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<document id="059E85A6BF54134188276170C1698962" ID-DOI="10.1080/00222933.2017.1344328" ID-ISSN="1464-5262" ID-Zenodo-Dep="5183763" ID-ZooBank="9ADC9001-1B11-4843-9A49-4B94675DFB33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628734269176" checkinUser="felipe" docAuthor="Fiers, Frank &amp; Van Damme, Kay" docDate="2017" docId="03C7290DEE627F15FE53C311FC98FB73" docLanguage="en" docName="JNatHist.51.41-42.2463-2507.pdf" docOrigin="Journal of Natural History 51 (41 - 42)" docSource="http://dx.doi.org/10.1080/00222933.2017.1344328" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Thalamocyclops pachypes Fiers &amp; Van Damme 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2489" masterDocId="FFFE5175EE667F0EFFECC261FF8EFFAA" masterDocTitle="Thalamocyclops pachypes gen. nov., sp. nov. (Copepoda: Cyclopoida: Cyclopidae), a crevicular cyclopine from Socotra Island (Yemen): tale of a remarkable survival drive" masterLastPageNumber="2507" masterPageNumber="2463" pageNumber="2466" updateTime="1699050578882" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="AFFACFFAB3796932DA5229473DC1E09C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="29D63A72451523EC959AEC9BE435CC11">
<mods:title id="E8763CFAB048F5F1D5FD8200FE418DD0">Thalamocyclops pachypes gen. nov., sp. nov. (Copepoda: Cyclopoida: Cyclopidae), a crevicular cyclopine from Socotra Island (Yemen): tale of a remarkable survival drive</mods:title>
</mods:titleInfo>
<mods:name id="611AE12F118F06AA8743E59B411201B8" type="personal">
<mods:role id="52A17E49FECE4DF727AC97E266E21F2E">
<mods:roleTerm id="D7AEB50D3BAE5F55EC18AE148574C936">Author</mods:roleTerm>
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<mods:namePart id="E60B51B8042ECF422D1458993011B2D6">Fiers, Frank</mods:namePart>
</mods:name>
<mods:name id="EF67B846296C2613C77145A008E22A05" type="personal">
<mods:role id="0EAC0571E617E3740659F09E627EEF8E">
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</mods:role>
<mods:namePart id="8C6F76524E52EC33896699408F0B0AC2">Van Damme, Kay</mods:namePart>
</mods:name>
<mods:typeOfResource id="64D08E807E7DE65E48C8A9948C992828">text</mods:typeOfResource>
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<mods:title id="212EBDBFC81F8DBD899B44DEE0BB2273">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="908824A32634439607D8CDEE23BA6341">
<mods:date id="BE1A82C9608BC822E9E25AA1D829F305">2017</mods:date>
<mods:detail id="BE413DA5EAF1E57EE17C2F49CFE06568" type="pubDate">
<mods:number id="1C7BA1B0816DE46B6D0BB9655438614D">2017-10-16</mods:number>
</mods:detail>
<mods:detail id="5DF381263F3D5242899AD22E9086982A" type="volume">
<mods:number id="CDA8CD2E9F2B52EE82DBF521A212A32F">51</mods:number>
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<mods:detail id="03A97D195ACD6D89A138F864C2BBCBEC" type="issue">
<mods:number id="803C0DD47799426A916EEF2AB0D80A6A">41 - 42</mods:number>
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<mods:start id="7D00CF589EDE89C870B926F6DB9B01B2">2463</mods:start>
<mods:end id="5D4ABBD8710B6CF5F484009459DCBFB0">2507</mods:end>
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<mods:url id="9B3095C36535AF6C05870E3ED6C4711E">http://dx.doi.org/10.1080/00222933.2017.1344328</mods:url>
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<mods:classification id="47367FBC19893162F8CE48D9940CF2A5">journal article</mods:classification>
<mods:identifier id="64FCEFE10BE453D610C733FCC0B908E0" type="DOI">10.1080/00222933.2017.1344328</mods:identifier>
<mods:identifier id="DE65A9EFD130C89FE8B9E31A17880089" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="7063C1E189C5AB46F4D855DA4F3FC179" type="Zenodo-Dep">5183763</mods:identifier>
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</mods:mods>
<treatment id="03C7290DEE627F15FE53C311FC98FB73" LSID="urn:lsid:plazi:treatment:03C7290DEE627F15FE53C311FC98FB73" httpUri="http://treatment.plazi.org/id/03C7290DEE627F15FE53C311FC98FB73" lastPageId="27" lastPageNumber="2489" pageId="4" pageNumber="2466">
<subSubSection id="C374CB90EE627F0AFE53C311FCEBFE23" box="[447,869,368,393]" pageId="4" pageNumber="2466" type="nomenclature">
<paragraph id="8BD1981BEE627F0AFE53C311FCEBFE23" blockId="4.[447,869,368,393]" box="[447,869,368,393]" pageId="4" pageNumber="2466">
<heading id="D0992F77EE627F0AFE53C311FCEBFE23" box="[447,869,368,393]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="2466" reason="5">
<taxonomicName id="4C6EE398EE627F0AFE53C311FC8EFE23" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[447,768,368,393]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="species" species="pachypes" status="sp. nov.">
<emphasis id="B91A4409EE627F0AFE53C311FC8EFE23" bold="true" box="[447,768,368,393]" italics="true" pageId="4" pageNumber="2466">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE627F0AFCE4C311FCEBFE23" box="[776,869,368,393]" pageId="4" pageNumber="2466" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE627F0AFDACC3F9FD6EFE1A" box="[576,736,408,432]" pageId="4" pageNumber="2466" type="description">
<paragraph id="8BD1981BEE627F0AFDACC3F9FD6EFE1A" blockId="4.[576,736,408,432]" box="[576,736,408,432]" pageId="4" pageNumber="2466">
(
<figureCitation id="1355849EEE627F0AFDAAC3F9FD56FE1A" box="[582,728,408,432]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-10="Figure 11" captionStart-11="Figure 12" captionStart-12="Figure 13" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="5.[160,225,1567,1590]" captionStartId-1="6.[160,225,1193,1216]" captionStartId-10="18.[160,225,1558,1581]" captionStartId-11="20.[160,225,1326,1349]" captionStartId-12="21.[160,225,1550,1573]" captionStartId-2="8.[160,225,1539,1562]" captionStartId-3="9.[160,225,1192,1215]" captionStartId-4="10.[160,225,1261,1284]" captionStartId-5="11.[160,225,1168,1191]" captionStartId-6="13.[160,225,1153,1176]" captionStartId-7="14.[160,225,1560,1583]" captionStartId-8="15.[160,225,1450,1473]" captionStartId-9="16.[160,225,1477,1500]" captionTargetBox-0="[163,1152,142,1534]" captionTargetBox-1="[163,1153,147,1162]" captionTargetBox-10="[163,1153,142,1527]" captionTargetBox-11="[163,1153,142,1295]" captionTargetBox-12="[163,1153,142,1519]" captionTargetBox-2="[163,1153,142,1508]" captionTargetBox-3="[163,1153,142,1161]" captionTargetBox-4="[163,1153,142,1230]" captionTargetBox-5="[163,1153,142,1137]" captionTargetBox-6="[163,1151,142,1122]" captionTargetBox-7="[163,1153,142,1528]" captionTargetBox-8="[163,1153,142,1419]" captionTargetBox-9="[163,1151,149,1446]" captionTargetId-0="figure-1@5.[163,1153,142,1536]" captionTargetId-1="figure-131@6.[163,1153,142,1162]" captionTargetId-10="figure-1@18.[163,1153,142,1527]" captionTargetId-11="figure-95@20.[163,1153,142,1295]" captionTargetId-12="figure-1@21.[163,1153,142,1519]" captionTargetId-2="figure-1@8.[163,1153,142,1508]" captionTargetId-3="figure-94@9.[163,1153,142,1161]" captionTargetId-4="figure-1@10.[163,1153,142,1230]" captionTargetId-5="figure-112@11.[163,1153,142,1137]" captionTargetId-6="figure-144@13.[163,1153,142,1122]" captionTargetId-7="figure-1@14.[163,1153,142,1529]" captionTargetId-8="figure-58@15.[163,1153,142,1419]" captionTargetId-9="figure-61@16.[163,1153,142,1446]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-10="18" captionTargetPageId-11="20" captionTargetPageId-12="21" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="16" captionText-0="Figure 1. Thalamocyclops pachypes gen. nov., sp. nov. (a) Habitus of ovigerous female in dorsal view; (b) idem, in lateral view; (c) habitus of adult male in dorsal view." captionText-1="Figure 2. Thalamocyclops pachypes gen. nov., sp. nov. (a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view." captionText-10="Figure 11. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral." captionText-11="Figure 12. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage I: (a) leg 1, caudal; (b) leg 2, caudal (endopodite illustrated separately); (c–e) Copepodid stage II: (c) leg 1, frontal; (d) leg 2, frontal; (e) leg 3, frontal." captionText-12="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." captionText-2="Figure 3. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennary coxobasis and first endopodite segment, frontal; (b) antenna, caudal; (c) female antennule, dorsal; (d) male antennule, dorsal (last segments detached; aesthetascs broken, insertion sockets indicated with asterisk)." captionText-3="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." captionText-4="Figure 5. Thalamocyclops pachypes gen. nov., sp. nov. (a) Leg 1, frontal (medial spine of basis depicted separately and enlarged, not to scale); (b) leg 2, caudal (endopodite detached); (c) female leg 3, frontal (second exopodite segment depicted separately); (d) pair of male third legs, frontal (second endopodite segment of right ramus and second exopodite segment of left exopodite not illustrated); (e) second endopodite segment of male leg 3, frontal, enlarged." captionText-5="Figure 6. Thalamocyclops pachypes gen. nov., sp. nov. (a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral." captionText-6="Figure 7. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Undefined naupliar stage, in dorsal and ventral view; (c–e) habitus of copepodid stages I to V, respectively, in dorsal view." captionText-7="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." captionText-8="Figure 9. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antenna in copepodid stage I, caudal; (b) idem, in copepodid stage II; (c) idem, in copepodid stage III; (d) antennule in copepodid stage I; (e) idem, in copepodid stage II." captionText-9="Figure 10. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage III; (b) idem, in copepodid stage IV (female); (c) idem, in copepodid stage V (female)." figureDoi-0="http://doi.org/10.5281/zenodo.5183765" figureDoi-1="http://doi.org/10.5281/zenodo.5183767" figureDoi-10="http://doi.org/10.5281/zenodo.5183785" figureDoi-11="http://doi.org/10.5281/zenodo.5183787" figureDoi-12="http://doi.org/10.5281/zenodo.5183789" figureDoi-2="http://doi.org/10.5281/zenodo.5183769" figureDoi-3="http://doi.org/10.5281/zenodo.5183771" figureDoi-4="http://doi.org/10.5281/zenodo.5183773" figureDoi-5="http://doi.org/10.5281/zenodo.5183775" figureDoi-6="http://doi.org/10.5281/zenodo.5183777" figureDoi-7="http://doi.org/10.5281/zenodo.5183779" figureDoi-8="http://doi.org/10.5281/zenodo.5183781" figureDoi-9="http://doi.org/10.5281/zenodo.5183783" httpUri-0="https://zenodo.org/record/5183765/files/figure.png" httpUri-1="https://zenodo.org/record/5183767/files/figure.png" httpUri-10="https://zenodo.org/record/5183785/files/figure.png" httpUri-11="https://zenodo.org/record/5183787/files/figure.png" httpUri-12="https://zenodo.org/record/5183789/files/figure.png" httpUri-2="https://zenodo.org/record/5183769/files/figure.png" httpUri-3="https://zenodo.org/record/5183771/files/figure.png" httpUri-4="https://zenodo.org/record/5183773/files/figure.png" httpUri-5="https://zenodo.org/record/5183775/files/figure.png" httpUri-6="https://zenodo.org/record/5183777/files/figure.png" httpUri-7="https://zenodo.org/record/5183779/files/figure.png" httpUri-8="https://zenodo.org/record/5183781/files/figure.png" httpUri-9="https://zenodo.org/record/5183783/files/figure.png" pageId="4" pageNumber="2466">Figures 1–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE627F0AFF4CC389FCB5FC20" pageId="4" pageNumber="2466" type="materials_examined">
<paragraph id="8BD1981BEE627F0AFF4CC389FE0EFDAB" blockId="4.[160,1156,488,687]" box="[160,384,488,513]" pageId="4" pageNumber="2466">
<emphasis id="B91A4409EE627F0AFF4CC389FE0EFDAB" bold="true" box="[160,384,488,513]" italics="true" pageId="4" pageNumber="2466">Material examined</emphasis>
</paragraph>
<paragraph id="8BD1981BEE627F0AFF4CC06DFC38FD05" blockId="4.[160,1156,488,687]" pageId="4" pageNumber="2466">
<materialsCitation id="3B069246EE627F0AFF4CC06DFC38FD05" collectionCode="A-I" pageId="4" pageNumber="2466" specimenCode="COP 5309, COP 5310, COP 5308, COP5311, COP 5312, COP 5313-5320" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D526B9EE627F0AFF4CC06DFE89FD8E" box="[160,263,524,548]" pageId="4" pageNumber="2466" type="holotype">Holotype</typeStatus>
female, adult, dissected with the appendages mounted on seven slides (RBINSc
<specimenCode id="DBC83060EE627F0AFF13C04EFEFEFDED" box="[255,368,559,583]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP 5309</specimenCode>
A–G),
<typeStatus id="54D526B9EE627F0AFE5AC04EFD9DFDED" box="[438,531,559,583]" pageId="4" pageNumber="2466" type="allotype">allotype</typeStatus>
, adult, dissected with appendages mounted on seven slides (RBINSc
<specimenCode id="DBC83060EE627F0AFEA5C030FE35FDC3" box="[329,443,593,617]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP 5310</specimenCode>
A–G);
<typeStatus id="54D526B9EE627F0AFDEDC030FDFCFDC3" box="[513,626,593,617]" pageId="4" pageNumber="2466" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D685392EE627F0AFD6CC030FC8BFDC3" box="[640,773,593,617]" pageId="4" pageNumber="2466" type="female">one female</specimenCount>
,
<specimenCount id="9D685392EE627F0AFCFEC030FC17FDC3" box="[786,921,593,617]" pageId="4" pageNumber="2466" type="male">three males</specimenCount>
and 70 copepodids and six nauplii ethanol-preserved (RBINSc
<specimenCode id="DBC83060EE627F0AFD6EC015FD7BFD26" box="[642,757,628,652]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP 5308</specimenCode>
),
<specimenCount id="9D685392EE627F0AFCEBC015FC1FFD26" box="[775,913,628,652]" pageId="4" pageNumber="2466" type="female">two females</specimenCount>
(RBINSc
<specimenCode id="DBC83060EE627F0AFC14C015FBECFD26" box="[1016,1122,628,652]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP5311</specimenCode>
<collectionCode id="ED7F00DEEE627F0AFB8EC015FB0FFD26" box="[1122,1153,628,652]" pageId="4" pageNumber="2466">A-I</collectionCode>
,
<specimenCode id="DBC83060EE627F0AFF4CC0F6FE9EFD05" box="[160,272,663,687]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP 5312</specimenCode>
A–G), eight copepodids dissected (RBINSc
<specimenCode id="DBC83060EE627F0AFD18C0F6FC22FD05" box="[756,940,663,687]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="4" pageNumber="2466">COP 5313–5320</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD1981BEE627F0AFF4CC086FEB9FCAA" blockId="4.[160,1155,743,906]" box="[160,311,743,768]" pageId="4" pageNumber="2466">
<emphasis id="B91A4409EE627F0AFF4CC086FEB9FCAA" bold="true" box="[160,311,743,768]" italics="true" pageId="4" pageNumber="2466">Type locality</emphasis>
</paragraph>
<paragraph id="8BD1981BEE627F0AFF4CC16BFCB5FC20" blockId="4.[160,1155,743,906]" pageId="4" pageNumber="2466">
<collectingCountry id="F379D88BEE627F0AFF4CC16BFEB1FC88" box="[160,319,778,802]" name="Socotra Island" pageId="4" pageNumber="2466">
<taxonomicName id="4C6EE398EE627F0AFF4CC16BFF7BFC88" box="[160,245,778,802]" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="2499" phylum="Arthropoda" rank="genus">Socotra</taxonomicName>
Island
</collectingCountry>
(
<collectingCountry id="F379D88BEE627F0AFEA2C16BFE2CFC88" box="[334,418,778,802]" name="Yemen" pageId="4" pageNumber="2466">Yemen</collectingCountry>
), Diksam Plateau (central coörd.:
<geoCoordinate id="EE5AFEDCEE627F0AFCF6C16BFC1CFC88" box="[794,914,778,802]" degrees="12" direction="north" minutes="31.50" orientation="latitude" pageId="4" pageNumber="2466" precision="9" value="12.525">
12°31.50
<emphasis id="B91A4409EE627F0AFC96C16BFC0EFC88" bold="true" box="[890,896,778,802]" italics="true" pageId="4" pageNumber="2466">ʹ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="EE5AFEDCEE627F0AFC76C16BFB81FC88" box="[922,1039,778,802]" degrees="53" direction="east" minutes="58.00" orientation="longitude" pageId="4" pageNumber="2466" precision="9" value="53.966667">
53°58.00
<emphasis id="B91A4409EE627F0AFC16C16BFB8EFC88" bold="true" box="[1018,1024,778,802]" italics="true" pageId="4" pageNumber="2466">ʹ</emphasis>
E
</geoCoordinate>
) at
<quantity id="4C9635FEEE627F0AFBD7C16BFB0DFC88" box="[1083,1155,778,802]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="2466" unit="m" value="700.0">700 m</quantity>
above sea level; limestone weathered crevices, inhabited by
<taxonomicName id="4C6EE398EE627F0AFC8BC14CFE6BFCC2" authority="Cumberlidge &amp; Wranik, 2002" authorityName="Cumberlidge &amp; Wranik" authorityYear="2002" class="Malacostraca" family="Potamidae" genus="Socotra" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="species" species="pseudocardisoma">
<emphasis id="B91A4409EE627F0AFC8BC14CFB0DFCEF" box="[871,1155,813,837]" italics="true" pageId="4" pageNumber="2466">Socotra pseudocardisoma</emphasis>
<bibRefCitation id="EFFFE5EAEE627F0AFF4CC131FE6BFCC2" author="Cumberlidge N &amp; Wranik W" box="[160,485,848,872]" pageId="4" pageNumber="2466" pagination="51 - 64" refId="ref17883" refString="Cumberlidge N, Wranik W. 2002. A new genus and new species of freshwater crab (Potamoidea, Potamidae) from Socotra Island, Yemen. J Nat Hist. 36: 51 - 64." type="journal article" year="2002">Cumberlidge &amp; Wranik, 2002</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C6EE398EE627F0AFE18C131FDE4FCC2" box="[500,618,848,872]" class="Malacostraca" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="order">Decapoda</taxonomicName>
: Brachyura:
<taxonomicName id="4C6EE398EE627F0AFD1FC131FCFEFCC2" authorityName="Ortmann" authorityYear="1896" box="[755,880,848,872]" class="Malacostraca" family="Potamidae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="family">Potamidae</taxonomicName>
), coll. M. Apel, (Museum Mensch und Natur, München,
<collectingCountry id="F379D88BEE627F0AFE16C113FDEAFC20" box="[506,612,882,906]" name="Germany" pageId="4" pageNumber="2466">Germany</collectingCountry>
),
<date id="FFD0BEDBEE627F0AFD9AC113FCB9FC20" box="[630,823,882,906]" pageId="4" pageNumber="2466" value="2000-10-26">26 October 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE627F0AFF4CC1A2FE44FBCC" pageId="4" pageNumber="2466" type="etymology">
<paragraph id="8BD1981BEE627F0AFF4CC1A2FE93FC76" blockId="4.[160,1156,963,1126]" box="[160,285,963,988]" pageId="4" pageNumber="2466">
<emphasis id="B91A4409EE627F0AFF4CC1A2FE93FC76" bold="true" box="[160,285,963,988]" italics="true" pageId="4" pageNumber="2466">Etymology</emphasis>
</paragraph>
<paragraph id="8BD1981BEE627F0AFF4CC187FE44FBCC" blockId="4.[160,1156,963,1126]" pageId="4" pageNumber="2466">
The specific epithet
<emphasis id="B91A4409EE627F0AFE69C187FD8CFC54" box="[389,514,998,1022]" italics="true" pageId="4" pageNumber="2466">
<taxonomicName id="4C6EE398EE627F0AFE69C187FE64FC54" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[389,490,998,1022]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="species" species="pachypes">pachypes</taxonomicName>
is
</emphasis>
a conjuction of the words
<emphasis id="B91A4409EE627F0AFCDAC187FCF5FC54" box="[822,891,998,1022]" italics="true" pageId="4" pageNumber="2466">pachis</emphasis>
(Greek meaning: thick) and
<emphasis id="B91A4409EE627F0AFF3EC668FF78FB8B" box="[210,246,1033,1057]" italics="true" pageId="4" pageNumber="2466">pes</emphasis>
(Latin meaning: leg), and refers to the unique swollen shape of the female leg 5 and of the leg 4 endopodite and protopod distinguishing the species from its congener
<taxonomicName id="4C6EE398EE627F0AFF4CC62FFECCFBCC" authorityName="Fiers &amp; Van Damme" authorityYear="2017" baseAuthorityName="Mirabdullayev, Van Damme &amp; Dumont" baseAuthorityYear="2002" box="[160,322,1102,1126]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="4" pageNumber="2466" phylum="Arthropoda" rank="species" species="soqotraensis" status="comb. nov.">
<emphasis id="B91A4409EE627F0AFF4CC62FFECCFBCC" box="[160,322,1102,1126]" italics="true" pageId="4" pageNumber="2466">T. soqotraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE627F0AFEA7C62FFE44FBCC" box="[331,458,1102,1126]" pageId="4" pageNumber="2466" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE627F18FF4CC6FFFD4DFDED" lastPageId="22" lastPageNumber="2484" pageId="4" pageNumber="2466" type="description">
<paragraph id="8BD1981BEE627F0AFF4CC6FFFE47FB1D" blockId="4.[160,1156,1182,1727]" box="[160,457,1182,1207]" pageId="4" pageNumber="2466">
<emphasis id="B91A4409EE627F0AFF4CC6FFFE47FB1D" bold="true" box="[160,457,1182,1207]" italics="true" pageId="4" pageNumber="2466">Description of the adults</emphasis>
</paragraph>
<paragraph id="8BD1981BEE627F0AFF4CC6A0FD92FA67" blockId="4.[160,1156,1182,1727]" pageId="4" pageNumber="2466">
<emphasis id="B91A4409EE627F0AFF4CC6A0FF72FB70" bold="true" box="[160,252,1217,1242]" italics="true" pageId="4" pageNumber="2466">Female.</emphasis>
Habitus (
<figureCitation id="1355849EEE627F0AFE9DC6A3FE49FB70" box="[369,455,1218,1242]" captionStart="Figure 1" captionStartId="5.[160,225,1567,1590]" captionTargetBox="[163,1152,142,1534]" captionTargetId="figure-1@5.[163,1153,142,1536]" captionTargetPageId="5" captionText="Figure 1. Thalamocyclops pachypes gen. nov., sp. nov. (a) Habitus of ovigerous female in dorsal view; (b) idem, in lateral view; (c) habitus of adult male in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5183765" httpUri="https://zenodo.org/record/5183765/files/figure.png" pageId="4" pageNumber="2466">Figure 1</figureCitation>
(a,b)) compact, dorsoventrally compressed, with clear division between prosome and urosome, both equally long; no pseudosomite defined; fifth leg pediger and genital double somite laterally expanded; body length between 815 and 840 μm (
<typeStatus id="54D526B9EE627F0AFEE4C74BFEE0FAE8" box="[264,366,1322,1346]" pageId="4" pageNumber="2466" type="holotype">holotype</typeStatus>
825 μm), with largest width (280–295 μm) near posterior margin of cephalothorax; posterior margins of all somites straight; integument without particular modifications (pits or refractile points), but apparently clothed with a dense carpet of short cilia (not illustrated); ovigerous females with pair of egg sacs positioned dorsolaterally, each containing
<specimenCount id="9D685392EE627F0AFE45C7D4FD96FA67" box="[425,536,1461,1485]" pageId="4" pageNumber="2466" type="egg">four eggs</specimenCount>
.
</paragraph>
<paragraph id="8BD1981BEE627F08FF2CC7B6FE45FAD8" blockId="4.[160,1156,1182,1727]" lastBlockId="6.[160,1156,1336,1706]" lastPageId="6" lastPageNumber="2468" pageId="4" pageNumber="2466">
Genital double somite (
<figureCitation id="1355849EEE627F0AFE02C7B6FDD9FA45" box="[494,599,1495,1519]" captionStart="Figure 2" captionStartId="6.[160,225,1193,1216]" captionTargetBox="[163,1153,147,1162]" captionTargetId="figure-131@6.[163,1153,142,1162]" captionTargetPageId="6" captionText="Figure 2. Thalamocyclops pachypes gen. nov., sp. nov. (a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5183767" httpUri="https://zenodo.org/record/5183767/files/figure.png" pageId="4" pageNumber="2466">Figure 2</figureCitation>
(d–f)) wider than long, with lateral margins expanded laterally and ventrally delineating a ventral space around copulatory area; vestigial leg 6 situated in middle of somite, dorsolaterally positioned; copulatory pore opening in caudal half of ventral surface, leading to receptacle with duct fitted in triangular shaped muscular mass; receptacle compacted centrally, anterior part expanded, posterior part apparently not present; urosomites 4, 5 and anal somite short, and parallel sided; anal operculum short, not extended, crescentic, with smooth border; anal somite set with few widely spaced small spinules along half of posteroventral margin.
</paragraph>
<caption id="DF11C893EE637F0BFF4CC47EFCBCF9FE" ID-DOI="http://doi.org/10.5281/zenodo.5183765" ID-Zenodo-Dep="5183765" httpUri="https://zenodo.org/record/5183765/files/figure.png" pageId="5" pageNumber="2467" startId="5.[160,225,1567,1590]" targetBox="[163,1152,142,1534]" targetPageId="5">
<paragraph id="8BD1981BEE637F0BFF4CC47EFCBCF9FE" blockId="5.[160,1156,1567,1620]" pageId="5" pageNumber="2467">
<emphasis id="B91A4409EE637F0BFF4CC47EFF74F99C" bold="true" box="[160,250,1567,1590]" pageId="5" pageNumber="2467">Figure 1.</emphasis>
<taxonomicName id="4C6EE398EE637F0BFEEFC47EFE75F99C" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[259,507,1567,1590]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="5" pageNumber="2467" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE637F0BFEEFC47EFE75F99C" box="[259,507,1567,1590]" italics="true" pageId="5" pageNumber="2467">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE637F0BFDEDC47EFD3AF99C" box="[513,692,1567,1590]" pageId="5" pageNumber="2467" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Habitus of ovigerous female in dorsal view; (b) idem, in lateral view; (c) habitus of adult male in dorsal view.
</paragraph>
</caption>
<caption id="DF11C893EE607F08FF4CC6C8FD14FB51" ID-DOI="http://doi.org/10.5281/zenodo.5183767" ID-Zenodo-Dep="5183767" httpUri="https://zenodo.org/record/5183767/files/figure.png" pageId="6" pageNumber="2468" startId="6.[160,225,1193,1216]" targetBox="[163,1153,147,1162]" targetPageId="6">
<paragraph id="8BD1981BEE607F08FF4CC6C8FD14FB51" blockId="6.[160,1156,1193,1275]" pageId="6" pageNumber="2468">
<emphasis id="B91A4409EE607F08FF4CC6C8FF73FB6A" bold="true" box="[160,253,1193,1216]" pageId="6" pageNumber="2468">Figure 2.</emphasis>
<taxonomicName id="4C6EE398EE607F08FEEAC6C8FD8FFB6A" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[262,513,1193,1216]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="6" pageNumber="2468" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE607F08FEEAC6C8FD8FFB6A" box="[262,513,1193,1216]" italics="true" pageId="6" pageNumber="2468">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE607F08FDE7C6C8FD47FB6A" box="[523,713,1193,1216]" pageId="6" pageNumber="2468" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view.
</paragraph>
</caption>
<paragraph id="8BD1981BEE607F08FF2CC71CFE83F9CF" blockId="6.[160,1156,1336,1706]" pageId="6" pageNumber="2468">
Caudal rami (
<figureCitation id="1355849EEE607F08FEB2C71CFE30FA3F" box="[350,446,1405,1429]" captionStart="Figure 2" captionStartId="6.[160,225,1193,1216]" captionTargetBox="[163,1153,147,1162]" captionTargetId="figure-131@6.[163,1153,142,1162]" captionTargetPageId="6" captionText="Figure 2. Thalamocyclops pachypes gen. nov., sp. nov. (a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5183767" httpUri="https://zenodo.org/record/5183767/files/figure.png" pageId="6" pageNumber="2468">Figure 2</figureCitation>
(d–f)) flattened, oval in cross-section, about twice as long as wide in dorsal view and parallel to each; six setae: anterolateral one inserted in median third, posterolateral one near outer distal edge; dorsal seta short, biarticulated, arising near to caudal margin; outer and inner terminal setae sturdy but well developed (outer 190 μm, middle 250 μm), bipinnate, with breaking plane; inner terminal seta, pinnate and slightly longer than half ramal length. Spinules present at insertion of posterolateral seta only.
</paragraph>
<paragraph id="8BD1981BEE607F08FF2CC411FCC2F900" blockId="6.[160,1156,1336,1706]" pageId="6" pageNumber="2468">
Rostrum (
<figureCitation id="1355849EEE607F08FEDCC411FE04F922" box="[304,394,1648,1672]" captionStart="Figure 1" captionStartId="5.[160,225,1567,1590]" captionTargetBox="[163,1152,142,1534]" captionTargetId="figure-1@5.[163,1153,142,1536]" captionTargetPageId="5" captionText="Figure 1. Thalamocyclops pachypes gen. nov., sp. nov. (a) Habitus of ovigerous female in dorsal view; (b) idem, in lateral view; (c) habitus of adult male in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5183765" httpUri="https://zenodo.org/record/5183765/files/figure.png" pageId="6" pageNumber="2468">Figure 1</figureCitation>
(a,b)) large, wide and linguiform; bent ventrally at right angle to longitudinal body axis; no integument modifications visible.
</paragraph>
<paragraph id="8BD1981BEE617F09FF2CC2EFFD6CFEFE" blockId="7.[160,1156,142,1726]" pageId="7" pageNumber="2469">
Antennule (
<figureCitation id="1355849EEE617F09FEA5C2EFFE24FF0C" box="[329,426,142,166]" captionStart="Figure 3" captionStartId="8.[160,225,1539,1562]" captionTargetBox="[163,1153,142,1508]" captionTargetId="figure-1@8.[163,1153,142,1508]" captionTargetPageId="8" captionText="Figure 3. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennary coxobasis and first endopodite segment, frontal; (b) antenna, caudal; (c) female antennule, dorsal; (d) male antennule, dorsal (last segments detached; aesthetascs broken, insertion sockets indicated with asterisk)." figureDoi="http://doi.org/10.5281/zenodo.5183769" httpUri="https://zenodo.org/record/5183769/files/figure.png" pageId="7" pageNumber="2469">Figure 3</figureCitation>
(c)) short, reaching only halfway along cephalothorax; 10-segmented appearance, but with segments 1 and 2 incompletely separated; integument of each segment smooth; segmental armament (first and second segment considered separated): 1[6]-2[7]-3[2]-4[1]-5[2]-6[3]-7[2 + Aesth]-8[2]-9[2 + Aesth]-10[7 + Aesth]; element on segment 4 short and conical; aesthetasc on segment 7 linguiform, on segment 9 filiform, and on segment 10 slender and filiform.
</paragraph>
<paragraph id="8BD1981BEE617F09FF2CC33FFC50FDAB" blockId="7.[160,1156,142,1726]" pageId="7" pageNumber="2469">
Antenna (
<figureCitation id="1355849EEE617F09FEDDC33FFE01FEDC" box="[305,399,350,374]" captionStart="Figure 3" captionStartId="8.[160,225,1539,1562]" captionTargetBox="[163,1153,142,1508]" captionTargetId="figure-1@8.[163,1153,142,1508]" captionTargetPageId="8" captionText="Figure 3. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennary coxobasis and first endopodite segment, frontal; (b) antenna, caudal; (c) female antennule, dorsal; (d) male antennule, dorsal (last segments detached; aesthetascs broken, insertion sockets indicated with asterisk)." figureDoi="http://doi.org/10.5281/zenodo.5183769" httpUri="https://zenodo.org/record/5183769/files/figure.png" pageId="7" pageNumber="2469">Figure 3</figureCitation>
(a,b)) four-segmented; coxobasis with single abexopodal seta, but without exopodite; endopodite with one, five and seven setae, respectively; outer apical element on second endopodite segment stout; coxobasis with three transverse lines of minute spinules in proximal half of frontal surface, caudal surface unadorned; endopodite segments with typical spiniform ornamentation along outer margin.
</paragraph>
<paragraph id="8BD1981BEE617F09FF2CC06DFED9FB1C" blockId="7.[160,1156,142,1726]" pageId="7" pageNumber="2469">
Labrum (
<figureCitation id="1355849EEE617F09FEC2C06DFE1DFD8E" box="[302,403,524,548]" captionStart="Figure 4" captionStartId="9.[160,225,1192,1215]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-94@9.[163,1153,142,1161]" captionTargetPageId="9" captionText="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="7" pageNumber="2469">Figure 4</figureCitation>
(f)) with prominent blunt corners, 10 blunt medial teeth, and pair of hairy pads. Mandible (
<figureCitation id="1355849EEE617F09FDEDC04FFDEAFDEC" box="[513,612,558,582]" captionStart="Figure 4" captionStartId="9.[160,225,1192,1215]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-94@9.[163,1153,142,1161]" captionTargetPageId="9" captionText="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="7" pageNumber="2469">Figure 4</figureCitation>
(a,b)) with slender gnathobasis, with undifferentiated palp, represented by a single robust seta; masticatory margin with sharp teeth in upper half, two cuspidate combs and one short seta; gnathal margin accompanied with a median row of long spinules. Maxillule (
<figureCitation id="1355849EEE617F09FC5FC0F7FB95FD04" box="[947,1051,662,686]" captionStart="Figure 4" captionStartId="9.[160,225,1192,1215]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-94@9.[163,1153,142,1161]" captionTargetPageId="9" captionText="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="7" pageNumber="2469">Figure 4</figureCitation>
(c)) with typical cyclopine appearance; arthrite with three robust smooth claws and a single robust seta along medial edge; lateral margin with six elements, outermost large and wide, bipinnate; maxillular palp with normal armament of three endopodal, one exopodal and three medial setae; endopodite articulating. Maxilla (
<figureCitation id="1355849EEE617F09FC10C140FBEEFC93" box="[1020,1120,801,825]" captionStart="Figure 4" captionStartId="9.[160,225,1192,1215]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-94@9.[163,1153,142,1161]" captionTargetPageId="9" captionText="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="7" pageNumber="2469">Figure 4</figureCitation>
(g– j)) with incompletely fused syncoxa, bearing three endites with normal armament; distal margin of syncoxa, on frontal side, with linguiform protuberance reaching beyond margin; basis typically with claw, a medial proximal and a frontal element; claw rather short and robust, with long spinule row on frontal side; endopodite two-segmented with two and three elements on proximal and distal segments, respectively. Maxilliped (
<figureCitation id="1355849EEE617F09FE26C190FDA0FBA3" box="[458,558,1009,1033]" captionStart="Figure 4" captionStartId="9.[160,225,1192,1215]" captionTargetBox="[163,1153,142,1161]" captionTargetId="figure-94@9.[163,1153,142,1161]" captionTargetPageId="9" captionText="Figure 4. Thalamocyclops pachypes gen. nov., sp. nov. (a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="7" pageNumber="2469">Figure 4</figureCitation>
(d,e)) four-segmented, compact, less than half as long as maxillar syncoxa; armament (proximal to distal segment): 2, 1, 1, 2; all setae, except one on proximal segment, conspicuously long and bipinnate; proximal segment with spinule row on frontal surface, second segment with several spinule rows near inner margin on frontal surface and one near outer distal edge on caudal face.
</paragraph>
<paragraph id="8BD1981BEE617F09FF2CC6A0FD96FA61" blockId="7.[160,1156,142,1726]" pageId="7" pageNumber="2469">
Legs 1–3 (
<figureCitation id="1355849EEE617F09FEDFC6A0FE1EFB73" box="[307,400,1217,1241]" captionStart="Figure 5" captionStartId="10.[160,225,1261,1284]" captionTargetBox="[163,1153,142,1230]" captionTargetId="figure-1@10.[163,1153,142,1230]" captionTargetPageId="10" captionText="Figure 5. Thalamocyclops pachypes gen. nov., sp. nov. (a) Leg 1, frontal (medial spine of basis depicted separately and enlarged, not to scale); (b) leg 2, caudal (endopodite detached); (c) female leg 3, frontal (second exopodite segment depicted separately); (d) pair of male third legs, frontal (second endopodite segment of right ramus and second exopodite segment of left exopodite not illustrated); (e) second endopodite segment of male leg 3, frontal, enlarged." figureDoi="http://doi.org/10.5281/zenodo.5183773" httpUri="https://zenodo.org/record/5183773/files/figure.png" pageId="7" pageNumber="2469">Figure 5</figureCitation>
(a–c)) with short sturdy two-segmented rami; prae-coxal flap in leg 1 robust, slender in legs 2–3 (not illustrated); intercoxal sclerite with large and acute processes with narrow median gap in leg 1 but widely spaced in legs 2 and 3; coxa of leg 1 with long bipinnate medial seta; coxa of legs 2 and 3 without medial element; surface of coxa smooth in leg 1, set with spinule row near distal outer edge on caudal surface in legs 2 and 3; medial margin of basis crescentic and set with hairs (leg 1) or slender spinules (legs 2 and 3); medial spine on basis of leg 1 present, serrate; leg armature summarized in
<tableCitation id="C6ECADA0EE617F09FE53C7D2FD9DFA61" box="[447,531,1459,1483]" captionStart="Table 1" captionStartId="10.[317,373,1486,1509]" captionText="Table 1. Armament of female legs 1–4." pageId="7" pageNumber="2469">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8BD1981BEE617F07FF2CC7B7FF6CFA67" blockId="7.[160,1156,142,1726]" lastBlockId="9.[160,1156,1391,1658]" lastPageId="9" lastPageNumber="2471" pageId="7" pageNumber="2469">
Leg 4 (
<figureCitation id="1355849EEE617F09FEFEC7B7FEFAFA44" box="[274,372,1494,1518]" captionStart="Figure 6" captionStartId="11.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure-112@11.[163,1153,142,1137]" captionTargetPageId="11" captionText="Figure 6. Thalamocyclops pachypes gen. nov., sp. nov. (a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral." figureDoi="http://doi.org/10.5281/zenodo.5183775" httpUri="https://zenodo.org/record/5183775/files/figure.png" pageId="7" pageNumber="2469">Figure 6</figureCitation>
(a,b)) with two-segmented exopodite, one-segmented endopodite and a robust general appearance; coxa without medial seta and smooth surfaces; intercoxal sclerite with large acute processes widely spaced medially; basis with large irregular ovate structure along distal margin between ramal insertion; medial margin crescentic, flat in anterior view, partially inflated on caudal surface; medial margin set with stiff spinules; exopodite as in the preceding legs, except for shorter aspect of inner setae; endopodite one-segmented, large, robust and roughly ovate; anterior surface flat, caudal surface with an inflated ovate region in proximal half; armament elements short and robust, all bipinnate, both terminal elements equally sized.
</paragraph>
<caption id="DF11C893EE6E7F06FF4CC462FC57F9FF" ID-DOI="http://doi.org/10.5281/zenodo.5183769" ID-Zenodo-Dep="5183769" httpUri="https://zenodo.org/record/5183769/files/figure.png" pageId="8" pageNumber="2470" startId="8.[160,225,1539,1562]" targetBox="[163,1153,142,1508]" targetPageId="8">
<paragraph id="8BD1981BEE6E7F06FF4CC462FC57F9FF" blockId="8.[160,1156,1539,1621]" pageId="8" pageNumber="2470">
<emphasis id="B91A4409EE6E7F06FF4CC462FF75F9B0" bold="true" box="[160,251,1539,1562]" pageId="8" pageNumber="2470">Figure 3.</emphasis>
<taxonomicName id="4C6EE398EE6E7F06FEE8C462FE73F9B0" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[260,509,1539,1562]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="8" pageNumber="2470" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE6E7F06FEE8C462FE73F9B0" box="[260,509,1539,1562]" italics="true" pageId="8" pageNumber="2470">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE6E7F06FDE8C462FD35F9B0" box="[516,699,1539,1562]" pageId="8" pageNumber="2470" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Antennary coxobasis and first endopodite segment, frontal; (b) antenna, caudal; (c) female antennule, dorsal; (d) male antennule, dorsal (last segments detached; aesthetascs broken, insertion sockets indicated with asterisk).
</paragraph>
</caption>
<caption id="DF11C893EE6F7F07FF4CC6C9FF69FABD" ID-DOI="http://doi.org/10.5281/zenodo.5183771" ID-Zenodo-Dep="5183771" httpUri="https://zenodo.org/record/5183771/files/figure.png" pageId="9" pageNumber="2471" startId="9.[160,225,1192,1215]" targetBox="[163,1153,142,1161]" targetPageId="9">
<paragraph id="8BD1981BEE6F7F07FF4CC6C9FF69FABD" blockId="9.[160,1157,1192,1303]" pageId="9" pageNumber="2471">
<emphasis id="B91A4409EE6F7F07FF4CC6C9FF74FB15" bold="true" box="[160,250,1192,1215]" pageId="9" pageNumber="2471">Figure 4.</emphasis>
<taxonomicName id="4C6EE398EE6F7F07FEEFC6C9FE75FB15" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[259,507,1192,1215]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="9" pageNumber="2471" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE6F7F07FEEFC6C9FE75FB15" box="[259,507,1192,1215]" italics="true" pageId="9" pageNumber="2471">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE6F7F07FDEEC6C9FD38FB15" box="[514,694,1192,1215]" pageId="9" pageNumber="2471" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Mandible; (b) mandibular biting edge, side view; (c) maxillule; (d) maxilliped, frontal; (e) maxilliped, caudal; (f) labrum; (g) maxillar endopodite, caudal; (h) maxillar basis, caudal; (i) maxillar distal endite, caudal; (j) maxillar syncoxa and basis, frontal.
</paragraph>
</caption>
<paragraph id="8BD1981BEE6F7F07FF2CC7B6FC86F9D0" blockId="9.[160,1156,1391,1658]" pageId="9" pageNumber="2471">
Leg 5 (
<figureCitation id="1355849EEE6F7F07FEE0C7B6FEE9FA45" box="[268,359,1495,1519]" captionStart="Figure 2" captionStartId="6.[160,225,1193,1216]" captionTargetBox="[163,1153,147,1162]" captionTargetId="figure-131@6.[163,1153,142,1162]" captionTargetPageId="6" captionText="Figure 2. Thalamocyclops pachypes gen. nov., sp. nov. (a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5183767" httpUri="https://zenodo.org/record/5183767/files/figure.png" pageId="9" pageNumber="2471">Figure 2</figureCitation>
(a–c)) represented by a large and wide lateral outgrowth of leg bearing somite, completely confluent with pediger, and inclined backward; segment with three rigid armature elements: two setiform ones (seta representing original basal segment and outer element of original distal segment) and one short conical element (homologous with inner element of original distal segment).
</paragraph>
<caption id="DF11C893EE6C7F04FF4CC68CFC11FAD0" ID-DOI="http://doi.org/10.5281/zenodo.5183773" ID-Zenodo-Dep="5183773" httpUri="https://zenodo.org/record/5183773/files/figure.png" pageId="10" pageNumber="2472" startId="10.[160,225,1261,1284]" targetBox="[163,1153,142,1230]" targetPageId="10">
<paragraph id="8BD1981BEE6C7F04FF4CC68CFC11FAD0" blockId="10.[160,1156,1261,1402]" pageId="10" pageNumber="2472">
<emphasis id="B91A4409EE6C7F04FF4CC68CFF71FAAE" bold="true" box="[160,255,1261,1284]" pageId="10" pageNumber="2472">Figure 5.</emphasis>
<taxonomicName id="4C6EE398EE6C7F04FEE4C68CFD8BFAAE" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[264,517,1261,1284]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="10" pageNumber="2472" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE6C7F04FEE4C68CFD8BFAAE" box="[264,517,1261,1284]" italics="true" pageId="10" pageNumber="2472">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE6C7F04FDFDC68CFD5BFAAE" box="[529,725,1261,1284]" pageId="10" pageNumber="2472" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Leg 1, frontal (medial spine of basis depicted separately and enlarged, not to scale); (b) leg 2, caudal (endopodite detached); (c) female leg 3, frontal (second exopodite segment depicted separately); (d) pair of male third legs, frontal (second endopodite segment of right ramus and second exopodite segment of left exopodite not illustrated); (e) second endopodite segment of male leg 3, frontal, enlarged.
</paragraph>
</caption>
<caption id="DF11C893EE6C7F04FED1C7AFFD45FA4F" box="[317,715,1485,1509]" pageId="10" pageNumber="2472" startId="10.[317,373,1486,1509]">
<paragraph id="8BD1981BEE6C7F04FED1C7AFFD45FA4F" blockId="10.[317,715,1485,1509]" box="[317,715,1485,1509]" pageId="10" pageNumber="2472">
<emphasis id="B91A4409EE6C7F04FED1C7AFFE1EFA4F" bold="true" box="[317,400,1486,1509]" pageId="10" pageNumber="2472">Table 1.</emphasis>
Armament of female legs 1
<emphasis id="B91A4409EE6C7F04FD40C7ACFD37FA4E" box="[684,697,1485,1508]" italics="true" pageId="10" pageNumber="2472">–</emphasis>
4.
</paragraph>
</caption>
<paragraph id="8BD1981BEE6C7F04FEABC793FC35F9C6" pageId="10" pageNumber="2472">
<table id="F96E6ABBEE6C80F1FEABC793FC5BF9C6" box="[327,981,1522,1644]" gridcols="5" gridrows="5" pageId="10" pageNumber="2472" rowsContinueIn="12.[331,985,1594,1716]">
<tr id="355E9A59EE6C80F1FEABC793FC5BF9AF" box="[327,981,1522,1541]" gridrow="0" pageId="10" pageNumber="2472">
<th id="768FF325EE6C80F1FEABC793FE0DF9AF" box="[327,387,1522,1541]" gridcol="0" gridrow="0" pageId="10" pageNumber="2472">Female</th>
<th id="768FF325EE6C80F1FE3BC793FE71F9AF" box="[471,511,1522,1541]" gridcol="1" gridrow="0" pageId="10" pageNumber="2472">Coxa</th>
<th id="768FF325EE6C80F1FDBFC793FDF5F9AF" box="[595,635,1522,1541]" gridcol="2" gridrow="0" pageId="10" pageNumber="2472">Basis</th>
<th id="768FF325EE6C80F1FD23C793FCACF9AF" box="[719,802,1522,1541]" gridcol="3" gridrow="0" pageId="10" pageNumber="2472">Exopodite</th>
<th id="768FF325EE6C80F1FC99C793FC5BF9AF" box="[885,981,1522,1541]" gridcol="4" gridrow="0" pageId="10" pageNumber="2472">Endopodite</th>
</tr>
<tr id="355E9A59EE6C80F1FEABC473FC5BF98F" box="[327,981,1554,1573]" gridrow="1" pageId="10" pageNumber="2472">
<th id="768FF325EE6C80F1FEABC473FE0DF98F" box="[327,387,1554,1573]" gridcol="0" gridrow="1" pageId="10" pageNumber="2472">leg 1</th>
<td id="768FF325EE6C80F1FE3BC473FE71F98F" box="[471,511,1554,1573]" gridcol="1" gridrow="1" pageId="10" pageNumber="2472">0-1</td>
<td id="768FF325EE6C80F1FDBFC473FDF5F98F" box="[595,635,1554,1573]" gridcol="2" gridrow="1" pageId="10" pageNumber="2472">1-I</td>
<td id="768FF325EE6C80F1FD23C473FCACF98F" box="[719,802,1554,1573]" gridcol="3" gridrow="1" pageId="10" pageNumber="2472">I-0; III-2-3</td>
<td id="768FF325EE6C80F1FC99C473FC5BF98F" box="[885,981,1554,1573]" gridcol="4" gridrow="1" pageId="10" pageNumber="2472">0-1; 1-I1-1</td>
</tr>
<tr id="355E9A59EE6C80F1FEABC44BFC5BF997" box="[327,981,1578,1597]" gridrow="2" pageId="10" pageNumber="2472">
<th id="768FF325EE6C80F1FEABC44BFE0DF997" box="[327,387,1578,1597]" gridcol="0" gridrow="2" pageId="10" pageNumber="2472">leg 2</th>
<td id="768FF325EE6C80F1FE3BC44BFE71F997" box="[471,511,1578,1597]" gridcol="1" gridrow="2" pageId="10" pageNumber="2472">0-0</td>
<td id="768FF325EE6C80F1FDBFC44BFDF5F997" box="[595,635,1578,1597]" gridcol="2" gridrow="2" pageId="10" pageNumber="2472">1-0</td>
<td id="768FF325EE6C80F1FD23C44BFCACF997" box="[719,802,1578,1597]" gridcol="3" gridrow="2" pageId="10" pageNumber="2472">I-0; II-I1-4</td>
<td id="768FF325EE6C80F1FC99C44BFC5BF997" box="[885,981,1578,1597]" gridcol="4" gridrow="2" pageId="10" pageNumber="2472">0-1; 1-I1-2</td>
</tr>
<tr id="355E9A59EE6C80F1FEABC423FC5BF9FF" box="[327,981,1602,1621]" gridrow="3" pageId="10" pageNumber="2472">
<th id="768FF325EE6C80F1FEABC423FE0DF9FF" box="[327,387,1602,1621]" gridcol="0" gridrow="3" pageId="10" pageNumber="2472">leg 3</th>
<td id="768FF325EE6C80F1FE3BC423FE71F9FF" box="[471,511,1602,1621]" gridcol="1" gridrow="3" pageId="10" pageNumber="2472">0-0</td>
<td id="768FF325EE6C80F1FDBFC423FDF5F9FF" box="[595,635,1602,1621]" gridcol="2" gridrow="3" pageId="10" pageNumber="2472">1-0</td>
<td id="768FF325EE6C80F1FD23C423FCACF9FF" box="[719,802,1602,1621]" gridcol="3" gridrow="3" pageId="10" pageNumber="2472">I-0; II-I1-4</td>
<td id="768FF325EE6C80F1FC99C423FC5BF9FF" box="[885,981,1602,1621]" gridcol="4" gridrow="3" pageId="10" pageNumber="2472">0-1; 1-I1-3</td>
</tr>
<tr id="355E9A59EE6C80F1FEABC438FC5BF9C6" box="[327,981,1625,1644]" gridrow="4" pageId="10" pageNumber="2472">
<th id="768FF325EE6C80F1FEABC438FE0DF9C6" box="[327,387,1625,1644]" gridcol="0" gridrow="4" pageId="10" pageNumber="2472">leg 4</th>
<td id="768FF325EE6C80F1FE3BC438FE71F9C6" box="[471,511,1625,1644]" gridcol="1" gridrow="4" pageId="10" pageNumber="2472">0-0</td>
<td id="768FF325EE6C80F1FDBFC438FDF5F9C6" box="[595,635,1625,1644]" gridcol="2" gridrow="4" pageId="10" pageNumber="2472">1-0</td>
<td id="768FF325EE6C80F1FD23C438FCACF9C6" box="[719,802,1625,1644]" gridcol="3" gridrow="4" pageId="10" pageNumber="2472">I-0; II-I1-3</td>
<td id="768FF325EE6C80F1FC99C438FC5BF9C6" box="[885,981,1625,1644]" gridcol="4" gridrow="4" pageId="10" pageNumber="2472">1-2-2</td>
</tr>
</table>
</paragraph>
<caption id="DF11C893EE6D7F05FF4CC6F1FCCFFB55" ID-DOI="http://doi.org/10.5281/zenodo.5183775" ID-Zenodo-Dep="5183775" httpUri="https://zenodo.org/record/5183775/files/figure.png" pageId="11" pageNumber="2473" startId="11.[160,225,1168,1191]" targetBox="[163,1153,142,1137]" targetPageId="11">
<paragraph id="8BD1981BEE6D7F05FF4CC6F1FCCFFB55" blockId="11.[160,1155,1168,1279]" pageId="11" pageNumber="2473">
<emphasis id="B91A4409EE6D7F05FF4CC6F1FE8FFB0D" bold="true" box="[160,257,1168,1191]" pageId="11" pageNumber="2473">Figure 6.</emphasis>
<taxonomicName id="4C6EE398EE6D7F05FEE6C6F1FD84FB0D" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[266,522,1168,1191]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="11" pageNumber="2473" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE6D7F05FEE6C6F1FD84FB0D" box="[266,522,1168,1191]" italics="true" pageId="11" pageNumber="2473">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE6D7F05FDFBC6F1FD6CFB0D" box="[535,738,1168,1191]" pageId="11" pageNumber="2473" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral.
</paragraph>
</caption>
<paragraph id="8BD1981BEE6D7F05FF2CC724FE8FFA08" blockId="11.[160,1156,1349,1442]" pageId="11" pageNumber="2473">
Leg 6 (
<figureCitation id="1355849EEE6D7F05FEE0C724FEE8FAF7" box="[268,358,1349,1373]" captionStart="Figure 2" captionStartId="6.[160,225,1193,1216]" captionTargetBox="[163,1153,147,1162]" captionTargetId="figure-131@6.[163,1153,142,1162]" captionTargetPageId="6" captionText="Figure 2. Thalamocyclops pachypes gen. nov., sp. nov. (a) Fifth leg bearing somite depicted from anterior to posterior; (b) idem, in dorsal view; (c) idem, in ventral view; (d) female abdomen, dorsal view; (e) idem, lateral view; (f) idem, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5183767" httpUri="https://zenodo.org/record/5183767/files/figure.png" pageId="11" pageNumber="2473">Figure 2</figureCitation>
(d,e)) rigidly sclerotized plate with three smooth elements along border: outer one longest, median one short and conical, and medial one half as long as outer element.
</paragraph>
<paragraph id="8BD1981BEE6D7F05FF4CC7B7FEBDF9D0" blockId="11.[160,1156,1494,1727]" pageId="11" pageNumber="2473">
<emphasis id="B91A4409EE6D7F05FF4CC7B7FF6EFA45" bold="true" box="[160,224,1494,1519]" italics="true" pageId="11" pageNumber="2473">Male.</emphasis>
Habitus (
<figureCitation id="1355849EEE6D7F05FEB6C7B6FE35FA45" box="[346,443,1495,1519]" captionStart="Figure 1" captionStartId="5.[160,225,1567,1590]" captionTargetBox="[163,1152,142,1534]" captionTargetId="figure-1@5.[163,1153,142,1536]" captionTargetPageId="5" captionText="Figure 1. Thalamocyclops pachypes gen. nov., sp. nov. (a) Habitus of ovigerous female in dorsal view; (b) idem, in lateral view; (c) habitus of adult male in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5183765" httpUri="https://zenodo.org/record/5183765/files/figure.png" pageId="11" pageNumber="2473">Figure 1</figureCitation>
(c)) dorsoventrally compressed, as in female, but metasomal somites tapering more sharply caudally; sixth leg bearing somite longer than wide, barrel-shaped; posterior urosomites parallel sided; caudal rami flattened as in female, less wide, and slightly tapering caudally; armature as in female; body length 800–840 μm.
</paragraph>
<paragraph id="8BD1981BEE6D7F02FF2CC4E4FC8FFE9B" blockId="11.[160,1156,1494,1727]" lastBlockId="12.[160,1156,142,305]" lastPageId="12" lastPageNumber="2474" pageId="11" pageNumber="2473">
Antennule (
<figureCitation id="1355849EEE6D7F05FEA0C4E4FE3EF937" box="[332,432,1669,1693]" captionStart="Figure 3" captionStartId="8.[160,225,1539,1562]" captionTargetBox="[163,1153,142,1508]" captionTargetId="figure-1@8.[163,1153,142,1508]" captionTargetPageId="8" captionText="Figure 3. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennary coxobasis and first endopodite segment, frontal; (b) antenna, caudal; (c) female antennule, dorsal; (d) male antennule, dorsal (last segments detached; aesthetascs broken, insertion sockets indicated with asterisk)." figureDoi="http://doi.org/10.5281/zenodo.5183769" httpUri="https://zenodo.org/record/5183769/files/figure.png" pageId="11" pageNumber="2473">Figure 3</figureCitation>
(d)) typically bigeniculate cyclopid shaped, composed of 15 segments each rigidly sclerotized; teminal segment with straight and blunt distal part; armature distribution as follows: 1(7 + 3Aesth)-2(4)-3(2)-4(2 + Aesth)-5(1)-6(2)-7(2)-8(1)- 9(2)-10(4)-11(2)-12(?)-13(1 + 1T)-14(2T)-15(11 + 2Aesth); aesthetascs on segments 1, 4 and 11 decomposed, represented by their pedestal only, aesthetascs on other segments not observed, except two filiform ones on ultimate segment; setae on segments 1–9 with remarkable sturdy appearance, sparsely pinnate.
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF4CC302FE3DFD19" blockId="12.[160,1156,355,899]" pageId="12" pageNumber="2474">
<emphasis id="B91A4409EE6A7F02FF4CC302FD23FED1" bold="true" box="[160,685,355,380]" italics="true" pageId="12" pageNumber="2474">Mouthparts and leg 1 and 2 as in female.</emphasis>
Leg 3 (
<figureCitation id="1355849EEE6A7F02FCFDC302FCFCFED1" box="[785,882,355,379]" captionStart="Figure 5" captionStartId="10.[160,225,1261,1284]" captionTargetBox="[163,1153,142,1230]" captionTargetId="figure-1@10.[163,1153,142,1230]" captionTargetPageId="10" captionText="Figure 5. Thalamocyclops pachypes gen. nov., sp. nov. (a) Leg 1, frontal (medial spine of basis depicted separately and enlarged, not to scale); (b) leg 2, caudal (endopodite detached); (c) female leg 3, frontal (second exopodite segment depicted separately); (d) pair of male third legs, frontal (second endopodite segment of right ramus and second exopodite segment of left exopodite not illustrated); (e) second endopodite segment of male leg 3, frontal, enlarged." figureDoi="http://doi.org/10.5281/zenodo.5183773" httpUri="https://zenodo.org/record/5183773/files/figure.png" pageId="12" pageNumber="2474">Figure 5</figureCitation>
(d,e)) with protopodite, exopodite, and proximal endopodite segment as in female; distal endopodite segment with five elements only: inner terminal female homologous element absent; outer terminal spine smooth, blunt and arched; inner subdistal element rigid, smooth and half as long terminal spine; outer and both inner proximal elements bipinnate; ancestral position of (lost) inner terminal armature element marked as a small internal dense spherical mass below distal margin of segment (not illustrated). Leg 4 (
<figureCitation id="1355849EEE6A7F02FF2EC037FEAAFDC4" box="[194,292,598,622]" captionStart="Figure 6" captionStartId="11.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure-112@11.[163,1153,142,1137]" captionTargetPageId="11" captionText="Figure 6. Thalamocyclops pachypes gen. nov., sp. nov. (a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral." figureDoi="http://doi.org/10.5281/zenodo.5183775" httpUri="https://zenodo.org/record/5183775/files/figure.png" pageId="12" pageNumber="2474">Figure 6</figureCitation>
(c)) unmodified, with two-segmented rami; second endopodite segment with short terminal outer spine and long inner bipinnate inner seta; armature of legs summarized in
<tableCitation id="C6ECADA0EE6A7F02FEBAC0FAFE23FD19" box="[342,429,667,691]" captionStart="Table 2" captionStartId="12.[320,376,1558,1581]" captionText="Table 2. Armament of male legs 1–4." pageId="12" pageNumber="2474">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF2CC0DFFEE4FC94" blockId="12.[160,1156,355,899]" pageId="12" pageNumber="2474">
Leg 5 (
<figureCitation id="1355849EEE6A7F02FEE0C0DFFEE7FD7C" box="[268,361,702,726]" captionStart="Figure 6" captionStartId="11.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure-112@11.[163,1153,142,1137]" captionTargetPageId="11" captionText="Figure 6. Thalamocyclops pachypes gen. nov., sp. nov. (a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral." figureDoi="http://doi.org/10.5281/zenodo.5183775" httpUri="https://zenodo.org/record/5183775/files/figure.png" pageId="12" pageNumber="2474">Figure 6</figureCitation>
(d)) basically as in female except for the more rectangular shape of the setae-bearing extension, and its ventrally positioned location on the pediger; apical elements of original distal segment of same nature as in female but distinctly longer, inner one serrate.
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF2CC128FC5BFC29" blockId="12.[160,1156,355,899]" pageId="12" pageNumber="2474">
Leg 6 (
<figureCitation id="1355849EEE6A7F02FEFEC128FEFEFCCB" box="[274,368,841,865]" captionStart="Figure 6" captionStartId="11.[160,225,1168,1191]" captionTargetBox="[163,1153,142,1137]" captionTargetId="figure-112@11.[163,1153,142,1137]" captionTargetPageId="11" captionText="Figure 6. Thalamocyclops pachypes gen. nov., sp. nov. (a) Pair of female fourth legs, frontal (endopodite of right leg not illustrated); (b) idem, caudal (exopodite of left leg not illustrated); (c) pair of male fourth legs, frontal (second segment of left endopodite and right exopodite not illustrated); (d) fifth and sixth leg bearing somites of male, ventral." figureDoi="http://doi.org/10.5281/zenodo.5183775" httpUri="https://zenodo.org/record/5183775/files/figure.png" pageId="12" pageNumber="2474">Figure 6</figureCitation>
(d)) represented as a rigid triangular smooth plate bearing 3 appendages: outer and median one setiform, inner one spiniform and serrate.
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF4CC1D4FEB0FC64" blockId="12.[160,1156,949,1182]" box="[160,318,949,974]" pageId="12" pageNumber="2474">
<emphasis id="B91A4409EE6A7F02FF4CC1D4FEB0FC64" bold="true" box="[160,318,949,974]" italics="true" pageId="12" pageNumber="2474">Development</emphasis>
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF4CC1B8FB9FFB34" blockId="12.[160,1156,949,1182]" pageId="12" pageNumber="2474">
Complete sets (stages I to V, females and males) of the copepodid stages were present, as well as several nauplii. However, the complete morphological development of the naupliar stages could not be reconstructed. An example is illustrated (
<figureCitation id="1355849EEE6A7F02FC06C67FFBC5FB9C" box="[1002,1099,1054,1078]" captionStart="Figure 7" captionStartId="13.[160,225,1153,1176]" captionTargetBox="[163,1151,142,1122]" captionTargetId="figure-144@13.[163,1153,142,1122]" captionTargetPageId="13" captionText="Figure 7. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Undefined naupliar stage, in dorsal and ventral view; (c–e) habitus of copepodid stages I to V, respectively, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5183777" httpUri="https://zenodo.org/record/5183777/files/figure.png" pageId="12" pageNumber="2474">Figure 7</figureCitation>
(a,b)) demonstrating the extreme flattened body shape and the presence and position of an integumental window on the head shield. The nauplius, at least the older stadia, is feeding. A thin-walled internal tube stretching from mouth to anus is visible.
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF4CC6B1FD89FB43" blockId="12.[160,1156,1232,1499]" box="[160,519,1232,1257]" pageId="12" pageNumber="2474">
<emphasis id="B91A4409EE6A7F02FF4CC6B1FD89FB43" bold="true" box="[160,519,1232,1257]" italics="true" pageId="12" pageNumber="2474">Description of the copepodids</emphasis>
</paragraph>
<paragraph id="8BD1981BEE6A7F02FF4CC693FBBCFA3C" blockId="12.[160,1156,1232,1499]" pageId="12" pageNumber="2474">
<emphasis id="B91A4409EE6A7F02FF4CC693FE37FAA1" bold="true" box="[160,441,1266,1291]" italics="true" pageId="12" pageNumber="2474">
Habitus (
<figureCitation id="1355849EEE6A7F02FEE1C693FEE1FAA1" box="[269,367,1266,1291]" captionStart="Figure 7" captionStartId="13.[160,225,1153,1176]" captionTargetBox="[163,1151,142,1122]" captionTargetId="figure-144@13.[163,1153,142,1122]" captionTargetPageId="13" captionText="Figure 7. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Undefined naupliar stage, in dorsal and ventral view; (c–e) habitus of copepodid stages I to V, respectively, in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5183777" httpUri="https://zenodo.org/record/5183777/files/figure.png" pageId="12" pageNumber="2474">Figure 7</figureCitation>
(a–e)).
</emphasis>
The body develops through the usual five copepodid stages: from stage I with five body somites to stage CV (pre-adult stage) composed by nine somites; length increases from 350 μm in stage CI to 960 μm in stage CV (maximum, CV stage somewhat longer than adult); both sexes (recognizable in stages CIV and CV) equally sized; prosome and urosome clearly differentiated in stages CIV and CV.
</paragraph>
<paragraph id="8BD1981BEE6A7F03FF2CC7C0FB0BFA4E" blockId="12.[160,1156,1232,1499]" lastBlockId="13.[160,1157,1276,1508]" lastPageId="13" lastPageNumber="2475" pageId="12" pageNumber="2474">
Caudal rami with six armature elements in all stages: two lateral and three terminal ones, and a biarticulate dorsal seta; rami rather short (1.5 times longer than wide) in stage CI (
<figureCitation id="1355849EEE6B7F03FEF5C69DFEF0FABE" box="[281,382,1276,1300]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="13" pageNumber="2475">Figure 8</figureCitation>
(a,b)), with irregular medial margin; terminal armature elements inserted along caudal margin: outer one shorter than ramus, middle one 1.5 times longer than ramus, and inner one longest (at least three times longer than middle one, broken in all specimens); breaking plane absent; caudal rami in stage CII (
<figureCitation id="1355849EEE6B7F03FF4BC7E6FE84FA35" box="[167,266,1415,1439]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="13" pageNumber="2475">Figure 8</figureCitation>
(c)) 1.5 times longer than wide, with outer terminal seta twice as long as ramus, middle one fully developed, and inner one only slightly longer than ramus; breaking plane present in middle seta only; setal armament as in adult in stages CIII–CV.
</paragraph>
<caption id="DF11C893EE6A7F02FEACC477FD35F987" box="[320,699,1557,1581]" pageId="12" pageNumber="2474" startId="12.[320,376,1558,1581]">
<paragraph id="8BD1981BEE6A7F02FEACC477FD35F987" blockId="12.[320,699,1557,1581]" box="[320,699,1557,1581]" pageId="12" pageNumber="2474">
<emphasis id="B91A4409EE6A7F02FEACC477FE1DF987" bold="true" box="[320,403,1558,1581]" pageId="12" pageNumber="2474">Table 2.</emphasis>
Armament of male legs 1
<emphasis id="B91A4409EE6A7F02FD70C474FD27F986" box="[668,681,1557,1580]" italics="true" pageId="12" pageNumber="2474">–</emphasis>
4.
</paragraph>
</caption>
<paragraph id="8BD1981BEE6A7F02FEA7C45BFC5AF91E" pageId="12" pageNumber="2474">
<table id="F96E6ABBEE6A80F1FEA7C45BFC57F91E" box="[331,985,1594,1716]" gridcols="5" gridrows="5" pageId="12" pageNumber="2474" rowsContinueFrom="10.[327,981,1522,1644]">
<tr id="355E9A59EE6A80F1FEA7C45BFC57F9E7" box="[331,985,1594,1613]" gridrow="0" pageId="12" pageNumber="2474">
<th id="768FF325EE6A80F1FEA7C45BFEFBF9E7" box="[331,373,1594,1613]" gridcol="0" gridrow="0" pageId="12" pageNumber="2474">Male</th>
<th id="768FF325EE6A80F1FE21C45BFE7BF9E7" box="[461,501,1594,1613]" gridcol="1" gridrow="0" pageId="12" pageNumber="2474">Coxa</th>
<th id="768FF325EE6A80F1FDA1C45BFDFBF9E7" box="[589,629,1594,1613]" gridcol="2" gridrow="0" pageId="12" pageNumber="2474">Basis</th>
<th id="768FF325EE6A80F1FD22C45BFCAFF9E7" box="[718,801,1594,1613]" gridcol="3" gridrow="0" pageId="12" pageNumber="2474">Exopodite</th>
<th id="768FF325EE6A80F1FC95C45BFC57F9E7" box="[889,985,1594,1613]" gridcol="4" gridrow="0" pageId="12" pageNumber="2474">Endopodite</th>
</tr>
<tr id="355E9A59EE6A80F1FEA7C438FC57F9C6" box="[331,985,1625,1644]" gridrow="1" pageId="12" pageNumber="2474">
<th id="768FF325EE6A80F1FEA7C438FEFBF9C6" box="[331,373,1625,1644]" gridcol="0" gridrow="1" pageId="12" pageNumber="2474">leg 1</th>
<td id="768FF325EE6A80F1FE21C438FE7BF9C6" box="[461,501,1625,1644]" gridcol="1" gridrow="1" pageId="12" pageNumber="2474">0-1</td>
<td id="768FF325EE6A80F1FDA1C438FDFBF9C6" box="[589,629,1625,1644]" gridcol="2" gridrow="1" pageId="12" pageNumber="2474">1-I</td>
<td id="768FF325EE6A80F1FD22C438FCAFF9C6" box="[718,801,1625,1644]" gridcol="3" gridrow="1" pageId="12" pageNumber="2474">I-0; III-2-4</td>
<td id="768FF325EE6A80F1FC95C438FC57F9C6" box="[889,985,1625,1644]" gridcol="4" gridrow="1" pageId="12" pageNumber="2474">0-1; 1-I1-1</td>
</tr>
<tr id="355E9A59EE6A80F1FEA7C413FC57F92F" box="[331,985,1650,1669]" gridrow="2" pageId="12" pageNumber="2474">
<th id="768FF325EE6A80F1FEA7C413FEFBF92F" box="[331,373,1650,1669]" gridcol="0" gridrow="2" pageId="12" pageNumber="2474">leg 2</th>
<td id="768FF325EE6A80F1FE21C413FE7BF92F" box="[461,501,1650,1669]" gridcol="1" gridrow="2" pageId="12" pageNumber="2474">0-0</td>
<td id="768FF325EE6A80F1FDA1C413FDFBF92F" box="[589,629,1650,1669]" gridcol="2" gridrow="2" pageId="12" pageNumber="2474">1-0</td>
<td id="768FF325EE6A80F1FD22C413FCAFF92F" box="[718,801,1650,1669]" gridcol="3" gridrow="2" pageId="12" pageNumber="2474">I-0; II-I1-4</td>
<td id="768FF325EE6A80F1FC95C413FC57F92F" box="[889,985,1650,1669]" gridcol="4" gridrow="2" pageId="12" pageNumber="2474">0-1; 1-I1-2</td>
</tr>
<tr id="355E9A59EE6A80F1FEA7C4EBFC57F937" box="[331,985,1674,1693]" gridrow="3" pageId="12" pageNumber="2474">
<th id="768FF325EE6A80F1FEA7C4EBFEFBF937" box="[331,373,1674,1693]" gridcol="0" gridrow="3" pageId="12" pageNumber="2474">leg 3</th>
<td id="768FF325EE6A80F1FE21C4EBFE7BF937" box="[461,501,1674,1693]" gridcol="1" gridrow="3" pageId="12" pageNumber="2474">0-0</td>
<td id="768FF325EE6A80F1FDA1C4EBFDFBF937" box="[589,629,1674,1693]" gridcol="2" gridrow="3" pageId="12" pageNumber="2474">1-0</td>
<td id="768FF325EE6A80F1FD22C4EBFCAFF937" box="[718,801,1674,1693]" gridcol="3" gridrow="3" pageId="12" pageNumber="2474">I-0; II-I1-4</td>
<td id="768FF325EE6A80F1FC95C4EBFC57F937" box="[889,985,1674,1693]" gridcol="4" gridrow="3" pageId="12" pageNumber="2474">0-1; 1-I0-3</td>
</tr>
<tr id="355E9A59EE6A80F1FEA7C4C0FC57F91E" box="[331,985,1697,1716]" gridrow="4" pageId="12" pageNumber="2474">
<th id="768FF325EE6A80F1FEA7C4C0FEFBF91E" box="[331,373,1697,1716]" gridcol="0" gridrow="4" pageId="12" pageNumber="2474">leg 4</th>
<td id="768FF325EE6A80F1FE21C4C0FE7BF91E" box="[461,501,1697,1716]" gridcol="1" gridrow="4" pageId="12" pageNumber="2474">0-0</td>
<td id="768FF325EE6A80F1FDA1C4C0FDFBF91E" box="[589,629,1697,1716]" gridcol="2" gridrow="4" pageId="12" pageNumber="2474">1-0</td>
<td id="768FF325EE6A80F1FD22C4C0FCAFF91E" box="[718,801,1697,1716]" gridcol="3" gridrow="4" pageId="12" pageNumber="2474">I-0; II-I1-3</td>
<td id="768FF325EE6A80F1FC95C4C0FC57F91E" box="[889,985,1697,1716]" gridcol="4" gridrow="4" pageId="12" pageNumber="2474">0-1; 1-I1-1</td>
</tr>
</table>
</paragraph>
<caption id="DF11C893EE6B7F03FF4CC6E0FC6EFB1F" ID-DOI="http://doi.org/10.5281/zenodo.5183777" ID-Zenodo-Dep="5183777" httpUri="https://zenodo.org/record/5183777/files/figure.png" pageId="13" pageNumber="2475" startId="13.[160,225,1153,1176]" targetBox="[163,1151,142,1122]" targetPageId="13">
<paragraph id="8BD1981BEE6B7F03FF4CC6E0FC6EFB1F" blockId="13.[160,1155,1152,1205]" pageId="13" pageNumber="2475">
<emphasis id="B91A4409EE6B7F03FF4CC6E0FF75FB32" bold="true" box="[160,251,1153,1176]" pageId="13" pageNumber="2475">Figure 7.</emphasis>
<taxonomicName id="4C6EE398EE6B7F03FEEFC6E0FE72FB32" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[259,508,1153,1176]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="13" pageNumber="2475" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE6B7F03FEEFC6E0FE72FB32" box="[259,508,1153,1176]" italics="true" pageId="13" pageNumber="2475">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE6B7F03FDEFC6E0FD34FB32" box="[515,698,1153,1176]" pageId="13" pageNumber="2475" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a, b) Undefined naupliar stage, in dorsal and ventral view; (c
<emphasis id="B91A4409EE6B7F03FED5C6FFFEC8FB1F" box="[313,326,1182,1205]" italics="true" pageId="13" pageNumber="2475">–</emphasis>
e) habitus of copepodid stages I to V, respectively, in dorsal view.
</paragraph>
</caption>
<paragraph id="8BD1981BEE6B7F03FF4CC447FEA8F90D" blockId="13.[160,1156,1574,1703]" pageId="13" pageNumber="2475">
<emphasis id="B91A4409EE6B7F03FF4CC447FD5BF995" bold="true" box="[160,725,1574,1599]" italics="true" pageId="13" pageNumber="2475">
Female antennule (
<figureCitation id="1355849EEE6B7F03FE66C447FE77F995" box="[394,505,1574,1599]" captionStart="Figure 9" captionStartId="15.[160,225,1450,1473]" captionTargetBox="[163,1153,142,1419]" captionTargetId="figure-58@15.[163,1153,142,1419]" captionTargetPageId="15" captionText="Figure 9. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antenna in copepodid stage I, caudal; (b) idem, in copepodid stage II; (c) idem, in copepodid stage III; (d) antennule in copepodid stage I; (e) idem, in copepodid stage II." figureDoi="http://doi.org/10.5281/zenodo.5183781" httpUri="https://zenodo.org/record/5183781/files/figure.png" pageId="13" pageNumber="2475">Figures 9</figureCitation>
(d,e) and 10(a–c)).
</emphasis>
Number of segments evolves in the successive copepodid stages from six-segmented (Stage CI) over seven-segmented (stage CII) and nine-segmented (stage CIII) to the adult ten-segmented shape in stages CIV and CV.
</paragraph>
<caption id="DF11C893EE687F00FF4CC479FBF2F92D" ID-DOI="http://doi.org/10.5281/zenodo.5183779" ID-Zenodo-Dep="5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="14" pageNumber="2476" startId="14.[160,225,1560,1583]" targetBox="[163,1153,142,1528]" targetPageId="14">
<paragraph id="8BD1981BEE687F00FF4CC479FBF2F92D" blockId="14.[160,1156,1559,1671]" pageId="14" pageNumber="2476">
<emphasis id="B91A4409EE687F00FF4CC479FF73F985" bold="true" box="[160,253,1560,1583]" pageId="14" pageNumber="2476">Figure 8.</emphasis>
<taxonomicName id="4C6EE398EE687F00FEEAC479FD8FF985" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[262,513,1560,1583]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="14" pageNumber="2476" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE687F00FEEAC479FD8FF985" box="[262,513,1560,1583]" italics="true" pageId="14" pageNumber="2476">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE687F00FDE7C479FD47F985" box="[523,713,1560,1583]" pageId="14" pageNumber="2476" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Postcephalic somites 2
<emphasis id="B91A4409EE687F00FC37C476FC66F984" box="[987,1000,1559,1582]" italics="true" pageId="14" pageNumber="2476">–</emphasis>
4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral.
</paragraph>
</caption>
<caption id="DF11C893EE697F01FF4CC7CBFE34FA56" ID-DOI="http://doi.org/10.5281/zenodo.5183781" ID-Zenodo-Dep="5183781" httpUri="https://zenodo.org/record/5183781/files/figure.png" pageId="15" pageNumber="2477" startId="15.[160,225,1450,1473]" targetBox="[163,1153,142,1419]" targetPageId="15">
<paragraph id="8BD1981BEE697F01FF4CC7CBFE34FA56" blockId="15.[160,1156,1450,1532]" pageId="15" pageNumber="2477">
<emphasis id="B91A4409EE697F01FF4CC7CBFF75FA6B" bold="true" box="[160,251,1450,1473]" pageId="15" pageNumber="2477">Figure 9.</emphasis>
<taxonomicName id="4C6EE398EE697F01FEE8C7CBFE73FA6B" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[260,509,1450,1473]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="15" pageNumber="2477" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE697F01FEE8C7CBFE73FA6B" box="[260,509,1450,1473]" italics="true" pageId="15" pageNumber="2477">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE697F01FDE9C7CBFD33FA6B" box="[517,701,1450,1473]" pageId="15" pageNumber="2477" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Antenna in copepodid stage I, caudal; (b) idem, in copepodid stage II; (c) idem, in copepodid stage III; (d) antennule in copepodid stage I; (e) idem, in copepodid stage II.
</paragraph>
</caption>
<paragraph id="8BD1981BEE697F1EFF2CC45DFB0DF904" blockId="15.[160,1155,1596,1724]" lastBlockId="16.[160,1155,1582,1710]" lastPageId="16" lastPageNumber="2478" pageId="15" pageNumber="2477">
Terminal (sixth) segment of the stage CI antennule possesses, apart from its setal complement, an apical hyaline lip-shaped structure that disappears during the moult to stage CII and remains absent in the following stages; aesthetascs present from stage C1 on (segments 3, 5, 6) being filiform; linguiform shape of aesthetascs appears first in stage CII on segment 4 (=segment
<quantity id="4C9635FEEE767F1EFE2CC44FFE66F9EC" box="[448,488,1582,1606]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="2478" unit="in" value="3.0">3 in</quantity>
stage C1) and on segment 7 (terminal segment); aesthetasc on segment 5 of stage CI remaining filiform through successive stages up to and including the adult; setal presence in successive stages summarized in
<tableCitation id="C6ECADA0EE767F1EFCA5C412FC14F921" box="[841,922,1651,1675]" captionStart="Table 3" captionStartId="17.[160,216,142,165]" captionText="Table 3. Antennulary armament in the successive female copepodid stages and the adult, projected against the ancestral antennulary segment sequences." pageId="16" pageNumber="2478">Table 3</tableCitation>
; growth by segment addition resulting from the subdividing of the antennular segments of stage CI: first
</paragraph>
<caption id="DF11C893EE767F1EFF4CC7A4FC36FA50" ID-DOI="http://doi.org/10.5281/zenodo.5183783" ID-Zenodo-Dep="5183783" httpUri="https://zenodo.org/record/5183783/files/figure.png" pageId="16" pageNumber="2478" startId="16.[160,225,1477,1500]" targetBox="[163,1151,149,1446]" targetPageId="16">
<paragraph id="8BD1981BEE767F1EFF4CC7A4FC36FA50" blockId="16.[160,1156,1477,1530]" pageId="16" pageNumber="2478">
<emphasis id="B91A4409EE767F1EFF4CC7A4FE85FA76" bold="true" box="[160,267,1477,1500]" pageId="16" pageNumber="2478">Figure 10.</emphasis>
<taxonomicName id="4C6EE398EE767F1EFEFFC7A4FD81FA76" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[275,527,1477,1500]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="16" pageNumber="2478" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE767F1EFEFFC7A4FD81FA76" box="[275,527,1477,1500]" italics="true" pageId="16" pageNumber="2478">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE767F1EFDF5C7A4FD57FA76" box="[537,729,1477,1500]" pageId="16" pageNumber="2478" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Antennule in copepodid stage III; (b) idem, in copepodid stage IV (female); (c) idem, in copepodid stage V (female).
</paragraph>
</caption>
<caption id="DF11C893EE777F1FFF4CC2EFFD4CFF68" pageId="17" pageNumber="2479" startId="17.[160,216,142,165]">
<paragraph id="8BD1981BEE777F1FFF4CC2EFFD4CFF68" blockId="17.[160,1156,142,194]" pageId="17" pageNumber="2479">
<emphasis id="B91A4409EE777F1FFF4CC2EFFF7FFF0F" bold="true" box="[160,241,142,165]" pageId="17" pageNumber="2479">Table 3.</emphasis>
Antennulary armament in the successive female copepodid stages and the adult, projected against the ancestral antennulary segment sequences.
</paragraph>
</caption>
<paragraph id="8BD1981BEE777F1FFF2BC2AEFBF6FE00" pageId="17" pageNumber="2479">
<table id="F96E6ABBEE7780F1FF4CC2AEFBF7FE00" box="[160,1145,207,426]" gridcols="12" gridrows="9" pageId="17" pageNumber="2479">
<tr id="355E9A59EE7780F1FF4CC2AEFBF7FF48" box="[160,1145,207,226]" gridrow="0" pageId="17" pageNumber="2479">
<th id="768FF325EE7780F1FF4CC2AEFED0FF48" box="[160,350,207,226]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="17" pageNumber="2479">Segm. no. in ad.</th>
<th id="768FF325EE7780F1FE4AC2AEFE3FFF48" box="[422,433,207,226]" gridcol="2" gridrow="0" pageId="17" pageNumber="2479">1</th>
<th id="768FF325EE7780F1FE31C2AEFE63FF48" box="[477,493,207,226]" gridcol="3" gridrow="0" pageId="17" pageNumber="2479">2</th>
<th id="768FF325EE7780F1FDF5C2AEFDBDFF48" box="[537,563,207,226]" gridcol="4" gridrow="0" pageId="17" pageNumber="2479">3</th>
<th id="768FF325EE7780F1FDB3C2AEFDF4FF48" box="[607,634,207,226]" gridcol="5" gridrow="0" pageId="17" pageNumber="2479">4</th>
<th id="768FF325EE7780F1FD4BC2AEFD4CFF48" box="[679,706,207,226]" gridcol="6" gridrow="0" pageId="17" pageNumber="2479">5</th>
<th id="768FF325EE7780F1FD02C2AEFC80FF48" box="[750,782,207,226]" gridcol="7" gridrow="0" pageId="17" pageNumber="2479">6</th>
<th id="768FF325EE7780F1FCD6C2AEFCE5FF48" box="[826,875,207,226]" gridcol="8" gridrow="0" pageId="17" pageNumber="2479">7</th>
<th id="768FF325EE7780F1FC74C2AEFC30FF48" box="[920,958,207,226]" gridcol="9" gridrow="0" pageId="17" pageNumber="2479">8</th>
<th id="768FF325EE7780F1FC06C2AEFB95FF48" box="[1002,1051,207,226]" gridcol="10" gridrow="0" pageId="17" pageNumber="2479">9</th>
<th id="768FF325EE7780F1FBA4C2AEFBF7FF48" box="[1096,1145,207,226]" gridcol="11" gridrow="0" pageId="17" pageNumber="2479">10</th>
</tr>
<tr id="355E9A59EE7780F1FF4CC28EFBF7FEA8" box="[160,1145,239,258]" gridrow="1" pageId="17" pageNumber="2479" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="768FF325EE7780F1FF4CC28EFF5DFEA8" box="[160,211,239,258]" gridcol="0" gridrow="1" pageId="17" pageNumber="2479">6</th>
<td id="768FF325EE7780F1FECBC28EFED0FEA8" box="[295,350,239,258]" gridcol="1" gridrow="1" pageId="17" pageNumber="2479">C I</td>
<td id="768FF325EE7780F1FE4AC28EFE3FFEA8" box="[422,433,239,258]" gridcol="2" gridrow="1" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FDF5C28EFDBDFEA8" box="[537,563,239,258]" gridcol="4" gridrow="1" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FCD6C28EFCE5FEA8" box="[826,875,239,258]" gridcol="8" gridrow="1" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C28EFC30FEA8" box="[920,958,239,258]" gridcol="9" gridrow="1" pageId="17" pageNumber="2479">1</td>
<td id="768FF325EE7780F1FC06C28EFB95FEA8" box="[1002,1051,239,258]" gridcol="10" gridrow="1" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C28EFBF7FEA8" box="[1096,1145,239,258]" gridcol="11" gridrow="1" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC366FBF7FEB0" box="[160,1145,263,282]" gridrow="2" pageId="17" pageNumber="2479" rowspan-3="1" rowspan-5="1" rowspan-6="1">
<th id="768FF325EE7780F1FF4CC366FF5DFEB0" box="[160,211,263,282]" gridcol="0" gridrow="2" pageId="17" pageNumber="2479">7</th>
<td id="768FF325EE7780F1FECBC366FED0FEB0" box="[295,350,263,282]" gridcol="1" gridrow="2" pageId="17" pageNumber="2479">C II</td>
<td id="768FF325EE7780F1FE4AC366FE3FFEB0" box="[422,433,263,282]" gridcol="2" gridrow="2" pageId="17" pageNumber="2479">6</td>
<td id="768FF325EE7780F1FDF5C366FDBDFEB0" box="[537,563,263,282]" gridcol="4" gridrow="2" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD02C366FC80FEB0" box="[750,782,263,282]" gridcol="7" gridrow="2" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FCD6C366FCE5FEB0" box="[826,875,263,282]" gridcol="8" gridrow="2" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C366FC30FEB0" box="[920,958,263,282]" gridcol="9" gridrow="2" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FC06C366FB95FEB0" box="[1002,1051,263,282]" gridcol="10" gridrow="2" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C366FBF7FEB0" box="[1096,1145,263,282]" gridcol="11" gridrow="2" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC37EFBF7FE98" box="[160,1145,287,306]" gridrow="3" pageId="17" pageNumber="2479" rowspan-5="1">
<th id="768FF325EE7780F1FF4CC37EFF5DFE98" box="[160,211,287,306]" gridcol="0" gridrow="3" pageId="17" pageNumber="2479">9</th>
<td id="768FF325EE7780F1FECBC37EFED0FE98" box="[295,350,287,306]" gridcol="1" gridrow="3" pageId="17" pageNumber="2479">C III</td>
<td id="768FF325EE7780F1FE4AC37EFE3FFE98" box="[422,433,287,306]" gridcol="2" gridrow="3" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FE31C37EFE63FE98" box="[477,493,287,306]" gridcol="3" gridrow="3" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FDF5C37EFDBDFE98" box="[537,563,287,306]" gridcol="4" gridrow="3" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD4BC37EFD4CFE98" box="[679,706,287,306]" gridcol="6" gridrow="3" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD02C37EFC80FE98" box="[750,782,287,306]" gridcol="7" gridrow="3" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FCD6C37EFCE5FE98" box="[826,875,287,306]" gridcol="8" gridrow="3" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C37EFC30FE98" box="[920,958,287,306]" gridcol="9" gridrow="3" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FC06C37EFB95FE98" box="[1002,1051,287,306]" gridcol="10" gridrow="3" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C37EFBF7FE98" box="[1096,1145,287,306]" gridcol="11" gridrow="3" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC356FBF7FEE0" box="[160,1145,311,330]" gridrow="4" pageId="17" pageNumber="2479">
<th id="768FF325EE7780F1FF4CC356FF5DFEE0" box="[160,211,311,330]" gridcol="0" gridrow="4" pageId="17" pageNumber="2479">10</th>
<td id="768FF325EE7780F1FECBC356FED0FEE0" box="[295,350,311,330]" gridcol="1" gridrow="4" pageId="17" pageNumber="2479">C IV</td>
<td id="768FF325EE7780F1FE4AC356FE3FFEE0" box="[422,433,311,330]" gridcol="2" gridrow="4" pageId="17" pageNumber="2479">5</td>
<td id="768FF325EE7780F1FE31C356FE63FEE0" box="[477,493,311,330]" gridcol="3" gridrow="4" pageId="17" pageNumber="2479">6</td>
<td id="768FF325EE7780F1FDF5C356FDBDFEE0" box="[537,563,311,330]" gridcol="4" gridrow="4" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FDB3C356FDF4FEE0" box="[607,634,311,330]" gridcol="5" gridrow="4" pageId="17" pageNumber="2479">1</td>
<td id="768FF325EE7780F1FD4BC356FD4CFEE0" box="[679,706,311,330]" gridcol="6" gridrow="4" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD02C356FC80FEE0" box="[750,782,311,330]" gridcol="7" gridrow="4" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FCD6C356FCE5FEE0" box="[826,875,311,330]" gridcol="8" gridrow="4" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C356FC30FEE0" box="[920,958,311,330]" gridcol="9" gridrow="4" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FC06C356FB95FEE0" box="[1002,1051,311,330]" gridcol="10" gridrow="4" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C356FBF7FEE0" box="[1096,1145,311,330]" gridcol="11" gridrow="4" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC32EFBF7FEC8" box="[160,1145,335,354]" gridrow="5" pageId="17" pageNumber="2479">
<th id="768FF325EE7780F1FF4CC32EFF5DFEC8" box="[160,211,335,354]" gridcol="0" gridrow="5" pageId="17" pageNumber="2479">10</th>
<td id="768FF325EE7780F1FECBC32EFED0FEC8" box="[295,350,335,354]" gridcol="1" gridrow="5" pageId="17" pageNumber="2479">C V</td>
<td id="768FF325EE7780F1FE4AC32EFE3FFEC8" box="[422,433,335,354]" gridcol="2" gridrow="5" pageId="17" pageNumber="2479">6</td>
<td id="768FF325EE7780F1FE31C32EFE63FEC8" box="[477,493,335,354]" gridcol="3" gridrow="5" pageId="17" pageNumber="2479">7</td>
<td id="768FF325EE7780F1FDF5C32EFDBDFEC8" box="[537,563,335,354]" gridcol="4" gridrow="5" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FDB3C32EFDF4FEC8" box="[607,634,335,354]" gridcol="5" gridrow="5" pageId="17" pageNumber="2479">1</td>
<td id="768FF325EE7780F1FD4BC32EFD4CFEC8" box="[679,706,335,354]" gridcol="6" gridrow="5" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD02C32EFC80FEC8" box="[750,782,335,354]" gridcol="7" gridrow="5" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FCD6C32EFCE5FEC8" box="[826,875,335,354]" gridcol="8" gridrow="5" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C32EFC30FEC8" box="[920,958,335,354]" gridcol="9" gridrow="5" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FC06C32EFB95FEC8" box="[1002,1051,335,354]" gridcol="10" gridrow="5" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C32EFBF7FEC8" box="[1096,1145,335,354]" gridcol="11" gridrow="5" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC306FBF7FED0" box="[160,1145,359,378]" gridrow="6" pageId="17" pageNumber="2479">
<th id="768FF325EE7780F1FF4CC306FF5DFED0" box="[160,211,359,378]" gridcol="0" gridrow="6" pageId="17" pageNumber="2479">
9
<emphasis id="B91A4409EE777F1FFF59C306FF4EFED0" box="[181,192,359,378]" italics="true" pageId="17" pageNumber="2479">–</emphasis>
10
</th>
<td id="768FF325EE7780F1FECBC306FED0FED0" box="[295,350,359,378]" gridcol="1" gridrow="6" pageId="17" pageNumber="2479">adult</td>
<td id="768FF325EE7780F1FE4AC306FE3FFED0" box="[422,433,359,378]" gridcol="2" gridrow="6" pageId="17" pageNumber="2479">6</td>
<td id="768FF325EE7780F1FE31C306FE63FED0" box="[477,493,359,378]" gridcol="3" gridrow="6" pageId="17" pageNumber="2479">7</td>
<td id="768FF325EE7780F1FDF5C306FDBDFED0" box="[537,563,359,378]" gridcol="4" gridrow="6" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FDB3C306FDF4FED0" box="[607,634,359,378]" gridcol="5" gridrow="6" pageId="17" pageNumber="2479">1</td>
<td id="768FF325EE7780F1FD4BC306FD4CFED0" box="[679,706,359,378]" gridcol="6" gridrow="6" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FD02C306FC80FED0" box="[750,782,359,378]" gridcol="7" gridrow="6" pageId="17" pageNumber="2479">3</td>
<td id="768FF325EE7780F1FCD6C306FCE5FED0" box="[826,875,359,378]" gridcol="8" gridrow="6" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FC74C306FC30FED0" box="[920,958,359,378]" gridcol="9" gridrow="6" pageId="17" pageNumber="2479">2</td>
<td id="768FF325EE7780F1FC06C306FB95FED0" box="[1002,1051,359,378]" gridcol="10" gridrow="6" pageId="17" pageNumber="2479">2 + A</td>
<td id="768FF325EE7780F1FBA4C306FBF7FED0" box="[1096,1145,359,378]" gridcol="11" gridrow="6" pageId="17" pageNumber="2479">7 + A</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC31EFBF7FE38" box="[160,1145,383,402]" gridrow="7" pageId="17" pageNumber="2479">
<th id="768FF325EE7780F1FF4CC31EFED0FE38" box="[160,350,383,402]" colspan="2" colspanRight="1" gridcol="0" gridrow="7" pageId="17" pageNumber="2479">Ancestral segment no.</th>
<td id="768FF325EE7780F1FE4AC31EFE3FFE38" box="[422,433,383,402]" gridcol="2" gridrow="7" pageId="17" pageNumber="2479">I</td>
<td id="768FF325EE7780F1FE31C31EFE63FE38" box="[477,493,383,402]" gridcol="3" gridrow="7" pageId="17" pageNumber="2479">VI</td>
<td id="768FF325EE7780F1FDF5C31EFDBDFE38" box="[537,563,383,402]" gridcol="4" gridrow="7" pageId="17" pageNumber="2479">XII</td>
<td id="768FF325EE7780F1FDB3C31EFDF4FE38" box="[607,634,383,402]" gridcol="5" gridrow="7" pageId="17" pageNumber="2479">XIV</td>
<td id="768FF325EE7780F1FD4BC31EFD4CFE38" box="[679,706,383,402]" gridcol="6" gridrow="7" pageId="17" pageNumber="2479">XV</td>
<td id="768FF325EE7780F1FD02C31EFC80FE38" box="[750,782,383,402]" gridcol="7" gridrow="7" pageId="17" pageNumber="2479">XVII</td>
<td id="768FF325EE7780F1FCD6C31EFCE5FE38" box="[826,875,383,402]" gridcol="8" gridrow="7" pageId="17" pageNumber="2479">XXI</td>
<td id="768FF325EE7780F1FC74C31EFC30FE38" box="[920,958,383,402]" gridcol="9" gridrow="7" pageId="17" pageNumber="2479">XXIV</td>
<td id="768FF325EE7780F1FC06C31EFB95FE38" box="[1002,1051,383,402]" gridcol="10" gridrow="7" pageId="17" pageNumber="2479">XXV</td>
<td id="768FF325EE7780F1FBA4C31EFBF7FE38" box="[1096,1145,383,402]" gridcol="11" gridrow="7" pageId="17" pageNumber="2479">XXVI</td>
</tr>
<tr id="355E9A59EE7780F1FF4CC3F6FBF7FE00" box="[160,1145,407,426]" gridrow="8" pageId="17" pageNumber="2479" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-5="1" rowspan-9="1">
<td id="768FF325EE7780F1FE4AC3F6FE3FFE00" box="[422,433,407,426]" gridcol="2" gridrow="8" pageId="17" pageNumber="2479">V</td>
<td id="768FF325EE7780F1FE31C3F6FE63FE00" box="[477,493,407,426]" gridcol="3" gridrow="8" pageId="17" pageNumber="2479">XI</td>
<td id="768FF325EE7780F1FDF5C3F6FDBDFE00" box="[537,563,407,426]" gridcol="4" gridrow="8" pageId="17" pageNumber="2479">XIII</td>
<td id="768FF325EE7780F1FD4BC3F6FD4CFE00" box="[679,706,407,426]" gridcol="6" gridrow="8" pageId="17" pageNumber="2479">XVI</td>
<td id="768FF325EE7780F1FD02C3F6FC80FE00" box="[750,782,407,426]" gridcol="7" gridrow="8" pageId="17" pageNumber="2479">XX</td>
<td id="768FF325EE7780F1FCD6C3F6FCE5FE00" box="[826,875,407,426]" gridcol="8" gridrow="8" pageId="17" pageNumber="2479">XXIII</td>
<td id="768FF325EE7780F1FBA4C3F6FBF7FE00" box="[1096,1145,407,426]" gridcol="11" gridrow="8" pageId="17" pageNumber="2479">XXVIII</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD1981BEE777F1FFF4CC3D8FE05FE66" blockId="17.[160,395,441,460]" box="[160,395,441,460]" pageId="17" pageNumber="2479">
<tableNote id="76889995EE777F1FFF4CC3D8FE05FE66" box="[160,395,441,460]" pageId="17" pageNumber="2479" targetBox="[160,1145,207,426]" targetPageId="17">Abbreviation: A, aesthetasc.</tableNote>
</paragraph>
<paragraph id="8BD1981BEE777F1FFF4CC069FE95FD22" blockId="17.[160,1156,520,648]" pageId="17" pageNumber="2479">
segment (in moult from stage CII to CIII) and stage I second segment (in moults from stage CI to CII, from stage CII to CIII, and from stage CIII to CIV); segmentation between segments 1 and
<quantity id="4C9635FEEE777F1FFF09C02CFE80FDCF" box="[229,270,589,613]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="17" pageNumber="2479" unit="in" value="2.0">2 in</quantity>
stage CV disappearing partially during terminal moult (CV to adult: dashed line in
<tableCitation id="C6ECADA0EE777F1FFF50C011FE80FD22" box="[188,270,624,648]" captionStart="Table 3" captionStartId="17.[160,216,142,165]" captionText="Table 3. Antennulary armament in the successive female copepodid stages and the adult, projected against the ancestral antennulary segment sequences." pageId="17" pageNumber="2479">Table 3</tableCitation>
).
</paragraph>
<paragraph id="8BD1981BEE777F1FFF4CC0D7FC17FC4E" blockId="17.[160,1156,694,996]" pageId="17" pageNumber="2479">
<emphasis id="B91A4409EE777F1FFF4CC0D7FDABFD65" bold="true" box="[160,549,694,719]" italics="true" pageId="17" pageNumber="2479">
Male antennule (
<figureCitation id="1355849EEE777F1FFE83C0D7FE52FD65" box="[367,476,694,719]" captionStart="Figure 11" captionStartId="18.[160,225,1558,1581]" captionTargetBox="[163,1153,142,1527]" captionTargetId="figure-1@18.[163,1153,142,1527]" captionTargetPageId="18" captionText="Figure 11. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183785" httpUri="https://zenodo.org/record/5183785/files/figure.png" pageId="17" pageNumber="2479">Figure 11</figureCitation>
(a,b)).
</emphasis>
Before stage CIV, no differences in shape and setation between male and female antennule copepodid antennules observed; stage CIV of male antennule resembling female stage CIV antennule, except for the slightly more robust appearance of the segments and the presence of a small additional (hyaline) protuberance on segment 6; moult between stage CIV and CV affecting the antennular shape and armament drastically: separation of stage CIV antennulary segments 3 and 4, and 5 and 6 is deleted (resulting in eight-segmented antennule), number of setae on segment 2 increases (see
<tableCitation id="C6ECADA0EE777F1FFE2AC1CBFD94FC68" box="[454,538,938,962]" captionStart="Table 4" captionStartId="19.[133,189,817,840]" captionText="Table 4. Segment and armament evolution of male antennule from stage III to adult, compared with the ancestral antennulary segment sequence." pageId="17" pageNumber="2479">Table 4</tableCitation>
), and protuberances on segments 4 and 5 increase in number and shape; armament evolution is summarized in
<tableCitation id="C6ECADA0EE777F1FFCACC1ADFC1AFC4E" box="[832,916,972,996]" captionStart="Table 4" captionStartId="19.[133,189,817,840]" captionText="Table 4. Segment and armament evolution of male antennule from stage III to adult, compared with the ancestral antennulary segment sequence." pageId="17" pageNumber="2479">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BD1981BEE777F1FFF4CC673FB0BFAB4" blockId="17.[160,1157,1042,1448]" pageId="17" pageNumber="2479">
<emphasis id="B91A4409EE777F1FFF4CC673FD07FB81" bold="true" box="[160,649,1042,1067]" italics="true" pageId="17" pageNumber="2479">
Antenna and mouthparts (
<figureCitation id="1355849EEE777F1FFE33C673FDB1FB81" box="[479,575,1042,1067]" captionStart="Figure 9" captionStartId="15.[160,225,1450,1473]" captionTargetBox="[163,1153,142,1419]" captionTargetId="figure-58@15.[163,1153,142,1419]" captionTargetPageId="15" captionText="Figure 9. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antenna in copepodid stage I, caudal; (b) idem, in copepodid stage II; (c) idem, in copepodid stage III; (d) antennule in copepodid stage I; (e) idem, in copepodid stage II." figureDoi="http://doi.org/10.5281/zenodo.5183781" httpUri="https://zenodo.org/record/5183781/files/figure.png" pageId="17" pageNumber="2479">Figure 9</figureCitation>
(a–c)).
</emphasis>
Antenna biramous at stage CI with a threesegmented endopodite and a non-segmented exopodal process composed of a cluster of cells forming a cylindrical outcrop with two slender setal elements inserted vaguely in the middle of cellular mass; basipodite and endopodite resembling adult morphology but latter with fewer setae (four on second endopodite segment, six on third segment); exopodal cellular mass and additional setal elements absent from stage CII onward; adult number of seta (five) on second endopodite segment present from stage CIII on, adult number of setae (seven) on terminal endopodite segment present from stage CIV.
</paragraph>
<paragraph id="8BD1981BEE777F1FFF2CC749FC4AFA02" blockId="17.[160,1157,1042,1448]" pageId="17" pageNumber="2479">Mandible, maxillule and maxilla and associated labrum completely shaped from stage C1 on, resembling the adult morphology in every detail; maxilliped in stage 1 represented as a small bud (about one-quarter of maxillary length) with four long setal appendages along inner side; maxilliped as in adult from stage C2 on.</paragraph>
<paragraph id="8BD1981BEE777F1FFF4CC7B7FE15F99F" blockId="17.[160,1155,1494,1727]" pageId="17" pageNumber="2479">
<emphasis id="B91A4409EE777F1FFF4CC7B7FD4BFA45" bold="true" box="[160,709,1494,1519]" italics="true" pageId="17" pageNumber="2479">
Development of legs 1–4 (
<figureCitation id="1355849EEE777F1FFE31C7B7FDD7FA45" box="[477,601,1494,1519]" captionStart="Figure 12" captionStartId="20.[160,225,1326,1349]" captionTargetBox="[163,1153,142,1295]" captionTargetId="figure-95@20.[163,1153,142,1295]" captionTargetPageId="20" captionText="Figure 12. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage I: (a) leg 1, caudal; (b) leg 2, caudal (endopodite illustrated separately); (c–e) Copepodid stage II: (c) leg 1, frontal; (d) leg 2, frontal; (e) leg 3, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183787" httpUri="https://zenodo.org/record/5183787/files/figure.png" pageId="17" pageNumber="2479">Figures 12</figureCitation>
and
<figureCitation id="1355849EEE777F1FFD76C7B7FD39FA45" box="[666,695,1494,1519]" captionStart="Figure 13" captionStartId="21.[160,225,1550,1573]" captionTargetBox="[163,1153,142,1519]" captionTargetId="figure-1@21.[163,1153,142,1519]" captionTargetPageId="21" captionText="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." figureDoi="http://doi.org/10.5281/zenodo.5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="17" pageNumber="2479">13</figureCitation>
).
</emphasis>
Leg development (i.e. segment differentiation and setal additions) truncated as in
<taxonomicName id="4C6EE398EE777F1FFD58C79BFBB6F9B8" authority="Bjornberg, 1985" authorityName="Bjornberg" authorityYear="1985" box="[692,1080,1530,1554]" class="Hexanauplia" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="17" pageNumber="2479" phylum="Arthropoda" rank="species" species="caroli">
<emphasis id="B91A4409EE777F1FFD58C79BFCF6F9B8" box="[692,888,1530,1554]" italics="true" pageId="17" pageNumber="2479">Bryocyclops caroli</emphasis>
<bibRefCitation id="EFFFE5EAEE777F1FFC6EC79BFBB6F9B8" author="Bjornberg MHGC" box="[898,1080,1530,1554]" pageId="17" pageNumber="2479" pagination="237 - 241" refId="ref17618" refString="Bjornberg MHGC. 1985. Bryocyclops caroli sp. n. (Crustacea, Copepoda, Cyclopoida), the first representative of the genus in South America. Hydrobiologia. 124: 237 - 241." type="journal article" year="1985">Bjornberg, 1985</bibRefCitation>
</taxonomicName>
, tabulated in
<bibRefCitation id="EFFFE5EAEE777F1FFF13C47CFE19F99F" author="Ferrari FD" box="[255,407,1565,1589]" pageId="17" pageNumber="2479" pagination="256 - 293" refId="ref18140" refString="Ferrari FD. 1988. Developmental patterns in numbers of ramal segments of copepod post-maxillipedal legs. Crustaceana. 54: 256 - 293." type="journal article" year="1988">Ferrari (1988)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD1981BEE777F1FFF2CC45EFD0DF915" blockId="17.[160,1155,1494,1727]" pageId="17" pageNumber="2479">
Subdistal outer spine on exopodite of leg 3 narrow and unarmed in stage CII (
<figureCitation id="1355849EEE777F1FFF4BC403FE97F9D0" box="[167,281,1634,1658]" captionStart="Figure 12" captionStartId="20.[160,225,1326,1349]" captionTargetBox="[163,1153,142,1295]" captionTargetId="figure-95@20.[163,1153,142,1295]" captionTargetPageId="20" captionText="Figure 12. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage I: (a) leg 1, caudal; (b) leg 2, caudal (endopodite illustrated separately); (c–e) Copepodid stage II: (c) leg 1, frontal; (d) leg 2, frontal; (e) leg 3, frontal." figureDoi="http://doi.org/10.5281/zenodo.5183787" httpUri="https://zenodo.org/record/5183787/files/figure.png" pageId="17" pageNumber="2479">Figure 12</figureCitation>
(e)), being reshaped as normal spine in stage IV (
<figureCitation id="1355849EEE777F1FFC94C403FC66F9D0" box="[888,1000,1634,1658]" captionStart="Figure 13" captionStartId="21.[160,225,1550,1573]" captionTargetBox="[163,1153,142,1519]" captionTargetId="figure-1@21.[163,1153,142,1519]" captionTargetPageId="21" captionText="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." figureDoi="http://doi.org/10.5281/zenodo.5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="17" pageNumber="2479">Figure 13</figureCitation>
(a)); subdistal exopodite spine of leg 4 displaying same development constraint (in stage CIII –
<figureCitation id="1355849EEE777F1FFF4CC4C6FE80F915" box="[160,270,1703,1727]" captionStart="Figure 13" captionStartId="21.[160,225,1550,1573]" captionTargetBox="[163,1153,142,1519]" captionTargetId="figure-1@21.[163,1153,142,1519]" captionTargetPageId="21" captionText="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." figureDoi="http://doi.org/10.5281/zenodo.5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="17" pageNumber="2479">Figure 13</figureCitation>
(b) and stage CIV –
<figureCitation id="1355849EEE777F1FFE03C4C6FDD7F915" box="[495,601,1703,1727]" captionStart="Figure 13" captionStartId="21.[160,225,1550,1573]" captionTargetBox="[163,1153,142,1519]" captionTargetId="figure-1@21.[163,1153,142,1519]" captionTargetPageId="21" captionText="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." figureDoi="http://doi.org/10.5281/zenodo.5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="17" pageNumber="2479">Figure 13</figureCitation>
(c)).
</paragraph>
<caption id="DF11C893EE747F1CFF4CC477FC8BF9C2" ID-DOI="http://doi.org/10.5281/zenodo.5183785" ID-Zenodo-Dep="5183785" httpUri="https://zenodo.org/record/5183785/files/figure.png" pageId="18" pageNumber="2480" startId="18.[160,225,1558,1581]" targetBox="[163,1153,142,1527]" targetPageId="18">
<paragraph id="8BD1981BEE747F1CFF4CC477FC8BF9C2" blockId="18.[160,1156,1558,1640]" pageId="18" pageNumber="2480">
<emphasis id="B91A4409EE747F1CFF4CC477FE86F987" bold="true" box="[160,264,1558,1581]" pageId="18" pageNumber="2480">Figure 11.</emphasis>
<taxonomicName id="4C6EE398EE747F1CFEFCC477FD84F987" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[272,522,1558,1581]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="2480" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE747F1CFEFCC477FD84F987" box="[272,522,1558,1581]" italics="true" pageId="18" pageNumber="2480">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE747F1CFDFDC477FD46F987" box="[529,712,1558,1581]" pageId="18" pageNumber="2480" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral.
</paragraph>
</caption>
<caption id="DF11C893EE757F1DFF69C150F9D8FCE2" box="[133,1622,817,840]" pageId="19" pageNumber="2481" startId="19.[133,189,817,840]">
<paragraph id="8BD1981BEE757F1DFF69C150F9D8FCE2" blockId="19.[133,1622,817,840]" box="[133,1622,817,840]" pageId="19" pageNumber="2481">
<emphasis id="B91A4409EE757F1DFF69C150FF56FCE2" bold="true" box="[133,216,817,840]" pageId="19" pageNumber="2481">Table 4.</emphasis>
Segment and armament evolution of male antennule from stage III to adult, compared with the ancestral antennulary segment sequence.
</paragraph>
</caption>
<paragraph id="8BD1981BEE757F1DFF7CC134FA6FFBAA" pageId="19" pageNumber="2481">
<table id="F96E6ABBEE7580F1FF7CC134F928FBAA" box="[144,1702,853,1024]" gridcols="19" gridrows="7" pageId="19" pageNumber="2481">
<tr id="355E9A59EE7580F1FF7CC134F928FCC2" box="[144,1702,853,872]" gridrow="0" pageId="19" pageNumber="2481" rowspan-1="1" rowspan-17="1" rowspan-18="1">
<th id="768FF325EE7580F1FF7CC134FF4CFCC2" box="[144,194,853,872]" gridcol="0" gridrow="0" pageId="19" pageNumber="2481">Ad. #.</th>
<th id="768FF325EE7580F1FED9C134FEFFFCC2" box="[309,369,853,872]" gridcol="2" gridrow="0" pageId="19" pageNumber="2481">1</th>
<th id="768FF325EE7580F1FE71C134FE3CFCC2" box="[413,434,853,872]" gridcol="3" gridrow="0" pageId="19" pageNumber="2481">2</th>
<th id="768FF325EE7580F1FE33C134FE77FCC2" box="[479,505,853,872]" gridcol="4" gridrow="0" pageId="19" pageNumber="2481">3</th>
<th id="768FF325EE7580F1FDC9C134FDD8FCC2" box="[549,598,853,872]" gridcol="5" gridrow="0" pageId="19" pageNumber="2481">4</th>
<th id="768FF325EE7580F1FD6EC134FD03FCC2" box="[642,653,853,872]" gridcol="6" gridrow="0" pageId="19" pageNumber="2481">5</th>
<th id="768FF325EE7580F1FD55C134FD47FCC2" box="[697,713,853,872]" gridcol="7" gridrow="0" pageId="19" pageNumber="2481">6</th>
<th id="768FF325EE7580F1FD1AC134FC9EFCC2" box="[758,784,853,872]" gridcol="8" gridrow="0" pageId="19" pageNumber="2481">7</th>
<th id="768FF325EE7580F1FCD0C134FCD9FCC2" box="[828,855,853,872]" gridcol="9" gridrow="0" pageId="19" pageNumber="2481">8</th>
<th id="768FF325EE7580F1FC6FC134FC4DFCC2" box="[899,963,853,872]" gridcol="10" gridrow="0" pageId="19" pageNumber="2481">9</th>
<th id="768FF325EE7580F1FC1CC134FB9EFCC2" box="[1008,1040,853,872]" gridcol="11" gridrow="0" pageId="19" pageNumber="2481">10</th>
<th id="768FF325EE7580F1FBD0C134FBEFFCC2" box="[1084,1121,853,872]" gridcol="12" gridrow="0" pageId="19" pageNumber="2481">11</th>
<th id="768FF325EE7580F1FB61C134FB26FCC2" box="[1165,1192,853,872]" gridcol="13" gridrow="0" pageId="19" pageNumber="2481">12</th>
<th id="768FF325EE7580F1FB39C134FA8AFCC2" box="[1237,1284,853,872]" gridcol="14" gridrow="0" pageId="19" pageNumber="2481">13</th>
<th id="768FF325EE7580F1FADCC134FAF4FCC2" box="[1328,1402,853,872]" gridcol="15" gridrow="0" pageId="19" pageNumber="2481">14</th>
<th id="768FF325EE7580F1FA4AC134FA62FCC2" box="[1446,1516,853,872]" gridcol="16" gridrow="0" pageId="19" pageNumber="2481">15</th>
</tr>
<tr id="355E9A59EE7580F1FF7CC114F928FC22" box="[144,1702,885,904]" gridrow="1" pageId="19" pageNumber="2481" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="768FF325EE7580F1FF7CC114FF4CFC22" box="[144,194,885,904]" gridcol="0" gridrow="1" pageId="19" pageNumber="2481">9</th>
<td id="768FF325EE7580F1FF02C114FE87FC22" box="[238,265,885,904]" gridcol="1" gridrow="1" pageId="19" pageNumber="2481">CIII</td>
<td id="768FF325EE7580F1FED9C114FEFFFC22" box="[309,369,885,904]" gridcol="2" gridrow="1" pageId="19" pageNumber="2481">3</td>
<td id="768FF325EE7580F1FE71C114FE3CFC22" box="[413,434,885,904]" gridcol="3" gridrow="1" pageId="19" pageNumber="2481">3</td>
<td id="768FF325EE7580F1FD1AC114FC9EFC22" box="[758,784,885,904]" gridcol="8" gridrow="1" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FC6FC114FC4DFC22" box="[899,963,885,904]" gridcol="10" gridrow="1" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FC1CC114FB9EFC22" box="[1008,1040,885,904]" gridcol="11" gridrow="1" pageId="19" pageNumber="2481">3</td>
<td id="768FF325EE7580F1FADCC114FAF4FC22" box="[1328,1402,885,904]" gridcol="15" gridrow="1" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1FA4AC114FA62FC22" box="[1446,1516,885,904]" gridcol="16" gridrow="1" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1F9F4C114F9C7FC22" box="[1560,1609,885,904]" gridcol="17" gridrow="1" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1F999C114F928FC22" box="[1653,1702,885,904]" gridcol="18" gridrow="1" pageId="19" pageNumber="2481">7 + A</td>
</tr>
<tr id="355E9A59EE7580F1FF7CC1ECF928FC0A" box="[144,1702,909,928]" gridrow="2" pageId="19" pageNumber="2481" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="768FF325EE7580F1FF7CC1ECFF4CFC0A" box="[144,194,909,928]" gridcol="0" gridrow="2" pageId="19" pageNumber="2481">10</th>
<td id="768FF325EE7580F1FF02C1ECFE87FC0A" box="[238,265,909,928]" gridcol="1" gridrow="2" pageId="19" pageNumber="2481">CIV</td>
<td id="768FF325EE7580F1FED9C1ECFEFFFC0A" box="[309,369,909,928]" gridcol="2" gridrow="2" pageId="19" pageNumber="2481">6</td>
<td id="768FF325EE7580F1FE71C1ECFE3CFC0A" box="[413,434,909,928]" gridcol="3" gridrow="2" pageId="19" pageNumber="2481">6</td>
<td id="768FF325EE7580F1FD1AC1ECFC9EFC0A" box="[758,784,909,928]" gridcol="8" gridrow="2" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FCD0C1ECFCD9FC0A" box="[828,855,909,928]" gridcol="9" gridrow="2" pageId="19" pageNumber="2481">1</td>
<td id="768FF325EE7580F1FC6FC1ECFC4DFC0A" box="[899,963,909,928]" gridcol="10" gridrow="2" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FC1CC1ECFB9EFC0A" box="[1008,1040,909,928]" gridcol="11" gridrow="2" pageId="19" pageNumber="2481">3</td>
<td id="768FF325EE7580F1FADCC1ECFAF4FC0A" box="[1328,1402,909,928]" gridcol="15" gridrow="2" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1FA4AC1ECFA62FC0A" box="[1446,1516,909,928]" gridcol="16" gridrow="2" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1F9F4C1ECF9C7FC0A" box="[1560,1609,909,928]" gridcol="17" gridrow="2" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1F999C1ECF928FC0A" box="[1653,1702,909,928]" gridcol="18" gridrow="2" pageId="19" pageNumber="2481">7 + A</td>
</tr>
<tr id="355E9A59EE7580F1FF7CC1C4F928FC12" box="[144,1702,933,952]" gridrow="3" pageId="19" pageNumber="2481" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="768FF325EE7580F1FF7CC1C4FF4CFC12" box="[144,194,933,952]" gridcol="0" gridrow="3" pageId="19" pageNumber="2481">8</th>
<td id="768FF325EE7580F1FF02C1C4FE87FC12" box="[238,265,933,952]" gridcol="1" gridrow="3" pageId="19" pageNumber="2481">CV</td>
<td id="768FF325EE7580F1FED9C1C4FEFFFC12" box="[309,369,933,952]" gridcol="2" gridrow="3" pageId="19" pageNumber="2481">6</td>
<td id="768FF325EE7580F1FE71C1C4FE3CFC12" box="[413,434,933,952]" gridcol="3" gridrow="3" pageId="19" pageNumber="2481">11</td>
<td id="768FF325EE7580F1FD1AC1C4FC9EFC12" box="[758,784,933,952]" gridcol="8" gridrow="3" pageId="19" pageNumber="2481">5</td>
<td id="768FF325EE7580F1FC6FC1C4FC4DFC12" box="[899,963,933,952]" gridcol="10" gridrow="3" pageId="19" pageNumber="2481">7 + 2pr</td>
<td id="768FF325EE7580F1FADCC1C4FAF4FC12" box="[1328,1402,933,952]" gridcol="15" gridrow="3" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1FA4AC1C4FA62FC12" box="[1446,1516,933,952]" gridcol="16" gridrow="3" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1F9F4C1C4F9C7FC12" box="[1560,1609,933,952]" gridcol="17" gridrow="3" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1F999C1C4F928FC12" box="[1653,1702,933,952]" gridcol="18" gridrow="3" pageId="19" pageNumber="2481">7 + A</td>
</tr>
<tr id="355E9A59EE7580F1FF7CC1DCF928FC7A" box="[144,1702,957,976]" gridrow="4" pageId="19" pageNumber="2481" rowspan-17="1" rowspan-18="1">
<th id="768FF325EE7580F1FF7CC1DCFF4CFC7A" box="[144,194,957,976]" gridcol="0" gridrow="4" pageId="19" pageNumber="2481">16</th>
<td id="768FF325EE7580F1FF02C1DCFE87FC7A" box="[238,265,957,976]" gridcol="1" gridrow="4" pageId="19" pageNumber="2481">ad.</td>
<td id="768FF325EE7580F1FED9C1DCFEFFFC7A" box="[309,369,957,976]" gridcol="2" gridrow="4" pageId="19" pageNumber="2481">7 + 3A</td>
<td id="768FF325EE7580F1FE71C1DCFE3CFC7A" box="[413,434,957,976]" gridcol="3" gridrow="4" pageId="19" pageNumber="2481">4</td>
<td id="768FF325EE7580F1FE33C1DCFE77FC7A" box="[479,505,957,976]" gridcol="4" gridrow="4" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FDC9C1DCFDD8FC7A" box="[549,598,957,976]" gridcol="5" gridrow="4" pageId="19" pageNumber="2481">2 + A</td>
<td id="768FF325EE7580F1FD6EC1DCFD03FC7A" box="[642,653,957,976]" gridcol="6" gridrow="4" pageId="19" pageNumber="2481">1</td>
<td id="768FF325EE7580F1FD55C1DCFD47FC7A" box="[697,713,957,976]" gridcol="7" gridrow="4" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FD1AC1DCFC9EFC7A" box="[758,784,957,976]" gridcol="8" gridrow="4" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FCD0C1DCFCD9FC7A" box="[828,855,957,976]" gridcol="9" gridrow="4" pageId="19" pageNumber="2481">1</td>
<td id="768FF325EE7580F1FC6FC1DCFC4DFC7A" box="[899,963,957,976]" gridcol="10" gridrow="4" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FC1CC1DCFB9EFC7A" box="[1008,1040,957,976]" gridcol="11" gridrow="4" pageId="19" pageNumber="2481">4</td>
<td id="768FF325EE7580F1FBD0C1DCFBEFFC7A" box="[1084,1121,957,976]" gridcol="12" gridrow="4" pageId="19" pageNumber="2481">2</td>
<td id="768FF325EE7580F1FB61C1DCFB26FC7A" box="[1165,1192,957,976]" gridcol="13" gridrow="4" pageId="19" pageNumber="2481">?</td>
<td id="768FF325EE7580F1FB39C1DCFA8AFC7A" box="[1237,1284,957,976]" gridcol="14" gridrow="4" pageId="19" pageNumber="2481">1 + T</td>
<td id="768FF325EE7580F1FADCC1DCFAF4FC7A" box="[1328,1402,957,976]" gridcol="15" gridrow="4" pageId="19" pageNumber="2481">2 + T</td>
<td id="768FF325EE7580F1FA4AC1DCFA62FC7A" box="[1446,1516,957,976]" gridcol="16" gridrow="4" pageId="19" pageNumber="2481">11 + 2A</td>
</tr>
<tr id="355E9A59EE7580F1FF7CC1B4F928FC42" box="[144,1702,981,1000]" gridrow="5" pageId="19" pageNumber="2481" rowspan-1="1" rowspan-17="1" rowspan-18="1">
<th id="768FF325EE7580F1FF7CC1B4FF4CFC42" box="[144,194,981,1000]" gridcol="0" gridrow="5" pageId="19" pageNumber="2481">anc. #</th>
<td id="768FF325EE7580F1FED9C1B4FEFFFC42" box="[309,369,981,1000]" gridcol="2" gridrow="5" pageId="19" pageNumber="2481">I</td>
<td id="768FF325EE7580F1FE71C1B4FE3CFC42" box="[413,434,981,1000]" gridcol="3" gridrow="5" pageId="19" pageNumber="2481">VI</td>
<td id="768FF325EE7580F1FE33C1B4FE77FC42" box="[479,505,981,1000]" gridcol="4" gridrow="5" pageId="19" pageNumber="2481">VIII</td>
<td id="768FF325EE7580F1FDC9C1B4FDD8FC42" box="[549,598,981,1000]" gridcol="5" gridrow="5" pageId="19" pageNumber="2481">IX</td>
<td id="768FF325EE7580F1FD6EC1B4FD03FC42" box="[642,653,981,1000]" gridcol="6" gridrow="5" pageId="19" pageNumber="2481">X</td>
<td id="768FF325EE7580F1FD55C1B4FD47FC42" box="[697,713,981,1000]" gridcol="7" gridrow="5" pageId="19" pageNumber="2481">XI</td>
<td id="768FF325EE7580F1FD1AC1B4FC9EFC42" box="[758,784,981,1000]" gridcol="8" gridrow="5" pageId="19" pageNumber="2481">XII</td>
<td id="768FF325EE7580F1FCD0C1B4FCD9FC42" box="[828,855,981,1000]" gridcol="9" gridrow="5" pageId="19" pageNumber="2481">XIV</td>
<td id="768FF325EE7580F1FC6FC1B4FC4DFC42" box="[899,963,981,1000]" gridcol="10" gridrow="5" pageId="19" pageNumber="2481">
XV
<emphasis id="B91A4409EE757F1DFC70C1B4FC29FC42" box="[924,935,981,1000]" italics="true" pageId="19" pageNumber="2481">–</emphasis>
XVI
</td>
<td id="768FF325EE7580F1FC1CC1B4FB9EFC42" box="[1008,1040,981,1000]" gridcol="11" gridrow="5" pageId="19" pageNumber="2481">XVII</td>
<td id="768FF325EE7580F1FBD0C1B4FBEFFC42" box="[1084,1121,981,1000]" gridcol="12" gridrow="5" pageId="19" pageNumber="2481">XVIII</td>
<td id="768FF325EE7580F1FB61C1B4FB26FC42" box="[1165,1192,981,1000]" gridcol="13" gridrow="5" pageId="19" pageNumber="2481">IXX</td>
<td id="768FF325EE7580F1FB39C1B4FA8AFC42" box="[1237,1284,981,1000]" gridcol="14" gridrow="5" pageId="19" pageNumber="2481">XX</td>
<td id="768FF325EE7580F1FADCC1B4FAF4FC42" box="[1328,1402,981,1000]" gridcol="15" gridrow="5" pageId="19" pageNumber="2481">
XXI
<emphasis id="B91A4409EE757F1DFAA7C1B4FAD8FC42" box="[1355,1366,981,1000]" italics="true" pageId="19" pageNumber="2481">–</emphasis>
XXIII
</td>
<td id="768FF325EE7580F1FA4AC1B4FA62FC42" box="[1446,1516,981,1000]" gridcol="16" gridrow="5" pageId="19" pageNumber="2481">XXIV</td>
</tr>
<tr id="355E9A59EE7580F1FF7CC18CF928FBAA" box="[144,1702,1005,1024]" gridrow="6" pageId="19" pageNumber="2481" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-15="1" rowspan-17="1" rowspan-18="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<td id="768FF325EE7580F1FED9C18CFEFFFBAA" box="[309,369,1005,1024]" gridcol="2" gridrow="6" pageId="19" pageNumber="2481">V</td>
<td id="768FF325EE7580F1FE71C18CFE3CFBAA" box="[413,434,1005,1024]" gridcol="3" gridrow="6" pageId="19" pageNumber="2481">VII</td>
<td id="768FF325EE7580F1FD1AC18CFC9EFBAA" box="[758,784,1005,1024]" gridcol="8" gridrow="6" pageId="19" pageNumber="2481">XIII</td>
<td id="768FF325EE7580F1FA4AC18CFA62FBAA" box="[1446,1516,1005,1024]" gridcol="16" gridrow="6" pageId="19" pageNumber="2481">XXVIII</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD1981BEE757F1DFF69C66FFD34FB8B" blockId="19.[133,698,1038,1057]" box="[133,698,1038,1057]" pageId="19" pageNumber="2481">
<tableNote id="76889995EE757F1DFF69C66FFD34FB8B" box="[133,698,1038,1057]" pageId="19" pageNumber="2481" targetBox="[144,1702,853,1024]" targetPageId="19">Abbreviation: A, aesthetasc; pr, protuberance; T, T-shaped process.</tableNote>
</paragraph>
<caption id="DF11C893EE727F1AFF4CC74FFE16FAD5" ID-DOI="http://doi.org/10.5281/zenodo.5183787" ID-Zenodo-Dep="5183787" httpUri="https://zenodo.org/record/5183787/files/figure.png" pageId="20" pageNumber="2482" startId="20.[160,225,1326,1349]" targetBox="[163,1153,142,1295]" targetPageId="20">
<paragraph id="8BD1981BEE727F1AFF4CC74FFE16FAD5" blockId="20.[160,1156,1326,1407]" pageId="20" pageNumber="2482">
<emphasis id="B91A4409EE727F1AFF4CC74FFE89FAEF" bold="true" box="[160,263,1326,1349]" pageId="20" pageNumber="2482">Figure 12.</emphasis>
<taxonomicName id="4C6EE398EE727F1AFEE3C74FFD89FAEF" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[271,519,1326,1349]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="20" pageNumber="2482" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE727F1AFEE3C74FFD89FAEF" box="[271,519,1326,1349]" italics="true" pageId="20" pageNumber="2482">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE727F1AFDE2C74FFD4CFAEF" box="[526,706,1326,1349]" pageId="20" pageNumber="2482" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a, b) Copepodid stage I: (a) leg 1, caudal; (b) leg 2, caudal (endopodite illustrated separately); (c
<emphasis id="B91A4409EE727F1AFD7AC72AFD2DFAC8" box="[662,675,1355,1378]" italics="true" pageId="20" pageNumber="2482">–</emphasis>
e) Copepodid stage II: (c) leg 1, frontal; (d) leg 2, frontal; (e) leg 3, frontal.
</paragraph>
</caption>
<paragraph id="8BD1981BEE727F1AFF2CC7B6FBB7F915" blockId="20.[160,1156,1495,1727]" pageId="20" pageNumber="2482">
Sexual differences of leg 3 endopodite apparently not expressed during development; terminal moult affecting male leg 3 only, including modification of terminal outer spine and deletion of terminal inner element; differences of leg 4 endopodite between male and female distinct in stage CV but not pronounced: female endopodite wider than in male, one-segmented but with clear remnant of articulation as transverse cuticular ridge between former proximal and distal segment, without spinule patterns along former articulation (
<figureCitation id="1355849EEE727F1AFE20C4C6FDB7F915" box="[460,569,1703,1727]" captionStart="Figure 13" captionStartId="21.[160,225,1550,1573]" captionTargetBox="[163,1153,142,1519]" captionTargetId="figure-1@21.[163,1153,142,1519]" captionTargetPageId="21" captionText="Figure 13. Thalamocyclops pachypes gen. nov., sp. nov. (a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal." figureDoi="http://doi.org/10.5281/zenodo.5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="20" pageNumber="2482">Figure 13</figureCitation>
(e)); armature elements similar in both sexes.
</paragraph>
<caption id="DF11C893EE737F1BFF4CC46FFC69F9CA" ID-DOI="http://doi.org/10.5281/zenodo.5183789" ID-Zenodo-Dep="5183789" httpUri="https://zenodo.org/record/5183789/files/figure.png" pageId="21" pageNumber="2483" startId="21.[160,225,1550,1573]" targetBox="[163,1153,142,1519]" targetPageId="21">
<paragraph id="8BD1981BEE737F1BFF4CC46FFC69F9CA" blockId="21.[160,1156,1550,1632]" pageId="21" pageNumber="2483">
<emphasis id="B91A4409EE737F1BFF4CC46FFE86F98F" bold="true" box="[160,264,1550,1573]" pageId="21" pageNumber="2483">Figure 13.</emphasis>
<taxonomicName id="4C6EE398EE737F1BFEFDC46FFD84F98F" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[273,522,1550,1573]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="21" pageNumber="2483" phylum="Arthropoda" rank="species" species="pachypes" status="gen. nov., sp. nov.">
<emphasis id="B91A4409EE737F1BFEFDC46FFD84F98F" box="[273,522,1550,1573]" italics="true" pageId="21" pageNumber="2483">Thalamocyclops pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE737F1BFDFEC46FFD44F98F" box="[530,714,1550,1573]" pageId="21" pageNumber="2483" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(a, b) Copepodid stage III: (a) leg 3, frontal; (b) leg 4, frontal; (c, d) Copepodid stage IV: (c) leg 4, frontal (second endopodite segment detached); (d) leg 4 second endopodite segment; (e) female copepodid stage V: leg 4, caudal.
</paragraph>
</caption>
<paragraph id="8BD1981BEE707F18FF4CC2EFFB0AFEDD" blockId="22.[160,1156,142,583]" pageId="22" pageNumber="2484">
<emphasis id="B91A4409EE707F18FF4CC2EFFC8FFF0D" bold="true" box="[160,769,142,167]" italics="true" pageId="22" pageNumber="2484">
Development of legs 5 and 6 (
<figureCitation id="1355849EEE707F18FDCCC2EFFD1EFF0D" box="[544,656,142,167]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="22" pageNumber="2484">Figures 8</figureCitation>
and
<figureCitation id="1355849EEE707F18FD3AC2EFFD7DFF0D" box="[726,755,142,167]" captionStart="Figure 11" captionStartId="18.[160,225,1558,1581]" captionTargetBox="[163,1153,142,1527]" captionTargetId="figure-1@18.[163,1153,142,1527]" captionTargetPageId="18" captionText="Figure 11. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183785" httpUri="https://zenodo.org/record/5183785/files/figure.png" pageId="22" pageNumber="2484">11</figureCitation>
).
</emphasis>
Leg 5 (precursor) appearing as a lateral triangular outgrowth in stage CII (
<figureCitation id="1355849EEE707F18FDB8C2D3FD25FF60" box="[596,683,178,202]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="22" pageNumber="2484">Figure 8</figureCitation>
(c,d)), reshaped in a large rectangular lateral lobe with a distal spine and a subdistal (dorsally located) seta in stage CIII (
<figureCitation id="1355849EEE707F18FC2BC2B5FBAEFF46" box="[967,1056,212,236]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="22" pageNumber="2484">Figure 8</figureCitation>
(e)); leg 5 represented by three armature elements in stage CIV and CV: two terminal elements (spine and seta) on a ventrolateral square outgrowth and one lateral setal element inserted on the posterior border of the somite (dorsally located); ventrolateral lobe reappearing almost unmodified in female copepodid CV, but being narrower and longer in male copepodid CV.
</paragraph>
<paragraph id="8BD1981BEE707F18FF2CC3E3FD4DFDED" blockId="22.[160,1156,142,583]" pageId="22" pageNumber="2484">
Leg 6 recognizable as lateral triangular outgrowth in stage CIII (
<figureCitation id="1355849EEE707F18FC72C3E3FC72FE30" box="[926,1020,386,410]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="22" pageNumber="2484">Figure 8</figureCitation>
(e)), present as a truncated triangular lobe, armed with a large spine and a short seta, in stage CIV (
<figureCitation id="1355849EEE707F18FF4BC3A6FE8AFE75" box="[167,260,455,479]" captionStart="Figure 8" captionStartId="14.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure-1@14.[163,1153,142,1529]" captionTargetPageId="14" captionText="Figure 8. Thalamocyclops pachypes gen. nov., sp. nov. (a) Postcephalic somites 2–4 in copepodid stage I, ventral; (b) idem, lateral view; (c) postcephalic somites 3 to 5 in copepodid stage II, ventral; (d) postcephalic somites 3 and 4 in copepodid stage II, lateral; (e) postcephalic somites 4 and 5 in copepodid stage III, lateral; (f) postcephalic somites 4 and 5 in female copepodid stage IV, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183779" httpUri="https://zenodo.org/record/5183779/files/figure.png" pageId="22" pageNumber="2484">Figure 8</figureCitation>
(f)), and bearing three armature elements in stage CV (one spine, two short setal elements); female leg
<quantity id="4C9635FEEE707F18FE77C38BFE48FDA8" box="[411,454,490,514]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="2484" unit="in" value="6.0">6 in</quantity>
stage CV confluent with border of somite and located dorsolaterally (
<figureCitation id="1355849EEE707F18FEE3C06DFE0EFD8E" box="[271,384,524,548]" captionStart="Figure 11" captionStartId="18.[160,225,1558,1581]" captionTargetBox="[163,1153,142,1527]" captionTargetId="figure-1@18.[163,1153,142,1527]" captionTargetPageId="18" captionText="Figure 11. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183785" httpUri="https://zenodo.org/record/5183785/files/figure.png" pageId="22" pageNumber="2484">Figure 11</figureCitation>
(c)); male leg
<quantity id="4C9635FEEE707F18FDDCC06DFDECFD8E" box="[560,610,524,548]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="22" pageNumber="2484" unit="in" value="6.0">6 in</quantity>
stage CV present as a distinct ventrolateral crescentic outgrowth in the male (
<figureCitation id="1355849EEE707F18FDC0C04EFD18FDED" box="[556,662,559,583]" captionStart="Figure 11" captionStartId="18.[160,225,1558,1581]" captionTargetBox="[163,1153,142,1527]" captionTargetId="figure-1@18.[163,1153,142,1527]" captionTargetPageId="18" captionText="Figure 11. Thalamocyclops pachypes gen. nov., sp. nov. (a) Antennule in copepodid stage IV (male); (b) antennule in copepodid stage V (male); (c) postcephalic somites 4 and 5 in female copepodid stage V, lateral; (d) idem, in male copepodid stage V, lateral." figureDoi="http://doi.org/10.5281/zenodo.5183785" httpUri="https://zenodo.org/record/5183785/files/figure.png" pageId="22" pageNumber="2484">Figure 11</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE707F14FF4CC015FD60FF58" lastPageId="26" lastPageNumber="2488" pageId="22" pageNumber="2484" type="discussion">
<paragraph id="8BD1981BEE707F18FF4CC015FE90FD27" blockId="22.[160,1156,628,1727]" box="[160,286,628,653]" pageId="22" pageNumber="2484">
<emphasis id="B91A4409EE707F18FF4CC015FE90FD27" bold="true" box="[160,286,628,653]" italics="true" pageId="22" pageNumber="2484">Comments</emphasis>
</paragraph>
<paragraph id="8BD1981BEE707F18FF4CC0F6FC38FCF6" blockId="22.[160,1156,628,1727]" pageId="22" pageNumber="2484">
The naupliar body shape is remarkably flattened compared with the nauplii known of other
<taxonomicName id="4C6EE398EE707F18FF0AC0DBFEECFD78" authorityName="Rafinesque, 1815 Sufamily CYCLOPINAE Kiefer" authorityYear="1927" box="[230,354,698,722]" class="Hexanauplia" family="Cyclopidae" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="2484" phylum="Arthropoda" rank="family">Cyclopidae</taxonomicName>
(see
<bibRefCitation id="EFFFE5EAEE707F18FE4DC0DBFD57FD78" author="Dahms H-U &amp; Fernando CH" box="[417,729,698,722]" pageId="22" pageNumber="2484" pagination="9 - 21" refId="ref17941" refString="Dahms H-U, Fernando CH. 1994. Redescription of female Macrocyclops fuscus (Jurine, 1820) (Copepoda, Cyclopoida) from Ontario with a description of naupliar stages. J Plankton Res. 16: 9 - 21." type="journal article" year="1994">Dahms and Fernando 1994</bibRefCitation>
;
<bibRefCitation id="EFFFE5EAEE707F18FD04C0DBFBA8FD78" author="Karaytug S &amp; Boxshall GA" box="[744,1062,698,722]" pageId="22" pageNumber="2484" pagination="41 - 70" refId="ref19072" refString="Karaytug S, Boxshall GA. 1996. The life cycle of Paracyclops fi mbriatus (Fischer, 1853) (Copepoda, Cyclopoida). Bull Natl Hist Museum Lond Zool. 62: 41 - 70." type="journal article" year="1996">Karaytug and Boxshall 1996</bibRefCitation>
;
<bibRefCitation id="EFFFE5EAEE707F18FBD9C0DBFF59FD5E" author="Dahms H-U" pageId="22" pageNumber="2484" pagination="91 - 99" refId="ref17921" refString="Dahms H-U. 2000. Phylogenetic implications of the Crustacean nauplius. Hydrobiologia. 417: 91 - 99." type="journal article" year="2000">Dahms 2000</bibRefCitation>
). The presence of a dorsal integumental window in cyclopine nauplii has not been documented previously. Body shape must be a particular adaptation to the cramped habitat in which it occurs. The window indicates extensive osmoregulatory activity as a response to the environmental conditions in the hole (
<bibRefCitation id="EFFFE5EAEE707F18FCFFC125FC27FCF6" author="Hosfeld B" box="[787,937,836,860]" pageId="22" pageNumber="2484" pagination="61 - 74" refId="ref18804" refString="Hosfeld B. 1999. Ultrastucture of ionocytes from osmoregulatory integumental windows of Tachidius discipes and Bryocamptus minutus (Crustacea, Copepoda, Harpacticoida) with remarks on the homology of nonsensory dorsal organs of crustaceans. Acta Zool. 80: 61 - 74." type="journal article" year="1999">Hosfeld 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8BD1981BEE707F18FF2CC106FB0AFACE" blockId="22.[160,1156,628,1727]" pageId="22" pageNumber="2484">
An antennary exopodal structure in the first copepodid (CI) has been documented for other cyclopines (see development of
<emphasis id="B91A4409EE707F18FDADC1EBFCA3FC08" box="[577,813,906,930]" italics="true" pageId="22" pageNumber="2484">
<taxonomicName id="4C6EE398EE707F18FDADC1EBFD32FC08" authorityName="Claus" authorityYear="1893" box="[577,700,906,930]" class="Hexanauplia" family="Cyclopidae" genus="Paracyclops" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="2484" phylum="Arthropoda" rank="genus">Paracyclops</taxonomicName>
<emphasis id="B91A4409EE707F18FD2FC1EBFD5EFC08" bold="true" box="[707,720,906,930]" italics="true" pageId="22" pageNumber="2484">fi</emphasis>
mbriatus
</emphasis>
(
<bibRefCitation id="EFFFE5EAEE707F18FCD6C1EBFC46FC08" author="Fischer S" box="[826,968,906,930]" pageId="22" pageNumber="2484" pagination="74 - 100" refId="ref18478" refString="Fischer S. 1853. Beitrage zur kenntnis der in der umgegend von st. Petersburg Sich Findenden Cyclopiden (Fortsetzung). Bull Soc Nat Moscou. 26: 74 - 100." type="journal article" year="1853">Fischer, 1853</bibRefCitation>
) in
<bibRefCitation id="EFFFE5EAEE707F18FC1EC1EBFEB6FC6E" author="Karaytug S &amp; Boxshall GA" pageId="22" pageNumber="2484" pagination="41 - 70" refId="ref19072" refString="Karaytug S, Boxshall GA. 1996. The life cycle of Paracyclops fi mbriatus (Fischer, 1853) (Copepoda, Cyclopoida). Bull Natl Hist Museum Lond Zool. 62: 41 - 70." type="journal article" year="1996">Karaytug and Boxshall 1996</bibRefCitation>
; personal observations) and is also present in the first copepodid stage of
<taxonomicName id="4C6EE398EE707F18FF4CC1AEFE95FC4D" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[160,283,975,999]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="2484" phylum="Arthropoda" rank="species" species="pachypes" status="sp. nov.">
<emphasis id="B91A4409EE707F18FF4CC1AEFE95FC4D" box="[160,283,975,999]" italics="true" pageId="22" pageNumber="2484">T. pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE707F18FECEC1AEFEF8FC4D" box="[290,374,975,999]" pageId="22" pageNumber="2484" rank="species">sp. nov.</taxonomicNameLabel>
The exopodal structure is an outgrowth of clustered cells, not wrapped within a chitinous membrane, with the two setae arising from an insertion location between the cellular masses. Mouthparts in stage CI cyclopine copepodids resemble the adult morphology closely but differ still in details of the ornaments, with the most conspicuous difference between stage CI and the successive stages, the presence of a mandibular palp structure (cellular mass) in the former. In contrast, the mandible, maxillule and maxilla in stage CI of
<taxonomicName id="4C6EE398EE707F18FF08C6FEFEEDFB1D" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[228,355,1183,1207]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="2484" phylum="Arthropoda" rank="species" species="pachypes" status="sp. nov.">
<emphasis id="B91A4409EE707F18FF08C6FEFEEDFB1D" box="[228,355,1183,1207]" italics="true" pageId="22" pageNumber="2484">T. pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE707F18FE83C6FEFE47FB1D" box="[367,457,1183,1207]" pageId="22" pageNumber="2484" rank="species">sp. nov.</taxonomicNameLabel>
are identical to those in the adults. The mandible lacks the undifferentiated cellular mass known to occur in other cyclopines in stage CI, bearing solely the short seta, as in the adults. The maxilliped, normally well developed in stage CI cyclopines and resembling the adult morphology (less one or two seta short), is far from being clearly differentiated in stage CI of
<taxonomicName id="4C6EE398EE707F18FE2AC74BFDB0FAE8" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[454,574,1322,1346]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="22" pageNumber="2484" phylum="Arthropoda" rank="species" species="pachypes" status="sp. nov.">
<emphasis id="B91A4409EE707F18FE2AC74BFDB0FAE8" box="[454,574,1322,1346]" italics="true" pageId="22" pageNumber="2484">T. pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE707F18FDA8C74BFD18FAE8" box="[580,662,1322,1346]" pageId="22" pageNumber="2484" rank="species">sp. nov.</taxonomicNameLabel>
From stage CII, the maxillipedal body is larger, equipped with an adult complement, but retains still the truncated appearance as in stage CI.
</paragraph>
<paragraph id="8BD1981BEE707F18FF2CC70EFEFBF915" blockId="22.[160,1156,628,1727]" pageId="22" pageNumber="2484">
The successive modification of the legs during the copepodid development is similar to the development in other cyclopines (
<bibRefCitation id="EFFFE5EAEE707F18FD3BC7F3FC7CFA00" author="Ferrari FD &amp; Dahms H-U" box="[727,1010,1426,1450]" pageId="22" pageNumber="2484" pagination="1 - 229" refId="ref18165" refString="Ferrari FD, Dahms H-U. 2007. Post-embryonic development of the Copepoda. Crustaceana Monogr. 8: 1 - 229." type="journal article" year="2007">Ferrari and Dahms 2007</bibRefCitation>
). Leg development, including the evolution of legs 3, and the vestigial legs 5 and 6, is similar in male and female copepodids although the expression of them is considerably different in the adults. The endopodite of the female leg 4 has a more robust and compact appearance in stage CV than in the male at the same stage. Although both segments are coalescent in the female CV, the (ancestral) division line between them is clearly present as an integument structure and the four armature elements still have the conventional structure (three long pinnate setae and a serrate spine) and location (i.e. 1.I.2).
</paragraph>
<caption id="DF11C893EE717F19FF4CC4ECFE53F975" ID-DOI="http://doi.org/10.5281/zenodo.5183791" ID-Zenodo-Dep="5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="23" pageNumber="2485" startId="23.[160,225,1677,1700]" targetBox="[212,1104,142,1646]" targetPageId="23">
<paragraph id="8BD1981BEE717F19FF4CC4ECFE53F975" blockId="23.[160,1157,1677,1759]" pageId="23" pageNumber="2485">
<emphasis id="B91A4409EE717F19FF4CC4ECFE86F90E" bold="true" box="[160,264,1677,1700]" pageId="23" pageNumber="2485">Figure 14.</emphasis>
<taxonomicName id="4C6EE398EE717F19FEFCC4ECFCB5F90E" authority="(Mirabdullayev et al., 2002)" authorityName="Fiers &amp; Van Damme" authorityYear="2017" baseAuthorityName="Mirabdullayev, Van Damme &amp; Dumont" baseAuthorityYear="2002" box="[272,827,1677,1700]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="23" pageNumber="2485" phylum="Arthropoda" rank="species" species="soqotraensis" status="comb. nov.">
<emphasis id="B91A4409EE717F19FEFCC4ECFDA8F90E" box="[272,550,1677,1700]" italics="true" pageId="23" pageNumber="2485">Thalamocyclops soqotraensis</emphasis>
(
<bibRefCitation id="EFFFE5EAEE717F19FDDFC4ECFCBAF90E" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[563,820,1677,1700]" pageId="23" pageNumber="2485" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">
Mirabdullayev
<emphasis id="B91A4409EE717F19FD2BC4ECFD7DF90E" box="[711,755,1677,1700]" italics="true" pageId="23" pageNumber="2485">et al</emphasis>
., 2002
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A229F972EE717F19FCAFC4ECFC3CF90E" box="[835,946,1677,1700]" pageId="23" pageNumber="2485" rank="species">comb. nov.</taxonomicNameLabel>
(a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492
<emphasis id="B91A4409EE717F19FE7FC4A9FE2EF975" box="[403,416,1736,1759]" italics="true" pageId="23" pageNumber="2485">–</emphasis>
4495).
</paragraph>
</caption>
<caption id="DF11C893EE7E7F16FF4CC784FC7BF99D" ID-DOI="http://doi.org/10.5281/zenodo.5183793" ID-Zenodo-Dep="5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="24" pageNumber="2486" startId="24.[160,225,1509,1532]" targetBox="[163,1153,142,1478]" targetPageId="24">
<paragraph id="8BD1981BEE7E7F16FF4CC784FC7BF99D" blockId="24.[160,1156,1509,1591]" pageId="24" pageNumber="2486">
<emphasis id="B91A4409EE7E7F16FF4CC784FE86FA56" bold="true" box="[160,264,1509,1532]" pageId="24" pageNumber="2486">Figure 15.</emphasis>
<taxonomicName id="4C6EE398EE7E7F16FEFDC784FCB3FA56" authority="(Mirabdullayev et al., 2002)" authorityName="Fiers &amp; Van Damme" authorityYear="2017" baseAuthorityName="Mirabdullayev, Van Damme &amp; Dumont" baseAuthorityYear="2002" box="[273,829,1509,1532]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="24" pageNumber="2486" phylum="Arthropoda" rank="species" species="soqotraensis" status="comb. nov.">
<emphasis id="B91A4409EE7E7F16FEFDC784FDA9FA56" box="[273,551,1509,1532]" italics="true" pageId="24" pageNumber="2486">Thalamocyclops soqotraensis</emphasis>
(
<bibRefCitation id="EFFFE5EAEE7E7F16FDD8C784FCB8FA56" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[564,822,1509,1532]" pageId="24" pageNumber="2486" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">
Mirabdullayev
<emphasis id="B91A4409EE7E7F16FD25C784FD7BFA56" box="[713,757,1509,1532]" italics="true" pageId="24" pageNumber="2486">et al</emphasis>
., 2002
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A229F972EE7E7F16FCA9C784FC3BFA56" box="[837,949,1509,1532]" pageId="24" pageNumber="2486" rank="species">comb. nov.</taxonomicNameLabel>
(a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492
<emphasis id="B91A4409EE7E7F16FC46C441FC39F99D" box="[938,951,1568,1591]" italics="true" pageId="24" pageNumber="2486">–</emphasis>
4495).
</paragraph>
</caption>
<caption id="DF11C893EE7F7F17FF4CC42DFE95F911" ID-DOI="http://doi.org/10.5281/zenodo.5183795" ID-Zenodo-Dep="5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="25" pageNumber="2487" startId="25.[160,225,1612,1635]" targetBox="[163,1153,142,1581]" targetPageId="25">
<paragraph id="8BD1981BEE7F7F17FF4CC42DFE95F911" blockId="25.[160,1156,1612,1723]" pageId="25" pageNumber="2487">
<emphasis id="B91A4409EE7F7F17FF4CC42DFE87F9C9" bold="true" box="[160,265,1612,1635]" pageId="25" pageNumber="2487">Figure 16.</emphasis>
<taxonomicName id="4C6EE398EE7F7F17FEFDC42DFCCFF9C9" authority="(Mirabdullayev et al., 2002)" authorityName="Fiers &amp; Van Damme" authorityYear="2017" baseAuthorityName="Mirabdullayev, Van Damme &amp; Dumont" baseAuthorityYear="2002" box="[273,833,1612,1635]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="25" pageNumber="2487" phylum="Arthropoda" rank="species" species="soqotraensis" status="comb. nov.">
<emphasis id="B91A4409EE7F7F17FEFDC42DFDA6F9C9" box="[273,552,1612,1635]" italics="true" pageId="25" pageNumber="2487">Thalamocyclops soqotraensis</emphasis>
(
<bibRefCitation id="EFFFE5EAEE7F7F17FDDAC42DFCB4F9C9" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[566,826,1612,1635]" pageId="25" pageNumber="2487" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">
Mirabdullayev
<emphasis id="B91A4409EE7F7F17FD27C42DFD79F9C9" box="[715,759,1612,1635]" italics="true" pageId="25" pageNumber="2487">et al</emphasis>
., 2002
</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A229F972EE7F7F17FCA5C42DFC34F9C9" box="[841,954,1612,1635]" pageId="25" pageNumber="2487" rank="species">comb. nov.</taxonomicNameLabel>
(a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492
<emphasis id="B91A4409EE7F7F17FF3CC4C5FF53F911" box="[208,221,1700,1723]" italics="true" pageId="25" pageNumber="2487">–</emphasis>
4495).
</paragraph>
</caption>
<paragraph id="8BD1981BEE7C7F14FF2AC2EFFD5BFF60" blockId="26.[198,1117,142,202]" pageId="26" pageNumber="2488">
<taxonomicName id="4C6EE398EE7C7F14FF2AC2EFFBD3FF0D" authority="(Mirabdullayev, Van Damme &amp; Dumont, 2002)" authorityName="Fiers &amp; Van Damme" authorityYear="2017" baseAuthorityName="Mirabdullayev, Van Damme &amp; Dumont" baseAuthorityYear="2002" box="[198,1117,142,167]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="2488" phylum="Arthropoda" rank="species" species="soqotraensis" status="comb. nov.">
<emphasis id="B91A4409EE7C7F14FF2AC2EFFDA1FF0D" bold="true" box="[198,559,142,167]" italics="true" pageId="26" pageNumber="2488">Thalamocyclops soqotraensis</emphasis>
(
<bibRefCitation id="EFFFE5EAEE7C7F14FDD2C2EFFBDBFF0D" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[574,1109,142,167]" pageId="26" pageNumber="2488" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">Mirabdullayev, Van Damme &amp; Dumont, 2002</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A229F972EE7C7F14FDA3C2D0FD5BFF60" box="[591,725,177,202]" pageId="26" pageNumber="2488" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BD1981BEE7C7F14FDACC2BBFD60FF58" blockId="26.[576,750,217,242]" box="[576,750,217,242]" pageId="26" pageNumber="2488">
(
<figureCitation id="1355849EEE7C7F14FDAAC2BBFD6BFF58" box="[582,741,217,242]" captionStart-0="Figure 14" captionStart-1="Figure 15" captionStart-2="Figure 16" captionStartId-0="23.[160,225,1677,1700]" captionStartId-1="24.[160,225,1509,1532]" captionStartId-2="25.[160,225,1612,1635]" captionTargetBox-0="[212,1104,142,1646]" captionTargetBox-1="[163,1153,142,1478]" captionTargetBox-2="[163,1153,142,1581]" captionTargetId-0="figure-1@23.[212,1104,142,1646]" captionTargetId-1="figure-1@24.[163,1153,142,1478]" captionTargetId-2="figure-1@25.[163,1153,142,1581]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." captionText-1="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." captionText-2="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi-0="http://doi.org/10.5281/zenodo.5183791" figureDoi-1="http://doi.org/10.5281/zenodo.5183793" figureDoi-2="http://doi.org/10.5281/zenodo.5183795" httpUri-0="https://zenodo.org/record/5183791/files/figure.png" httpUri-1="https://zenodo.org/record/5183793/files/figure.png" httpUri-2="https://zenodo.org/record/5183795/files/figure.png" pageId="26" pageNumber="2488">Figures 14–16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE7C7F14FF4CC36FFD88FE24" pageId="26" pageNumber="2488" type="reference_group">
<paragraph id="8BD1981BEE7C7F14FF4CC36FFE15FEE2" blockId="26.[160,1156,269,398]" pageId="26" pageNumber="2488">
<treatmentCitationGroup id="AB7EBF35EE7C7F14FF4CC36FFE15FEE2" pageId="26" pageNumber="2488">
<taxonomicName id="4C6EE398EE7C7F14FF4CC36FFE3DFE8C" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[160,435,270,294]" class="Hexanauplia" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="2488" phylum="Arthropoda" rank="species" species="soqotraensis" status="sp. nov.">
<emphasis id="B91A4409EE7C7F14FF4CC36FFE3DFE8C" box="[160,435,270,294]" italics="true" pageId="26" pageNumber="2488">Bryocyclops soqotraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE7C7F14FE2DC36FFD90FE8C" box="[449,542,270,294]" pageId="26" pageNumber="2488" rank="species">sp. nov.</taxonomicNameLabel>
– in
<treatmentCitation id="0ACFBE0AEE7C7F14FD85C36FFEACFEE2" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" page="264" pageId="26" pageNumber="2488" year="2002">
<bibRefCitation id="EFFFE5EAEE7C7F14FD85C36FFEACFEE2" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" pageId="26" pageNumber="2488" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">Mirabdullayev, Van Damme &amp; Dumont, 2002: 264–266</bibRefCitation>
</treatmentCitation>
, figs 2–23.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1981BEE7C7F14FF4CC332FD88FE24" blockId="26.[160,1156,269,398]" pageId="26" pageNumber="2488">
<treatmentCitationGroup id="AB7EBF35EE7C7F14FF4CC332FD88FE24" pageId="26" pageNumber="2488">
<taxonomicName id="4C6EE398EE7C7F14FF4CC332FBF0FEC1" authority="Mirabdullayev, Van Damme &amp; Dumont, 2002" authorityName="Mirabdullayev, Van Damme &amp; Dumont" authorityYear="2002" box="[160,1150,339,363]" class="Hexanauplia" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="2488" phylum="Arthropoda" rank="species" species="soqotraensis" subGenus="Bryocyclops">
<emphasis id="B91A4409EE7C7F14FF4CC332FE90FEC1" box="[160,286,339,363]" italics="true" pageId="26" pageNumber="2488">Bryocyclops</emphasis>
(
<emphasis id="B91A4409EE7C7F14FED9C332FE3DFEC1" box="[309,435,339,363]" italics="true" pageId="26" pageNumber="2488">Bryocyclops</emphasis>
)
<emphasis id="B91A4409EE7C7F14FE26C332FDDFFEC1" box="[458,593,339,363]" italics="true" pageId="26" pageNumber="2488">soqotraensis</emphasis>
<bibRefCitation id="EFFFE5EAEE7C7F14FD8DC332FBF0FEC1" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[609,1150,339,363]" pageId="26" pageNumber="2488" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">Mirabdullayev, Van Damme &amp; Dumont, 2002</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="0ACFBE0AEE7C7F14FF2CC317FD8CFE24" author="Dussart BH &amp; Defaye D" box="[192,514,374,398]" page="151" pageId="26" pageNumber="2488" year="2006">
<bibRefCitation id="EFFFE5EAEE7C7F14FF2CC317FD8CFE24" author="Dussart BH &amp; Defaye D" box="[192,514,374,398]" pageId="26" pageNumber="2488" refId="ref18117" refString="Dussart BH, Defaye D. 2006. World directory of Crustacea Copepoda of Inland waters II-cyclopiformes. Leiden: Backhuys Publishers." type="book" year="2006">Dussart &amp; Defaye, 2006: 151</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374CB90EE7C7F15FF4CC3A1FC98FB73" lastPageId="27" lastPageNumber="2489" pageId="26" pageNumber="2488" type="materials_examined">
<paragraph id="8BD1981BEE7C7F14FF4CC3A1FE0EFE73" blockId="26.[160,1156,448,681]" box="[160,384,448,473]" pageId="26" pageNumber="2488">
<emphasis id="B91A4409EE7C7F14FF4CC3A1FE0EFE73" bold="true" box="[160,384,448,473]" italics="true" pageId="26" pageNumber="2488">Material examined</emphasis>
</paragraph>
<paragraph id="8BD1981BEE7C7F14FF4CC385FCBAFDEB" blockId="26.[160,1156,448,681]" pageId="26" pageNumber="2488">
<materialsCitation id="3B069246EE7C7F14FF4CC385FCA1FDEB" collectionCode="COP" pageId="26" pageNumber="2488" specimenCode="COP 4492, COP 4493-4495" specimenCount="3" typeStatus="holotype">
Female
<typeStatus id="54D526B9EE7C7F14FF17C385FED1FE56" box="[251,351,484,508]" pageId="26" pageNumber="2488" type="holotype">holotype</typeStatus>
dissected and mounted on a single slide (RBINSc
<specimenCode id="DBC83060EE7C7F14FC46C385FB93FE56" box="[938,1053,484,508]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="26" pageNumber="2488">COP 4492</specimenCode>
, labelled ‘type’),
<typeStatus id="54D526B9EE7C7F14FF14C067FED2FDB4" box="[248,348,518,542]" pageId="26" pageNumber="2488" type="paratype">paratype</typeStatus>
female and
<typeStatus id="54D526B9EE7C7F14FE12C067FDECFDB4" box="[510,610,518,542]" pageId="26" pageNumber="2488" type="paratype">paratype</typeStatus>
male (RBINSc
<specimenCode id="DBC83060EE7C7F14FCF4C067FC58FDB4" box="[792,982,518,542]" collectionCode="COP" country="Venezuela" httpUri="http://grbio.org/cool/i9n1-e075" name="Coleccion Ornitologica Phelps" pageId="26" pageNumber="2488">COP 4493–4495</specimenCode>
), dissected; all glycerine mounted and sealed with polyurethane varnish
</materialsCitation>
.
</paragraph>
<paragraph id="8BD1981BEE7C7F14FF2CC02DFD5DFD03" blockId="26.[160,1156,448,681]" pageId="26" pageNumber="2488">
The complete description of this species has to be consulted in
<bibRefCitation id="EFFFE5EAEE7C7F14FC48C02DFF69FD2C" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" pageId="26" pageNumber="2488" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">Mirabdullayev et al. (2002)</bibRefCitation>
. The following amendments deal with certain morphological details in order to compare
<taxonomicName id="4C6EE398EE7C7F14FEE0C0F0FE20FD03" authorityName="Mirabdullayev, Van Damme &amp; Dumont" authorityYear="2002" box="[268,430,657,681]" class="Hexanauplia" family="Cyclopidae" genus="Bryocyclops" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="2488" phylum="Arthropoda" rank="species" species="soqotraensis">
<emphasis id="B91A4409EE7C7F14FEE0C0F0FE20FD03" box="[268,430,657,681]" italics="true" pageId="26" pageNumber="2488">T. soqotraensis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C6EE398EE7C7F14FE1DC0F0FDFFFD03" authority="Fiers &amp; Van Damme, 2017" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[497,625,657,681]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="26" pageNumber="2488" phylum="Arthropoda" rank="species" species="pachypes" status="sp. nov.">
<emphasis id="B91A4409EE7C7F14FE1DC0F0FDFFFD03" box="[497,625,657,681]" italics="true" pageId="26" pageNumber="2488">T. pachypes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229F972EE7C7F14FD96C0F0FD5DFD03" box="[634,723,657,681]" pageId="26" pageNumber="2488" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BD1981BEE7C7F14FF4CC0BDFD50FC4D" blockId="26.[160,1156,732,1727]" pageId="26" pageNumber="2488">
<emphasis id="B91A4409EE7C7F14FF4CC0BDFF72FD5E" bold="true" box="[160,252,732,757]" italics="true" pageId="26" pageNumber="2488">Female.</emphasis>
Genital double-somite (
<figureCitation id="1355849EEE7C7F14FDF6C0BDFD09FD5E" box="[538,647,732,756]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="26" pageNumber="2488">Figure 14</figureCitation>
(a,b)) wide, about 1.4 times wider than long (ratio of mounted urosome), and longer than following urosomites combined; lateral margins arched; lateral margins; posterior margins of genital double-somite and following urosomites with wide, irregularly undulated hyaline fringe; integument of cephalothorax and following somites, including caudal rami naked, devoid of structural ornaments; copulatory pore situated in middle of ventral surface of double-somite, directly leading to ovate field of compact tissue; anterior receptacle wide, only slightly expanded anteriorly; caudal receptacle not visible.
</paragraph>
<paragraph id="8BD1981BEE7C7F14FF2CC193FC59FBE5" blockId="26.[160,1156,732,1727]" pageId="26" pageNumber="2488">Anal operculum triangular, reaching beyond anal area, with serrate margins and transverse row of spinules on underside; posterior margin of anal somite without ornamentation dorsally, set with rigid spinules laterally and ventrally.</paragraph>
<paragraph id="8BD1981BEE7C7F14FF2CC63BFB0AFAE8" blockId="26.[160,1156,732,1727]" pageId="26" pageNumber="2488">
Caudal rami (
<figureCitation id="1355849EEE7C7F14FE8DC63BFE5CFBD8" box="[353,466,1114,1138]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="26" pageNumber="2488">Figure 14</figureCitation>
(a,b)) cylindrical, only slightly tapering caudally (cylindrical appearance in
<figureCitation id="1355849EEE7C7F14FEA9C61DFE3FFB3E" box="[325,433,1148,1172]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="26" pageNumber="2488">Figure 14</figureCitation>
(b) results from inclined position on slide), rami widely separated from each medially, and diverting; lateral seta inserted in middle of margin, without spinules at its insertion; distal lateral element spiniform, serrate along its outer margin, plumose at the inner margin; insertion of distal lateral element set with robust spinules; terminal principal ones with breaking plane; medial terminal seta somewhat shorter than inner medial margin of ramus, setiform; dorsal seta about twice as long as ramus.
</paragraph>
<paragraph id="8BD1981BEE7C7F15FF2CC72DFCAEFF63" blockId="26.[160,1156,732,1727]" lastBlockId="27.[160,1156,142,617]" lastPageId="27" lastPageNumber="2489" pageId="26" pageNumber="2488">
Antennule (
<figureCitation id="1355849EEE7C7F14FEAEC72DFE24FACE" box="[322,426,1356,1380]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="26" pageNumber="2488">Figure 15</figureCitation>
(b)) 10-segmented (ancestral segments
<collectionCode id="ED7F00DEEE7C7F14FCB2C72DFCFDFACE" box="[862,883,1356,1380]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="26" pageNumber="2488">VI</collectionCode>
to IX fused) with following armature:1[6]-2[7]-3[2]-4[1]-5[2]-6[3]-7[2 + Aesth]-8[2]-9[2 + Aesth]-10[7 + Aesth]; aesthetascs on segments 7 and 10 lingiform, aesthetasc on segment 9 filiform; setal armament flexible, finely pinnate; first segment without spinular ornamentation. Antenna (
<figureCitation id="1355849EEE7C7F14FF4BC7B6FE83FA45" box="[167,269,1495,1519]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="26" pageNumber="2488">Figure 15</figureCitation>
(a)) rather slim, not compact, without exopodite and a single abexopodal element; first endopodite segment with one seta, second segment with five setae, and terminal segment with six terminal setae; mandible (
<figureCitation id="1355849EEE7C7F14FD6AC47DFD61F99E" box="[646,751,1564,1588]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="26" pageNumber="2488">Figure 15</figureCitation>
(c)) with multi-cuspidate masticatory margin; palp absent, represented by single short seta. Maxillule (
<figureCitation id="1355849EEE7C7F14FC71C45EFB85F9FD" box="[925,1035,1599,1623]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="26" pageNumber="2488">Figure 15</figureCitation>
(e)) typical cyclopid with separate endopodite and with five smooth and one pinnate element along lateral margin of arthrite. Maxilla (
<figureCitation id="1355849EEE7C7F14FDF7C4E5FD0BF936" box="[539,645,1668,1692]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="26" pageNumber="2488">Figure 15</figureCitation>
(d)) with two-segmented endopodite, bearing two spiniform elements on proximal segment, one spiniform and two short naked elements on distal segment. Maxilliped (
<figureCitation id="1355849EEE7D7F15FE03C2EFFDD9FF0C" box="[495,599,142,166]" captionStart="Figure 15" captionStartId="24.[160,225,1509,1532]" captionTargetBox="[163,1153,142,1478]" captionTargetId="figure-1@24.[163,1153,142,1478]" captionTargetPageId="24" captionText="Figure 15. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Antenna, caudal; (b) female antennules; (c) mandible; (d) maxilla; (e) maxillule; (f) maxilliped, caudal; (g) proximal maxillipedal segment, frontal (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183793" httpUri="https://zenodo.org/record/5183793/files/figure.png" pageId="27" pageNumber="2489">Figure 15</figureCitation>
(f,g)) four-segmented, penultimate one with single, pinnate element, ultimate segment with two naked setae.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF2CC2B5FE1AFE9B" blockId="27.[160,1156,142,617]" pageId="27" pageNumber="2489">
Legs 1–4 as described in
<bibRefCitation id="EFFFE5EAEE7D7F15FE08C2B5FC9AFF46" author="Mirabdullayev IM &amp; Van Damme K &amp; Dumont HJ" box="[484,788,212,236]" pageId="27" pageNumber="2489" pagination="261 - 271" refId="ref19581" refString="Mirabdullayev IM, Van Damme K, Dumont HJ. 2002. Freshwater cyclopoids (Crustacea: Copepoda) from the Socotra Archipelago, Yemen with the description of a new species of Bryocyclops. Fauna Arab. 19: 261 - 271." type="journal article" year="2002">Mirabdullayev et al. (2002)</bibRefCitation>
; setal armament sturdy, lanceolate; ornaments along inner margin of exopodites and outer margins of endopodites (
<figureCitation id="1355849EEE7D7F15FF44C378FE9BFE9B" box="[168,277,281,305]" captionStart="Figure 16" captionStartId="25.[160,225,1612,1635]" captionTargetBox="[163,1153,142,1581]" captionTargetId="figure-1@25.[163,1153,142,1581]" captionTargetPageId="25" captionText="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="27" pageNumber="2489">Figure 16</figureCitation>
(b,d)) hairy.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF2CC35DFF57FDAB" blockId="27.[160,1156,142,617]" pageId="27" pageNumber="2489">
Leg 5 (
<figureCitation id="1355849EEE7D7F15FEFBC35DFE0BFEFE" box="[279,389,316,340]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="27" pageNumber="2489">Figure 14</figureCitation>
(a)) completely confluent with the pediger, represented by three armature elements; seta representing basal segment inserted laterally on somite; armament representing original distal segment arising from large truncated outgrowth of pediger; ventrally located and caudally directed; outer element, slightly subdistally inserted, setiform, inner one robust, spiniform and serrate; both elements both equally long.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF2CC06DFDC5FDC3" blockId="27.[160,1156,142,617]" pageId="27" pageNumber="2489">
Leg 6 vestiges (
<figureCitation id="1355849EEE7D7F15FE82C06DFE57FD8E" box="[366,473,524,548]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="27" pageNumber="2489">Figure 14</figureCitation>
(b)) located dorsally in anterior half of genital double-somite; leg vestige rigidly sclerotized, bearing three smooth and short armature elements, middle one shorter than both others.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF4CC0F7FD5DFC93" blockId="27.[160,1155,662,1241]" pageId="27" pageNumber="2489">
<emphasis id="B91A4409EE7D7F15FF4CC0F7FF6EFD04" bold="true" box="[160,224,662,687]" italics="true" pageId="27" pageNumber="2489">Male.</emphasis>
Urosomites ornamentated as in female, but second and third ones not fused. Leg 5 (
<figureCitation id="1355849EEE7D7F15FF50C0D8FEA9FD7B" box="[188,295,697,721]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="27" pageNumber="2489">Figure 14</figureCitation>
(c)) basically as in female, process representing distal segment less wide. Leg 6 vestiges (
<figureCitation id="1355849EEE7D7F15FEC1C0BDFE10FD5E" box="[301,414,732,756]" captionStart="Figure 14" captionStartId="23.[160,225,1677,1700]" captionTargetBox="[212,1104,142,1646]" captionTargetId="figure-1@23.[212,1104,142,1646]" captionTargetPageId="23" captionText="Figure 14. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Female urosome, dorsal; (b) female abdominal somites, ventral; (c) male urosome, ventral (illustrations from type series, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183791" httpUri="https://zenodo.org/record/5183791/files/figure.png" pageId="27" pageNumber="2489">Figure 14</figureCitation>
(c)) symmetrical, rectangular, rigid with smooth surface; outer posterior edge slightly expanded, bearing three elements: medial one spiniform (smooth?), middle and outer one sturdy setiform.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF2CC125FF78FB86" blockId="27.[160,1155,662,1241]" pageId="27" pageNumber="2489">
Antennule (
<figureCitation id="1355849EEE7D7F15FEA7C125FE35FCF6" box="[331,443,836,860]" captionStart="Figure 16" captionStartId="25.[160,225,1612,1635]" captionTargetBox="[163,1153,142,1581]" captionTargetId="figure-1@25.[163,1153,142,1581]" captionTargetPageId="25" captionText="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="27" pageNumber="2489">Figure 16</figureCitation>
(a)) 17-segmented with terminal segment complexes [XXIV– XV] and [XXVI–XXVIII] separated; setal armament as in
<taxonomicName id="4C6EE398EE7D7F15FCABC107FC5EFCD4" authorityName="Fiers &amp; Van Damme" authorityYear="2017" box="[839,976,870,894]" class="Hexanauplia" family="Cyclopidae" genus="Thalamocyclops" kingdom="Animalia" order="Cyclopoida" pageId="27" pageNumber="2489" phylum="Arthropoda" rank="species" species="pachypes">
<emphasis id="B91A4409EE7D7F15FCABC107FC5EFCD4" box="[839,976,870,894]" italics="true" pageId="27" pageNumber="2489">T. pachypes</emphasis>
</taxonomicName>
, but elements less sturdy; first segment with three linguiform aesthetascs, segments 4, 9 and 12 each with one linguiform aesthetasc, all of moderate length; segment 16 and 17 each with one short filiform aesthetasc (bithek of ultimate segment lost on antennule in
<figureCitation id="1355849EEE7D7F15FF11C190FEE1FBA3" box="[253,367,1009,1033]" captionStart="Figure 16" captionStartId="25.[160,225,1612,1635]" captionTargetBox="[163,1153,142,1581]" captionTargetId="figure-1@25.[163,1153,142,1581]" captionTargetPageId="25" captionText="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="27" pageNumber="2489">Figure 16</figureCitation>
(a)). Mouthparts, leg 1 and 2, and exopodite of legs 3 and 4 as in female.
</paragraph>
<paragraph id="8BD1981BEE7D7F15FF2CC657FC98FB73" blockId="27.[160,1155,662,1241]" pageId="27" pageNumber="2489">
Leg 3 endopodite (
<figureCitation id="1355849EEE7D7F15FE4AC657FD9BFBE4" box="[422,533,1078,1102]" captionStart="Figure 16" captionStartId="25.[160,225,1612,1635]" captionTargetBox="[163,1153,142,1581]" captionTargetId="figure-1@25.[163,1153,142,1581]" captionTargetPageId="25" captionText="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="27" pageNumber="2489">Figure 16</figureCitation>
(c)) inner proximal pair of setae on second segment setiform, proportionally shorter than in female; inner subdistal and inner distal one setiform armed with short sturdy setules; outer distal element long, hook-shaped, with serrate rounded expansion in distal half of outer margin. Leg 4 endopodite (
<figureCitation id="1355849EEE7D7F15FC1AC6FFFBEEFB1C" box="[1014,1120,1182,1206]" captionStart="Figure 16" captionStartId="25.[160,225,1612,1635]" captionTargetBox="[163,1153,142,1581]" captionTargetId="figure-1@25.[163,1153,142,1581]" captionTargetPageId="25" captionText="Figure 16. Thalamocyclops soqotraensis (Mirabdullayev et al., 2002) comb. nov. (a) Male antennule; (b), leg 3 endopodite of female, frontal; (c) leg 3 endopodite of male, caudal; (d) leg 4 endopodite of female, frontal; (e) leg 4 endopodite of male, caudal (illustrations from types, RBINSc COP 4492–4495)." figureDoi="http://doi.org/10.5281/zenodo.5183795" httpUri="https://zenodo.org/record/5183795/files/figure.png" pageId="27" pageNumber="2489">Figure 16</figureCitation>
(e)) distinctly two-segmented, with armament as in female.
</paragraph>
</subSubSection>
</treatment>
</document>
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