735 lines
83 KiB
XML
735 lines
83 KiB
XML
<document id="87C741D0532DDCA864807AAD3999AEC4" ID-CLB-Dataset="45392" ID-DOI="10.11646/zootaxa.3821.4.7" ID-GBIF-Dataset="35d5dd40-f2f5-4553-8c8d-8ac4c308e8e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="228521" ID-ZooBank="8DC046AC-C047-44FE-86E0-B4B785F0BFFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460442744190" checkinUser="plazi" docAuthor="Grismer, L. Lee, Riyanto, Awal, Iskandar, Djoko T. & Mcguire, Jimmy A." docDate="2014" docId="03A0F001C65EFFA4FF01FBE4FD3CFB5E" docLanguage="en" docName="zt03821p495.pdf" docOrigin="Zootaxa 3821 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemiphyllodactylus engganoensis Grismer, Riyanto, Iskandar & Mcguire, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="494" masterDocId="FF998879C65CFFADFF96FF9FFFA6FFFE" masterDocTitle="A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia" masterLastPageNumber="495" masterPageNumber="485" pageNumber="487" updateTime="1698333035416" updateUser="ExternalLinkService">
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<mods:mods id="F2EB13987CD1B30110A6DFB80C0AC6EC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="82014DD1F8DF33C4808E6DBF96F3C277">
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<mods:title id="5CF19C2328C7A14F3391ED6C99E00CA2">A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Pulau Enggano, southwestern Sumatra, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="CFC5FB0254BE0D8B0EFC77EDF5B72456" type="personal">
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<mods:role id="E8952C396A60F3A1353CE88FDE1A4E6E">
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<mods:roleTerm id="506F7F2C3D4283C8D1505A2106C5D001">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ACC70D8CE8EA298A894E72204DF55A89">Grismer, L. Lee</mods:namePart>
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</mods:name>
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<mods:name id="76B971FA78036FD5EA8FA10B51E2429C" type="personal">
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<mods:role id="5A9B3EE126F5189D3A59065A5ADFC34E">
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<mods:roleTerm id="A3911D23781E831E011C02CB2482D254">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="23048B3812009A54A0C2EA24DC94D47D">Riyanto, Awal</mods:namePart>
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</mods:name>
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<mods:name id="BBEA7AB0CA55AA36D601744F57BD784D" type="personal">
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<mods:roleTerm id="5566C57CC54E34D2AA287CD744132E79">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35ADF349F477212D56969D6FE96E4379">Iskandar, Djoko T.</mods:namePart>
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</mods:name>
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<mods:name id="E76D4D556FE62B85C64921649DF05598" type="personal">
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<mods:role id="D5087182CE11E3EF5C9AA523B8E35325">
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<mods:roleTerm id="192A54AC90C199B460160B8E956E2CB1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4E5574116C5A06354FF2C47795DDF51C">Mcguire, Jimmy A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7396D7EABCFD3EC024258DC8D80A5E2C">text</mods:typeOfResource>
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<mods:titleInfo id="727BEA7E9C4D678001A0614D0BB5A7C8">
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<mods:title id="7ADEF3B995D202C2B1A9FA3B96EE9246">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1FD46DD5F1D9717C01AF4E40322A91CC">
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<mods:date id="007A63E48DAB90076652379B1CAD40F0">2014</mods:date>
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<mods:detail id="7AB29C1CB4A5F4671503808DB3B5E617" type="volume">
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<mods:number id="63405ED69DEB2986FCCCF939FB96F1FB">3821</mods:number>
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</mods:detail>
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<mods:detail id="D5FFB2FF1E61AF2C411D5B484BA0FE5F" type="issue">
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<mods:number id="A51969916FB136E955529B58A9147192">4</mods:number>
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<mods:start id="EE224850F141E6AEA52885DB1A96E413">485</mods:start>
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<mods:end id="6257332C0C53F267C92B5B34090281AD">495</mods:end>
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<mods:classification id="28C07996B4533E0569DD7A0D24041DC6">journal article</mods:classification>
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<mods:identifier id="82BF41719FF0C74E92A488F131512104" type="CLB-Dataset">45392</mods:identifier>
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<mods:identifier id="F517E60144E3F749B067F697E628D01F" type="DOI">10.11646/zootaxa.3821.4.7</mods:identifier>
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<mods:identifier id="DF75848F1A3112E92A3F234E69F13903" type="GBIF-Dataset">35d5dd40-f2f5-4553-8c8d-8ac4c308e8e5</mods:identifier>
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<mods:identifier id="8001FF8328F2C78D0F2B1CCBE36CF9D4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A0F001C65EFFA4FF01FBE4FD3CFB5E" ID-DOI="http://doi.org/10.5281/zenodo.5682094" ID-GBIF-Taxon="119508804" ID-Zenodo-Dep="5682094" LSID="urn:lsid:plazi:treatment:03A0F001C65EFFA4FF01FBE4FD3CFB5E" httpUri="http://treatment.plazi.org/id/03A0F001C65EFFA4FF01FBE4FD3CFB5E" lastPageId="9" lastPageNumber="494" pageId="2" pageNumber="487">
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<subSubSection id="C313129CC65EFFAFFF01FBE4FF59FAFE" pageId="2" pageNumber="487" type="nomenclature">
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<paragraph id="8BB64117C65EFFAFFF01FBE4FD24FB6A" blockId="2.[151,642,1147,1280]" box="[151,642,1147,1172]" pageId="2" pageNumber="487">
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<heading id="D0FEF67BC65EFFAFFF01FBE4FD24FB6A" bold="true" box="[151,642,1147,1172]" fontSize="10" level="2" pageId="2" pageNumber="487" reason="2">
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<emphasis id="B97D9D05C65EFFAFFF01FBE4FD24FB6A" bold="true" box="[151,642,1147,1172]" pageId="2" pageNumber="487">
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<taxonomicName id="4C093A94C65EFFAFFF01FBE4FD87FB6A" box="[151,545,1147,1172]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="487" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
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<emphasis id="B97D9D05C65EFFAFFF01FBE4FD87FB6A" bold="true" box="[151,545,1147,1172]" italics="true" pageId="2" pageNumber="487">Hemiphyllodactylus engganoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E207EC65EFFAFFDBEFBE3FD24FB6A" box="[552,642,1148,1172]" pageId="2" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="8BB64117C65EFFAFFF01FB00FE34FB23" blockId="2.[151,642,1147,1280]" pageId="2" pageNumber="487">
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<collectingCountry id="F31E0187C65EFFAFFF01FB00FF71FB46" box="[151,215,1183,1208]" name="Iceland" pageId="2" pageNumber="487">Pulau</collectingCountry>
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Enggano Dwarf Gecko Cicak Kerdil Enggano
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</paragraph>
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||
<paragraph id="8BB64117C65EFFAFFF01FB78FF59FAFE" blockId="2.[151,642,1147,1280]" box="[151,255,1255,1280]" pageId="2" pageNumber="487">
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<figureCitation id="13325D92C65EFFAFFF01FB78FF42FAFE" box="[151,228,1255,1280]" captionStart="FIGURE 3" captionStartId="6.[151,250,1051,1073]" captionTargetBox="[313,1273,395,1029]" captionTargetId="figure@6.[313,1273,395,1029]" captionTargetPageId="6" captionText="FIGURE 3. Paratype MVZ 239345 (subadult female) of Hemiphyllodactylus engganoensis sp. nov. from Pulau Enggano, Indonesia." httpUri="https://zenodo.org/record/228524/files/figure.png" pageId="2" pageNumber="487">Figs. 3</figureCitation>
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,
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<figureCitation id="13325D92C65EFFAFFF67FB77FF59FAFE" box="[241,255,1256,1280]" captionStart="FIGURE 4" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[179,1407,265,1934]" captionTargetId="figure@8.[179,1408,265,1934]" captionTargetPageId="8" captionText="FIGURE 4. Type series of Hemiphyllodactylus engganoensis sp. nov from Pulau Enggano, Indonesia. Upper: holotype MZB. Lace 4568 (adult female). Middle: paratype MVZ 239346 (adult male). Lower: paratype MVZ 239345 (subadult female)." httpUri="https://zenodo.org/record/228525/files/figure.png" pageId="2" pageNumber="487">4</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313129CC65EFFAFFF01FAAFFEA6FA26" pageId="2" pageNumber="487" type="materials_examined">
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<paragraph id="8BB64117C65EFFAFFF01FAAFFA95FA6E" blockId="2.[151,1437,1327,2037]" pageId="2" pageNumber="487">
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<emphasis id="B97D9D05C65EFFAFFF01FAAFFEADFAB6" bold="true" box="[151,267,1328,1352]" pageId="2" pageNumber="487">
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<typeStatus id="54B2FFB5C65EFFAFFF01FAAFFEA0FAB6" box="[151,262,1328,1352]" pageId="2" pageNumber="487" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult female (
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<collectionCode id="ED18D9D2C65EFFAFFE25FAAFFE54FAB6" box="[435,498,1328,1352]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="2" pageNumber="487">MZB</collectionCode>
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||
.Lace 4568) collected at 2220 hours on
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<date id="FFB767D7C65EFFAFFC3DFAAFFBA4FAB6" box="[939,1026,1328,1352]" pageId="2" pageNumber="487" value="2011-05">11 May</date>
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, 2013 by Jimmy A. McGuire, Djoko T. Iskandar, Awal Riyanto, and Mulyadi near the village of Malakoni on the island of Enggano (Kecamatan Enggano, Kabupaten Bengkulu, Propinsi Bengkulu,
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<collectingCountry id="F31E0187C65EFFAFFD75FAE8FCF2FA6E" box="[739,852,1399,1424]" name="Indonesia" pageId="2" pageNumber="487">Indonesia</collectingCountry>
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;
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||
<geoCoordinate id="EE3D27D0C65EFFAFFCC9FAE7FC46FA6E" box="[863,992,1399,1424]" direction="south" orientation="latitude" pageId="2" pageNumber="487" precision="1" value="-0.089215">S 0 5.35290</geoCoordinate>
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,
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<geoCoordinate id="EE3D27D0C65EFFAFFC7AFAE7FBD8FA6E" box="[1004,1150,1400,1424]" direction="east" orientation="longitude" pageId="2" pageNumber="487" precision="1" value="102.27742">E 102.27742</geoCoordinate>
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,
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<quantity id="4CF1ECF2C65EFFAFFB1FFAE7FB12FA71" box="[1161,1204,1400,1424]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="487" unit="m" value="1.0">1 m</quantity>
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elevation).
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||
</paragraph>
|
||
<paragraph id="8BB64117C65EFFAFFF51FA03FEA6FA26" blockId="2.[151,1437,1327,2037]" pageId="2" pageNumber="487">
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<typeStatus id="54B2FFB5C65EFFAFFF51FA03FE92FA4A" box="[199,308,1436,1460]" pageId="2" pageNumber="487" type="paratype">
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||
<emphasis id="B97D9D05C65EFFAFFF51FA03FE92FA4A" bold="true" box="[199,308,1436,1460]" pageId="2" pageNumber="487">Paratype</emphasis>
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||
</typeStatus>
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||
. Subadult female (
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||
<collectionCode id="ED18D9D2C65EFFAFFD91FA03FDE1FA4A" LSID="urn:lsid:biocol.org:col:34777" box="[519,583,1436,1460]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="487">MVZ</collectionCode>
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239345) and adult male (
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||
<collectionCode id="ED18D9D2C65EFFAFFCF3FA03FC03FA4A" LSID="urn:lsid:biocol.org:col:34777" box="[869,933,1436,1460]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="2" pageNumber="487">MVZ</collectionCode>
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239346) bear the same collection data as the
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<typeStatus id="54B2FFB5C65EFFAFFF01FA20FF5BFA26" box="[151,253,1471,1496]" pageId="2" pageNumber="487" type="holotype">holotype</typeStatus>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313129CC65EFFAFFF51FA7CFEA5F906" pageId="2" pageNumber="487" type="diagnosis">
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||
<paragraph id="8BB64117C65EFFAFFF51FA7CFEA5F906" blockId="2.[151,1437,1327,2037]" pageId="2" pageNumber="487">
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||
<emphasis id="B97D9D05C65EFFAFFF51FA7CFEE7FA02" bold="true" box="[199,321,1507,1532]" pageId="2" pageNumber="487">Diagnosis.</emphasis>
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<taxonomicName id="4C093A94C65EFFAFFEDEFA7BFD6EFA02" box="[328,712,1508,1532]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="487" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
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||
<emphasis id="B97D9D05C65EFFAFFEDEFA7BFD6EFA02" box="[328,712,1508,1532]" italics="true" pageId="2" pageNumber="487">Hemiphyllodactylus engganoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24E207EC65EFFAFFD59FA7BFC8FFA02" box="[719,809,1508,1532]" pageId="2" pageNumber="487" rank="species">
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<emphasis id="B97D9D05C65EFFAFFD59FA7BFC8FFA02" bold="true" box="[719,809,1508,1532]" pageId="2" pageNumber="487">sp. nov.</emphasis>
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</taxonomicNameLabel>
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is differentiated from all other congeners by having the unique combination of a maximum SVL of
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<quantity id="4CF1ECF2C65EFFAFFD19F997FD5EF9E1" box="[655,760,1544,1568]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="2" pageNumber="487" unit="mm" value="37.3">37.3 mm</quantity>
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; six chin scales; no enlarged postmentals; five circumnasal scales; three or four scales between the supranasals; 12 supralabials; 24 or 25 dorsal scales; 14 ventral scales; a lamellar hand formula of 4554 or 4454; a lamellar foot formula of 4555; four or five subdigital lamellae on the first finger; four or five subdigital lamellae on the first toe; a contiguous femoroprecloacal pore series of 42; five cloacal spurs in males; no enlarged subcaudal scales; no dark postorbital stripes or striping on body; small dark blotches on dorsum; a yellowish postsacral mark bearing anteriorly projecting arms; and a pigmented caecum and gonads. (Table 1).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C313129CC65EFFAAFF51F89CFD97F80D" lastPageId="7" lastPageNumber="492" pageId="2" pageNumber="487" type="description">
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<paragraph id="8BB64117C65EFFABFF51F89CFBB8FE9B" blockId="2.[151,1437,1327,2037]" lastBlockId="6.[151,1436,151,357]" lastPageId="6" lastPageNumber="491" pageId="2" pageNumber="487">
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||
<emphasis id="B97D9D05C65EFFAFFF51F89CFE4EF8E2" bold="true" box="[199,488,1795,1820]" pageId="2" pageNumber="487">
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||
Description of
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||
<typeStatus id="54B2FFB5C65EFFAFFEECF89BFE45F8E2" box="[378,483,1796,1820]" pageId="2" pageNumber="487" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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Adult female; head triangular in dorsal profile depressed, distinct from neck; lores and interorbital regions flat; rostrum relatively long (NarEye/ HeadL = 0.30); prefrontal region flat to weakly concave; canthus rostralis smoothly rounded, barely discernable; snout moderate, rounded in dorsal profile; eye large; ear opening oval, small; eye to ear distance greater than diameter of eye; rostral wider than high, partially divided dorsally, bordered posteriorly by large supranasals; three internasals (=postnasals); external nares bordered anteriorly by rostral, dorsally by supranasal, posteriorly by two postnasals, ventrally by first supralabial (collectively the circumnasals 5 R, L); 12 (R, L) square supralabials tapering to below posterior margin of orbit; 12 (R, L) square infralabials tapering to below posterior margin of orbit; scales of rostrum and lores, raised; scales on top of head and occiput small, granular; dorsal superciliaries raised, rectangular; mental triangular, bordered laterally by first infralabials and posteriorly by two non-enlarged postmentals; each postmental bordered laterally by a single sublabial; row of smaller scales extending transversely from juncture of second and third infralabials and contacting mental; gular scales triangular, small, granular, grading posteriorly into slightly larger, subimbricate throat and pectoral scales which grade into slightly larger, subimbricate ventrals.
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||
</paragraph>
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||
<caption id="DF76119FC659FFA8FF01F83EFBBEF828" httpUri="https://zenodo.org/record/228523/files/figure.png" pageId="5" pageNumber="490" targetBox="[181,1420,202,1923]" targetPageId="5">
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||
<paragraph id="8BB64117C659FFA8FF01F83EFBBEF828" blockId="5.[151,1436,1953,2006]" pageId="5" pageNumber="490">
|
||
<emphasis id="B97D9D05C659FFA8FF01F83EFEBEF848" bold="true" box="[151,280,1953,1975]" pageId="5" pageNumber="490">FIGURE 2.</emphasis>
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Maximum-likelihood phylogram (–lnL 22097.690183) of the genus
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<taxonomicName id="4C093A94C659FFA8FC6DF83EFAC2F849" authority="Bleeker, 1860" authorityName="Bleeker" authorityYear="1860" box="[1019,1380,1953,1975]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="490" phylum="Chordata" rank="genus">
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||
<emphasis id="B97D9D05C659FFA8FC6DF83EFB60F849" box="[1019,1222,1953,1975]" italics="true" pageId="5" pageNumber="490">Hemiphyllodactylus</emphasis>
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||
Bleeker, 1860
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</taxonomicName>
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with Bayesian posterior probabilities and maximum-likelihood bootstrap values, respectively.
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||
</paragraph>
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||
</caption>
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||
<caption id="DF76119FC65AFFABFF01FB84FEA7FBB1" httpUri="https://zenodo.org/record/228524/files/figure.png" pageId="6" pageNumber="491" targetBox="[313,1273,395,1029]" targetPageId="6">
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||
<paragraph id="8BB64117C65AFFABFF01FB84FEA7FBB1" blockId="6.[151,1436,1051,1103]" pageId="6" pageNumber="491">
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||
<emphasis id="B97D9D05C65AFFABFF01FB84FEB0FBCE" bold="true" box="[151,278,1051,1073]" pageId="6" pageNumber="491">FIGURE 3.</emphasis>
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Paratype MVZ 239345 (subadult female) of
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||
<taxonomicName id="4C093A94C65AFFABFD61FB84FBF0FBCF" box="[759,1110,1051,1073]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="491" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C65AFFABFD61FB84FBF0FBCF" box="[759,1110,1051,1073]" italics="true" pageId="6" pageNumber="491">Hemiphyllodactylus engganoensis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A24E207EC65AFFABFBC9FB84FB15FBCE" box="[1119,1203,1051,1072]" pageId="6" pageNumber="491" rank="species">
|
||
<emphasis id="B97D9D05C65AFFABFBC9FB84FB15FBCE" bold="true" box="[1119,1203,1051,1072]" pageId="6" pageNumber="491">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
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||
from Pulau Enggano, Indonesia.
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||
</paragraph>
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||
</caption>
|
||
<paragraph id="8BB64117C65AFFABFF51FBE9FCD1F955" blockId="6.[151,1437,1142,2031]" pageId="6" pageNumber="491">
|
||
Body elongate, dorsoventrally compressed; ventrolateral folds absent; dorsal scales small, granular, 25 scales contained within one eye diameter; ventral scales flat, subimbricate, much larger than dorsal scales, 14 scales contained within one eye diameter; no enlarged, precloacal scales; no pore-bearing scales femoral or precloacal scales; forelimbs short, robust in stature, covered with granular scales dorsally and slightly larger, flat, subimbricate scales ventrally; palmar scales flat, subimbricate; all digits except digit I well-developed; digit I vestigial, clawless; distal, subdigital lamellae of digits II–V divided, angular and U-shaped; lamellae proximal to these transversely expanded, undivided; lamellar formula of digits
|
||
<date id="FFB767D7C65AFFABFC15FACFFC43FA99" box="[899,997,1358,1383]" pageId="6" pageNumber="491">II–V 4-4</date>
|
||
-5-4 (R, L); four transversely expanded lamellae on digit I; claws on digits II–V well-developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; hind limbs short, more robust than forelimbs, covered with slightly pointed, juxtaposed scales dorsally and by larger, flat, subimbricate scales ventrally; plantar scales low, flat, subimbricate; all digits except digit I well-developed; digit I vestigial, clawless; distal, subdigital lamellae of digits II–V divided, angular and U-shaped; lamellae proximal to U-shaped lamellae transversely expanded, undivided; lamellar formula of digits
|
||
<date id="FFB767D7C65AFFABFD27F9B7FCB5F9C1" box="[689,787,1574,1599]" pageId="6" pageNumber="491">II–V 4-5</date>
|
||
-5-5 (R, L); five transversely expanded lamellae on digit I; claws on digits II–V well-developed, unsheathed; distal portions of digits strongly curved, terminal joint free, arising from central portion of lamellar pad; all caudal scales flat, imbricate, not occurring in caudal segments, no enlarged subcaudals. Morphometric data are presented in
|
||
<tableCitation id="C68B74ACC65AFFABFC8BF90CFCD5F955" box="[797,883,1683,1708]" captionStart="TABLE 2" captionStartId="7.[151,239,332,355]" captionTargetBox="[159,1239,456,1594]" captionTargetPageId="7" captionText="TABLE 2. Scale counts, color pattern characteristics, and morphometric ratios of the type series of Hemiphyllodactylus engganoensis sp. nov. (variable features only)." httpUri="http://table.plazi.org/id/DF76119FC65BFFAAFF01FED3FD2CFE7D" pageId="6" pageNumber="491" tableUuid="DF76119FC65BFFAAFF01FED3FD2CFE7D">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB64117C65AFFABFF51F929FBE5F8A1" blockId="6.[151,1437,1142,2031]" pageId="6" pageNumber="491">
|
||
<emphasis id="B97D9D05C65AFFABFF51F929FE6DF931" bold="true" box="[199,459,1718,1743]" pageId="6" pageNumber="491">Coloration in alcohol.</emphasis>
|
||
Top of head, body and limbs nearly unicolor tan; ground color of dorsal surface of tail tan bearing nine, dark-colored, diffuse bands; lores slightly darker; no postorbital striping; faint, diffuse lightcolored spots on dorsum barely discernable; light-colored, postsacral marking bearing faint, anteriorly projecting arms; ground color of gular region beige with small, dark-brown spots; ground color of ventral surfaces of body and limbs beige, immaculate; dorsal coloration invades lateral sections of abdomen.
|
||
</paragraph>
|
||
<paragraph id="8BB64117C65AFFAAFF51F8F5FD45FEE2" blockId="6.[151,1437,1142,2031]" lastBlockId="7.[151,1436,151,285]" lastPageId="7" lastPageNumber="492" pageId="6" pageNumber="491">
|
||
<emphasis id="B97D9D05C65AFFABFF51F8F5FE9DF87D" bold="true" box="[199,315,1898,1923]" pageId="6" pageNumber="491">Variation.</emphasis>
|
||
The
|
||
<typeStatus id="54B2FFB5C65AFFABFEECF8F3FE7CF87A" box="[378,474,1900,1924]" pageId="6" pageNumber="491" type="paratype">paratype</typeStatus>
|
||
MVZ 239345 approaches the
|
||
<typeStatus id="54B2FFB5C65AFFABFCA1F8F4FC3FF87A" box="[823,921,1899,1924]" pageId="6" pageNumber="491" type="holotype">holotype</typeStatus>
|
||
in general dorsal coloration and pattern (
|
||
<figureCitation id="13325D92C65AFFABFAF9F8F4FF01F859" captionStart="FIGURE 3" captionStartId="6.[151,250,1051,1073]" captionTargetBox="[313,1273,395,1029]" captionTargetId="figure@6.[313,1273,395,1029]" captionTargetPageId="6" captionText="FIGURE 3. Paratype MVZ 239345 (subadult female) of Hemiphyllodactylus engganoensis sp. nov. from Pulau Enggano, Indonesia." httpUri="https://zenodo.org/record/228524/files/figure.png" pageId="6" pageNumber="491">Fig. 3</figureCitation>
|
||
,
|
||
<figureCitation id="13325D92C65AFFABFF25F810FF62F859" box="[179,196,1935,1959]" captionStart="FIGURE 4" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[179,1407,265,1934]" captionTargetId="figure@8.[179,1408,265,1934]" captionTargetPageId="8" captionText="FIGURE 4. Type series of Hemiphyllodactylus engganoensis sp. nov from Pulau Enggano, Indonesia. Upper: holotype MZB. Lace 4568 (adult female). Middle: paratype MVZ 239346 (adult male). Lower: paratype MVZ 239345 (subadult female)." httpUri="https://zenodo.org/record/228525/files/figure.png" pageId="6" pageNumber="491">4</figureCitation>
|
||
). In life, the dorsal ground color of the head is dull-yellow and that of the body, limbs, and tail is tan with faint, darker bands. All dorsal surfaces are overlain with a reticulum of darker markings that tend to form thin, zig-zag lines across the body and wider bands on the tail. The light-colored, postsacral marking is more vivid. The iris is silver. The tail is complete, original, and round in cross-section (
|
||
<figureCitation id="13325D92C65BFFAAFC01FF08FC47FF4E" box="[919,993,151,176]" captionStart="FIGURE 4" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[179,1407,265,1934]" captionTargetId="figure@8.[179,1408,265,1934]" captionTargetPageId="8" captionText="FIGURE 4. Type series of Hemiphyllodactylus engganoensis sp. nov from Pulau Enggano, Indonesia. Upper: holotype MZB. Lace 4568 (adult female). Middle: paratype MVZ 239346 (adult male). Lower: paratype MVZ 239345 (subadult female)." httpUri="https://zenodo.org/record/228525/files/figure.png" pageId="7" pageNumber="492">Fig. 4</figureCitation>
|
||
). The male
|
||
<typeStatus id="54B2FFB5C65BFFAAFBE2FF07FB72FF4E" box="[1140,1236,152,176]" pageId="7" pageNumber="492" type="paratype">paratype</typeStatus>
|
||
MVZ 239346 is missing a tail and has a uniform tan dorsal ground color (
|
||
<figureCitation id="13325D92C65BFFAAFCF3FF23FC16FF2A" box="[869,944,188,213]" captionStart="FIGURE 4" captionStartId="8.[151,250,1956,1978]" captionTargetBox="[179,1407,265,1934]" captionTargetId="figure@8.[179,1408,265,1934]" captionTargetPageId="8" captionText="FIGURE 4. Type series of Hemiphyllodactylus engganoensis sp. nov from Pulau Enggano, Indonesia. Upper: holotype MZB. Lace 4568 (adult female). Middle: paratype MVZ 239346 (adult male). Lower: paratype MVZ 239345 (subadult female)." httpUri="https://zenodo.org/record/228525/files/figure.png" pageId="7" pageNumber="492">Fig. 4</figureCitation>
|
||
). It also has a continuous series of 42 femoroprecloacal pores and five cloacal spurs, both of which are lacking in the two female specimens. Differences in meristics and morphometrics are listed in
|
||
<tableCitation id="C68B74ACC65BFFAAFD1FFE9BFD79FEE2" box="[649,735,260,285]" captionStart="TABLE 2" captionStartId="7.[151,239,332,355]" captionTargetBox="[159,1239,456,1594]" captionTargetPageId="7" captionText="TABLE 2. Scale counts, color pattern characteristics, and morphometric ratios of the type series of Hemiphyllodactylus engganoensis sp. nov. (variable features only)." httpUri="http://table.plazi.org/id/DF76119FC65BFFAAFF01FED3FD2CFE7D" pageId="7" pageNumber="492" tableUuid="DF76119FC65BFFAAFF01FED3FD2CFE7D">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF76119FC65BFFAAFF01FED3FD2CFE7D" ID-Table-UUID="DF76119FC65BFFAAFF01FED3FD2CFE7D" httpUri="http://table.plazi.org/id/DF76119FC65BFFAAFF01FED3FD2CFE7D" pageId="7" pageNumber="492" targetBox="[159,1239,456,1594]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8BB64117C65BFFAAFF01FED3FD2CFE7D" blockId="7.[151,1436,331,387]" pageId="7" pageNumber="492">
|
||
<emphasis id="B97D9D05C65BFFAAFF01FED3FEACFE9D" bold="true" box="[151,266,332,355]" pageId="7" pageNumber="492">TABLE 2.</emphasis>
|
||
Scale counts, color pattern characteristics, and morphometric ratios of the type series of
|
||
<taxonomicName id="4C093A94C65BFFAAFB51FED4FE8EFE7D" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="492" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C65BFFAAFB51FED4FE8EFE7D" italics="true" pageId="7" pageNumber="492">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC65BFFAAFEB8FEF3FE22FE7D" box="[302,388,364,387]" pageId="7" pageNumber="492" rank="species">
|
||
<emphasis id="B97D9D05C65BFFAAFEB8FEF3FE22FE7D" bold="true" box="[302,388,364,387]" pageId="7" pageNumber="492">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(variable features only).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB64117C65BFFAAFDF5FE00FD06F905" blockId="7.[611,1208,415,436]" lastBlockId="7.[151,1437,1655,2035]" pageId="7" pageNumber="492">
|
||
MZB.Lace MVZ MVZ
|
||
<emphasis id="B97D9D05C65BFFAAFF51F9E8FEC6F96E" bold="true" box="[199,352,1655,1680]" pageId="7" pageNumber="492">Distribution.</emphasis>
|
||
<taxonomicName id="4C093A94C65BFFAAFEF1F9E7FD40F971" box="[359,742,1656,1680]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="492" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C65BFFAAFEF1F9E7FD40F971" box="[359,742,1656,1680]" italics="true" pageId="7" pageNumber="492">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC65BFFAAFD78F9E7FCEEF96E" box="[750,840,1656,1680]" pageId="7" pageNumber="492" rank="species">
|
||
<emphasis id="B97D9D05C65BFFAAFD78F9E7FCEEF96E" bold="true" box="[750,840,1656,1680]" pageId="7" pageNumber="492">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is known only from the
|
||
<typeStatus id="54B2FFB5C65BFFAAFBCAF9E7FB2AF96E" box="[1116,1164,1656,1680]" pageId="7" pageNumber="492">type</typeStatus>
|
||
locality near the village of Malakoni on the island of Enggano. We presume it to be endemic to the entire island, though it may be restricted to Enggano’s coastal perimeter. All three specimens in the original series were collected within a few hundred meters of one another in beachside vegetation.
|
||
</paragraph>
|
||
<paragraph id="8BB64117C65BFFAAFDF5FE57FB0AF9C4" pageId="7" pageNumber="492">
|
||
<table id="F909B3B7C65B0052FF09FE57FB71F9C4" box="[159,1239,456,1594]" gridcols="4" gridrows="28" pageId="7" pageNumber="492">
|
||
<tr id="35394355C65B0052FF09FE57FB71FE20" box="[159,1239,456,478]" gridrow="0" pageId="7" pageNumber="492" rowspan-0="1">
|
||
<th id="76E82A29C65B0052FDF5FE57FD62FE20" box="[611,708,456,478]" gridcol="1" gridrow="0" pageId="7" pageNumber="492">4568</th>
|
||
<th id="76E82A29C65B0052FC14FE57FC7CFE20" box="[898,986,456,478]" gridcol="2" gridrow="0" pageId="7" pageNumber="492">239345</th>
|
||
<th id="76E82A29C65B0052FB16FE57FB71FE20" box="[1152,1239,456,478]" gridcol="3" gridrow="0" pageId="7" pageNumber="492">239246</th>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FE6EFB71FDF9" box="[159,1239,497,519]" gridrow="1" pageId="7" pageNumber="492" rowspan-0="1">
|
||
<td id="76E82A29C65B0052FDF5FE6EFD62FDF9" box="[611,708,497,519]" gridcol="1" gridrow="1" pageId="7" pageNumber="492">holotype</td>
|
||
<td id="76E82A29C65B0052FC14FE6EFC7CFDF9" box="[898,986,497,519]" gridcol="2" gridrow="1" pageId="7" pageNumber="492">paratype</td>
|
||
<td id="76E82A29C65B0052FB16FE6EFB71FDF9" box="[1152,1239,497,519]" gridcol="3" gridrow="1" pageId="7" pageNumber="492">paratype</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FD85FB71FDCE" box="[159,1239,538,560]" gridrow="2" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FD85FE71FDCE" box="[159,471,538,560]" gridcol="0" gridrow="2" pageId="7" pageNumber="492">Sex</th>
|
||
<td id="76E82A29C65B0052FDF5FD85FD62FDCE" box="[611,708,538,560]" gridcol="1" gridrow="2" pageId="7" pageNumber="492">f</td>
|
||
<td id="76E82A29C65B0052FC14FD85FC7CFDCE" box="[898,986,538,560]" gridcol="2" gridrow="2" pageId="7" pageNumber="492">f</td>
|
||
<td id="76E82A29C65B0052FB16FD85FB71FDCE" box="[1152,1239,538,560]" gridcol="3" gridrow="2" pageId="7" pageNumber="492">m</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FDDBFB71FDA4" box="[159,1239,580,602]" gridrow="3" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FDDBFE71FDA4" box="[159,471,580,602]" gridcol="0" gridrow="3" pageId="7" pageNumber="492">SVL</th>
|
||
<td id="76E82A29C65B0052FDF5FDDBFD62FDA4" box="[611,708,580,602]" gridcol="1" gridrow="3" pageId="7" pageNumber="492">34.6</td>
|
||
<td id="76E82A29C65B0052FC14FDDBFC7CFDA4" box="[898,986,580,602]" gridcol="2" gridrow="3" pageId="7" pageNumber="492">30.6</td>
|
||
<td id="76E82A29C65B0052FB16FDDBFB71FDA4" box="[1152,1239,580,602]" gridcol="3" gridrow="3" pageId="7" pageNumber="492">37.3</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FDF2FB71FD7D" box="[159,1239,621,643]" gridrow="4" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FDF2FE71FD7D" box="[159,471,621,643]" gridcol="0" gridrow="4" pageId="7" pageNumber="492">dorsal scales</th>
|
||
<td id="76E82A29C65B0052FDF5FDF2FD62FD7D" box="[611,708,621,643]" gridcol="1" gridrow="4" pageId="7" pageNumber="492">25</td>
|
||
<td id="76E82A29C65B0052FC14FDF2FC7CFD7D" box="[898,986,621,643]" gridcol="2" gridrow="4" pageId="7" pageNumber="492">25</td>
|
||
<td id="76E82A29C65B0052FB16FDF2FB71FD7D" box="[1152,1239,621,643]" gridcol="3" gridrow="4" pageId="7" pageNumber="492">24</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FD09FB71FD52" box="[159,1239,662,684]" gridrow="5" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FD09FE71FD52" box="[159,471,662,684]" gridcol="0" gridrow="5" pageId="7" pageNumber="492">ventral scales</th>
|
||
<td id="76E82A29C65B0052FDF5FD09FD62FD52" box="[611,708,662,684]" gridcol="1" gridrow="5" pageId="7" pageNumber="492">14</td>
|
||
<td id="76E82A29C65B0052FC14FD09FC7CFD52" box="[898,986,662,684]" gridcol="2" gridrow="5" pageId="7" pageNumber="492">14</td>
|
||
<td id="76E82A29C65B0052FB16FD09FB71FD52" box="[1152,1239,662,684]" gridcol="3" gridrow="5" pageId="7" pageNumber="492">15</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FD5FFB71FD28" box="[159,1239,704,726]" gridrow="6" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FD5FFE71FD28" box="[159,471,704,726]" gridcol="0" gridrow="6" pageId="7" pageNumber="492">lamellar formula on hand</th>
|
||
<td id="76E82A29C65B0052FDF5FD5FFD62FD28" box="[611,708,704,726]" gridcol="1" gridrow="6" pageId="7" pageNumber="492">4-4-5-4</td>
|
||
<td id="76E82A29C65B0052FC14FD5FFC7CFD28" box="[898,986,704,726]" gridcol="2" gridrow="6" pageId="7" pageNumber="492">4-5-5-4</td>
|
||
<td id="76E82A29C65B0052FB16FD5FFB71FD28" box="[1152,1239,704,726]" gridcol="3" gridrow="6" pageId="7" pageNumber="492">4-4-5-4</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FD76FB71FD01" box="[159,1239,745,767]" gridrow="7" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FD76FE71FD01" box="[159,471,745,767]" gridcol="0" gridrow="7" pageId="7" pageNumber="492">subdigital lamellae on first toe</th>
|
||
<td id="76E82A29C65B0052FDF5FD76FD62FD01" box="[611,708,745,767]" gridcol="1" gridrow="7" pageId="7" pageNumber="492">5</td>
|
||
<td id="76E82A29C65B0052FC14FD76FC7CFD01" box="[898,986,745,767]" gridcol="2" gridrow="7" pageId="7" pageNumber="492">5</td>
|
||
<td id="76E82A29C65B0052FB16FD76FB71FD01" box="[1152,1239,745,767]" gridcol="3" gridrow="7" pageId="7" pageNumber="492">4</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FC8DFB71FCD6" box="[159,1239,786,808]" gridrow="8" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FC8DFE71FCD6" box="[159,471,786,808]" gridcol="0" gridrow="8" pageId="7" pageNumber="492">dark dorsal transverse blotches</th>
|
||
<td id="76E82A29C65B0052FDF5FC8DFD62FCD6" box="[611,708,786,808]" gridcol="1" gridrow="8" pageId="7" pageNumber="492">very faint</td>
|
||
<td id="76E82A29C65B0052FC14FC8DFC7CFCD6" box="[898,986,786,808]" gridcol="2" gridrow="8" pageId="7" pageNumber="492">faint</td>
|
||
<td id="76E82A29C65B0052FB16FC8DFB71FCD6" box="[1152,1239,786,808]" gridcol="3" gridrow="8" pageId="7" pageNumber="492">absent</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FCA3FB71FCAC" box="[159,1239,828,850]" gridrow="9" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FCA3FE71FCAC" box="[159,471,828,850]" gridcol="0" gridrow="9" pageId="7" pageNumber="492">Trunk/SVL</th>
|
||
<td id="76E82A29C65B0052FDF5FCA3FD62FCAC" box="[611,708,828,850]" gridcol="1" gridrow="9" pageId="7" pageNumber="492">0.47</td>
|
||
<td id="76E82A29C65B0052FC14FCA3FC7CFCAC" box="[898,986,828,850]" gridcol="2" gridrow="9" pageId="7" pageNumber="492">0.51</td>
|
||
<td id="76E82A29C65B0052FB16FCA3FB71FCAC" box="[1152,1239,828,850]" gridcol="3" gridrow="9" pageId="7" pageNumber="492">0.49</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FCFAFB71FC85" box="[159,1239,869,891]" gridrow="10" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FCFAFE71FC85" box="[159,471,869,891]" gridcol="0" gridrow="10" pageId="7" pageNumber="492">HeadL/SVL</th>
|
||
<td id="76E82A29C65B0052FDF5FCFAFD62FC85" box="[611,708,869,891]" gridcol="1" gridrow="10" pageId="7" pageNumber="492">0.25</td>
|
||
<td id="76E82A29C65B0052FC14FCFAFC7CFC85" box="[898,986,869,891]" gridcol="2" gridrow="10" pageId="7" pageNumber="492">0.26</td>
|
||
<td id="76E82A29C65B0052FB16FCFAFB71FC85" box="[1152,1239,869,891]" gridcol="3" gridrow="10" pageId="7" pageNumber="492">0.24</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FC11FB71FC5A" box="[159,1239,910,932]" gridrow="11" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FC11FE71FC5A" box="[159,471,910,932]" gridcol="0" gridrow="11" pageId="7" pageNumber="492">HeadW/SVL</th>
|
||
<td id="76E82A29C65B0052FDF5FC11FD62FC5A" box="[611,708,910,932]" gridcol="1" gridrow="11" pageId="7" pageNumber="492">0.16</td>
|
||
<td id="76E82A29C65B0052FC14FC11FC7CFC5A" box="[898,986,910,932]" gridcol="2" gridrow="11" pageId="7" pageNumber="492">0.16</td>
|
||
<td id="76E82A29C65B0052FB16FC11FB71FC5A" box="[1152,1239,910,932]" gridcol="3" gridrow="11" pageId="7" pageNumber="492">0.16</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FC27FB71FC30" box="[159,1239,952,974]" gridrow="12" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FC27FE71FC30" box="[159,471,952,974]" gridcol="0" gridrow="12" pageId="7" pageNumber="492">HeadW/HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FC27FD62FC30" box="[611,708,952,974]" gridcol="1" gridrow="12" pageId="7" pageNumber="492">0.62</td>
|
||
<td id="76E82A29C65B0052FC14FC27FC7CFC30" box="[898,986,952,974]" gridcol="2" gridrow="12" pageId="7" pageNumber="492">0.62</td>
|
||
<td id="76E82A29C65B0052FB16FC27FB71FC30" box="[1152,1239,952,974]" gridcol="3" gridrow="12" pageId="7" pageNumber="492">0.67</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FC7EFB71FC09" box="[159,1239,993,1015]" gridrow="13" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FC7EFE71FC09" box="[159,471,993,1015]" gridcol="0" gridrow="13" pageId="7" pageNumber="492">SnEye/HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FC7EFD62FC09" box="[611,708,993,1015]" gridcol="1" gridrow="13" pageId="7" pageNumber="492">0.41</td>
|
||
<td id="76E82A29C65B0052FC14FC7EFC7CFC09" box="[898,986,993,1015]" gridcol="2" gridrow="13" pageId="7" pageNumber="492">0.41</td>
|
||
<td id="76E82A29C65B0052FB16FC7EFB71FC09" box="[1152,1239,993,1015]" gridcol="3" gridrow="13" pageId="7" pageNumber="492">0.43</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FB95FB71FBDE" box="[159,1239,1034,1056]" gridrow="14" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FB95FE71FBDE" box="[159,471,1034,1056]" gridcol="0" gridrow="14" pageId="7" pageNumber="492">NarEye/HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FB95FD62FBDE" box="[611,708,1034,1056]" gridcol="1" gridrow="14" pageId="7" pageNumber="492">0.30</td>
|
||
<td id="76E82A29C65B0052FC14FB95FC7CFBDE" box="[898,986,1034,1056]" gridcol="2" gridrow="14" pageId="7" pageNumber="492">0.29</td>
|
||
<td id="76E82A29C65B0052FB16FB95FB71FBDE" box="[1152,1239,1034,1056]" gridcol="3" gridrow="14" pageId="7" pageNumber="492">0.3</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FBABFB71FBB4" box="[159,1239,1076,1098]" gridrow="15" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FBABFE71FBB4" box="[159,471,1076,1098]" gridcol="0" gridrow="15" pageId="7" pageNumber="492">EyeD/HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FBABFD62FBB4" box="[611,708,1076,1098]" gridcol="1" gridrow="15" pageId="7" pageNumber="492">0.24</td>
|
||
<td id="76E82A29C65B0052FC14FBABFC7CFBB4" box="[898,986,1076,1098]" gridcol="2" gridrow="15" pageId="7" pageNumber="492">0.24</td>
|
||
<td id="76E82A29C65B0052FB16FBABFB71FBB4" box="[1152,1239,1076,1098]" gridcol="3" gridrow="15" pageId="7" pageNumber="492">0.23</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FBC2FB71FB8D" box="[159,1239,1117,1139]" gridrow="16" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FBC2FE71FB8D" box="[159,471,1117,1139]" gridcol="0" gridrow="16" pageId="7" pageNumber="492">SnW/HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FBC2FD62FB8D" box="[611,708,1117,1139]" gridcol="1" gridrow="16" pageId="7" pageNumber="492">0.14</td>
|
||
<td id="76E82A29C65B0052FC14FBC2FC7CFB8D" box="[898,986,1117,1139]" gridcol="2" gridrow="16" pageId="7" pageNumber="492">0.15</td>
|
||
<td id="76E82A29C65B0052FB16FBC2FB71FB8D" box="[1152,1239,1117,1139]" gridcol="3" gridrow="16" pageId="7" pageNumber="492">0.13</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FB19FB71FB62" box="[159,1239,1158,1180]" gridrow="17" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FB19FE71FB62" box="[159,471,1158,1180]" gridcol="0" gridrow="17" pageId="7" pageNumber="492">EyeD/NarEye</th>
|
||
<td id="76E82A29C65B0052FDF5FB19FD62FB62" box="[611,708,1158,1180]" gridcol="1" gridrow="17" pageId="7" pageNumber="492">0.80</td>
|
||
<td id="76E82A29C65B0052FC14FB19FC7CFB62" box="[898,986,1158,1180]" gridcol="2" gridrow="17" pageId="7" pageNumber="492">0.82</td>
|
||
<td id="76E82A29C65B0052FB16FB19FB71FB62" box="[1152,1239,1158,1180]" gridcol="3" gridrow="17" pageId="7" pageNumber="492">0.77</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FB2FFB71FB38" box="[159,1239,1200,1222]" gridrow="18" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FB2FFE71FB38" box="[159,471,1200,1222]" gridcol="0" gridrow="18" pageId="7" pageNumber="492">SnW/HeadW</th>
|
||
<td id="76E82A29C65B0052FDF5FB2FFD62FB38" box="[611,708,1200,1222]" gridcol="1" gridrow="18" pageId="7" pageNumber="492">0.22</td>
|
||
<td id="76E82A29C65B0052FC14FB2FFC7CFB38" box="[898,986,1200,1222]" gridcol="2" gridrow="18" pageId="7" pageNumber="492">0.24</td>
|
||
<td id="76E82A29C65B0052FB16FB2FFB71FB38" box="[1152,1239,1200,1222]" gridcol="3" gridrow="18" pageId="7" pageNumber="492">0.2</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FB46FB71FB11" box="[159,1239,1241,1263]" gridrow="19" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FB46FE71FB11" box="[159,471,1241,1263]" gridcol="0" gridrow="19" pageId="7" pageNumber="492">SVL</th>
|
||
<td id="76E82A29C65B0052FDF5FB46FD62FB11" box="[611,708,1241,1263]" gridcol="1" gridrow="19" pageId="7" pageNumber="492">34.6</td>
|
||
<td id="76E82A29C65B0052FC14FB46FC7CFB11" box="[898,986,1241,1263]" gridcol="2" gridrow="19" pageId="7" pageNumber="492">30.6</td>
|
||
<td id="76E82A29C65B0052FB16FB46FB71FB11" box="[1152,1239,1241,1263]" gridcol="3" gridrow="19" pageId="7" pageNumber="492">37.3</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FA9DFB71FAE6" box="[159,1239,1282,1304]" gridrow="20" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FA9DFE71FAE6" box="[159,471,1282,1304]" gridcol="0" gridrow="20" pageId="7" pageNumber="492">trunk</th>
|
||
<td id="76E82A29C65B0052FDF5FA9DFD62FAE6" box="[611,708,1282,1304]" gridcol="1" gridrow="20" pageId="7" pageNumber="492">16.4</td>
|
||
<td id="76E82A29C65B0052FC14FA9DFC7CFAE6" box="[898,986,1282,1304]" gridcol="2" gridrow="20" pageId="7" pageNumber="492">15.7</td>
|
||
<td id="76E82A29C65B0052FB16FA9DFB71FAE6" box="[1152,1239,1282,1304]" gridcol="3" gridrow="20" pageId="7" pageNumber="492">18.3</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FAB3FB71FABC" box="[159,1239,1324,1346]" gridrow="21" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FAB3FE71FABC" box="[159,471,1324,1346]" gridcol="0" gridrow="21" pageId="7" pageNumber="492">HeadL</th>
|
||
<td id="76E82A29C65B0052FDF5FAB3FD62FABC" box="[611,708,1324,1346]" gridcol="1" gridrow="21" pageId="7" pageNumber="492">8.7</td>
|
||
<td id="76E82A29C65B0052FC14FAB3FC7CFABC" box="[898,986,1324,1346]" gridcol="2" gridrow="21" pageId="7" pageNumber="492">7.9</td>
|
||
<td id="76E82A29C65B0052FB16FAB3FB71FABC" box="[1152,1239,1324,1346]" gridcol="3" gridrow="21" pageId="7" pageNumber="492">8.9</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FACAFB71FA95" box="[159,1239,1365,1387]" gridrow="22" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FACAFE71FA95" box="[159,471,1365,1387]" gridcol="0" gridrow="22" pageId="7" pageNumber="492">HeadW</th>
|
||
<td id="76E82A29C65B0052FDF5FACAFD62FA95" box="[611,708,1365,1387]" gridcol="1" gridrow="22" pageId="7" pageNumber="492">5.4</td>
|
||
<td id="76E82A29C65B0052FC14FACAFC7CFA95" box="[898,986,1365,1387]" gridcol="2" gridrow="22" pageId="7" pageNumber="492">4.9</td>
|
||
<td id="76E82A29C65B0052FB16FACAFB71FA95" box="[1152,1239,1365,1387]" gridcol="3" gridrow="22" pageId="7" pageNumber="492">6</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FAE1FB71FA6A" box="[159,1239,1406,1428]" gridrow="23" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FAE1FE71FA6A" box="[159,471,1406,1428]" gridcol="0" gridrow="23" pageId="7" pageNumber="492">SnEye</th>
|
||
<td id="76E82A29C65B0052FDF5FAE1FD62FA6A" box="[611,708,1406,1428]" gridcol="1" gridrow="23" pageId="7" pageNumber="492">3.6</td>
|
||
<td id="76E82A29C65B0052FC14FAE1FC7CFA6A" box="[898,986,1406,1428]" gridcol="2" gridrow="23" pageId="7" pageNumber="492">3.2</td>
|
||
<td id="76E82A29C65B0052FB16FAE1FB71FA6A" box="[1152,1239,1406,1428]" gridcol="3" gridrow="23" pageId="7" pageNumber="492">3.8</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FA37FB71FA40" box="[159,1239,1448,1470]" gridrow="24" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FA37FE71FA40" box="[159,471,1448,1470]" gridcol="0" gridrow="24" pageId="7" pageNumber="492">NarEye</th>
|
||
<td id="76E82A29C65B0052FDF5FA37FD62FA40" box="[611,708,1448,1470]" gridcol="1" gridrow="24" pageId="7" pageNumber="492">2.6</td>
|
||
<td id="76E82A29C65B0052FC14FA37FC7CFA40" box="[898,986,1448,1470]" gridcol="2" gridrow="24" pageId="7" pageNumber="492">2.3</td>
|
||
<td id="76E82A29C65B0052FB16FA37FB71FA40" box="[1152,1239,1448,1470]" gridcol="3" gridrow="24" pageId="7" pageNumber="492">2.7</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FA4EFB71FA19" box="[159,1239,1489,1511]" gridrow="25" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FA4EFE71FA19" box="[159,471,1489,1511]" gridcol="0" gridrow="25" pageId="7" pageNumber="492">EyeD</th>
|
||
<td id="76E82A29C65B0052FDF5FA4EFD62FA19" box="[611,708,1489,1511]" gridcol="1" gridrow="25" pageId="7" pageNumber="492">2.1</td>
|
||
<td id="76E82A29C65B0052FC14FA4EFC7CFA19" box="[898,986,1489,1511]" gridcol="2" gridrow="25" pageId="7" pageNumber="492">1.9</td>
|
||
<td id="76E82A29C65B0052FB16FA4EFB71FA19" box="[1152,1239,1489,1511]" gridcol="3" gridrow="25" pageId="7" pageNumber="492">2.1</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09FA65FB71F9EE" box="[159,1239,1530,1552]" gridrow="26" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09FA65FE71F9EE" box="[159,471,1530,1552]" gridcol="0" gridrow="26" pageId="7" pageNumber="492">SnW</th>
|
||
<td id="76E82A29C65B0052FDF5FA65FD62F9EE" box="[611,708,1530,1552]" gridcol="1" gridrow="26" pageId="7" pageNumber="492">1.2</td>
|
||
<td id="76E82A29C65B0052FC14FA65FC7CF9EE" box="[898,986,1530,1552]" gridcol="2" gridrow="26" pageId="7" pageNumber="492">1.2</td>
|
||
<td id="76E82A29C65B0052FB16FA65FB71F9EE" box="[1152,1239,1530,1552]" gridcol="3" gridrow="26" pageId="7" pageNumber="492">1.2</td>
|
||
</tr>
|
||
<tr id="35394355C65B0052FF09F9BBFB71F9C4" box="[159,1239,1572,1594]" gridrow="27" pageId="7" pageNumber="492">
|
||
<th id="76E82A29C65B0052FF09F9BBFE71F9C4" box="[159,471,1572,1594]" gridcol="0" gridrow="27" pageId="7" pageNumber="492">TaiL</th>
|
||
<td id="76E82A29C65B0052FDF5F9BBFD62F9C4" box="[611,708,1572,1594]" gridcol="1" gridrow="27" pageId="7" pageNumber="492">30.0</td>
|
||
<td id="76E82A29C65B0052FC14F9BBFC7CF9C4" box="[898,986,1572,1594]" gridcol="2" gridrow="27" pageId="7" pageNumber="492">/</td>
|
||
<td id="76E82A29C65B0052FB16F9BBFB71F9C4" box="[1152,1239,1572,1594]" gridcol="3" gridrow="27" pageId="7" pageNumber="492">28.3</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB64117C65BFFAAFF51F899FD97F80D" blockId="7.[151,1437,1655,2035]" pageId="7" pageNumber="492">
|
||
<emphasis id="B97D9D05C65BFFAAFF51F899FE2FF8E0" bold="true" box="[199,393,1797,1822]" pageId="7" pageNumber="492">Natural history.</emphasis>
|
||
The three specimens of
|
||
<taxonomicName id="4C093A94C65BFFAAFD38F899FB95F8E0" box="[686,1075,1798,1822]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="492" phylum="Chordata" rank="species" species="engganoeneis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C65BFFAAFD38F899FB95F8E0" box="[686,1075,1798,1822]" italics="true" pageId="7" pageNumber="492">Hemiphyllodactylus engganoeneis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC65BFFAAFBA8F899FB3AF8E0" box="[1086,1180,1798,1822]" pageId="7" pageNumber="492" rank="species">
|
||
<emphasis id="B97D9D05C65BFFAAFBA8F899FB3AF8E0" bold="true" box="[1086,1180,1798,1822]" pageId="7" pageNumber="492">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were collected where lowland forest habitat interfaced with a sandy beach near the mouth of a small river along the Enggano coastline. The
|
||
<typeStatus id="54B2FFB5C65BFFAAFF5DF8D3FE8BF89B" box="[203,301,1868,1893]" pageId="7" pageNumber="492" type="holotype">holotype</typeStatus>
|
||
was collected approximately
|
||
<quantity id="4CF1ECF2C65BFFAAFD15F8D2FD16F89A" box="[643,688,1869,1893]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="492" unit="m" value="1.0">1 m</quantity>
|
||
above the ground on the leaf of a sapling, and the two
|
||
<typeStatus id="54B2FFB5C65BFFAAFAA7F8D2FA3AF89B" box="[1329,1436,1869,1893]" pageId="7" pageNumber="492" type="paratype">paratypes</typeStatus>
|
||
were collected about
|
||
<quantity id="4CF1ECF2C65BFFAAFE1AF8EFFE1CF876" box="[396,442,1904,1928]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="492" unit="m" value="1.0">1 m</quantity>
|
||
above the ground on and between
|
||
<taxonomicName id="4C093A94C65BFFAAFCC6F8EFFC66F876" box="[848,960,1904,1928]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="492" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97D9D05C65BFFAAFCC6F8EFFC66F876" box="[848,960,1904,1928]" italics="true" pageId="7" pageNumber="492">Pandanus</emphasis>
|
||
</taxonomicName>
|
||
leaves. The specimens were all collected between
|
||
<date id="FFB767D7C65BFFAAFF6AF80BFE03F852" box="[252,421,1939,1964]" pageId="7" pageNumber="492" value="2200" valueMax="2225">2200 and 2225</date>
|
||
hrs.
|
||
<taxonomicName id="4C093A94C65BFFAAFE48F80BFDE8F852" box="[478,590,1940,1964]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="7" pageNumber="492" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B97D9D05C65BFFAAFE48F80BFDE8F852" box="[478,590,1940,1964]" italics="true" pageId="7" pageNumber="492">Pandanus</emphasis>
|
||
</taxonomicName>
|
||
was abundant along the forest edge, and the crevices between its serrated, strap-shaped leaves were also inhabited by two other gecko species,
|
||
<taxonomicName id="4C093A94C65BFFAAFC36F828FA3AF82E" authority="Dumeril & Bibron" authorityName="Dumeril & Bibron" box="[928,1436,1975,2000]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="492" phylum="Chordata" rank="species" species="lugubris">
|
||
<emphasis id="B97D9D05C65BFFAAFC36F828FB16F831" box="[928,1200,1975,1999]" italics="true" pageId="7" pageNumber="492">Lepidodactylus lugubris</emphasis>
|
||
(Duméril & Bibron)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C093A94C65BFFAAFF50F844FD97F80D" authority="Schlegel." authorityName="Schlegel." box="[198,561,2010,2035]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="492" phylum="Chordata" rank="species" species="frenatus">
|
||
<emphasis id="B97D9D05C65BFFAAFF50F844FE64F80D" box="[198,450,2011,2035]" italics="true" pageId="7" pageNumber="492">Hemidactylus frenatus</emphasis>
|
||
Schlegel.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C313129CC654FFA4FF51FF07FD3CFB5E" lastPageId="9" lastPageNumber="494" pageId="8" pageNumber="493" type="etymology">
|
||
<paragraph id="8BB64117C654FFA5FF51FF07FE57FF2B" blockId="8.[151,1436,151,213]" pageId="8" pageNumber="493">
|
||
<emphasis id="B97D9D05C654FFA5FF51FF07FEE8FF4E" bold="true" box="[199,334,152,176]" pageId="8" pageNumber="493">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C093A94C654FFA5FDACFF07FD74FF51" box="[570,722,152,175]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="493" phylum="Chordata" rank="species" species="engganoensis">
|
||
<emphasis id="B97D9D05C654FFA5FDACFF07FD74FF51" box="[570,722,152,175]" italics="true" pageId="8" pageNumber="493">engganoensis</emphasis>
|
||
</taxonomicName>
|
||
is an adjective in reference to the
|
||
<typeStatus id="54B2FFB5C654FFA5FBC1FF07FB21FF4E" box="[1111,1159,152,176]" pageId="8" pageNumber="493">type</typeStatus>
|
||
locality
|
||
<collectingCountry id="F31E0187C654FFA5FB7CFF08FA8CFF4E" box="[1258,1322,151,176]" name="Iceland" pageId="8" pageNumber="493">Pulau</collectingCountry>
|
||
Enggano, Bengkulu Province,
|
||
<collectingCountry id="F31E0187C654FFA5FEEAFF23FE4BFF2B" box="[380,493,188,213]" name="Indonesia" pageId="8" pageNumber="493">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF76119FC654FFA5FF01F83BFA3BF826" httpUri="https://zenodo.org/record/228525/files/figure.png" pageId="8" pageNumber="493" targetBox="[179,1407,265,1934]" targetPageId="8">
|
||
<paragraph id="8BB64117C654FFA5FF01F83BFA3BF826" blockId="8.[151,1437,1956,2008]" pageId="8" pageNumber="493">
|
||
<emphasis id="B97D9D05C654FFA5FF01F83BFEBEF847" bold="true" box="[151,280,1956,1978]" pageId="8" pageNumber="493">FIGURE 4.</emphasis>
|
||
Type series of
|
||
<taxonomicName id="4C093A94C654FFA5FE5EF83BFC8EF844" box="[456,808,1956,1978]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="493" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C654FFA5FE5EF83BFC8EF844" box="[456,808,1956,1978]" italics="true" pageId="8" pageNumber="493">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC654FFA5FCA2F83AFC24F844" box="[820,898,1957,1978]" pageId="8" pageNumber="493" rank="species">sp. nov</taxonomicNameLabel>
|
||
from Pulau Enggano, Indonesia. Upper: holotype MZB.Lace 4568 (adult female). Middle: paratype MVZ 239346 (adult male). Lower: paratype MVZ 239345 (subadult female).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB64117C655FFA4FF51FF08FD3CFB5E" blockId="9.[151,1437,151,1184]" pageId="9" pageNumber="494">
|
||
<emphasis id="B97D9D05C655FFA4FF51FF08FECBFF4E" bold="true" box="[199,365,151,176]" pageId="9" pageNumber="494">Comparisons.</emphasis>
|
||
The taxonomy of Grismer
|
||
<emphasis id="B97D9D05C655FFA4FD0BFF06FD73FF4E" box="[669,725,152,176]" italics="true" pageId="9" pageNumber="494">et al.</emphasis>
|
||
(2013, 2014),
|
||
<bibRefCitation id="EF983CE6C655FFA4FCECFF07FB98FF4E" box="[890,1086,152,176]" pageId="9" pageNumber="494" refString="Ngo, V. T., Grismer, L. L., Thai, P. H. & Wood, P. L. (2014) A new species of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from Ba Na-Nui Nature Reserve, central Vietnam. Zootaxa, 3760 (4), 539 - 552. http: // dx. doi. org / 10.11646 / zootaxa. 3760.4.3" type="journal article">
|
||
Ngo
|
||
<emphasis id="B97D9D05C655FFA4FC27FF06FC4FFF4E" box="[945,1001,152,176]" italics="true" pageId="9" pageNumber="494">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF983CE6C655FFA4FBDFFF07FA92FF4E" box="[1097,1332,152,176]" pageId="9" pageNumber="494" refString="Nguyen T., Lehmann T., Le M. D., Duong H. T., Bonkowski, M. & Ziegler, T. (2013) A new species of Hemiphyllodactylus (Reptilia: Gekkonidae) from northern Vietnam. Zootaxa, 3736 (1), 89 - 98. http: // dx. doi. org / 10.11646 / zootaxa. 3736.1.5" type="journal article">
|
||
Nguyen
|
||
<emphasis id="B97D9D05C655FFA4FB3EFF06FB47FF4E" box="[1192,1249,152,176]" italics="true" pageId="9" pageNumber="494">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EF983CE6C655FFA4FAF9FF07FF47FF2A" author="Zug" pageId="9" pageNumber="494" refString="Zug, G. R. (2010) Speciation and dispersal in a low diversity taxon: the Slender geckos Hemiphyllodactylus (Reptilia, Gekkonidae). Smithsonian Contributions to Zoology, 631, 1 - 70." type="journal article" year="2010">Zug (2010)</bibRefCitation>
|
||
is used in the comparisons below for
|
||
<taxonomicName id="4C093A94C655FFA4FD2DFF22FC37FF2A" box="[699,913,189,212]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B97D9D05C655FFA4FD2DFF22FC37FF2A" box="[699,913,189,212]" italics="true" pageId="9" pageNumber="494">H. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
Zug,
|
||
<taxonomicName id="4C093A94C655FFA4FC49FF22FB00FF2B" authority="Bleeker" authorityName="Bleeker" box="[991,1190,188,213]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="typus">
|
||
<emphasis id="B97D9D05C655FFA4FC49FF22FBE4FF2A" box="[991,1090,189,212]" italics="true" pageId="9" pageNumber="494">
|
||
H.
|
||
<typeStatus id="54B2FFB5C655FFA4FB91FF22FBE4FF2A" box="[1031,1090,189,212]" pageId="9" pageNumber="494">typus</typeStatus>
|
||
</emphasis>
|
||
Bleeker
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C093A94C655FFA4FB7FFF22FEB8FF06" authority="Boulenger" authorityName="Boulenger" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunannensis">
|
||
<emphasis id="B97D9D05C655FFA4FB7FFF22FA3AFF2A" box="[1257,1436,189,212]" italics="true" pageId="9" pageNumber="494">H. yunannensis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C093A94C655FFA4FEBAFF7FFD0DFF09" box="[300,683,224,248]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FEBAFF7FFD0DFF09" box="[300,683,224,248]" italics="true" pageId="9" pageNumber="494">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FD27FF7FFCACFF06" box="[689,778,224,248]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FD27FF7FFCACFF06" bold="true" box="[689,778,224,248]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is one of the smallest species of the genus and differs from
|
||
<taxonomicName id="4C093A94C655FFA4FF01FE9AFDAAFEE3" authority="Ngo, Grismer, Thai" authorityName="Ngo, Grismer, Thai" box="[151,524,260,285]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">
|
||
<emphasis id="B97D9D05C655FFA4FF01FE9AFE8FFEE2" box="[151,297,260,284]" italics="true" pageId="9" pageNumber="494">H. banaensis</emphasis>
|
||
Ngo, Grismer,
|
||
<collectingCountry id="F31E0187C655FFA4FE40FE9BFDAAFEE3" box="[470,524,260,285]" name="Thailand" pageId="9" pageNumber="494">Thai</collectingCountry>
|
||
</taxonomicName>
|
||
, & Wood;
|
||
<taxonomicName id="4C093A94C655FFA4FD13FE9AFBACFEE3" authority="Grismer, Wood" authorityName="Grismer, Wood" box="[645,1034,260,285]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="chiangmaiensis">
|
||
<emphasis id="B97D9D05C655FFA4FD13FE9AFCF3FEE2" box="[645,853,260,284]" italics="true" pageId="9" pageNumber="494">H. chiangmaiensis</emphasis>
|
||
Grismer, Wood
|
||
</taxonomicName>
|
||
, & Cota;
|
||
<taxonomicName id="4C093A94C655FFA4FBE3FE9AFA3EFEE3" authority="Boulenger" authorityName="Boulenger" box="[1141,1432,260,285]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="larutensis">
|
||
<emphasis id="B97D9D05C655FFA4FBE3FE9AFAA3FEE2" box="[1141,1285,260,284]" italics="true" pageId="9" pageNumber="494">H. larutensis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FF01FEB6FE45FEBE" authority="Brongersma" authorityName="Brongersma" box="[151,483,296,320]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="margarethae">
|
||
<emphasis id="B97D9D05C655FFA4FF01FEB6FEEAFEBE" box="[151,332,296,320]" italics="true" pageId="9" pageNumber="494">H. margarethae</emphasis>
|
||
Brongersma
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FE64FEB6FD63FEC1" box="[498,709,296,320]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="titiwangsaensis">
|
||
<emphasis id="B97D9D05C655FFA4FE64FEB6FD63FEC1" box="[498,709,296,320]" italics="true" pageId="9" pageNumber="494">H. titiwangsaensis</emphasis>
|
||
</taxonomicName>
|
||
Zug;
|
||
<taxonomicName id="4C093A94C655FFA4FC86FEB6FC71FEBE" authority="Bleeker" authorityName="Bleeker" box="[784,983,295,320]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="typus">
|
||
<emphasis id="B97D9D05C655FFA4FC86FEB6FCD7FEBE" box="[784,881,297,320]" italics="true" pageId="9" pageNumber="494">
|
||
H.
|
||
<typeStatus id="54B2FFB5C655FFA4FCA0FEB6FCD7FEBE" box="[822,881,297,320]" pageId="9" pageNumber="494">typus</typeStatus>
|
||
</emphasis>
|
||
Bleeker
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FC70FEB6FB31FEC1" box="[998,1175,296,320]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunnanensis">
|
||
<emphasis id="B97D9D05C655FFA4FC70FEB6FB31FEC1" box="[998,1175,296,320]" italics="true" pageId="9" pageNumber="494">H. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C093A94C655FFA4FB4AFEB6FE53FE9B" authority="Nguyen, Lehmann, Le Duc, Bonkowski" authorityName="Nguyen, Lehmann, Le Duc, Bonkowski" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="zugi">
|
||
<emphasis id="B97D9D05C655FFA4FB4AFEB6FA94FEC1" box="[1244,1330,296,320]" italics="true" pageId="9" pageNumber="494">H. zugi</emphasis>
|
||
Nguyen, Lehmann, Le
|
||
<collectingCountry id="F31E0187C655FFA4FEA3FED2FEC1FE9A" box="[309,359,333,356]" name="Germany" pageId="9" pageNumber="494">Duc</collectingCountry>
|
||
, Bonkowski
|
||
</taxonomicName>
|
||
, & Ziegler by its maximum SVL of
|
||
<quantity id="4CF1ECF2C655FFA4FC04FED3FC51FE9A" box="[914,1015,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="3.7299999999999995" pageId="9" pageNumber="494" unit="mm" value="37.3">37.3 mm</quantity>
|
||
vs a SVL>41.0 mm. Its postmentals are not distinctly enlarged which separates it from
|
||
<emphasis id="B97D9D05C655FFA4FD66FEEEFBD1FE76" box="[752,1143,368,392]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FD66FEEEFC2FFE76" box="[752,905,368,392]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">H. banaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FC0FFEEEFBC9FE76" box="[921,1135,368,392]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="chiangmaiensis">H. chiangmaiensis</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<taxonomicName id="4C093A94C655FFA4FB14FEEEFAD4FE76" authority="(Werner)" authorityName="Werner" baseAuthorityName="Werner" box="[1154,1394,368,392]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="harterti">
|
||
<emphasis id="B97D9D05C655FFA4FB14FEEEFB58FE76" box="[1154,1278,368,392]" italics="true" pageId="9" pageNumber="494">H. harterti</emphasis>
|
||
(Werner)
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B97D9D05C655FFA4FA17FEEEFB24FE52" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FA17FEEEFEA1FE52" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="larutensis">H. larutensis</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FE8CFE0AFE74FE52" box="[282,466,404,428]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="margarethae">H. margarethae</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FE73FE0AFD15FE52" box="[485,691,405,428]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="titwangsaensis">H. titwangsaensis</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FD50FE0AFC8CFE52" box="[710,810,405,428]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="typus">
|
||
H.
|
||
<typeStatus id="54B2FFB5C655FFA4FD7BFE0AFC8CFE52" box="[749,810,405,428]" pageId="9" pageNumber="494">typus</typeStatus>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FCABFE0AFC57FE52" box="[829,1009,405,428]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunnanensis">H. yunnanensis</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C093A94C655FFA4FB92FE0AFB24FE52" authority="Grismer, Wood, Anuar, Muin, Quah, McGuire, Brown, Ngo" authorityName="Grismer, Wood, Anuar, Muin, Quah, McGuire, Brown, Ngo" box="[1028,1154,404,428]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="tehtarik">H. tehtarik</taxonomicName>
|
||
</emphasis>
|
||
Grismer, Wood, Anuar, Muin, Quah, McGuire, Brown, Ngo, &
|
||
<collectingCountry id="F31E0187C655FFA4FDE7FE28FD0EFE2E" box="[625,680,439,464]" name="Thailand" pageId="9" pageNumber="494">Thai</collectingCountry>
|
||
; and
|
||
<emphasis id="B97D9D05C655FFA4FD7AFE26FB7FFE31" box="[748,1241,440,464]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FD7AFE26FCE3FE31" box="[748,837,440,464]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="zugi">H. zugi</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C093A94C655FFA4FCC3FE27FB7FFE31" box="[853,1241,440,464]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">Hemiphyllodactylus engganoensis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FB73FE27FAE5FE2E" box="[1253,1347,440,464]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FB73FE27FAE5FE2E" bold="true" box="[1253,1347,440,464]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be differentiated from
|
||
<emphasis id="B97D9D05C655FFA4FEEEFE42FC30FE0A" box="[376,918,476,500]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FEEEFE42FDB7FE0A" box="[376,529,476,500]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">H. banaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FD89FE42FD10FE0A" box="[543,694,476,500]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="larutensis">H. larutensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FD52FE42FC34FE0A" box="[708,914,477,500]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="titwangsaensis">H. titwangsaensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C093A94C655FFA4FC45FE42FB80FE0A" box="[979,1062,477,500]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="zugi">
|
||
<emphasis id="B97D9D05C655FFA4FC45FE42FB80FE0A" box="[979,1062,477,500]" italics="true" pageId="9" pageNumber="494">H. zugi</emphasis>
|
||
</taxonomicName>
|
||
by having five vs. two or three circumnasal scales. It differs from all other species except
|
||
<taxonomicName id="4C093A94C655FFA4FCB9FD9EFC26FDE9" box="[815,896,512,536]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="zugi">
|
||
<emphasis id="B97D9D05C655FFA4FCB9FD9EFC26FDE9" box="[815,896,512,536]" italics="true" pageId="9" pageNumber="494">H. zugi</emphasis>
|
||
</taxonomicName>
|
||
in having 24 or 25 dorsal scales (as opposed to having fewer than 22) and having 14 ventral scales separates it from
|
||
<emphasis id="B97D9D05C655FFA4FC34FDBAFB9AFDC2" box="[930,1084,548,572]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FC34FDBAFB9EFDC2" box="[930,1080,548,572]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">H. banaensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C093A94C655FFA4FBD2FDBAFAD3FDC3" authority="(Beddome)" authorityName="Beddome" baseAuthorityName="Beddome" box="[1092,1397,548,573]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="aurantiacus">
|
||
<emphasis id="B97D9D05C655FFA4FBD2FDBAFB4BFDC2" box="[1092,1261,549,572]" italics="true" pageId="9" pageNumber="494">H. aurantiacus</emphasis>
|
||
(Beddome)
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B97D9D05C655FFA4FA17FDBAFE50FD9E" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FA17FDBAFEEEFD9E" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="chiangmaiensis">H. chiangmaiensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FEC0FDD6FE50FD9E" box="[342,502,584,608]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="ganoklonis">H. ganoklonis</taxonomicName>
|
||
</emphasis>
|
||
Zug,
|
||
<emphasis id="B97D9D05C655FFA4FDA8FDD6FB33FDA1" box="[574,1173,584,608]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FDA8FDD6FD53FD9E" box="[574,757,584,608]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="margarethae">H. margarethae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FC95FDD6FC76FDA1" box="[771,976,584,608]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="titwangsaensis">H. titwangsaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FC48FDD6FB36FDA1" box="[990,1168,584,608]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunnanensis">H. yunnanensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C093A94C655FFA4FB47FDD6FAEBFD9E" box="[1233,1357,584,608]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="tehtarik">
|
||
<emphasis id="B97D9D05C655FFA4FB47FDD6FAEBFD9E" box="[1233,1357,584,608]" italics="true" pageId="9" pageNumber="494">H. tehtarik</emphasis>
|
||
</taxonomicName>
|
||
which have 12 or less.
|
||
<taxonomicName id="4C093A94C655FFA4FED9FDF3FD76FD7A" box="[335,720,620,644]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FED9FDF3FD76FD7A" box="[335,720,620,644]" italics="true" pageId="9" pageNumber="494">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FD41FDF3FC94FD7A" box="[727,818,620,644]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FD41FDF3FC94FD7A" bold="true" box="[727,818,620,644]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has a unique lamellar formula on the hand (4-5-5-4 or 4-4-5-4) that separate if from all other species except
|
||
<taxonomicName id="4C093A94C655FFA4FD78FD0EFC26FD56" box="[750,896,656,680]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">
|
||
<emphasis id="B97D9D05C655FFA4FD78FD0EFC26FD56" box="[750,896,656,680]" italics="true" pageId="9" pageNumber="494">H. banaensis</emphasis>
|
||
</taxonomicName>
|
||
that have various other combinations (see Table 1). Having a series of femoral and precloacal pores that are not contiguous separates
|
||
<taxonomicName id="4C093A94C655FFA4FBD3FD2AFB59FD32" box="[1093,1279,693,716]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FBD3FD2AFB59FD32" box="[1093,1279,693,716]" italics="true" pageId="9" pageNumber="494">H. engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FA90FD2BFAF9FD32" box="[1286,1375,692,716]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FA90FD2BFAF9FD32" bold="true" box="[1286,1375,692,716]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from
|
||
<emphasis id="B97D9D05C655FFA4FF01FD46FD26FD0E" box="[151,640,728,752]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FF01FD46FEE4FD11" box="[151,322,728,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="aurantiacus">H. aurantiacus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FED8FD46FE49FD0E" box="[334,495,728,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="ganoklonis">H. ganoklonis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FE6DFD46FD26FD0E" authority="Taylor" authorityName="Taylor" box="[507,640,728,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="insularis">H. insularis</taxonomicName>
|
||
</emphasis>
|
||
Taylor,
|
||
<emphasis id="B97D9D05C655FFA4FD48FD46FC31FD0E" box="[734,919,728,752]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FD48FD46FC34FD0E" box="[734,914,728,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="margarethae">H. margarethae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
some
|
||
<emphasis id="B97D9D05C655FFA4FC77FD46FBE0FD0E" box="[993,1094,729,752]" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FC77FD46FBE5FD0E" box="[993,1091,729,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="typus">
|
||
H.
|
||
<typeStatus id="54B2FFB5C655FFA4FB93FD46FBE5FD0E" box="[1029,1091,729,752]" pageId="9" pageNumber="494">typus</typeStatus>
|
||
</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C093A94C655FFA4FBE8FD46FA8AFD11" box="[1150,1324,728,752]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunnanensis">
|
||
<emphasis id="B97D9D05C655FFA4FBE8FD46FA8AFD11" box="[1150,1324,728,752]" italics="true" pageId="9" pageNumber="494">H. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
which all have a contiguous pore series. Additionally, having a total pore count of 42 further differentiates
|
||
<taxonomicName id="4C093A94C655FFA4FB76FD62FA3AFCEA" box="[1248,1436,765,788]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FB76FD62FA3AFCEA" box="[1248,1436,765,788]" italics="true" pageId="9" pageNumber="494">H. engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FF01FCBFFF57FCC6" box="[151,241,800,824]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FF01FCBFFF57FCC6" bold="true" box="[151,241,800,824]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from all other species (except
|
||
<taxonomicName id="4C093A94C655FFA4FDD8FCBEFD60FCC6" box="[590,710,800,824]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="harterti">
|
||
<emphasis id="B97D9D05C655FFA4FDD8FCBEFD60FCC6" box="[590,710,800,824]" italics="true" pageId="9" pageNumber="494">H. harterti</emphasis>
|
||
</taxonomicName>
|
||
) who have less than 40 (0–39 collectively).
|
||
<taxonomicName id="4C093A94C655FFA4FB2AFCBFFE89FCA2" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FB2AFCBFFE89FCA2" italics="true" pageId="9" pageNumber="494">Hemiphyllodactylus engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FEA0FCDBFE36FCA2" box="[310,400,836,860]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FEA0FCDBFE36FCA2" bold="true" box="[310,400,836,860]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has a high number (5) of cloacal spurs which at least distinguishes it from
|
||
<emphasis id="B97D9D05C655FFA4FB4FFCDAFB00FC81" italics="true" pageId="9" pageNumber="494">
|
||
<taxonomicName id="4C093A94C655FFA4FB4FFCDAFAD0FCA2" box="[1241,1398,837,860]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="auratiacus">H. auratiacus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FA17FCDAFEACFC7E" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="banaensis">H. banaensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FE88FCF6FE07FC7E" box="[286,417,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="harterti">H. harterti</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FE20FCF6FDE0FC7E" box="[438,582,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="insularis">H. insularis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FDCCFCF6FD50FC7E" box="[602,758,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="larutensis">H. larutensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FC9CFCF6FC61FC7E" box="[778,967,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="margarethae">H. margarethae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C093A94C655FFA4FC4DFCF6FB07FC81" box="[987,1185,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="yunnanaensis">H. yunnanaensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C093A94C655FFA4FB78FCF6FAE1FC81" box="[1262,1351,872,896]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="zugi">
|
||
<emphasis id="B97D9D05C655FFA4FB78FCF6FAE1FC81" box="[1262,1351,872,896]" italics="true" pageId="9" pageNumber="494">H. zugi</emphasis>
|
||
</taxonomicName>
|
||
which collectively have 0–3 cloacal spurs. There are a number of morphometric ratios concerning various structures that are potentially diagnostic (Table 1). However, due to the incomplete sample sizes across all age classes for all species, we consider their diagnostic utility as tentative at this point. Additionally,
|
||
<taxonomicName id="4C093A94C655FFA4FBCCFC4AFAB1FC12" box="[1114,1303,981,1004]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FBCCFC4AFAB1FC12" box="[1114,1303,981,1004]" italics="true" pageId="9" pageNumber="494">H. engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FAB4FC4BFAD9FC12" box="[1314,1407,980,1004]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FAB4FC4BFAD9FC12" bold="true" box="[1314,1407,980,1004]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is possibly separated from various other species on the basis of a number color pattern characters (Table1). However, because the color variation in the
|
||
<typeStatus id="54B2FFB5C655FFA4FD8CFB82FDECFBCB" box="[538,586,1053,1077]" pageId="9" pageNumber="494">type</typeStatus>
|
||
series (three specimens) is extensive (
|
||
<figureCitation id="13325D92C655FFA4FB97FB83FBE0FBCA" box="[1025,1094,1052,1077]" captionStart="FIGURE 3" captionStartId="6.[151,250,1051,1073]" captionTargetBox="[313,1273,395,1029]" captionTargetId="figure@6.[313,1273,395,1029]" captionTargetPageId="6" captionText="FIGURE 3. Paratype MVZ 239345 (subadult female) of Hemiphyllodactylus engganoensis sp. nov. from Pulau Enggano, Indonesia." httpUri="https://zenodo.org/record/228524/files/figure.png" pageId="9" pageNumber="494">Fig. 3</figureCitation>
|
||
), we defer judgment as to the utility of these characters as well until the acquisition of additional material. Based on the mitochondrial ND2 gene, Grismer
|
||
<emphasis id="B97D9D05C655FFA4FF6CFBFAFE92FB82" box="[250,308,1124,1148]" italics="true" pageId="9" pageNumber="494">et al.</emphasis>
|
||
(2013) noted that
|
||
<taxonomicName id="4C093A94C655FFA4FD93FBFAFD67FB82" box="[517,705,1125,1148]" class="Reptilia" family="Gekkonidae" genus="Hemiphyllodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="494" phylum="Chordata" rank="species" species="engganoensis" status="sp. nov.">
|
||
<emphasis id="B97D9D05C655FFA4FD93FBFAFD67FB82" box="[517,705,1125,1148]" italics="true" pageId="9" pageNumber="494">H. engganoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24E207EC655FFA4FD5EFBFBFC85FB82" box="[712,803,1124,1148]" pageId="9" pageNumber="494" rank="species">
|
||
<emphasis id="B97D9D05C655FFA4FD5EFBFBFC85FB82" bold="true" box="[712,803,1124,1148]" pageId="9" pageNumber="494">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has an uncorrected sequence divergence of 17.5% from its closest relative from
|
||
<collectingCountry id="F31E0187C655FFA4FE32FB18FE42FB5E" box="[420,484,1159,1184]" name="Iceland" pageId="9" pageNumber="494">Pulau</collectingCountry>
|
||
Sibu,
|
||
<collectingCountry id="F31E0187C655FFA4FDBAFB18FD30FB5E" box="[556,662,1159,1184]" name="Malaysia" pageId="9" pageNumber="494">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |