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<document id="27E1D491723EB665B37D2E3ECA2A6E29" ID-DOI="10.5281/zenodo.5375016" ID-ISSN="1638-9395" ID-Zenodo-Dep="5375016" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630277386616" checkinUser="marcus" docAuthor="Modzalevskaya, Tatiana L." docDate="2003" docId="03A0162AE95EFFB3D95EFB577F6E1894" docLanguage="en" docName="Geodiversitas.2003.25.1.73-107.pdf" docOrigin="Geodiversitas 25 (1)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Grebenella laciniatus Modzalevskaya 2003, n. sp." docType="treatment" docVersion="3" lastPageNumber="99" masterDocId="FF996E52E947FFA9DA1EFFC77B251E11" masterDocTitle="Silurian and Devonian brachiopods from Severnaya Zemlya (Russian Arctic)" masterLastPageNumber="107" masterPageNumber="73" pageNumber="98" updateTime="1699210675571" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="0876FDFEE2BBEB2C62CCCA07F4A09C1B">Silurian and Devonian brachiopods from Severnaya Zemlya (Russian Arctic)</mods:title>
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<mods:namePart id="E7C02C89C3F3C1D42038913C8A67A4B3">Modzalevskaya, Tatiana L.</mods:namePart>
<mods:affiliation id="4E389EF2D1C83811783E8B926C10451A">All-Russia Geological Research Institute (VSEGEI), Srednij pr. 74, St. Petersburg, 199106 (Russia) modz @ IB 2567. spb. edu.</mods:affiliation>
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<mods:title id="929B0481CB9C3809CCB7A2F24851297C">Geodiversitas</mods:title>
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<mods:date id="79CD1154293E06E47E1F7FAC444B51A8">2003</mods:date>
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<mods:identifier id="697756A1DF64ADCC3D97A671BFBDCC12" type="DOI">10.5281/zenodo.5375016</mods:identifier>
<mods:identifier id="E22AA014C8FDD02605C7F5FDD246B4A0" type="ISSN">1638-9395</mods:identifier>
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<subSubSection id="C313F4B7E95EFFB0D95EFB577F3F1ADB" pageId="25" pageNumber="98" type="nomenclature">
<paragraph id="8BB6A73CE95EFFB0D95EFB577F3F1ADB" blockId="25.[832,1129,1168,1226]" pageId="25" pageNumber="98">
<heading id="D0FE1050E95EFFB0D95EFB577F3F1ADB" centered="true" fontSize="11" level="2" pageId="25" pageNumber="98" reason="2">
<taxonomicName id="4C09DCBFE95EFFB0D95EFB577F021ABB" authority="Modzalevskaya, 2003" authorityName="Modzalevskaya" authorityYear="2003" box="[832,1063,1168,1194]" class="Rhynchonellata" family="Reticulariidae" genus="Grebenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="25" pageNumber="98" phylum="Brachiopoda" rank="species" species="laciniatus" status="sp. nov.">
<emphasis id="B97D7B2EE95EFFB0D95EFB577F021ABB" bold="true" box="[832,1063,1168,1194]" italics="true" pageId="25" pageNumber="98">Grebenella laciniatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC655E95EFFB0DE30FB577F4C1ABB" box="[1070,1129,1168,1194]" pageId="25" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="1332BBB9E95EFFB0D988FB7778CC1ADB" box="[918,1001,1200,1226]" captionStart="FIG" captionStartId="25.[151,162,899,916]" captionTargetBox="[264,1117,168,861]" captionTargetId="figure-394@25.[289,1083,169,861]" captionTargetPageId="25" captionText="FIG. 28. — Grebenella laciniatus n. sp.; A, ventral view; B, dorsal view; C, lateral view; D, posterior view of a complete shell (N 79/12991), loc. 21581, Severnaya Zemlya or Podmnaya formations; E, ventral view; F, dorsal view; G, lateral view; H, anterior view of a complete shell (holotype N 78/12991); I, ventral interior with dental lamellae and muscle field (N 80/12991), Severnaya Zemlya or Podemnaya formations; J, enlarged external surface of a ventral valve, showing concentric growth lines with the impressions of the spines (N 81/12991), loc. 20399, Rusanov or Albanov formations. Scale bars: A-D, 5 mm; E-H, 3 mm; I, J, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5375071" httpUri="https://zenodo.org/record/5375071/files/figure.png" pageId="25" pageNumber="98">Figs 28</figureCitation>
;
<figureCitation id="1332BBB9E95EFFB0D9EBFB777F371ADB" box="[1013,1042,1200,1226]" captionStart="FIG" captionStartId="26.[151,162,803,820]" captionTargetBox="[335,1073,173,762]" captionTargetId="figure-377@26.[333,1089,169,782]" captionTargetPageId="26" captionText="FIG. 29. — Selected transverse serial sections, illustrating in the development of internal structures in Grebenella laciniatus n. sp. (N 82/12991). Measurements: length 6.8 mm, width?, thickness?; loc. 20399, Rusanov or Albanov formations. Numerical designations are the same as on Fig. 11. Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5375073" httpUri="https://zenodo.org/record/5375073/files/figure.png" pageId="25" pageNumber="98">29</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95EFFB0D8D4FB297F151B29" pageId="25" pageNumber="98" type="materials_examined">
<paragraph id="8BB6A73CE95EFFB0D8D4FB297F151B29" blockId="25.[714,1247,1262,1338]" pageId="25" pageNumber="98">
<materialsCitation id="3B61AD61E95EFFB0D8D4FB297F151B29" collectionCode="CNIGR, T" location="Museum" pageId="25" pageNumber="98" specimenCount="1" stateProvince="Saint Petersburg" typeStatus="holotype">
<typeStatus id="54B2199EE95EFFB0D8D4FB29781D1B15" box="[714,824,1262,1285]" pageId="25" pageNumber="98" type="holotype">HOLOTYPE</typeStatus>
. — N 78/12991,
<collectionCode id="ED183FF9E95EFFB0D9E2FB287F751B16" box="[1020,1104,1263,1287]" pageId="25" pageNumber="98">CNIGR</collectionCode>
<location id="8ED6F1E7E95EFFB0DE47FB297F931B14" LSID="urn:lsid:plazi:treatment:03A0162AE95EFFB3D95EFB577F6E1894:8ED6F1E7E95EFFB0DE47FB297F931B14" box="[1113,1206,1262,1285]" name="Museum" pageId="25" pageNumber="98" stateProvince="Saint Petersburg">Museum</location>
,
<collectingRegion id="49CD69DEE95EFFB0DEDDFB2878651B30" country="Russia" name="Saint Petersburg" pageId="25" pageNumber="98">St. Petersburg</collectingRegion>
(
<figureCitation id="1332BBB9E95EFFB0D94CFACE78831B30" box="[850,934,1289,1313]" captionStart="FIG" captionStartId="25.[151,162,899,916]" captionTargetBox="[264,1117,168,861]" captionTargetId="figure-394@25.[289,1083,169,861]" captionTargetPageId="25" captionText="FIG. 28. — Grebenella laciniatus n. sp.; A, ventral view; B, dorsal view; C, lateral view; D, posterior view of a complete shell (N 79/12991), loc. 21581, Severnaya Zemlya or Podmnaya formations; E, ventral view; F, dorsal view; G, lateral view; H, anterior view of a complete shell (holotype N 78/12991); I, ventral interior with dental lamellae and muscle field (N 80/12991), Severnaya Zemlya or Podemnaya formations; J, enlarged external surface of a ventral valve, showing concentric growth lines with the impressions of the spines (N 81/12991), loc. 20399, Rusanov or Albanov formations. Scale bars: A-D, 5 mm; E-H, 3 mm; I, J, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5375071" httpUri="https://zenodo.org/record/5375071/files/figure.png" pageId="25" pageNumber="98">Fig. 28</figureCitation>
E-H), conjoined valves; L = 6.0 mm, W = 8.0 mm,
<collectionCode id="ED183FF9E95EFFB0D9AFFAE678E71B29" box="[945,962,1313,1336]" name="Tavera, Department of Geology and Geophysics" pageId="25" pageNumber="98">T</collectionCode>
=
<quantity id="4CF10AD9E95EFFB0D9C5FAE57F091B29" box="[987,1068,1313,1338]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="25" pageNumber="98" unit="mm" value="4.1">4.1 mm</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95EFFB0D8D4FA887F991B79" box="[714,1212,1359,1384]" pageId="25" pageNumber="98" type="etymology">
<paragraph id="8BB6A73CE95EFFB0D8D4FA887F991B79" blockId="25.[714,1212,1359,1384]" box="[714,1212,1359,1384]" pageId="25" pageNumber="98">
ETYMOLOGY. — From
<emphasis id="B97D7B2EE95EFFB0D9B2FA977F201B79" box="[940,1029,1360,1384]" italics="true" pageId="25" pageNumber="98">laciniosus</emphasis>
(Latin): shredded.
</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95EFFB0D8D4FABA7F971BCC" pageId="25" pageNumber="98" type="materials_examined">
<paragraph id="8BB6A73CE95EFFB0D8D4FABA7F9B1BBE" blockId="25.[714,1246,1405,1455]" pageId="25" pageNumber="98">
<collectionCode id="ED183FF9E95EFFB0D8D4FABA78781B82" box="[714,861,1405,1428]" pageId="25" pageNumber="98">TYPE LOCALITY</collectionCode>
. — Loc. 20399, source of the left tributary of the Snegovaya River, Komsomolets Island.
</paragraph>
<paragraph id="8BB6A73CE95EFFB0D8D4FA027F971BCC" blockId="25.[714,1202,1477,1501]" box="[714,1202,1477,1501]" pageId="25" pageNumber="98">FORMATION. — Rusanov or Albanov formations.</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95EFFB0D8D4FA3478EE181A" box="[714,971,1523,1547]" pageId="25" pageNumber="98" type="description">
<paragraph id="8BB6A73CE95EFFB0D8D4FA3478EE181A" blockId="25.[714,971,1523,1547]" box="[714,971,1523,1547]" pageId="25" pageNumber="98">AGE. — Lower Devonian.</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95EFFB3D8D4F9E679061DCF" lastPageId="26" lastPageNumber="99" pageId="25" pageNumber="98" type="materials_examined">
<paragraph id="8BB6A73CE95EFFB3D8D4F9E679061DCF" blockId="25.[714,1248,1569,1670]" lastBlockId="26.[151,685,940,1176]" lastPageId="26" lastPageNumber="99" pageId="25" pageNumber="98">
MATERIAL EXAMINED. — Shelly limestone, two conjoined and 10 separated valves from loc. 20399 (source of the left tributary of the Snegovaya River) and loc. 21581 (southern bank of Krasnaya Bay), Komsomolets Island, Severnaya Zemlya (
<bibRefCitation id="EF98DACDE95DFFB3DBF8FC6A798C1DD4" author="MANNIK P. &amp; MENNER V. V. &amp; MATUKHIN R. G. &amp; KURSS V." box="[486,681,940,965]" pageId="26" pageNumber="99" pagination="99 - 122" refId="ref17313" refString="MANNIK P., MENNER V. V., MATUKHIN R. G. &amp; KURSS V. 2002. - Silurian and Devonian strata on the Severnaya Zemlya and Sedov archipelagos (Russia). Geodiversitas 24 (1): 99 - 122." type="journal article" year="2002">
Männik
<emphasis id="B97D7B2EE95DFFB3D823FC6B79401DD4" box="[573,613,940,965]" italics="true" pageId="26" pageNumber="99">et al</emphasis>
. 2002
</bibRefCitation>
: fig. 1); Rusanov or Albanov formations.
</paragraph>
</subSubSection>
<caption id="DF76F7B4E95DFFB3DA89FCE4796E1D4E" ID-DOI="http://doi.org/10.5281/zenodo.5375073" ID-Zenodo-Dep="5375073" httpUri="https://zenodo.org/record/5375073/files/figure.png" pageId="26" pageNumber="99" startId="26.[151,162,803,820]" targetBox="[335,1073,173,762]" targetPageId="26">
<paragraph id="8BB6A73CE95DFFB3DA89FCE4796E1D4E" blockId="26.[151,1249,803,863]" pageId="26" pageNumber="99">
FIG. 29. — Selected transverse serial sections, illustrating in the development of internal structures in
<taxonomicName id="4C09DCBFE95DFFB3D9E4FCE47F8C1D25" authority="Modzalevskaya, 2003" authorityName="Modzalevskaya" authorityYear="2003" box="[1018,1193,803,820]" class="Rhynchonellata" family="Reticulariidae" genus="Grebenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="26" pageNumber="99" phylum="Brachiopoda" rank="species" species="laciniatus" status="sp. nov.">
<emphasis id="B97D7B2EE95DFFB3D9E4FCE47F8C1D25" box="[1018,1193,803,820]" italics="true" pageId="26" pageNumber="99">Grebenella laciniatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24EC655E95DFFB3DEB1FCE47FC41D25" box="[1199,1249,803,820]" pageId="26" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
(N 82/12991). Measurements: length 6.8 mm, width?, thickness?; loc. 20399, Rusanov or Albanov formations. Numerical designations are the same as on Fig. 11. Scale bar: 1.6 mm.
</paragraph>
</caption>
<subSubSection id="C313F4B7E95DFFB3DA89FC287A351A33" pageId="26" pageNumber="99" type="description">
<paragraph id="8BB6A73CE95DFFB3DA89FC287A351A33" blockId="26.[151,685,940,1176]" pageId="26" pageNumber="99">MEASUREMENTS (IN MM). — L = 4.8-8.5, W = 6.6-8.0, T = 3.6-4.3.</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95DFFB3DA89FBF47A241A89" pageId="26" pageNumber="99" type="diagnosis">
<paragraph id="8BB6A73CE95DFFB3DA89FBF47A241A89" blockId="26.[151,685,940,1176]" pageId="26" pageNumber="99">DIAGNOSIS. — Small shells, transversally oval in outline and ventribiconvex in lateral profile; dorsal fold with deep narrow furrows on the flanks; plicae weakly developed.</paragraph>
</subSubSection>
<subSubSection id="C313F4B7E95DFFB3DA89FB0D7F6E1894" pageId="26" pageNumber="99" type="description">
<paragraph id="8BB6A73CE95DFFB3DA89FB0D7A0D1AF3" blockId="26.[151,685,1226,1669]" box="[151,296,1226,1252]" pageId="26" pageNumber="99">DESCRIPTION</paragraph>
<paragraph id="8BB6A73CE95DFFB3DA89FB2C785D1B34" blockId="26.[151,685,1226,1669]" lastBlockId="26.[714,1248,938,1317]" pageId="26" pageNumber="99">
Shells small, transversally oval, ventribiconvex with developed sinus and fold, and with smooth flanks. Ventral valve is the most convex in its umbonal part. Apex acute, hanging over the dorsal valve, delthyrium partly closed by deltidial plates, interarea triangular, low. Cardinal extremities rounded, sinus is well limited, arises from the beak, widening anteriorly, anterior tongue pronounced. Dorsal valve with sharply outlined fold bordered by deep narrow furrows. Fold originates at the beak. Broad plicae on flanks arise at about one third of valve length. Concentric growth lamellae are rhythmical, and bear the impressions of spines on their anterior edges in the mould. Ventral umbonal chambers filled by callus. Dental plates high, long, strongly thickened, widely situated and divergent; teeth massive. Ventral muscle field triangular with a triangular furrow near its anterior margin. Callus also well developed in dorsal valve, where thickened crural plates are widely divergent posteriorly and rest on short septum at the valve floor. Crura extend to near the valve floor. Dental sockets well limited medially by inner socket ridges (
<figureCitation id="1332BBB9E95DFFB3D907FACC78491B34" box="[793,876,1291,1317]" captionStart="FIG" captionStartId="26.[151,162,803,820]" captionTargetBox="[335,1073,173,762]" captionTargetId="figure-377@26.[333,1089,169,782]" captionTargetPageId="26" captionText="FIG. 29. — Selected transverse serial sections, illustrating in the development of internal structures in Grebenella laciniatus n. sp. (N 82/12991). Measurements: length 6.8 mm, width?, thickness?; loc. 20399, Rusanov or Albanov formations. Numerical designations are the same as on Fig. 11. Scale bar: 1.6 mm." figureDoi="http://doi.org/10.5281/zenodo.5375073" httpUri="https://zenodo.org/record/5375073/files/figure.png" pageId="26" pageNumber="99">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8BB6A73CE95DFFB3D8D4FA8C787F1B73" blockId="26.[714,1247,1355,1669]" box="[714,858,1355,1381]" pageId="26" pageNumber="99">COMPARISON</paragraph>
<paragraph id="8BB6A73CE95DFFB3D8D4FAAC7F6E1894" blockId="26.[714,1247,1355,1669]" pageId="26" pageNumber="99">
The flattened dorsal valve, transverse outline, and weakly developed plications suggest the comparison with
<taxonomicName id="4C09DCBFE95DFFB3D922FA6C78561BF4" authority="T. Modzalevskaya, 1994" authorityName="T. Modzalevskaya" authorityYear="1994" class="Rhynchonellata" family="Reticulariidae" genus="Grebenella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spiriferida" pageId="26" pageNumber="99" phylum="Brachiopoda" rank="species" species="impalpabilis">
<emphasis id="B97D7B2EE95DFFB3D922FA6C7F711BD4" box="[828,1108,1451,1477]" italics="true" pageId="26" pageNumber="99">Grebenella impalpabilis</emphasis>
T.
<bibRefCitation id="EF98DACDE95DFFB3DE9DFA6C78561BF4" author="MODZALEVSKAYA T. L. &amp; CHERKESOVA S. V." pageId="26" pageNumber="99" pagination="33 - 90" refId="ref17688" refString="MODZALEVSKAYA T. L. &amp; CHERKESOVA S. V. 1994. - [Brachiopods], in CHERKESOVA S. V., KARA- TAJUTE- TALIMAA V. N. &amp; MATUKHIN R. G. (eds), Stratigrafiya i fauna nizhnedevonskikh otlozhenji Tarejskogo opornogo razreza (Tajmyr). Nedra, St. Petersburg: 33 - 90 (in Russian)." type="book chapter" year="1994">Modzalevskaya, 1994</bibRefCitation>
</taxonomicName>
from the Tolbat Beds, Tajmyr (
<bibRefCitation id="EF98DACDE95DFFB3D8CFFA2C7FB71814" author="MODZALEVSKAYA T. L. &amp; CHERKESOVA S. V." box="[721,1170,1515,1541]" pageId="26" pageNumber="99" pagination="33 - 90" refId="ref17688" refString="MODZALEVSKAYA T. L. &amp; CHERKESOVA S. V. 1994. - [Brachiopods], in CHERKESOVA S. V., KARA- TAJUTE- TALIMAA V. N. &amp; MATUKHIN R. G. (eds), Stratigrafiya i fauna nizhnedevonskikh otlozhenji Tarejskogo opornogo razreza (Tajmyr). Nedra, St. Petersburg: 33 - 90 (in Russian)." type="book chapter" year="1994">Modzalevskaya &amp; Cherkesova 1994: 87</bibRefCitation>
, pl. X, figs 11-14). However, the fewer plicae, comparatively weak dorsal fold with deep narrow furrows on the flanks support assignment of Severnaya Zemlya specimens to a new species.
</paragraph>
</subSubSection>
</treatment>
</document>