB31774AF975FCD4F96E" box="[791,855,1748,1772]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="162" unit="m" value="0.3">0.3 m</quantity>
of calling sites (
<tableCitation id="C6EC032CFFD3EB31704DF972FBE4F969" box="[1040,1127,1747,1772]" captionStart="TABLE 5" captionStartId="11.[151,239,404,427]" captionText="TABLE 5. Call parameters for two individuals each of Litoria gracilenta and Litoria bella sp. nov." pageId="9" pageNumber="162">Table 5</tableCitation>
;
<figureCitation id="13552A12FFD3EB317029F972FB34F969" box="[1140,1207,1747,1772]" captionStart="FIGURE 5" captionStartId="10.[151,250,1407,1429]" captionTargetBox="[152,1423,199,1381]" captionTargetId="figure@10.[152,1435,195,1385]" captionTargetPageId="10" captionText="FIGURE 5. Advertisement call of (A) Litoria bella sp. nov. recorded at 24.1 ºC, (i) waveform of relative amplitude over time of a single call of QM J 74468, (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM J 74468, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM J 74467. (B) Litoria gracilenta recorded at 21.7 21.8 ºC, (i) waveform of relative amplitude over time of a single call of AMS R 184768 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184768, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184777. (C) Litoria chloris recorded at 21.8 ºC, (i) waveform of relative amplitude over time of a single call of AMS R 184770 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184770." httpUri="https://zenodo.org/record/268265/files/figure.png" pageId="9" pageNumber="162">Fig. 5</figureCitation>
). The advertisement call of
<taxonomicName id="4C6E4D14FFD3EB3174B5F959FEF5F88A" box="[232,374,1783,1807]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="9" pageNumber="162" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD3EB3174B5F959FEF5F88A" box="[232,374,1783,1807]" italics="true" pageId="9" pageNumber="162">Litoria bella</emphasis>
</taxonomicName>
consists of a single note, although three of the eight recorded calls of QMJ 74467 were only separated by a short interval. The duration of each call varied from 1.42
<geoCoordinate id="EE5A5050FFD3EB3177B4F8BAFBACF8B6" box="[1001,1071,1819,1844]" direction="south" orientation="latitude" pageId="9" pageNumber="162" precision="555" value="-2.62">2.62 s</geoCoordinate>
and calls were separated by an interval of 0.88
<geoCoordinate id="EE5A5050FFD3EB31750BF89EFE1DF8D2" box="[342,414,1855,1880]" direction="south" orientation="latitude" pageId="9" pageNumber="162" precision="555" value="-5.78">5.78 s</geoCoordinate>
. Each call had 89156 uniformly spaced pulses repeated at a rate of around 60 pulses/s. Calls were amplitude modulated, increasing gradually to a peak at approximately 70% of the call duration (except for the three calls separated by a short interval). The dominant frequency was 2.62.8 Hz, but frequency increased over the duration of the call about 0.2 kHz, from 2.52.6 to 2.72.8 kHz.
</paragraph>
<caption id="DF11661FFFD0EB3274CAFADEFBAFF90F" httpUri="https://zenodo.org/record/268265/files/figure.png" pageId="10" pageNumber="163" targetBox="[152,1423,199,1381]" targetPageId="10">
<paragraph id="8BD13697FFD0EB3274CAFADEFBAFF90F" blockId="10.[151,1436,1407,1674]" pageId="10" pageNumber="163">
<emphasis id="B91AEA85FFD0EB3274CAFADEFE90FA11" bold="true" box="[151,275,1407,1429]" pageId="10" pageNumber="163">FIGURE 5.</emphasis>
Advertisement call of (A)
<taxonomicName id="4C6E4D14FFD0EB327675FA21FD2BFA10" box="[552,680,1407,1429]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFD0EB327675FA21FD2BFA10" box="[552,680,1407,1429]" italics="true" pageId="10" pageNumber="163">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFD0EB3276F2FADEFC83FA11" box="[687,768,1407,1428]" pageId="10" pageNumber="163" rank="species">
<emphasis id="B91AEA85FFD0EB3276F2FADEFC83FA11" bold="true" box="[687,768,1407,1428]" pageId="10" pageNumber="163">sp. nov.</emphasis>
</taxonomicNameLabel>
recorded at 24.1ºC, (i) waveform of relative amplitude over time of a single call of QM
<accessionNumber id="943DAB74FFD0EB3275DFFA3CFE4DFA36" box="[386,462,1437,1459]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74468" pageId="10" pageNumber="163" type="EnaNcbi">J74468</accessionNumber>
, (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM
<accessionNumber id="943DAB74FFD0EB3275BFFA1DFDADFA57" box="[482,558,1468,1490]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74468" pageId="10" pageNumber="163" type="EnaNcbi">J74468</accessionNumber>
, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM
<accessionNumber id="943DAB74FFD0EB32762CFA7AFD3DFA74" box="[625,702,1499,1521]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74467" pageId="10" pageNumber="163" type="EnaNcbi">J74467</accessionNumber>
. (B)
<taxonomicName id="4C6E4D14FFD0EB3276B2FA7DFC20FA74" box="[751,931,1499,1521]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD0EB3276B2FA7DFC20FA74" box="[751,931,1499,1521]" italics="true" pageId="10" pageNumber="163">Litoria gracilenta</emphasis>
</taxonomicName>
recorded at 21.721.8ºC, (i) waveform of relative amplitude over time of a single call of AMS R184768 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R184768, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R184777. (C)
<taxonomicName id="4C6E4D14FFD0EB327061F999FB4CF9C8" box="[1084,1231,1591,1613]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD0EB327061F999FB4CF9C8" box="[1084,1231,1591,1613]" italics="true" pageId="10" pageNumber="163">Litoria chloris</emphasis>
</taxonomicName>
recorded at 21.8ºC, (i) waveform of relative amplitude over time of a single call of AMS R184770 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R184770.
</paragraph>
</caption>
<paragraph id="8BD13697FFD0EB32749AF917FAACF8DA" blockId="10.[151,1437,1718,2032]" pageId="10" pageNumber="163">
<emphasis id="B91AEA85FFD0EB32749AF917FEA8F94A" bold="true" box="[199,299,1718,1743]" pageId="10" pageNumber="163">Ecology.</emphasis>
<taxonomicName id="4C6E4D14FFD0EB32756EF919FE42F94A" box="[307,449,1719,1743]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD0EB32756EF919FE42F94A" box="[307,449,1719,1743]" italics="true" pageId="10" pageNumber="163">Litoria bella</emphasis>
</taxonomicName>
inhabits moist monsoon rainforest and vine thicket on hills and plains. The species has been observed breeding in ephemeral pools adjacent to permanent streams and observed (but not breeding) around ephemeral pools in gullies flowing into permanent streams. In
<date id="FFD01057FFD0EB3277DBF95FFBA4F892" box="[902,1063,1790,1815]" pageId="10" pageNumber="163" value="2000-10">October 2000</date>
, breeding was observed after a thunderstorm. Two pairs of frogs in inguinal amplexus (female QM
<accessionNumber id="943DAB74FFD0EB3277C4F882FC6AF8B9" box="[921,1001,1827,1852]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74465" pageId="10" pageNumber="163" type="EnaNcbi">J74465</accessionNumber>
and male QM J4466, and QM
<accessionNumber id="943DAB74FFD0EB327116F882FA1EF8B9" box="[1355,1437,1827,1852]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74469" pageId="10" pageNumber="163" type="EnaNcbi">J74469</accessionNumber>
and QM
<accessionNumber id="943DAB74FFD0EB3274A7F8E6FECEF8DA" box="[250,333,1863,1887]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74470" pageId="10" pageNumber="163" type="EnaNcbi">J74470</accessionNumber>
) laid spawn with 606 and 844 dorsally brown-pigmented eggs in clear jelly respectively.
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFD0EB33749AF8CAFAA9FB92" lastPageId="11" lastPageNumber="164" pageId="10" pageNumber="163" type="distribution">
<paragraph id="8BD13697FFD0EB33749AF8CAFDECFEE1" blockId="10.[151,1437,1718,2032]" lastBlockId="11.[151,1436,151,357]" lastPageId="11" lastPageNumber="164" pageId="10" pageNumber="163">
<emphasis id="B91AEA85FFD0EB32749AF8CAFD0FF801" bold="true" box="[199,652,1899,1924]" pageId="10" pageNumber="163">Distribution and conservation status.</emphasis>
<taxonomicName id="4C6E4D14FFD0EB3276CAF8CDFCABF806" box="[663,808,1899,1923]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD0EB3276CAF8CDFCABF806" box="[663,808,1899,1923]" italics="true" pageId="10" pageNumber="163">Litoria bella</emphasis>
</taxonomicName>
is known from 5560 metres elevation between Moa
<collectingCountry id="F3797607FFD0EB3274CAF82FFF58F822" box="[151,219,1934,1959]" name="Iceland" pageId="10" pageNumber="163">Island</collectingCountry>
in the north and about
<quantity id="4C969B72FFD0EB3275A6F82EFDC5F822" box="[507,582,1934,1959]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="163" unit="km" value="20.0">20 km</quantity>
south of Coen in the south (
<figureCitation id="13552A12FFD0EB3277FEF82FFC6AF822" box="[931,1001,1934,1959]" captionStart="FIGURE 1" captionStartId="1.[151,250,1642,1664]" captionTargetBox="[154,1432,520,1615]" captionTargetId="figure@1.[151,1436,518,1617]" captionTargetPageId="1" captionText="FIGURE 1. Localities of specimens and tissues of Litoria auae (pale yellow), Litoria bella sp. nov. (dark yellow), Litoria gracilenta (dark orange), and L. chloris (dark red) used in this study. DNA only = circle, specimen only = diamond, DNA and specimen = square. The type localities of the first three species are indicated by stars." httpUri="https://zenodo.org/record/268261/files/figure.png" pageId="10" pageNumber="163">Fig. 1</figureCitation>
). The species appears to be patchily distributed within the Cape
<collectingRegion id="49AAF875FFD0EB3275B9F812FD98F849" box="[484,539,1971,1996]" country="United Kingdom" name="York" pageId="10" pageNumber="163">York</collectingRegion>
Peninsula Bioregion (
<bibRefCitation id="EFFF4B66FFD0EB327779F812FBCFF84E" author="Sattler" box="[804,1100,1971,1996]" pageId="10" pageNumber="163" refString="Sattler, P. S. &amp; Williams, R. D. (1999) The Conservation Status of Queensland Bioregional Ecosystems. Environmental Protection Agency, Brisbane." type="book" year="1999">Sattler &amp; Williams 1999</bibRefCitation>
), while
<taxonomicName id="4C6E4D14FFD0EB3270F2F815FAFEF84E" box="[1199,1405,1971,1995]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="10" pageNumber="163" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD0EB3270F2F815FAFEF84E" box="[1199,1405,1971,1995]" italics="true" pageId="10" pageNumber="163">Litoria gracilenta</emphasis>
</taxonomicName>
is restricted to the Wet Tropics, Brigalow Belt,
<collectingRegion id="49AAF875FFD0EB3276D0F876FD5CF875" box="[653,735,2007,2032]" country="Solomon Islands" name="Central" pageId="10" pageNumber="163">Central</collectingRegion>
<collectingRegion id="49AAF875FFD0EB3276BBF876FCE9F875" box="[742,874,2007,2032]" country="Australia" name="Queensland" pageId="10" pageNumber="163">Queensland</collectingRegion>
Coast and South East
<collectingRegion id="49AAF875FFD0EB32703AF876FB68F875" box="[1127,1259,2007,2032]" country="Australia" name="Queensland" pageId="10" pageNumber="163">Queensland</collectingRegion>
Bioregions and the
<collectingRegion id="49AAF875FFD1EB3374E2FF39FE90FF2A" box="[191,275,152,175]" country="American Samoa" name="Eastern" pageId="11" pageNumber="164">Eastern</collectingRegion>
Australian Temperate Forests (
<bibRefCitation id="EFFF4B66FFD1EB337632FF36FC07FF35" author="Sattler" box="[623,900,151,176]" pageId="11" pageNumber="164" refString="Sattler, P. S. &amp; Williams, R. D. (1999) The Conservation Status of Queensland Bioregional Ecosystems. Environmental Protection Agency, Brisbane." type="book" year="1999">Sattler &amp; Williams 1999</bibRefCitation>
). The southern-most location of the new species is separated by
<quantity id="4C969B72FFD1EB33751EFF1DFE14FF50" box="[323,407,188,213]" metricMagnitude="5" metricUnit="m" metricValue="2.2" pageId="11" pageNumber="164" unit="km" value="220.0">220 km</quantity>
from the most northern record of
<taxonomicName id="4C6E4D14FFD1EB33774CFF1CFC5BFF51" box="[785,984,188,212]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB33774CFF1CFC5BFF51" box="[785,984,188,212]" italics="true" pageId="11" pageNumber="164">Litoria gracilenta</emphasis>
</taxonomicName>
in the Endeavour Valley.
<taxonomicName id="4C6E4D14FFD1EB3370ABFF1CFA00FF51" box="[1270,1411,188,212]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD1EB3370ABFF1CFA00FF51" box="[1270,1411,188,212]" italics="true" pageId="11" pageNumber="164">Litoria bella</emphasis>
</taxonomicName>
is known to occur within at least three protected areas, the Kulla National Park (CYPAL), Kutini-Payamu (Iron Range) National Park (CYPAL) and the Oyala Thumotang National Park (CYPAL). Given its relatively broad distribution, no known threats, and its occurrence within at least three protected areas the species likely falls under the IUCN classification of Least Concern.
</paragraph>
<caption id="DF11661FFFD1EB3374CAFE35FB3AFE2E" box="[151,1209,403,427]" pageId="11" pageNumber="164">
<paragraph id="8BD13697FFD1EB3374CAFE35FB3AFE2E" blockId="11.[151,1209,403,427]" box="[151,1209,403,427]" pageId="11" pageNumber="164">
<emphasis id="B91AEA85FFD1EB3374CAFE35FE89FE2E" bold="true" box="[151,266,404,427]" pageId="11" pageNumber="164">TABLE 5.</emphasis>
Call parameters for two individuals each of
<taxonomicName id="4C6E4D14FFD1EB3376B8FE35FC27FE2E" box="[741,932,403,427]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3376B8FE35FC27FE2E" box="[741,932,403,427]" italics="true" pageId="11" pageNumber="164">Litoria gracilenta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D14FFD1EB33778AFE35FBDEFE2E" box="[983,1117,403,427]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFD1EB33778AFE35FBDEFE2E" box="[983,1117,403,427]" italics="true" pageId="11" pageNumber="164">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFD1EB337039FE35FB3AFE2E" box="[1124,1209,404,427]" pageId="11" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD13697FFD1EB3375BDFE69FB26FE59" blockId="11.[159,1395,454,810]" box="[480,1189,454,477]" pageId="11" pageNumber="164">
<taxonomicName id="4C6E4D14FFD1EB3375BDFE69FDDCFE59" box="[480,607,454,477]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="bella" status="sp. nov.">
<emphasis id="B91AEA85FFD1EB3375BDFE69FDDCFE59" box="[480,607,454,477]" italics="true" pageId="11" pageNumber="164">Litoria bella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22957FEFFD1EB337638FE66FD35FE59" box="[613,694,455,476]" pageId="11" pageNumber="164" rank="species">
<emphasis id="B91AEA85FFD1EB337638FE66FD35FE59" bold="true" box="[613,694,455,476]" pageId="11" pageNumber="164">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C6E4D14FFD1EB3377ADFE69FB26FE59" box="[1008,1189,454,477]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3377ADFE69FB26FE59" box="[1008,1189,454,477]" italics="true" pageId="11" pageNumber="164">Litoria gracilenta</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BD13697FFD1EB3375BDFE51FAE6FD83" blockId="11.[159,1395,454,810]" box="[480,1381,496,518]" pageId="11" pageNumber="164">
QM
<accessionNumber id="943DAB74FFD1EB337652FE51FDDBFD83" box="[527,600,496,518]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74467" pageId="11" pageNumber="164" type="EnaNcbi">J74467</accessionNumber>
QM
<accessionNumber id="943DAB74FFD1EB337766FE51FC07FD83" box="[827,900,496,518]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J74468" pageId="11" pageNumber="164" type="EnaNcbi">J74468</accessionNumber>
AMS R184768 AMS R184777
</paragraph>
<paragraph id="8BD13697FFD1EB3374C2FDB8FA8FFDAA" blockId="11.[159,1395,454,810]" box="[159,1292,537,559]" pageId="11" pageNumber="164">Measured 8 calls (3 double=5 sets) 6 calls (all single) 6 calls 6 calls</paragraph>
<paragraph id="8BD13697FFD1EB3374C2FDE3FB74FCAF" blockId="11.[159,1395,454,810]" pageId="11" pageNumber="164">Call length (s) 1.94 (1.422.62) 2.34 (2.292.43) 2.06 (2.012.09) 2.20 (1.312.64) Intercall interval (s) 2.85 (0.884.54) 5.78 (5.055.78) 3.38 (2.624.49) 5.39 (4.187.33) Pulses 116.5(89156) 140.8 (135149) 180.2 (176191) 183 (116225) Pulse/s 60.6(55.864.0) 59.9(57.763.6) 85.7(85.793.0) 84.2(74.788.8) Dominant Frequency (Hz) 2.7 (2.72.8) 2.6 (2.62.7) 2.4 2.7 (2.62.8) Temperature (ºC) 24.1 24.1 21.8 21.7</paragraph>
<paragraph id="8BD13697FFD1EB33749AFCCEFAA9FB92" blockId="11.[151,1437,879,2020]" pageId="11" pageNumber="164">
<emphasis id="B91AEA85FFD1EB33749AFCCEFEEFFC0D" bold="true" box="[199,364,879,904]" pageId="11" pageNumber="164">Comparisons.</emphasis>
From the eight species of the
<taxonomicName id="4C6E4D14FFD1EB3376E6FCD1FC01FC02" box="[699,898,879,903]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3376E6FCD1FC01FC02" box="[699,898,879,903]" italics="true" pageId="11" pageNumber="164">Litoria gracilenta</emphasis>
</taxonomicName>
group currently known, the new species may be distinguished by the combination of (1) male body size moderately large (
<quantity id="4C969B72FFD1EB33707BFC32FB51FC2E" box="[1062,1234,915,940]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="11" pageNumber="164" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL), (2) nearimmaculate green dorsum, (3) orange venter, (4) bright orange digits and webbing, (5), iridescent bluish purple lateral surfaces of thigh, (6) no pale canthal stripe, (7) white bones, and (8) a highly-pulsed, single-note, male advertisement call with a pulse rate of 5664 pulses/s and dominant frequency of 2.62.8 kHz at 21.4ºC.
</paragraph>
</subSubSection>
<subSubSection id="C374651CFFD1EB35749AFB85FD9BFF51" lastPageId="13" lastPageNumber="166" pageId="11" pageNumber="164" type="description">
<paragraph id="8BD13697FFD1EB34749AFB85FA87FE0D" blockId="11.[151,1437,879,2020]" lastBlockId="12.[151,1437,151,968]" lastPageId="12" lastPageNumber="165" pageId="11" pageNumber="164">
From
<taxonomicName id="4C6E4D14FFD1EB337553FB85FE1FFBBE" box="[270,412,1060,1083]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auruensis">
<emphasis id="B91AEA85FFD1EB337553FB85FE1FFBBE" box="[270,412,1060,1083]" italics="true" pageId="11" pageNumber="164">L. auruensis</emphasis>
</taxonomicName>
, the new species further differs by lacking broad lateral fringes on fingers and toes (vs present in
<taxonomicName id="4C6E4D14FFD1EB337556FBE9FE04FBDA" box="[267,391,1096,1119]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="aruensis">
<emphasis id="B91AEA85FFD1EB337556FBE9FE04FBDA" box="[267,391,1096,1119]" italics="true" pageId="11" pageNumber="164">L. aruensis</emphasis>
</taxonomicName>
), and in having white bones (versus green in
<taxonomicName id="4C6E4D14FFD1EB3377DFFBE9FC7DFBDA" box="[898,1022,1096,1119]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="aruensis">
<emphasis id="B91AEA85FFD1EB3377DFFBE9FC7DFBDA" box="[898,1022,1096,1119]" italics="true" pageId="11" pageNumber="164">L. aruensis</emphasis>
</taxonomicName>
). It further differs from
<taxonomicName id="4C6E4D14FFD1EB337151FBE9FADCFBDA" box="[1292,1375,1096,1119]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB337151FBE9FADCFBDA" box="[1292,1375,1096,1119]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
(
<figureCitation id="13552A12FFD1EB337133FBE6FF26FB01" captionStart="FIGURE 6" captionStartId="12.[151,250,1775,1797]" captionTargetBox="[151,1436,1021,1753]" captionTargetId="figure@12.[151,1436,1021,1753]" captionTargetPageId="12" captionText="FIGURE 6. (A) Litoria auae from the Muller Range, Papua New Guinea; Litoria gracilenta from (B) Mareeba and (C) Kirrama, QLD, Australia, and (D) Richmond Range, NSW Australia; (E) Litoria chloris from Loadstone, NSW, Australia (AMS R 184770); and (F) Litoria xanthomera from Tully Gorge, QLD, Australia." httpUri="https://zenodo.org/record/268266/files/figure.png" pageId="11" pageNumber="164">Fig. 6</figureCitation>
A) by having a larger (although slightly overlapping) male body size (
<quantity id="4C969B72FFD1EB3377E9FBCAFBDBFB06" box="[948,1112,1131,1156]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="11" pageNumber="164" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs
<quantity id="4C969B72FFD1EB3370E7FBCAFADEFB06" box="[1210,1373,1131,1156]" metricMagnitude="-2" metricUnit="m" metricValue="3.3049999999999997" metricValueMax="3.62" metricValueMin="2.9899999999999998" pageId="11" pageNumber="164" unit="mm" value="33.05" valueMax="36.2" valueMin="29.9">29.936.2 mm</quantity>
in
<specimenCount id="9D68FD1EFFD1EB3371DDFBCDFF2DFB22" pageId="11" pageNumber="164" type="generic">
37
<taxonomicName id="4C6E4D14FFD1EB3374CAFB31FF2DFB22" box="[151,174,1168,1191]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB3374CAFB31FF2DFB22" box="[151,174,1168,1191]" italics="true" pageId="11" pageNumber="164">L.</emphasis>
</taxonomicName>
</specimenCount>
auae), smaller relative eye size (0.100.12 EYE/SVL vs
<quantity id="4C969B72FFD1EB337705FB2EFC6AFB2D" box="[856,1001,1167,1192]" metricMagnitude="-3" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="11" pageNumber="164" unit="in" value="0.14" valueMax="0.16" valueMin="0.12">0.120.16 in</quantity>
<taxonomicName id="4C6E4D14FFD1EB3377A9FB31FBCEFB22" box="[1012,1101,1168,1191]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB3377A9FB31FBCEFB22" box="[1012,1101,1168,1191]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), bright orange webbing (vs golden yellow in
<taxonomicName id="4C6E4D14FFD1EB337501FB15FE32FB4E" box="[348,433,1204,1227]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB337501FB15FE32FB4E" box="[348,433,1204,1227]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), immaculate dorsum in most individuals, with only a few small spots on the dorsum on five of
<specimenCount id="9D68FD1EFFD1EB3374B1FB76FECFFB75" box="[236,332,1239,1264]" pageId="11" pageNumber="164" type="generic" typeStatus="types">13 types</specimenCount>
(vs few to many small, scattered pale yellow spots in all but a few of the
<specimenCount id="9D68FD1EFFD1EB3370F6FB76FA8FFB75" box="[1195,1292,1239,1264]" pageId="11" pageNumber="164" type="generic" typeStatus="types">40 types</specimenCount>
of
<taxonomicName id="4C6E4D14FFD1EB337168FB79FA08FB6A" box="[1333,1419,1240,1263]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB337168FB79FA08FB6A" box="[1333,1419,1240,1263]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), bluish purple dorsal thigh coloration (vs yellow in
<taxonomicName id="4C6E4D14FFD1EB3376B8FB5DFCB8FA96" box="[741,827,1276,1299]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB3376B8FB5DFCB8FA96" box="[741,827,1276,1299]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), no pale canthal stripe (vs indistinct yellow canthal stripe in
<taxonomicName id="4C6E4D14FFD1EB3374A7FA81FECDFAB2" box="[250,334,1312,1335]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB3374A7FA81FECDFAB2" box="[250,334,1312,1335]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), relatively longer (although slightly overlapping) male advertisement call (1.42
<geoCoordinate id="EE5A5050FFD1EB3370BAFABEFAAFFAB2" box="[1255,1324,1311,1336]" direction="south" orientation="latitude" pageId="11" pageNumber="164" precision="555" value="-2.62">2.62 s</geoCoordinate>
at 24.1ºC vs 0.48
<geoCoordinate id="EE5A5050FFD1EB3374A5FAE5FEBAFADE" box="[248,313,1348,1372]" direction="south" orientation="latitude" pageId="11" pageNumber="164" precision="555" value="-1.78">1.78 s</geoCoordinate>
at 24ºC in
<taxonomicName id="4C6E4D14FFD1EB3375E5FAE5FD8FFADE" box="[440,524,1348,1371]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD1EB3375E5FAE5FD8FFADE" box="[440,524,1348,1371]" italics="true" pageId="11" pageNumber="164">L. auae</emphasis>
</taxonomicName>
), and molecular divergence (see below). The new species differs from
<taxonomicName id="4C6E4D14FFD1EB337171FAE5FA1FFADE" box="[1324,1436,1347,1371]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB337171FAE5FA1FFADE" box="[1324,1436,1347,1371]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
by having a larger male body size (
<quantity id="4C969B72FFD1EB337641FAC6FD43FAFA" box="[540,704,1383,1408]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="11" pageNumber="164" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs
<quantity id="4C969B72FFD1EB33777CFAC6FC40FAFA" box="[801,963,1383,1408]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.45" metricValueMin="3.15" pageId="11" pageNumber="164" unit="mm" value="33.0" valueMax="34.5" valueMin="31.5">31.534.5 mm</quantity>
in
<taxonomicName id="4C6E4D14FFD1EB3377B8FAC9FBD6FAFA" box="[997,1109,1383,1407]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB3377B8FAC9FBD6FAFA" box="[997,1109,1383,1407]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
), orange venter (vs yellow
<emphasis id="B91AEA85FFD1EB3371DBFAC9FE8BFA26" italics="true" pageId="11" pageNumber="164">
in
<taxonomicName id="4C6E4D14FFD1EB3374CAFA2DFE8BFA26" box="[151,264,1419,1443]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">L. callista</taxonomicName>
</emphasis>
), bluish purple dorsal thigh coloration (vs orange in
<taxonomicName id="4C6E4D14FFD1EB337739FA2DFC56FA26" box="[868,981,1419,1443]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB337739FA2DFC56FA26" box="[868,981,1419,1443]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
), no pale canthal stripe (vs pale canthal stripe in
<taxonomicName id="4C6E4D14FFD1EB3374A0FA11FEF3FA42" box="[253,368,1455,1479]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB3374A0FA11FEF3FA42" box="[253,368,1455,1479]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
), bright orange webbing (vs yellow in
<taxonomicName id="4C6E4D14FFD1EB337769FA11FC25FA42" box="[820,934,1455,1479]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB337769FA11FC25FA42" box="[820,934,1455,1479]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
), and a male advertisement call of a longer duration (1.42
<geoCoordinate id="EE5A5050FFD1EB33751EFA72FE0AFA6E" box="[323,393,1491,1516]" direction="south" orientation="latitude" pageId="11" pageNumber="164" precision="555" value="-2.62">2.62 s</geoCoordinate>
at 24.1ºC vs 0.045
<geoCoordinate id="EE5A5050FFD1EB33762CFA72FD47FA6E" box="[625,708,1491,1516]" direction="south" orientation="latitude" pageId="11" pageNumber="164" precision="55" value="-0.19">0.190 s</geoCoordinate>
at 24ºC in
<taxonomicName id="4C6E4D14FFD1EB33771BFA75FC34FA6E" box="[838,951,1491,1515]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB33771BFA75FC34FA6E" box="[838,951,1491,1515]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
), greater average number of notes per call (~117141 vs
<quantity id="4C969B72FFD1EB33756AFA56FEF4F995" box="[311,375,1527,1552]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" pageId="11" pageNumber="164" unit="in" value="9.5">9.5 in</quantity>
<taxonomicName id="4C6E4D14FFD1EB337520FA59FE6DF98A" box="[381,494,1527,1551]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB337520FA59FE6DF98A" box="[381,494,1527,1551]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
) and a higher dominant frequency (2.62.8 kHz at 24.1ºC vs 1.282.39 kHz at 24ºC in
<taxonomicName id="4C6E4D14FFD1EB3374E8F9BDFEA4F9B6" box="[181,295,1563,1587]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="callista">
<emphasis id="B91AEA85FFD1EB3374E8F9BDFEA4F9B6" box="[181,295,1563,1587]" italics="true" pageId="11" pageNumber="164">L. callista</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C6E4D14FFD1EB3375D9F9BDFE69F9B6" box="[388,490,1563,1587]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB3375D9F9BDFE69F9B6" box="[388,490,1563,1587]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
, the new species differs by having a larger male body size (
<quantity id="4C969B72FFD1EB3370C5F9BAFABDF9B6" box="[1176,1342,1563,1588]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="11" pageNumber="164" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs
<quantity id="4C969B72FFD1EB3374CAF99EFEB9F9D2" box="[151,314,1599,1624]" metricMagnitude="-2" metricUnit="m" metricValue="2.895" metricValueMax="3.04" metricValueMin="2.75" pageId="11" pageNumber="164" unit="mm" value="28.95" valueMax="30.4" valueMin="27.5">27.530.4 mm</quantity>
in
<taxonomicName id="4C6E4D14FFD1EB337503F9E1FE40F9D2" box="[350,451,1599,1623]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB337503F9E1FE40F9D2" box="[350,451,1599,1623]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
), immaculately green dorsum in most individuals (vs pale green with indistinct whitish dorsal spotting in
<taxonomicName id="4C6E4D14FFD1EB337502F9C5FE47F9FE" box="[351,452,1635,1659]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB337502F9C5FE47F9FE" box="[351,452,1635,1659]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
), orange venter (vs cream in
<taxonomicName id="4C6E4D14FFD1EB337756F9C5FCF3F9FE" box="[779,880,1635,1659]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB337756F9C5FCF3F9FE" box="[779,880,1635,1659]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
), no pale canthal stripe (vs white canthal stripe in
<taxonomicName id="4C6E4D14FFD1EB3374CAF929FF7DF91A" box="[151,254,1671,1695]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB3374CAF929FF7DF91A" box="[151,254,1671,1695]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
), and a male advertisement call with a single note with a relatively slow pulse rate (5664 pulses/s at 24.1ºC vs a biphasic call with 80101 pulses/s in
<taxonomicName id="4C6E4D14FFD1EB33769EF90DFCA4F946" box="[707,807,1707,1731]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="elkeae">
<emphasis id="B91AEA85FFD1EB33769EF90DFCA4F946" box="[707,807,1707,1731]" italics="true" pageId="11" pageNumber="164">L. elkeae</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C6E4D14FFD1EB3377DEF90DFC75F946" box="[899,1014,1707,1731]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="eschata">
<emphasis id="B91AEA85FFD1EB3377DEF90DFC75F946" box="[899,1014,1707,1731]" italics="true" pageId="11" pageNumber="164">L. eschata</emphasis>
</taxonomicName>
, the new species differs by having an orange venter (vs yellow in
<taxonomicName id="4C6E4D14FFD1EB33758AF971FDC8F962" box="[471,587,1743,1767]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="eschata">
<emphasis id="B91AEA85FFD1EB33758AF971FDC8F962" box="[471,587,1743,1767]" italics="true" pageId="11" pageNumber="164">L. eschata</emphasis>
</taxonomicName>
), bluish purple dorsal thigh coloration (vs caramel brown in
<taxonomicName id="4C6E4D14FFD1EB33715FF971FAF5F962" box="[1282,1398,1743,1767]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="eschata">
<emphasis id="B91AEA85FFD1EB33715FF971FAF5F962" box="[1282,1398,1743,1767]" italics="true" pageId="11" pageNumber="164">L. eschata</emphasis>
</taxonomicName>
), a male advertisement call with a slower pulse rate (5664 pulses/s at 21.4ºC vs
<quantity id="4C969B72FFD1EB337075F955FB0DF889" box="[1064,1166,1779,1804]" metricMagnitude="0" metricUnit="m" metricValue="1.9304" metricValueMax="2.032" metricValueMin="1.8288" pageId="11" pageNumber="164" unit="in" value="76.0" valueMax="80.0" valueMin="72.0">7280 in</quantity>
<taxonomicName id="4C6E4D14FFD1EB3370C5F955FA8DF88E" box="[1176,1294,1779,1803]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="eschata">
<emphasis id="B91AEA85FFD1EB3370C5F955FA8DF88E" box="[1176,1294,1779,1803]" italics="true" pageId="11" pageNumber="164">L. eschata</emphasis>
</taxonomicName>
) and higher dominant frequency (2.62.8 kHz at 24.1ºC vs 2.22.5 kHz in
<taxonomicName id="4C6E4D14FFD1EB337731F8B9FC62F8AA" box="[876,993,1815,1839]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="eschata">
<emphasis id="B91AEA85FFD1EB337731F8B9FC62F8AA" box="[876,993,1815,1839]" italics="true" pageId="11" pageNumber="164">L. eschata</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C6E4D14FFD1EB33701FF8B9FB56F8AA" box="[1090,1237,1815,1839]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB33701FF8B9FB56F8AA" box="[1090,1237,1815,1839]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
(
<figureCitation id="13552A12FFD1EB3370BBF8B6FAA8F8B5" box="[1254,1323,1815,1840]" captionStart="FIGURE 6" captionStartId="12.[151,250,1775,1797]" captionTargetBox="[151,1436,1021,1753]" captionTargetId="figure@12.[151,1436,1021,1753]" captionTargetPageId="12" captionText="FIGURE 6. (A) Litoria auae from the Muller Range, Papua New Guinea; Litoria gracilenta from (B) Mareeba and (C) Kirrama, QLD, Australia, and (D) Richmond Range, NSW Australia; (E) Litoria chloris from Loadstone, NSW, Australia (AMS R 184770); and (F) Litoria xanthomera from Tully Gorge, QLD, Australia." httpUri="https://zenodo.org/record/268266/files/figure.png" pageId="11" pageNumber="164">Fig. 6</figureCitation>
BD), the new species differs by having an orange venter (vs white to yellow in
<taxonomicName id="4C6E4D14FFD1EB337795F89DFBD8F8D6" box="[968,1115,1851,1875]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB337795F89DFBD8F8D6" box="[968,1115,1851,1875]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
), bluish purple dorsal thigh coloration (versus purplish red in
<taxonomicName id="4C6E4D14FFD1EB33767AF8C1FD3FF8F2" box="[551,700,1887,1911]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB33767AF8C1FD3FF8F2" box="[551,700,1887,1911]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
), no pale canthal stripe (vs pale yellowish canthal stripe in
<taxonomicName id="4C6E4D14FFD1EB3371D8F8C1FE89F81E" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3371D8F8C1FE89F81E" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
), orange webbing (vs yellow in
<taxonomicName id="4C6E4D14FFD1EB3376D1F825FCA3F81E" box="[652,800,1923,1947]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3376D1F825FCA3F81E" box="[652,800,1923,1947]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
), bright golden iris with thick black iris periphery (vs brownish gold iris, often with blue or green at borders and a narrow black periphery in
<taxonomicName id="4C6E4D14FFD1EB3370C2F809FAB7F83A" box="[1183,1332,1959,1983]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB3370C2F809FAB7F83A" box="[1183,1332,1959,1983]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
), a male advertisement call with fewer pulses (~120140 at 24.1ºC vs ~180 pulses in
<taxonomicName id="4C6E4D14FFD1EB337055F86DFB18F866" box="[1032,1179,1995,2019]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="164" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD1EB337055F86DFB18F866" box="[1032,1179,1995,2019]" italics="true" pageId="11" pageNumber="164">L. gracilenta</emphasis>
</taxonomicName>
at 21.721.8ºC) and a slower pulse rate (~60 at 24.1ºC vs ~85 pulses/s in
<taxonomicName id="4C6E4D14FFD6EB3476B8FF38FCFAFF35" box="[741,889,152,176]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD6EB3476B8FF38FCFAFF35" box="[741,889,152,176]" italics="true" pageId="12" pageNumber="165">L. gracilenta</emphasis>
</taxonomicName>
at 21.721.8ºC), and molecular divergence (see below). From
<taxonomicName id="4C6E4D14FFD6EB347565FF1CFE23FF51" box="[312,416,188,212]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="kumae">
<emphasis id="B91AEA85FFD6EB347565FF1CFE23FF51" box="[312,416,188,212]" italics="true" pageId="12" pageNumber="165">L. kumae</emphasis>
</taxonomicName>
, the new species differs by having a larger male body size (
<quantity id="4C969B72FFD6EB347017FF1DFB73FF51" box="[1098,1264,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="12" pageNumber="165" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs
<quantity id="4C969B72FFD6EB34710BFF1DFF40FF72" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.0" metricValueMin="2.4" pageId="12" pageNumber="165" unit="mm" value="27.0" valueMax="30.0" valueMin="24.0">2430 mm</quantity>
in
<taxonomicName id="4C6E4D14FFD6EB3474B6FF40FED6FF7D" box="[235,341,224,248]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="kumae">
<emphasis id="B91AEA85FFD6EB3474B6FF40FED6FF7D" box="[235,341,224,248]" italics="true" pageId="12" pageNumber="165">L. kumae</emphasis>
</taxonomicName>
), an orange venter (vs a venter that is white anteriorally and yellowish gold posteriorally in
<taxonomicName id="4C6E4D14FFD6EB3471D8FF40FF63FE99" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="kumae">
<emphasis id="B91AEA85FFD6EB3471D8FF40FF63FE99" italics="true" pageId="12" pageNumber="165">L. kumae</emphasis>
</taxonomicName>
), bluish purple dorsal thigh coloration (vs reddish gold in
<taxonomicName id="4C6E4D14FFD6EB3477D3FEA4FC7BFE99" box="[910,1016,260,284]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="kumae">
<emphasis id="B91AEA85FFD6EB3477D3FEA4FC7BFE99" box="[910,1016,260,284]" italics="true" pageId="12" pageNumber="165">L. kumae</emphasis>
</taxonomicName>
) and no pale canthal stripe (vs pale canthal stripe in
<taxonomicName id="4C6E4D14FFD6EB347509FE88FE38FEC5" box="[340,443,296,320]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="kumae">
<emphasis id="B91AEA85FFD6EB347509FE88FE38FEC5" box="[340,443,296,320]" italics="true" pageId="12" pageNumber="165">L. kumae</emphasis>
</taxonomicName>
). From
<taxonomicName id="4C6E4D14FFD6EB347645FE88FD30FEC5" box="[536,691,296,320]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="robinsonae">
<emphasis id="B91AEA85FFD6EB347645FE88FD30FEC5" box="[536,691,296,320]" italics="true" pageId="12" pageNumber="165">L. robinsonae</emphasis>
</taxonomicName>
, the new species differs by having a larger male body size (
<quantity id="4C969B72FFD6EB347106FE86FF79FEE1" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="12" pageNumber="165" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.5 41.8 mm</quantity>
SVL vs
<quantity id="4C969B72FFD6EB347501FEEDFE7DFEE1" box="[348,510,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="2.87" metricValueMin="2.83" pageId="12" pageNumber="165" unit="mm" value="28.5" valueMax="28.7" valueMin="28.3">28.328.7 mm</quantity>
in
<taxonomicName id="4C6E4D14FFD6EB34767CFEECFD38FEE1" box="[545,699,332,356]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="robinsonae">
<emphasis id="B91AEA85FFD6EB34767CFEECFD38FEE1" box="[545,699,332,356]" italics="true" pageId="12" pageNumber="165">L. robinsonae</emphasis>
</taxonomicName>
), orange webbing and digits (vs bluish-grey webbing with bluishgrey to translucent anterior digits) and no pale canthal stripe (vs pale canthal stripe in
<taxonomicName id="4C6E4D14FFD6EB347004FED0FB70FE0D" box="[1113,1267,368,392]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="robinsonae">
<emphasis id="B91AEA85FFD6EB347004FED0FB70FE0D" box="[1113,1267,368,392]" italics="true" pageId="12" pageNumber="165">L. robinsonae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BD13697FFD6EB34749AFE35FD41FD75" blockId="12.[151,1437,151,968]" pageId="12" pageNumber="165">
The new species differs from
<taxonomicName id="4C6E4D14FFD6EB34764FFE34FD37FE29" box="[530,692,404,428]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB34764FFE34FD37FE29" box="[530,692,404,428]" italics="true" pageId="12" pageNumber="165">Litoria chloris</emphasis>
</taxonomicName>
(
<figureCitation id="13552A12FFD6EB34769EFE35FC86FE28" box="[707,773,404,429]" captionStart="FIGURE 6" captionStartId="12.[151,250,1775,1797]" captionTargetBox="[151,1436,1021,1753]" captionTargetId="figure@12.[151,1436,1021,1753]" captionTargetPageId="12" captionText="FIGURE 6. (A) Litoria auae from the Muller Range, Papua New Guinea; Litoria gracilenta from (B) Mareeba and (C) Kirrama, QLD, Australia, and (D) Richmond Range, NSW Australia; (E) Litoria chloris from Loadstone, NSW, Australia (AMS R 184770); and (F) Litoria xanthomera from Tully Gorge, QLD, Australia." httpUri="https://zenodo.org/record/268266/files/figure.png" pageId="12" pageNumber="165">Fig. 6</figureCitation>
E) by having a smaller male body size (
<quantity id="4C969B72FFD6EB3470E0FE35FAE2FE29" box="[1213,1377,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="12" pageNumber="165" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs&gt;
<quantity id="4C969B72FFD6EB347494FE16FE9CFE55" box="[201,287,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="165" unit="mm" value="45.0">45 mm</quantity>
), an orange venter (versus yellowish white in
<taxonomicName id="4C6E4D14FFD6EB347768FE18FC20FE55" box="[821,931,440,464]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB347768FE18FC20FE55" box="[821,931,440,464]" italics="true" pageId="12" pageNumber="165">L. chloris</emphasis>
</taxonomicName>
), orange webbing (vs yellow in
<taxonomicName id="4C6E4D14FFD6EB347140FE18FA08FE55" box="[1309,1419,440,464]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB347140FE18FA08FE55" box="[1309,1419,440,464]" italics="true" pageId="12" pageNumber="165">L. chloris</emphasis>
</taxonomicName>
), bright golden iris with thick black iris periphery (vs reddish-orange with thick black periphery in
<taxonomicName id="4C6E4D14FFD6EB34708CFE7CFABFFE71" box="[1233,1340,476,500]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB34708CFE7CFABFFE71" box="[1233,1340,476,500]" italics="true" pageId="12" pageNumber="165">L. chloris</emphasis>
</taxonomicName>
), a male advertisement call of a single, pulsed note of more than 1s duration (vs a multi-note call with each note &lt;1s in duration in
<taxonomicName id="4C6E4D14FFD6EB347541FD84FE09FDB9" box="[284,394,548,572]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB347541FD84FE09FDB9" box="[284,394,548,572]" italics="true" pageId="12" pageNumber="165">L. chloris</emphasis>
</taxonomicName>
; see
<figureCitation id="13552A12FFD6EB347594FD85FD86FDB8" box="[457,517,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1407,1429]" captionTargetBox="[152,1423,199,1381]" captionTargetId="figure@10.[152,1435,195,1385]" captionTargetPageId="10" captionText="FIGURE 5. Advertisement call of (A) Litoria bella sp. nov. recorded at 24.1 ºC, (i) waveform of relative amplitude over time of a single call of QM J 74468, (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM J 74468, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for QM J 74467. (B) Litoria gracilenta recorded at 21.7 21.8 ºC, (i) waveform of relative amplitude over time of a single call of AMS R 184768 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184768, and (iii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184777. (C) Litoria chloris recorded at 21.8 ºC, (i) waveform of relative amplitude over time of a single call of AMS R 184770 (ii) waveform of relative amplitude over time and corresponding spectrogram (frequency over time) below over 40 s for AMS R 184770." httpUri="https://zenodo.org/record/268265/files/figure.png" pageId="12" pageNumber="165">Fig 5</figureCitation>
C), and molecular divergence (see below). The new species differs from
<taxonomicName id="4C6E4D14FFD6EB347113FD84FE9EFDE5" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xanthomera">
<emphasis id="B91AEA85FFD6EB347113FD84FE9EFDE5" italics="true" pageId="12" pageNumber="165">Litoria xanthomera</emphasis>
</taxonomicName>
(
<figureCitation id="13552A12FFD6EB347571FDE6FEEDFDE5" box="[300,366,583,608]" captionStart="FIGURE 6" captionStartId="12.[151,250,1775,1797]" captionTargetBox="[151,1436,1021,1753]" captionTargetId="figure@12.[151,1436,1021,1753]" captionTargetPageId="12" captionText="FIGURE 6. (A) Litoria auae from the Muller Range, Papua New Guinea; Litoria gracilenta from (B) Mareeba and (C) Kirrama, QLD, Australia, and (D) Richmond Range, NSW Australia; (E) Litoria chloris from Loadstone, NSW, Australia (AMS R 184770); and (F) Litoria xanthomera from Tully Gorge, QLD, Australia." httpUri="https://zenodo.org/record/268266/files/figure.png" pageId="12" pageNumber="165">Fig. 6</figureCitation>
F) by having a smaller male body size (
<quantity id="4C969B72FFD6EB347778FDE6FC4AFDDA" box="[805,969,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="3.815" metricValueMax="4.18" metricValueMin="3.45" pageId="12" pageNumber="165" unit="mm" value="38.15" valueMax="41.8" valueMin="34.5">34.541.8 mm</quantity>
SVL vs&gt;
<quantity id="4C969B72FFD6EB347067FDE6FB0EFDE5" box="[1082,1165,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="165" unit="mm" value="45.0">45 mm</quantity>
), bright orange webbing (vs yellow to pale orange-yellow in
<taxonomicName id="4C6E4D14FFD6EB34766CFDCCFD55FD01" box="[561,726,620,644]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xanthomera">
<emphasis id="B91AEA85FFD6EB34766CFDCCFD55FD01" box="[561,726,620,644]" italics="true" pageId="12" pageNumber="165">L. xanthomera</emphasis>
</taxonomicName>
), bright golden iris with thick black iris periphery (vs reddishorange with thick black periphery in
<taxonomicName id="4C6E4D14FFD6EB347600FD30FD79FD2D" box="[605,762,656,680]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xathomera">
<emphasis id="B91AEA85FFD6EB347600FD30FD79FD2D" box="[605,762,656,680]" italics="true" pageId="12" pageNumber="165">L. xathomera</emphasis>
</taxonomicName>
), bluish purple dorsal thigh coloration (vs orange in
<taxonomicName id="4C6E4D14FFD6EB3471D8FD30FE9EFD49" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xanthomera">
<emphasis id="B91AEA85FFD6EB3471D8FD30FE9EFD49" italics="true" pageId="12" pageNumber="165">L. xanthomera</emphasis>
</taxonomicName>
), and a male advertisement call of a single, pulsed note of more than 1s duration (vs a multi-note call with each note &lt;1s in duration in
<taxonomicName id="4C6E4D14FFD6EB347653FD78FD31FD75" box="[526,690,728,752]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xanthomera">
<emphasis id="B91AEA85FFD6EB347653FD78FD31FD75" box="[526,690,728,752]" italics="true" pageId="12" pageNumber="165">L. xanthomera</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BD13697FFD6EB34749AFD5DFCC8FC4D" blockId="12.[151,1437,151,968]" pageId="12" pageNumber="165">
<bibRefCitation id="EFFF4B66FFD6EB34749AFD5DFE67FC91" author="Tyler" box="[199,484,764,789]" pageId="12" pageNumber="165" refString="Tyler, M. J. &amp; Davies, M. (1978) Species-groups within the Australopapuan hulid frog genus Litoria Tschudi. Australian Journal of Zoology, 26, 1 - 47. http: // dx. doi. org / 10.1071 / AJZS 063" type="journal article" year="1978">Tyler and Davies (1978)</bibRefCitation>
included the New
<collectingCountry id="F3797607FFD6EB347697FD5DFC98FC90" box="[714,795,764,789]" name="Guinea-Bissau" pageId="12" pageNumber="165">Guinea</collectingCountry>
species
<taxonomicName id="4C6E4D14FFD6EB3477DEFD5CFBC4FC91" box="[899,1095,765,788]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B91AEA85FFD6EB3477DEFD5CFBC4FC91" box="[899,1095,765,788]" italics="true" pageId="12" pageNumber="165">Litoria graminea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D14FFD6EB3470DBFD5CFA99FC91" box="[1158,1306,764,788]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="multiplica">
<emphasis id="B91AEA85FFD6EB3470DBFD5CFA99FC91" box="[1158,1306,764,788]" italics="true" pageId="12" pageNumber="165">L. multiplica</emphasis>
</taxonomicName>
in their
<taxonomicName id="4C6E4D14FFD6EB3471D8FD5CFF75FCB2" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="aruensis">
<emphasis id="B91AEA85FFD6EB3471D8FD5CFF75FCB2" italics="true" pageId="12" pageNumber="165">L. aruensis</emphasis>
</taxonomicName>
group.
<taxonomicName id="4C6E4D14FFD6EB34750CFC81FD90FCB2" box="[337,531,800,823]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="graminea">
<emphasis id="B91AEA85FFD6EB34750CFC81FD90FCB2" box="[337,531,800,823]" italics="true" pageId="12" pageNumber="165">Litoria graminea</emphasis>
</taxonomicName>
and the closely related
<taxonomicName id="4C6E4D14FFD6EB347777FC80FCF0FCBD" box="[810,883,800,824]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="dux">
<emphasis id="B91AEA85FFD6EB347777FC80FCF0FCBD" box="[810,883,800,824]" italics="true" pageId="12" pageNumber="165">L. dux</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D14FFD6EB3477F3FC80FBA1FCB2" box="[942,1058,800,824]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="sauroni">
<emphasis id="B91AEA85FFD6EB3477F3FC80FBA1FCB2" box="[942,1058,800,824]" italics="true" pageId="12" pageNumber="165">L. sauroni</emphasis>
</taxonomicName>
are large (male SVL&gt;
<quantity id="4C969B72FFD6EB347166FCBEFA13FCBD" box="[1339,1424,799,824]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="165" unit="mm" value="60.0">60 mm</quantity>
), robust canopy dwelling frogs with prominent white dermal ridges on the arms and legs and are restricted to New
<collectingCountry id="F3797607FFD6EB3474CAFCC6FF6BFC05" box="[151,232,871,896]" name="Guinea-Bissau" pageId="12" pageNumber="165">Guinea</collectingCountry>
(
<bibRefCitation id="EFFF4B66FFD6EB3474AAFCC6FD8FFC05" author="Richards" box="[247,524,871,896]" pageId="12" pageNumber="165" refString="Richards, S. J. &amp; Oliver, P. (2006) Two new species of large green canopy-dwelling frogs (Anura: Hylidae: Litoria) from Papua New Guinea. Zootaxa, 1295, 41 - 60." type="journal article" year="2006">Richards &amp; Oliver 2006</bibRefCitation>
).
<taxonomicName id="4C6E4D14FFD6EB347642FCC9FD65FC05" box="[543,742,872,896]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="multiplica">
<emphasis id="B91AEA85FFD6EB347642FCC9FD65FC05" box="[543,742,872,896]" italics="true" pageId="12" pageNumber="165">Litoria multiplica</emphasis>
</taxonomicName>
is a similar size to
<taxonomicName id="4C6E4D14FFD6EB3477E2FCC8FB95FC05" box="[959,1046,872,896]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD6EB3477E2FCC8FB95FC05" box="[959,1046,872,896]" italics="true" pageId="12" pageNumber="165">L. bella</emphasis>
</taxonomicName>
but has a prominent white bilobed dermal ridge around the vent, white dermal ridges around the knees and along the outer edge of the tarsus, and dark purplish black lateral spots or mottling (
<bibRefCitation id="EFFF4B66FFD6EB34760AFC0EFCBCFC4D" author="Richards" box="[599,831,943,968]" pageId="12" pageNumber="165" refString="Richards, S. J., Oliver, P., Krey, K. &amp; Tjaturadi, B. (2009) A new species of Litoria (Amphibia: Anura: Hylidae) from the foothills of the Foja Mountains, Papua Province, Indonesia. Zootaxa, 2277, 1 - 13." type="journal article" year="2009">
Richards
<emphasis id="B91AEA85FFD6EB34769EFC10FD78FC4D" box="[707,763,944,968]" italics="true" pageId="12" pageNumber="165">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
<caption id="DF11661FFFD6EB3474CAF94EFC4CF8C7" httpUri="https://zenodo.org/record/268266/files/figure.png" pageId="12" pageNumber="165" targetBox="[151,1436,1021,1753]" targetPageId="12">
<paragraph id="8BD13697FFD6EB3474CAF94EFC4CF8C7" blockId="12.[151,1436,1775,1858]" pageId="12" pageNumber="165">
<emphasis id="B91AEA85FFD6EB3474CAF94EFE95F881" bold="true" box="[151,278,1775,1797]" pageId="12" pageNumber="165">FIGURE 6.</emphasis>
(A)
<taxonomicName id="4C6E4D14FFD6EB347513F951FE53F880" box="[334,464,1776,1797]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD6EB347513F951FE53F880" box="[334,464,1776,1797]" italics="true" pageId="12" pageNumber="165">Litoria auae</emphasis>
</taxonomicName>
from the Muller Range, Papua New Guinea;
<taxonomicName id="4C6E4D14FFD6EB3477E0F951FBF5F880" box="[957,1142,1775,1797]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD6EB3477E0F951FBF5F880" box="[957,1142,1775,1797]" italics="true" pageId="12" pageNumber="165">Litoria gracilenta</emphasis>
</taxonomicName>
from (B) Mareeba and (C) Kirrama, QLD, Australia, and (D) Richmond Range, NSW Australia; (E)
<taxonomicName id="4C6E4D14FFD6EB3477F9F8AEFBB8F8A6" box="[932,1083,1805,1828]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB3477F9F8AEFBB8F8A6" box="[932,1083,1805,1828]" italics="true" pageId="12" pageNumber="165">Litoria chloris</emphasis>
</taxonomicName>
from Loadstone, NSW, Australia (AMS R184770); and (F)
<taxonomicName id="4C6E4D14FFD6EB3475C2F88CFDE6F8C7" box="[415,613,1836,1858]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="xanthomera">
<emphasis id="B91AEA85FFD6EB3475C2F88CFDE6F8C7" box="[415,613,1836,1858]" italics="true" pageId="12" pageNumber="165">Litoria xanthomera</emphasis>
</taxonomicName>
from Tully Gorge, QLD, Australia.
</paragraph>
</caption>
<paragraph id="8BD13697FFD6EB34749AF8CEFA1FF871" blockId="12.[151,1437,1903,2036]" pageId="12" pageNumber="165">
<emphasis id="B91AEA85FFD6EB34749AF8CEFE45F80D" bold="true" box="[199,454,1903,1928]" pageId="12" pageNumber="165">Sequence divergence.</emphasis>
Uncorrected sequence divergences between
<taxonomicName id="4C6E4D14FFD6EB3477E3F8D1FB97F802" box="[958,1044,1903,1927]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD6EB3477E3F8D1FB97F802" box="[958,1044,1903,1927]" italics="true" pageId="12" pageNumber="165">L. bella</emphasis>
</taxonomicName>
and the two most morphologically similar species,
<taxonomicName id="4C6E4D14FFD6EB34750DF835FE67F82E" box="[336,484,1939,1963]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="gracilenta">
<emphasis id="B91AEA85FFD6EB34750DF835FE67F82E" box="[336,484,1939,1963]" italics="true" pageId="12" pageNumber="165">L. gracilenta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D14FFD6EB34767DF835FDF4F82E" box="[544,631,1940,1963]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="auae">
<emphasis id="B91AEA85FFD6EB34767DF835FDF4F82E" box="[544,631,1940,1963]" italics="true" pageId="12" pageNumber="165">L. auae</emphasis>
</taxonomicName>
, was 20.123.0% and 13.014.5% at the more rapidly evolving ND4 gene fragment analysed and 5.26.1% and 2.0% at the relatively conservative 16S gene fragment analysed. Uncorrected sequence divergences between
<taxonomicName id="4C6E4D14FFD6EB3476D0F87DFD67F876" box="[653,740,2011,2035]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="bella">
<emphasis id="B91AEA85FFD6EB3476D0F87DFD67F876" box="[653,740,2011,2035]" italics="true" pageId="12" pageNumber="165">L. bella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E4D14FFD6EB347740F87DFC09F876" box="[797,906,2011,2035]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="12" pageNumber="165" phylum="Chordata" rank="species" species="chloris">
<emphasis id="B91AEA85FFD6EB347740F87DFC09F876" box="[797,906,2011,2035]" italics="true" pageId="12" pageNumber="165">L. chloris</emphasis>
</taxonomicName>
was 20.521.5 % (ND4) and 4.85.0% (16S).
</paragraph>
<paragraph id="8BD13697FFD7EB3574CAFF36FD9BFF51" blockId="13.[151,1436,151,213]" pageId="13" pageNumber="166">
These values are comparable to those reported among other recognised
<taxonomicName id="4C6E4D14FFD7EB3577E4FF39FB84FF2A" box="[953,1031,152,175]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="13" pageNumber="166" phylum="Chordata" rank="genus">
<emphasis id="B91AEA85FFD7EB3577E4FF39FB84FF2A" box="[953,1031,152,175]" italics="true" pageId="13" pageNumber="166">Litoria</emphasis>
</taxonomicName>
species (
<bibRefCitation id="EFFF4B66FFD7EB357030FF36FACEFF35" author="Mahony" box="[1133,1357,151,176]" pageId="13" pageNumber="166" refString="Mahony, M., Knowles, R., Foster, R. &amp; Donnellan, S. (2001) Systematics of the Litoria citropa (Anura: Hylidae) complex in northern New South Wales and southern Queensland, Australia, with the description of a new species. Records of the Australian Museum, 53, 37 - 48. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.53.2001.1322" type="journal article" year="2001">
Mahony
<emphasis id="B91AEA85FFD7EB35708EFF38FA8FFF35" box="[1235,1292,152,176]" italics="true" pageId="13" pageNumber="166">et al.</emphasis>
2001
</bibRefCitation>
;
<bibRefCitation id="EFFF4B66FFD7EB357104FF39FEB9FF50" author="Burns" pageId="13" pageNumber="166" refString="Burns, E. L. &amp; Crayn, D. M. (2006) Phylogenetics and evolution of bell frogs (Litoria aurea species-group, Anura: Hylidae) based on mitochondrial ND 4 sequences. Molecular Phylogenetics and Evolution, 39, 573 - 579. http: // dx. doi. org / 10.1016 / j. ympev. 2005.11.017" type="journal article" year="2006">Burns &amp; Crayn 2006</bibRefCitation>
;
<bibRefCitation id="EFFF4B66FFD7EB35751BFF1DFD8FFF51" author="Smith" box="[326,524,188,213]" pageId="13" pageNumber="166" refString="Smith, K. L., Hale, J. M., Kearney, M. R., Austin, J. J. &amp; Melville, J. (2013) Molecular patterns of introgression in a classic hybrid zone between the Australian tree frogs, Litoria ewingii and L. paraewingi: evidence of a tension zone. Molecular Ecology, 22, 1869 - 1883. http: // dx. doi. org / 10.1111 / mec. 12176" type="journal article" year="2013">
Smith
<emphasis id="B91AEA85FFD7EB3575CDFF1CFE42FF51" box="[400,449,188,212]" italics="true" pageId="13" pageNumber="166">et al</emphasis>
. 2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>